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<document id="4F6062E05C549EFAC1888432DE499DD4" ID-CLB-Dataset="7130" ID-DOI="10.12782/sd.18.1.023" ID-GBIF-Dataset="0e6ff786-9ade-4aa8-940f-3346a7f26866" ID-ISSN="2189-7301" ID-Zenodo-Dep="4584815" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614946392566" checkinUser="felipe" docAuthor="Komai, Tomoyuki" docDate="2013" docId="03FF87B3E910300CFEB30DA5FD0D3904" docLanguage="en" docName="SpeciesDiversity.18.1.23-32.pdf" docOrigin="Species Diversity 18 (1)" docSource="http://dx.doi.org/10.12782/sd.18.1.023" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Pliopagurus curvimanus Komai 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FFC6FFCBE9123004FF960D63FFF23844" masterDocTitle="A New Genus and New Species of Paguridae (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines" masterLastPageNumber="32" masterPageNumber="23" pageNumber="25" updateTime="1698932712113" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F2B74DAC8B69085E8CC9EBE2A5FD8AA1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D74C201C86221B936060D2E800FE676">
<mods:title id="0C00964D41D00900B21CE2E008FEA195">A New Genus and New Species of Paguridae (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="AE8C4D185AF2C5250D37C3534D1C2291" type="personal">
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<mods:namePart id="2AF8A77CDD108EC6E17EF38A2F012CF0">Komai, Tomoyuki</mods:namePart>
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<mods:title id="76D0DF33F84CFD1997AF87354473E3E6">Species Diversity</mods:title>
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<mods:part id="B35F689CE1C4017F67E0E5DEC847C961">
<mods:date id="53FC0DFF1805CF566579AD57C6BDA2DF">2013</mods:date>
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<mods:number id="2C75CD2A23864C63C6D13B682AEBAECE">2013-05-25</mods:number>
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<mods:number id="62EBB8D2A87473FC33E6EF6D799679A0">1</mods:number>
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<mods:classification id="72EF645CE0BE4E87D9FC562E4DC25023">journal article</mods:classification>
<mods:identifier id="DC8A086F3AF68130E299D45B3F8A56C1" type="CLB-Dataset">7130</mods:identifier>
<mods:identifier id="2BF05E0A2A81F5782D4F3A1AF970CD25" type="DOI">10.12782/sd.18.1.023</mods:identifier>
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<treatment id="03FF87B3E910300CFEB30DA5FD0D3904" ID-DOI="http://doi.org/10.5281/zenodo.4730819" ID-GBIF-Taxon="181761969" ID-Zenodo-Dep="4730819" LSID="urn:lsid:plazi:treatment:03FF87B3E910300CFEB30DA5FD0D3904" httpUri="http://treatment.plazi.org/id/03FF87B3E910300CFEB30DA5FD0D3904" lastPageId="8" lastPageNumber="31" pageId="2" pageNumber="25">
<subSubSection id="C34C652EE9103006FEB30DA5FD85389A" box="[293,631,198,222]" pageId="2" pageNumber="25" type="nomenclature">
<paragraph id="8BE936A5E9103006FEB30DA5FD85389A" blockId="2.[293,631,198,222]" box="[293,631,198,222]" pageId="2" pageNumber="25">
<heading id="D0A181C9E9103006FEB30DA5FD85389A" box="[293,631,198,222]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="25" reason="2">
<taxonomicName id="4C564D26E9103006FEB30DA5FDD7389A" ID-CoL="fa70211b-be4f-42cd-913b-0c0800f2ac06" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[293,549,198,222]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="curvimanus" status="sp. nov.">
<emphasis id="B922EAB7E9103006FEB30DA5FDD7389A" bold="true" box="[293,549,198,222]" italics="true" pageId="2" pageNumber="25">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9103006FDBC0DA4FD85389A" box="[554,631,199,222]" pageId="2" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C652EE9103006FE350D84FE0A38BB" box="[419,504,231,255]" pageId="2" pageNumber="25" type="description">
<paragraph id="8BE936A5E9103006FE350D84FE0A38BB" blockId="2.[419,504,231,255]" box="[419,504,231,255]" pageId="2" pageNumber="25">
<heading id="D0A181C9E9103006FE350D84FE0A38BB" box="[419,504,231,255]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="25" reason="2">
<figureCitation id="136D2A20E9103006FE350D84FE0A38BB" box="[419,504,231,255]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="3.[156,187,1820,1842]" captionStartId-1="4.[157,188,838,860]" captionStartId-2="5.[156,187,1817,1839]" captionStartId-3="6.[156,187,1061,1083]" captionStartId-4="7.[156,187,1554,1576]" captionTargetBox-0="[259,1330,204,1798]" captionTargetBox-1="[178,1409,204,816]" captionTargetBox-2="[306,1281,205,1795]" captionTargetBox-3="[258,1329,204,1039]" captionTargetBox-4="[246,1344,204,1532]" captionTargetId-0="figure-4@3.[258,1330,204,1798]" captionTargetId-1="figure-650@4.[178,1409,204,816]" captionTargetId-2="figure-79@5.[306,1281,204,1795]" captionTargetId-3="figure-885@6.[258,1330,204,1039]" captionTargetId-4="figure-429@7.[244,1344,204,1532]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." captionText-1="Fig. 2. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view.Scale bar: 0.5mm." captionText-2="Fig. 3. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view.Scale bar: 1mm." captionText-3="Fig. 4. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view.Scale bar: 1mm." captionText-4="Fig. 5. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left second pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown); C, left third pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown). Scale bars: 1mm for A, C; 0.5mm for B, D." figureDoi-0="http://doi.org/10.5281/zenodo.4584817" figureDoi-1="http://doi.org/10.5281/zenodo.4584819" figureDoi-2="http://doi.org/10.5281/zenodo.4584821" figureDoi-3="http://doi.org/10.5281/zenodo.4584823" figureDoi-4="http://doi.org/10.5281/zenodo.4584825" httpUri-0="https://zenodo.org/record/4584817/files/figure.png" httpUri-1="https://zenodo.org/record/4584819/files/figure.png" httpUri-2="https://zenodo.org/record/4584821/files/figure.png" httpUri-3="https://zenodo.org/record/4584823/files/figure.png" httpUri-4="https://zenodo.org/record/4584825/files/figure.png" pageId="2" pageNumber="25">Figs 1–5</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C652EE9103006FF230C4BFDC639A6" pageId="2" pageNumber="25" type="materials_examined">
<paragraph id="8BE936A5E9103006FF230C4BFD4D39C5" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
<emphasis id="B922EAB7E9103006FF230C4BFE763904" bold="true" box="[181,388,296,320]" pageId="2" pageNumber="25">Material examined</emphasis>
.
<materialsCitation id="3B3E3CF8E9103006FE320C4BFD4D39C5" ID-GBIF-Occurrence="3095328302" collectingDate="2004-07-02" elevation="85" latitude="9.52" location="Sea" longLatPrecision="129" longitude="123.67834" municipality="Balicasag Island" pageId="2" pageNumber="25" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54ED8807E9103006FE320C4BFDF73904" box="[420,517,296,320]" pageId="2" pageNumber="25" type="holotype">Holotype</typeStatus>
: PANGLAO 2004, stn L42,
<collectingMunicipality id="6B8DACDFE9103006FF430C2BFE723924" box="[213,384,328,352]" pageId="2" pageNumber="25">Balicasag Island</collectingMunicipality>
,
<collectingRegion id="4992F847E9103006FE070C2BFE3C3924" box="[401,462,328,352]" country="Philippines" name="Bohol" pageId="2" pageNumber="25">Bohol</collectingRegion>
<location id="8E89607EE9103006FE4D0C2AFE0D3925" LSID="urn:lsid:plazi:treatment:03FF87B3E910300CFEB30DA5FD0D3904:8E89607EE9103006FE4D0C2AFE0D3925" box="[475,511,329,353]" latitude="9.52" longLatPrecision="129" longitude="123.67834" municipality="Balicasag Island" name="Sea" pageId="2" pageNumber="25" stateProvince="Bohol">Sea</location>
,
<geoCoordinate id="EE625062E9103006FD870C2AFD8B3924" box="[529,633,327,353]" degrees="09" direction="north" minutes="31.2" orientation="latitude" pageId="2" pageNumber="25" precision="92" value="9.52">09°31.2′N</geoCoordinate>
,
<geoCoordinate id="EE625062E9103006FD1D0C2AFD093924" box="[651,763,327,353]" degrees="123" direction="east" minutes="40.7" orientation="longitude" pageId="2" pageNumber="25" precision="92" value="123.67834">123°40.7′E</geoCoordinate>
,
<quantity id="4CAE9B40E9103006FF0A0C0AFF0139C5" box="[156,243,361,385]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="25" unit="m" value="85.0" valueMax="90.0" valueMin="80.0">
<elevation id="007BD196E9103006FF0A0C0AFF0139C5" box="[156,243,361,385]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="25" unit="m" value="85.0" valueMax="90.0" valueMin="80.0">80–90 m</elevation>
</quantity>
,
<date id="FFE81065E9103006FF6B0C0AFE9D39C5" box="[253,367,361,385]" pageId="2" pageNumber="25" value="2004-07-02">
<collectingDate id="EFACE98DE9103006FF6B0C0AFE9D39C5" box="[253,367,361,385]" pageId="2" pageNumber="25" value="2004-07-02">2 July 2004</collectingDate>
</date>
, male (sl
<quantity id="4CAE9B40E9103006FE5B0C0AFDE939C5" box="[461,539,361,385]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="25" unit="mm" value="2.5">2.5 mm</quantity>
), NMCR 39083.
</materialsCitation>
</paragraph>
<paragraph id="8BE936A5E9103006FF230CE9FDC639A6" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
<materialsCitation id="3B3E3CF8E9103006FF230CE9FDDD39A6" ID-GBIF-Occurrence="3095328301" collectingDate="2004-07-02" collectionCode="ZRC" elevation="85" latitude="9.52" location="Sea" longLatPrecision="129" longitude="123.67834" municipality="Balicasag Island" pageId="2" pageNumber="25" specimenCode="ZRC 2013.0578" specimenCount="2" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="54ED8807E9103006FF230CE9FEEB39E6" box="[181,281,394,418]" pageId="2" pageNumber="25" type="paratype">Paratypes</typeStatus>
: same data as holotype, 1 ovigerous female (sl
<quantity id="4CAE9B40E9103006FF0A0CC9FF183986" box="[156,234,426,450]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="25" unit="mm" value="1.3">1.3 mm</quantity>
),
<specimenCode id="DBF09EDEE9103006FF680CC9FE523986" box="[254,416,426,450]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="25">ZRC 2013.0578</specimenCode>
; same data as
<typeStatus id="54ED8807E9103006FDAD0CC9FD643986" box="[571,662,426,450]" pageId="2" pageNumber="25" type="holotype">holotype</typeStatus>
, 1 ovigerous female (sl
<quantity id="4CAE9B40E9103006FEBA0CA9FE8839A7" box="[300,378,458,483]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="25" unit="mm" value="1.5">1.5 mm</quantity>
), CBM-ZC 11571
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EE9103002FF230C88FF173C8B" lastPageId="6" lastPageNumber="29" pageId="2" pageNumber="25" type="description">
<paragraph id="8BE936A5E9103006FF230C88FD4A3AA3" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
<emphasis id="B922EAB7E9103006FF230C88FEC13A47" bold="true" box="[181,307,491,515]" pageId="2" pageNumber="25">Description</emphasis>
. Shield (
<figureCitation id="136D2A20E9103006FE080C88FE193A47" box="[414,491,491,515]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1A</figureCitation>
) about 1.1 times as long as wide; anterior margin between rostrum and lateral projections concave; dorsal surface with some tufts of short setae laterally, paragastric grooves delimited only on posterior parts; anterolateral margins sloping. Rostrum broadly triangular, subacutely pointed, extending beyond lateral projections. Lateral projections broadly rounded, moderately separated from rostrum, each with 1 submarginal spinule.
</paragraph>
<paragraph id="8BE936A5E9103006FF230F8CFDF43C48" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
Ocular peduncles (
<figureCitation id="136D2A20E9103006FEE90F8CFE223B43" box="[383,464,751,775]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1A</figureCitation>
) about 0.8 times as long as shield, moderately slender, faintly constricted at mid-length, each with longitudinal row of tufts of short setae on dorsal surface; cornea slightly dilated; basal width of peduncle subequal to corneal width. Ocular acicles narrowly triangular, separated basally by about basal width of 1 acicle, each with acutely or subacutely pointed tip, without submarginal spine distally; distal part of ocular acicle nearly flat on dorsal surface. Interocular lobe clearly visible.
</paragraph>
<paragraph id="8BE936A5E9103006FF230977FEA03C8B" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
Antennular peduncles (
<figureCitation id="136D2A20E9103006FE320977FE023C68" box="[420,496,1044,1068]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1A</figureCitation>
) slightly overreaching distal corneal margins. Ultimate segment with single short seta near dorsolateral distal angle, dorsal surface with few additional short setae. Basal segment with moderately inflated statocyst lobe bearing small lateral spine; ventrodistal angle slightly produced.
</paragraph>
<paragraph id="8BE936A5E9103006FF2309B4FE183EBC" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
Antennal peduncles (
<figureCitation id="136D2A20E9103006FE1909B4FE2F3CAB" box="[399,477,1239,1263]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1A</figureCitation>
) slightly overreaching distal corneal margins. Fifth segment with row of short, stiff setae on mesial margin. Fourth segment with some short, stiff setae. Third segment with tiny spine at ventromesial distal angle, although this spine not visible in dorsal view. Second segment with dorsolateral distal angle produced into strong spine not reaching midlength of fourth segment, mesial margin without additional spinules; dorsomesial distal angle with tiny spine. First segment unarmed on lateral surface; ventrodistal margin strongly produced, with minute terminal spine. Antennal acicle slightly arcuate, reaching distal corneal margin, terminating in tiny spine, with row of short setae dorsomesially (right antennal acicle of
<typeStatus id="54ED8807E9103006FDCA0B3DFD473E32" box="[604,693,1630,1654]" pageId="2" pageNumber="25" type="holotype">holotype</typeStatus>
abnormally short, only reaching corneal base). Antennal flagellum about 5 times as long as shield, overreaching extended right cheliped; each article with 1–3 short setae, with 1 additional mesial long setae every 2 articles.
</paragraph>
<paragraph id="8BE936A5E9103006FF230A63FD1D3058" blockId="2.[156,768,296,2077]" pageId="2" pageNumber="25">
Mandible without distinguishing characters. Maxillule (
<figureCitation id="136D2A20E9103006FF350A42FF073F7D" box="[163,245,1825,1849]" captionStart="Fig" captionStartId="4.[157,188,838,860]" captionTargetBox="[178,1409,204,816]" captionTargetId="figure-650@4.[178,1409,204,816]" captionTargetPageId="4" captionText="Fig. 2. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view.Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4584819" httpUri="https://zenodo.org/record/4584819/files/figure.png" pageId="2" pageNumber="25">Fig. 2A</figureCitation>
) with proximal endite subquadrate; distal endite slightly widened distally, mesial margin truncate; endopod with obsolete inner lobe bearing 2 bristle-like setae and slender, non-recurved outer lobe. Maxilla (
<figureCitation id="136D2A20E9103006FDFB0AE1FD4C3FDE" box="[621,702,1922,1946]" captionStart="Fig" captionStartId="4.[157,188,838,860]" captionTargetBox="[178,1409,204,816]" captionTargetId="figure-650@4.[178,1409,204,816]" captionTargetPageId="4" captionText="Fig. 2. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view.Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4584819" httpUri="https://zenodo.org/record/4584819/files/figure.png" pageId="2" pageNumber="25">Fig. 2B</figureCitation>
) with anterior lobe of scaphognathite not reaching level of distal margin of distal endite. First maxilliped (
<figureCitation id="136D2A20E9103006FDD40AA0FD623F98" box="[578,656,1987,2012]" captionStart="Fig" captionStartId="4.[157,188,838,860]" captionTargetBox="[178,1409,204,816]" captionTargetId="figure-650@4.[178,1409,204,816]" captionTargetPageId="4" captionText="Fig. 2. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view.Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4584819" httpUri="https://zenodo.org/record/4584819/files/figure.png" pageId="2" pageNumber="25">Fig. 2C</figureCitation>
) with exopod moderately expanded proximolaterally. Second maxilliped (
<figureCitation id="136D2A20E9103006FF4D0567FEDA3059" box="[219,296,2052,2077]" captionStart="Fig" captionStartId="4.[157,188,838,860]" captionTargetBox="[178,1409,204,816]" captionTargetId="figure-650@4.[178,1409,204,816]" captionTargetPageId="4" captionText="Fig. 2. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view.Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4584819" httpUri="https://zenodo.org/record/4584819/files/figure.png" pageId="2" pageNumber="25">Fig. 2D</figureCitation>
) with endopod relatively short; exopod long.
</paragraph>
<paragraph id="8BE936A5E9103006FCD80DA5FB493924" blockId="2.[820,1432,198,2076]" pageId="2" pageNumber="25">
Third maxilliped (
<figureCitation id="136D2A20E9103006FB800DA5FB95389A" box="[1046,1127,198,222]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1B</figureCitation>
) moderately stout. Ischium (
<figureCitation id="136D2A20E9103006FCAA0D84FC7938BB" box="[828,907,231,255]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="2" pageNumber="25">Fig. 1C</figureCitation>
) bearing well-developed crista dentata consisting of row of small corneous teeth increasing in size proximally and 1 accessory tooth. Merus and carpus unarmed. Exopod overreaching distal margin of ischium.
</paragraph>
<paragraph id="8BE936A5E9103006FCD80C0AFBE63EF2" blockId="2.[820,1432,198,2076]" pageId="2" pageNumber="25">
Chelipeds appreciably unequal and dissimilar. Right cheliped (
<figureCitation id="136D2A20E9103006FC0A0CEAFC2B39E5" box="[924,985,393,417]" captionStart="Fig" captionStartId="5.[156,187,1817,1839]" captionTargetBox="[306,1281,205,1795]" captionTargetId="figure-79@5.[306,1281,204,1795]" captionTargetPageId="5" captionText="Fig. 3. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view.Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4584821" httpUri="https://zenodo.org/record/4584821/files/figure.png" pageId="2" pageNumber="25">Fig. 3</figureCitation>
A–D) thick, suboperculiform; angle of articulation of chela and carpus about 30°. Dactylus distinctly shorter than palm, strongly curved ventrally; dorsal surface with distinct median ridge accompanied by shallow sulcus mesially, otherwise almost smooth; dorsomesial margin carinate, with row of small blunt denticles decreasing in size distally; mesial surface with broad, shallow sulcus medially, flanked by rows of short setae; cutting edge with row of rounded calcareous teeth, terminating in small corneous claw. Palm slightly widened distally, slightly longer than carpus; dorsal surface gently convex, unarmed, with sparse single setae or tufts of short setae; dorsolateral margin with tuberculate carina extending to midlength of fixed finger but not to proximal margin of palm; dorsomesial margin slightly elevated, with single row of small spines or tubercles proximally decreasing in size and acuteness and with row of moderately long stiff setae, dorsomesial distal angle slightly produced; lateral surface almost glabrous, slightly rugose with scattered short, oblique ridges; mesial surface nearly flat, with scattered granules and sparse setae, and with low, oblique ridge proximoventrally; ventral surface almost glabrous, with shallow concavity at base of fixed finger. Fixed finger with large, subtriangular tooth on median part of cutting edge, terminating in small corneous claw. Carpus subequal in length to merus, distinctly widened distally; dorsomesial margin with row of long setae and small spines at least in distal half; dorsal midline slightly elevated, with 4–6 tiny to moderately small spines and 2 irregular rows of long setae; dorsolateral surface sloping, almost glabrous; mesial surface shallowly depressed distally, with long setae dorsally and ventrally; ventral surface strongly convex, with scattered short to long setae. Merus unarmed on dorsodistal margin; dorsal surface also unarmed, with moderately long setae in distal half; lateral surface glabrous, distal half of ventrolateral margin slightly concave and with row of some tiny spines; mesial surface also almost glabrous, distal half of ventromesial margin with row of small spines; ventral surface convex in proximal half, with numerous long setae, distal half forming concavity accommodating proximal part of carpus when entire cheliped flexed. Ischium unarmed, with short setae dorsally and ventrally.
</paragraph>
<paragraph id="8BE936A5E9103000FCD80BDCFD7A3E11" blockId="2.[820,1432,198,2076]" lastBlockId="4.[156,767,947,2077]" lastPageId="4" lastPageNumber="27" pageId="2" pageNumber="25">
Left cheliped (
<figureCitation id="136D2A20E9103006FC7D0BDCFBD93E93" box="[1003,1067,1727,1751]" captionStart="Fig" captionStartId="6.[156,187,1061,1083]" captionTargetBox="[258,1329,204,1039]" captionTargetId="figure-885@6.[258,1330,204,1039]" captionTargetPageId="6" captionText="Fig. 4. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view.Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4584823" httpUri="https://zenodo.org/record/4584823/files/figure.png" pageId="2" pageNumber="25">Fig. 4</figureCitation>
A–D) with chela curving laterally and ventrally, like a bird beak in general shape, subequal in length to carpus and merus combined; angle of articulation of chela and carpus about 45°. Dactylus about twice as long as palm, surfaces unarmed, bearing scattered long setae; dorsal surface with shallow median sulcus extending beyond midlength; cutting edge with row of closely-set corneous teeth (several teeth partially fused), terminating in small corneous claw. Palm short, about half length of carpus; dorsomesial margin with row of 4 or 5 small spines; dorsal surface sloping to lateral surface, with row of long setae on midline; lateral and mesial faces with sparse short setae; ventral surface slightly convex, with few short setae; fixed finger with scattered tufts of long setae, cutting edge with row of small corneous teeth, terminating in small corneous claw. Carpus subequal in length to merus; dorsolateral margin with row of 3 or 4 spines (dorsodistal spine strong, other 2 spines small) in distal half, dorsomesial margin with row of 4 or 5 spines decreasing in size proximally, each margin also bearing row of long, stiff setae; dorsodistal margin with 1 additional minute spine laterally; lateral surface with short to long setae dorsally, otherwise glabrous, ventrolateral distal angle with minute spine; mesial surface smooth, with long setae dorsally and ventrally; ventral surface convex, with several long setae. Merus unarmed on dorsodistal margin; dorsal surface with row of short setae; lateral surface nearly glabrous, generally convex ventrolateral margin with row of prominent spines in distal half; mesial surface also smooth, with few setae dorsally and ventrally, ventromesial margin with row of tiny spines or tubercles in distal two-thirds; ventral surface with numerous setae. Ischium unarmed, with short setae dorsally and ventrally.
</paragraph>
<caption id="DF29662DE9113007FF0A0A7FFB423FE6" ID-DOI="http://doi.org/10.5281/zenodo.4584817" ID-Zenodo-Dep="4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="3" pageNumber="26" startId="3.[156,187,1820,1842]" targetBox="[259,1330,204,1798]" targetPageId="3">
<paragraph id="8BE936A5E9113007FF0A0A7FFB423FE6" blockId="3.[156,1433,1820,1954]" pageId="3" pageNumber="26">
Fig. 1.
<taxonomicName id="4C564D26E9113007FF660A7FFE3B3F76" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[240,457,1820,1842]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="curvimanus" status="gen. nov., sp. nov.">
<emphasis id="B922EAB7E9113007FF660A7FFE3B3F76" box="[240,457,1820,1842]" italics="true" pageId="3" pageNumber="26">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9113007FE460A7EFD863F76" box="[464,628,1821,1842]" pageId="3" pageNumber="26" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
A–C, E, G, H, holotype, male (sl 2.5 mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3 mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5 mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5 mm for B–H.
</paragraph>
</caption>
<caption id="DF29662DE9163000FF0B0E25FE863BD0" ID-DOI="http://doi.org/10.5281/zenodo.4584819" ID-Zenodo-Dep="4584819" httpUri="https://zenodo.org/record/4584819/files/figure.png" pageId="4" pageNumber="27" startId="4.[157,188,838,860]" targetBox="[178,1409,204,816]" targetPageId="4">
<paragraph id="8BE936A5E9163000FF0B0E25FE863BD0" blockId="4.[156,1432,838,916]" pageId="4" pageNumber="27">
Fig. 2.
<taxonomicName id="4C564D26E9163000FF660E25FE3A3B18" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[240,456,838,860]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="curvimanus" status="gen. nov., sp. nov.">
<emphasis id="B922EAB7E9163000FF660E25FE3A3B18" box="[240,456,838,860]" italics="true" pageId="4" pageNumber="27">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9163000FE580E24FD9D3B18" box="[462,623,839,860]" pageId="4" pageNumber="27" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083, left mouthparts. A, maxillule, ventral view; inset, endopod, lateral view; B, maxilla, ventral view; C, first maxilliped, ventral view; D, second maxilliped, ventral view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BE936A5E9163000FF230B3DFC573D74" blockId="4.[156,767,947,2077]" lastBlockId="4.[820,1433,946,2076]" pageId="4" pageNumber="27">
Ambulatory legs (
<figureCitation id="136D2A20E9163000FEE70B3DFE113E32" box="[369,483,1630,1654]" captionStart="Fig" captionStartId="7.[156,187,1554,1576]" captionTargetBox="[246,1344,204,1532]" captionTargetId="figure-429@7.[244,1344,204,1532]" captionTargetPageId="7" captionText="Fig. 5. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left second pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown); C, left third pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown). Scale bars: 1mm for A, C; 0.5mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.4584825" httpUri="https://zenodo.org/record/4584825/files/figure.png" pageId="4" pageNumber="27">Fig. 5A, D</figureCitation>
) generally similar on right and left. Dactyli slightly curved ventrally in lateral view, nearly straight in dorsal view, 1.5–1.6 times as long as propodi; dorsal margins each with row of sparse tufts of moderately long setae decreasing in length distally; lateral and mesial surfaces without sulcus or groove, mesial surfaces each with 2 min corneous spinules adjacent to dorsal margin in distal half (second;
<figureCitation id="136D2A20E9163000FE200A22FDF13F1D" box="[438,515,1857,1881]" captionStart="Fig" captionStartId="7.[156,187,1554,1576]" captionTargetBox="[246,1344,204,1532]" captionTargetId="figure-429@7.[244,1344,204,1532]" captionTargetPageId="7" captionText="Fig. 5. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left second pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown); C, left third pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown). Scale bars: 1mm for A, C; 0.5mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.4584825" httpUri="https://zenodo.org/record/4584825/files/figure.png" pageId="4" pageNumber="27">Fig. 5B</figureCitation>
) or double row of 6 or 7 min corneous spinules adjacent to dorsal margin (third;
<figureCitation id="136D2A20E9163000FF0A0AE1FF1C3FDE" box="[156,238,1922,1946]" captionStart="Fig" captionStartId="7.[156,187,1554,1576]" captionTargetBox="[246,1344,204,1532]" captionTargetId="figure-429@7.[244,1344,204,1532]" captionTargetPageId="7" captionText="Fig. 5. Pliopagurus curvimanus gen. nov., sp. nov., holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left second pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown); C, left third pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown). Scale bars: 1mm for A, C; 0.5mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.4584825" httpUri="https://zenodo.org/record/4584825/files/figure.png" pageId="4" pageNumber="27">Fig. 5D</figureCitation>
); ventral margins each with row of 8–11 slender corneous spines. Propodi with numerous tufts of long setae on dorsal margins and fewer tufts of long setae on ventral margins; lateral and mesial faces with few tufts of short setae ventrally; ventrodistal margins each with 1 slender corneous spine. Carpi each with tiny dorsodistal spine; dorsal surfaces each with row of long setae; ventral surfaces with few short and long setae (second) or almost glabrous (third). Meri with tufts of moderately long setae on dorsal margins; lateral and mesial faces smooth, bearing row of short or very short setae ventrally; ventral margins each with tufts of long setae (second) or few short to long setae (third), ventrolateral distal margin slightly concave and armed with tiny subterminal spine (second) or straight and unarmed (third). Ischia about half length of meri (second) or subequal in length to it (third). Female with paired gonopores on coxae of third pereopods.
</paragraph>
<paragraph id="8BE936A5E9163000FCD8085AFBEC3DF6" blockId="4.[820,1433,946,2076]" pageId="4" pageNumber="27">
Fourth pereopods (
<figureCitation id="136D2A20E9163000FBB6085AFB863D15" box="[1056,1140,1337,1361]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1D</figureCitation>
) semichelate. Dactyli each with row of minute corneous teeth on ventral margins; no preungual process. Propodi with rasp consisting of single row of corneous scales.
</paragraph>
<paragraph id="8BE936A5E9163000FCD808D8FBF33EF3" blockId="4.[820,1433,946,2076]" pageId="4" pageNumber="27">
Coxae of fifth pereopods in male (
<figureCitation id="136D2A20E9163000FB2E08D8FAF13D97" box="[1208,1283,1467,1491]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1E</figureCitation>
) symmetrical, bearing tufts of moderately short setae at anterolateral and anteromesial angles, each with very short, abruptly tapering sexual tube directed anteroventrally. Coxae of female fifth pereopods (
<figureCitation id="136D2A20E9163000FC3B0B5EFC073E11" box="[941,1013,1597,1621]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1F</figureCitation>
) unequal with left larger, right with tufts of setae at anterolateral and anteromesial angles, left with numerous longer setae extending from anterior margin to posterolateral angle.
</paragraph>
<paragraph id="8BE936A5E9163000FCD80BDCFC553F1D" blockId="4.[820,1433,946,2076]" pageId="4" pageNumber="27">
Third thoracic sternite unarmed on anterior margin. Sixth thoracic sternite (
<figureCitation id="136D2A20E9163000FBB30B83FB813EBC" box="[1061,1139,1760,1784]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1G</figureCitation>
) with anterior lobe subtrapezoidal; anterior margin rounded, with several long setae. Eighth thoracic sternite (
<figureCitation id="136D2A20E9163000FBA10A42FB703F7D" box="[1079,1154,1825,1849]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1E</figureCitation>
) slightly bilobed, with few short setae.
</paragraph>
<paragraph id="8BE936A5E9163001FCD80A01FDCB3FFF" blockId="4.[820,1433,946,2076]" lastBlockId="5.[156,767,1923,2077]" lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27">
Pleon dextrally twisted. Male with 3 (third to fifth) unpaired left pleopods, decreasing in size posteriorly, third unequally biramous, fourth and fifth uniramous. Female with paired first pleopods modified as gonopods, each indistinctly bi-articulated (
<figureCitation id="136D2A20E9163000FC750A87FBDE3FB8" box="[995,1068,2020,2044]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="4" pageNumber="27">Fig. 1F</figureCitation>
); unpaired, left second to fifth pleopods present, second and third subequally biramous, fourth slightly unequally biramous, and fifth strongly unequally biramous. Uropods strongly asymmetrical.
</paragraph>
<caption id="DF29662DE9173001FF0A0A7AFB9E3F0F" ID-DOI="http://doi.org/10.5281/zenodo.4584821" ID-Zenodo-Dep="4584821" httpUri="https://zenodo.org/record/4584821/files/figure.png" pageId="5" pageNumber="28" startId="5.[156,187,1817,1839]" targetBox="[306,1281,205,1795]" targetPageId="5">
<paragraph id="8BE936A5E9173001FF0A0A7AFB9E3F0F" blockId="5.[156,1432,1817,1867]" pageId="5" pageNumber="28">
Fig. 3.
<taxonomicName id="4C564D26E9173001FF660A7AFE3B3F6B" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[240,457,1817,1839]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="curvimanus" status="gen. nov., sp. nov.">
<emphasis id="B922EAB7E9173001FF660A7AFE3B3F6B" box="[240,457,1817,1839]" italics="true" pageId="5" pageNumber="28">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9173001FE590A79FD813F6B" box="[463,627,1818,1839]" pageId="5" pageNumber="28" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, holotype, male (sl 2.5 mm), PANGLAO 2004, stn L42, NMCR 39083. A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BE936A5E9173001FF230AA7FC3B3FFF" blockId="5.[156,767,1923,2077]" lastBlockId="5.[820,1432,1923,2077]" pageId="5" pageNumber="28">
Telson (
<figureCitation id="136D2A20E9173001FE9E0AA7FEAB3F98" box="[264,345,1988,2012]" captionStart="Fig" captionStartId="3.[156,187,1820,1842]" captionTargetBox="[259,1330,204,1798]" captionTargetId="figure-4@3.[258,1330,204,1798]" captionTargetPageId="3" captionText="Fig. 1. Pliopagurus curvimanus gen. nov., sp. nov. A–C, E, G, H, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083; D, paratype, ovigerous female (sl 1.3mm), same station, ZRC 39083; F, paratype, ovigerous female (sl 1.5mm), same station, CBM-ZC 11571. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium and basis, ventral view; D, left fourth pereopod, lateral view; E, coxae of fifth pereopods and eighth thoracic sternite, ventral view; F, coxae of fifth pereopods and first pleopods, ventral view; G, sixth thoracic sternite, ventral view; H, telson, dorsal view. Scale bars: 1mm for A; 0.5mm for B–H." figureDoi="http://doi.org/10.5281/zenodo.4584817" httpUri="https://zenodo.org/record/4584817/files/figure.png" pageId="5" pageNumber="28">Fig. 1H</figureCitation>
) with faint (
<typeStatus id="54ED8807E9173001FE490AA7FDCE3F98" box="[479,572,1988,2012]" pageId="5" pageNumber="28" type="holotype">holotype</typeStatus>
) or shallow (
<typeStatus id="54ED8807E9173001FD5E0AA7FF213FB8" pageId="5" pageNumber="28" type="paratype">paratypes</typeStatus>
) lateral indentations; posterior lobes slightly asymmetrical, subtriangular, each with few short setae proximally on lateral margin; terminal margins strongly oblique, each with some spinules.
</paragraph>
<paragraph id="8BE936A5E9173002FCD80AA0FF173C8B" blockId="5.[820,1432,1923,2077]" lastBlockId="6.[156,768,1142,1426]" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28">
<emphasis id="B922EAB7E9173001FCD80AA0FB9B3F9F" bold="true" box="[846,1129,1987,2011]" pageId="5" pageNumber="28">Coloration in preservative</emphasis>
. No distinct markings seen on shield, antennae, chelipeds, or ambulatory legs. Ocular peduncle with orange and white stripes, white stripes on dorsal and ventral midline and on lateral and mesial faces, white stripe on dorsal mid-line much narrower than orange stripes.
</paragraph>
</subSubSection>
<caption id="DF29662DE9143002FF0A0946FB9E3C13" ID-DOI="http://doi.org/10.5281/zenodo.4584823" ID-Zenodo-Dep="4584823" httpUri="https://zenodo.org/record/4584823/files/figure.png" pageId="6" pageNumber="29" startId="6.[156,187,1061,1083]" targetBox="[258,1329,204,1039]" targetPageId="6">
<paragraph id="8BE936A5E9143002FF0A0946FB9E3C13" blockId="6.[156,1432,1061,1111]" pageId="6" pageNumber="29">
Fig. 4.
<taxonomicName id="4C564D26E9143002FF660946FE383C7F" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[240,458,1061,1083]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="curvimanus" status="gen. nov., sp. nov.">
<emphasis id="B922EAB7E9143002FF660946FE383C7F" box="[240,458,1061,1083]" italics="true" pageId="6" pageNumber="29">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9143002FE470945FD8A3C7F" box="[465,632,1062,1083]" pageId="6" pageNumber="29" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, holotype, male (sl 2.5 mm), PANGLAO 2004, stn L42, NMCR 39083. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C34C652EE9143002FF2309B4FE263D74" pageId="6" pageNumber="29" type="distribution">
<paragraph id="8BE936A5E9143002FF2309B4FD343D54" blockId="6.[156,768,1142,1426]" pageId="6" pageNumber="29">
<emphasis id="B922EAB7E9143002FF2309B4FEC83CAB" bold="true" box="[181,314,1239,1263]" pageId="6" pageNumber="29">Distribution</emphasis>
. Known only from the
<typeStatus id="54ED8807E9143002FDD209BBFD823CB4" box="[580,624,1240,1264]" pageId="6" pageNumber="29">type</typeStatus>
locality, Balicasag Island,
<collectingRegion id="4992F847E9143002FEB7099BFEAC3D54" box="[289,350,1272,1296]" country="Philippines" name="Bohol" pageId="6" pageNumber="29">Bohol</collectingRegion>
Sea, the
<collectingCountry id="F3417635E9143002FE21099BFDD93D54" box="[439,555,1272,1296]" name="Philippines" pageId="6" pageNumber="29">Philippines</collectingCountry>
,
<quantity id="4CAE9B40E9143002FDA3099BFD793D4B" box="[565,651,1272,1296]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="29" unit="m" value="85.0" valueMax="90.0" valueMin="80.0">80–90 m</quantity>
deep.
</paragraph>
<paragraph id="8BE936A5E9143002FF23087BFE263D74" blockId="6.[156,768,1142,1426]" box="[181,468,1304,1328]" pageId="6" pageNumber="29">
<emphasis id="B922EAB7E9143002FF23087BFEF73D74" bold="true" box="[181,261,1304,1328]" pageId="6" pageNumber="29">Habitat</emphasis>
. Gastropod shells.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EE9143002FF23085AFD3E3DD6" pageId="6" pageNumber="29" type="etymology">
<paragraph id="8BE936A5E9143002FF23085AFD3E3DD6" blockId="6.[156,768,1142,1426]" pageId="6" pageNumber="29">
<emphasis id="B922EAB7E9143002FF23085AFEDA3D15" bold="true" box="[181,296,1337,1361]" pageId="6" pageNumber="29">Etymology</emphasis>
. From the combination of the Latin,
<emphasis id="B922EAB7E9143002FD280859FF413D35" italics="true" pageId="6" pageNumber="29">curvatis</emphasis>
(=curved) and
<emphasis id="B922EAB7E9143002FEC90839FE503D35" box="[351,418,1370,1393]" italics="true" pageId="6" pageNumber="29">manus</emphasis>
(=hand), in reference to the noticeably curved left chela. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34C652EE914300CFF3108D8FD0D3904" lastPageId="8" lastPageNumber="31" pageId="6" pageNumber="29" type="key">
<paragraph id="8BE936A5E9143002FF3108D8FD293DB7" blockId="6.[167,756,1467,1524]" pageId="6" pageNumber="29">
<emphasis id="B922EAB7E9143002FF3108D8FE883D97" bold="true" box="[167,378,1467,1491]" pageId="6" pageNumber="29">Key to genera of the</emphasis>
“
<emphasis id="B922EAB7E9143002FE1F08D8FD563D97" bold="true" box="[393,676,1467,1491]" italics="true" pageId="6" pageNumber="29">Pylopagurus-Tomopagurus</emphasis>
”
<emphasis id="B922EAB7E9143002FD2208D8FD293DB7" bold="true" pageId="6" pageNumber="29">
group [emended from
<bibRefCitation id="EFC74B54E9143002FEFF08B8FD263DB7" author="Lemaitre, R. &amp; McLaughlin, P. A." box="[361,724,1499,1524]" pageId="6" pageNumber="29" pagination="464 - 486" refId="ref6172" refString="Lemaitre, R. and McLaughlin, P. A. 2003 a. Revision of Pylopagurus and Tomopagurus (Crustacea: Decapoda: Paguridae), with descriptions of new genera and species. Addendum and taxonomic summary. Proceedings of the Biological Society of Washington 116: 464 - 486." type="journal article" year="2003">Lemaitre and McLaughlin (2003a)</bibRefCitation>
]
</emphasis>
</paragraph>
<key id="FFC76194E914300CFF0A0B7EFD0D3904" lastPageId="8" lastPageNumber="31" pageId="6" pageNumber="29">
<keyStep id="30A22E00E9143002FF0A0B7EFD0D3E93" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FF0A0B7EFD0D3E31" blockId="6.[156,767,1565,2077]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FF0A0B7EFD0D3E31" pageId="6" pageNumber="29">
1. Pleon reduced; males without unpaired pleopods; females with unpaired second to fourth pleopods.........................................
<taxonomicName id="4C564D26E9143002FDCC0B3DFD0D3E31" ID-CoL="478a9dd5-2135-45ee-82f6-ec171bfa317b" authorityName="Lemaitre and McLaughlin" authorityYear="1996" box="[602,767,1630,1653]" class="Malacostraca" family="Paguridae" genus="Protoniopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FDCC0B3DFD0D3E31" box="[602,767,1630,1653]" italics="true" pageId="6" pageNumber="29">Protoniopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FF0A0B1CFD0D3E93" blockId="6.[156,767,1565,2077]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FF0A0B1CFD0D3E93" pageId="6" pageNumber="29">– Pleon not reduced; males with some unpaired pleopods; females with unpaired second to fifth pleopods.. ............................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FF0A0B83FD0D3F98" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FF0A0B83FD0D3F1D" blockId="6.[156,767,1565,2077]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FF0A0B83FD0D3F1D" pageId="6" pageNumber="29">
2. Protopods of uropods prominently produced posteriorly; dorsal surface of right chela commonly with characteristic covering of mushroom-shaped tubercles........................................
<taxonomicName id="4C564D26E9143002FDE10A22FD0D3F1D" ID-CoL="41a85561-4afd-48bd-9d56-63068c3d31d6" authorityName="McLaughlin" authorityYear="1981" box="[631,767,1857,1881]" class="Malacostraca" family="Paguridae" genus="Agaricochirus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FDE10A22FD0D3F1D" box="[631,767,1857,1881]" italics="true" pageId="6" pageNumber="29">Agaricochirus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FF0A0A01FD0D3F98" blockId="6.[156,767,1565,2077]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FF0A0A01FD0D3F98" pageId="6" pageNumber="29">– Protopods of uropods not prominently produced posteriorly; dorsal surface of right chela usually without characteristic covering of mushroom-shaped tubercles ............................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FF0A0A87FA653CEA" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FF0A0A87FD0D3059" blockId="6.[156,767,1565,2077]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FF0A0A87FD0D3059" pageId="6" pageNumber="29">
3. Spines on dorsal surfaces of chelae with basal rosettes.....................................
<taxonomicName id="4C564D26E9143002FD140566FD0D3059" ID-CoL="bb044813-3600-4b9c-bae9-29353c080de8" authorityName="McLaughlin" authorityYear="1981" box="[642,767,2053,2077]" class="Malacostraca" family="Paguridae" genus="Rhodochirus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FD140566FD0D3059" box="[642,767,2053,2077]" italics="true" pageId="6" pageNumber="29">Rhodochirus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA20915FA653CEA" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20915FA653CEA" pageId="6" pageNumber="29">– Spines on dorsal surfaces of chelae without basal rosettes.......................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FCA209D4FA6A3D74" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA209D4FA6A3CAB" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA209D4FA6A3CAB" pageId="6" pageNumber="29">4. Propodal rasps of fourth pereopods each with more than one row of corneous scales................... 5</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA2099BFA6A3D74" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA2099BFA6A3D74" pageId="6" pageNumber="29">– Propodal rasps of fourth pereopods each with only one row of corneous scales........................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FCA2085AFA6A3DB7" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA2085AFA6A3DD6" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA2085AFA6A3DD6" pageId="6" pageNumber="29">5. Left chela triangular or subtriangular in cross-section, dactylus and fixed finger not dorsoventrally flattened.. ............................................... 6</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA208F8FA6A3DB7" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA208F8FA6A3DB7" pageId="6" pageNumber="29">
– Left chela not triangular or subtriangular in cross-section, dactylus and fixed finger dorsoventrally flattened...................................
<taxonomicName id="4C564D26E9143002FB6408B8FA6A3DB7" ID-CoL="81b5af87-4283-4751-bf54-d7e7f69922e7" authorityName="McLaughlin" authorityYear="1981" box="[1266,1432,1499,1523]" class="Malacostraca" family="Paguridae" genus="Manucomplanus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FB6408B8FA6A3DB7" box="[1266,1432,1499,1523]" italics="true" pageId="6" pageNumber="29">Manucomplanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FCA2089FFA6A3E32" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA2089FFA653E70" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA2089FFA653E70" pageId="6" pageNumber="29">
6. Telson with lateral indentations suggesting division into anterior and posterior portions.....
<taxonomicName id="4C564D26E9143002FA990B7EFA653E70" ID-CoL="6888f3e1-fde3-4b51-9a25-de7080d0c87e" authorityName="McLaughlin" authorityYear="1981" box="[1295,1431,1565,1588]" class="Malacostraca" family="Paguridae" genus="Anisopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FA990B7EFA653E70" box="[1295,1431,1565,1588]" italics="true" pageId="6" pageNumber="29">Anisopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA20B5DFA6A3E32" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20B5DFA6A3E32" pageId="6" pageNumber="29">
– Telson without lateral indentations suggesting division into anterior and posterior portions..
<taxonomicName id="4C564D26E9143002FB660B3DFA6A3E32" ID-CoL="fea1bc48-f27e-4d12-bb75-22084f32c7c5" authorityName="McLaughlin" authorityYear="1981" box="[1264,1432,1630,1654]" class="Malacostraca" family="Paguridae" genus="Enallopaguropsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FB660B3DFA6A3E32" box="[1264,1432,1630,1654]" italics="true" pageId="6" pageNumber="29">Enallopaguropsis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FCA20B1CFA6A3EBC" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA20B1CFA6A3EF3" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20B1CFA6A3EF3" pageId="6" pageNumber="29">
7. Ocular acicles multispinose; coxae of male fifth pereopods asymmetrical.................
<taxonomicName id="4C564D26E9143002FB6E0BFCFA6A3EF3" ID-CoL="11d92cad-fea4-46de-887f-9b685b6e43e7" authorityName="McLaughlin and Lemaitre" authorityYear="2001" box="[1272,1432,1695,1719]" class="Malacostraca" family="Paguridae" genus="Pylopaguridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FB6E0BFCFA6A3EF3" box="[1272,1432,1695,1719]" italics="true" pageId="6" pageNumber="29">Pylopaguridium</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA20BA3FA6A3EBC" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20BA3FA6A3EBC" pageId="6" pageNumber="29">– Ocular acicles simple; coxae of male fifth pereopods symmetrical.................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143002FCA20A63FA6A3F3E" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA20A63FA6A3F7D" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20A63FA6A3F7D" pageId="6" pageNumber="29">8. Telson with lateral indentations suggesting division into anterior and posterior portions................ 9</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143002FCA20A21FA6A3F3E" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20A21FA6A3F3E" pageId="6" pageNumber="29">
– Telson without lateral indentations suggesting division into anterior and posterior portions.....
<taxonomicName id="4C564D26E9143002FA9F0A01FA6A3F3E" ID-CoL="8503997f-5a4d-48b9-9d11-8574b42ffd5e" authorityName="McLaughlin" authorityYear="1981" box="[1289,1432,1890,1914]" class="Malacostraca" family="Paguridae" genus="Enallopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9143002FA9F0A01FA6A3F3E" box="[1289,1432,1890,1914]" italics="true" pageId="6" pageNumber="29">Enallopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9143003FCA20AE1FD0D3ED2" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
<paragraph id="8BE936A5E9143002FCA20AE1FA653FB8" blockId="6.[820,1432,1142,2076]" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143002FCA20AE1FA653FB8" pageId="6" pageNumber="29">9. Chela of right cheliped subovate to subcircular, margins unarmed, weakly tuberculate or minutely crenulate and/or serrate, but never armed with prominent, blunt or acute spines............................ 10</keyLead>
</paragraph>
<paragraph id="8BE936A5E9143003FCA20566FD0D3ED2" blockId="6.[820,1432,1142,2076]" lastBlockId="7.[156,767,1662,2077]" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
<keyLead id="30A79590E9143003FCA20566FD0D3ED2" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">– Chela of right cheliped variable, margins armed with prominent, blunt or acute spines or tubercles....... 11</keyLead>
</paragraph>
</keyStep>
<caption id="DF29662DE9153003FF0A0B71FCB33E1B" ID-DOI="http://doi.org/10.5281/zenodo.4584825" ID-Zenodo-Dep="4584825" httpUri="https://zenodo.org/record/4584825/files/figure.png" pageId="7" pageNumber="30" startId="7.[156,187,1554,1576]" targetBox="[246,1344,204,1532]" targetPageId="7">
<paragraph id="8BE936A5E9153003FF0A0B71FCB33E1B" blockId="7.[156,1432,1554,1632]" pageId="7" pageNumber="30">
Fig. 5.
<taxonomicName id="4C564D26E9153003FF660B71FE3B3E63" ID-CoL="fa70211b-be4f-42cd-913b-0c0800f2ac06" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[240,457,1554,1575]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="curvimanus" status="gen. nov., sp. nov.">
<emphasis id="B922EAB7E9153003FF660B71FE3B3E63" box="[240,457,1554,1575]" italics="true" pageId="7" pageNumber="30">Pliopagurus curvimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9153003FE590B71FD813E63" box="[463,627,1554,1575]" pageId="7" pageNumber="30" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, holotype, male (sl 2.5mm), PANGLAO 2004, stn L42, NMCR 39083. A, left second pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown); C, left third pereopod, lateral view; D, same, dactylus, mesial view (only mesial setae shown). Scale bars: 1 mm for A, C; 0.5 mm for B, D.
</paragraph>
</caption>
<keyStep id="30A22E00E9153003FF0A0BFCFD0D3F5C" pageId="7" pageNumber="30">
<paragraph id="8BE936A5E9153003FF0A0BFCFD0D3E93" blockId="7.[156,767,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FF0A0BFCFD0D3E93" pageId="7" pageNumber="30">
10. Fourth pereopods with large, very prominent preungual process at base of claw.............
<taxonomicName id="4C564D26E9153003FD140BDCFD0D3E93" ID-CoL="820dc039-8543-4606-b469-9dad37a81797" authorityName="McLaughlin" authorityYear="1981" box="[642,767,1727,1751]" class="Malacostraca" family="Paguridae" genus="Phimochirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9153003FD140BDCFD0D3E93" box="[642,767,1727,1751]" italics="true" pageId="7" pageNumber="30">Phimochirus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9153003FF0A0B83FD0D3F5C" blockId="7.[156,767,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FF0A0B83FD0D3F5C" pageId="7" pageNumber="30">
– Fourth pereopods without prominent preungual process at base of claw.....................
<taxonomicName id="4C564D26E9153003FD120A63FD0D3F5C" ID-CoL="e2e685c7-508a-484e-b563-479985a2a59a" authorityName="A. Milne-Edwards and Bouvier" authorityYear="1891" box="[644,767,1792,1816]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9153003FD120A63FD0D3F5C" box="[644,767,1792,1816]" italics="true" pageId="7" pageNumber="30">Pylopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9153003FF0A0A42FD0D3F98" pageId="7" pageNumber="30">
<paragraph id="8BE936A5E9153003FF0A0A42FD0D3F3E" blockId="7.[156,767,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FF0A0A42FD0D3F3E" pageId="7" pageNumber="30">
11. Dactylus and fixed finger of left chela excavated ventrally and presenting spoon-shaped appearance...........................................
<taxonomicName id="4C564D26E9153003FDE10A00FD0D3F3E" ID-CoL="8cdc5535-f27f-4afd-b308-8f11cb83e9a9" authorityName="A.Milne-Edwards &amp; Bouvier" authorityYear="1893" box="[631,767,1891,1914]" class="Malacostraca" family="Paguridae" genus="Tomopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E9153003FDE10A00FD0D3F3E" box="[631,767,1891,1914]" italics="true" pageId="7" pageNumber="30">Tomopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9153003FF0A0AE0FD0D3F98" blockId="7.[156,767,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FF0A0AE0FD0D3F98" pageId="7" pageNumber="30">– Dactylus and fixed finger of left chela not excavated ventrally and not presenting spoon-shaped appearance .............................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9153003FF0A0A87FA6B3F7C" pageId="7" pageNumber="30">
<paragraph id="8BE936A5E9153003FF0A0A87FA653EF3" blockId="7.[156,767,1662,2077]" lastBlockId="7.[820,1433,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FF0A0A87FA653EF3" pageId="7" pageNumber="30">12. Left chela with midline elevated into prominent keel or crest and/or with lateral margin expanded and distinctly carinate; male without sexual tubes or papilla-like protrusions of vasa deferentia.................... 13</keyLead>
</paragraph>
<paragraph id="8BE936A5E9153003FCA20BA3FA6B3F7C" blockId="7.[820,1433,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FCA20BA3FA6B3F7C" pageId="7" pageNumber="30">
– Left chela with midline not elevated into prominent keel and lateral margin not delimited; male with very short but prominent sexual tubes.................................................
<taxonomicName id="4C564D26E9153003FB540A42FACA3F7D" ID-CoL="f90750ac-18b7-4c98-8a04-d533d2dcb3d3" authority="Komai, 2013" authorityName="Komai" authorityYear="2013" box="[1218,1336,1825,1849]" class="Malacostraca" family="Paguridae" genus="Pliopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B922EAB7E9153003FB540A42FACA3F7D" box="[1218,1336,1825,1849]" italics="true" pageId="7" pageNumber="30">Pliopagurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157CCE9153003FAA80A42FA6B3F7C" box="[1342,1433,1825,1849]" pageId="7" pageNumber="30" rank="genus">gen. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E9153003FCA20A21FA653F98" pageId="7" pageNumber="30">
<paragraph id="8BE936A5E9153003FCA20A21FA6A3FDF" blockId="7.[820,1433,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FCA20A21FA6A3FDF" pageId="7" pageNumber="30">
13. Right chela circumscribed by row of dorsomesial, dorsoproximal and dorsolateral marginal spines..............................
<taxonomicName id="4C564D26E9153003FBF20AE1FA6A3FDF" authority="" box="[1124,1432,1922,1947]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="30" phylum="Arthropoda" rank="subGenus" subGenus="Australeremus">
<emphasis id="B922EAB7E9153003FBF20AE1FB003FDE" box="[1124,1266,1922,1946]" italics="true" pageId="7" pageNumber="30">Lophopagurus</emphasis>
(
<emphasis id="B922EAB7E9153003FA960AE1FA7D3FDE" box="[1280,1423,1922,1946]" italics="true" pageId="7" pageNumber="30">Australeremus</emphasis>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E9153003FCA20AC7FA653F98" blockId="7.[820,1433,1662,2077]" pageId="7" pageNumber="30">
<keyLead id="30A79590E9153003FCA20AC7FA653F98" pageId="7" pageNumber="30">– Right chela not circumscribed by row of dorsomesial, dorsoproximal and dorsolateral spines............. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A22E00E915300CFCA20A87FD0D3904" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30">
<paragraph id="8BE936A5E915300CFCA20A87FD0D389A" blockId="7.[820,1433,1662,2077]" lastBlockId="8.[156,767,198,320]" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30">
<keyLead id="30A79590E915300CFCA20A87FD0D389A" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30">
14. Left chela with midline elevated into prominent keel or crest, lateral margin at most granulate or minutely tuberculate.............
<taxonomicName id="4C564D26E91A300CFE5B0DA5FD0D389A" authority="(Lophopagurus)" box="[461,767,198,222]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="31" phylum="Arthropoda" rank="subGenus" subGenus="Lophopagurus">
<emphasis id="B922EAB7E91A300CFE5B0DA5FDA9389A" box="[461,603,198,222]" italics="true" pageId="8" pageNumber="31">Lophopagurus</emphasis>
(
<emphasis id="B922EAB7E91A300CFDFF0DA5FD05389A" box="[617,759,198,222]" italics="true" pageId="8" pageNumber="31">Lophopagurus</emphasis>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE936A5E91A300CFF0A0D84FD0D3904" blockId="8.[156,767,198,320]" pageId="8" pageNumber="31">
<keyLead id="30A79590E91A300CFF0A0D84FD0D3904" pageId="8" pageNumber="31">
– Left chela with midline slightly elevated, but not into prominent keel or crest, lateral margin strongly spinose...................................
<taxonomicName id="4C564D26E91A300CFDFB0C4AFD0D3904" ID-CoL="e3f28d9b-77e0-442f-934a-cf79c9c77fe8" authorityName="McLaughlin" authorityYear="2005" box="[621,767,297,320]" class="Malacostraca" family="Paguridae" genus="Haigiopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB7E91A300CFDFB0C4AFD0D3904" box="[621,767,297,320]" italics="true" pageId="8" pageNumber="31">Haigiopagurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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