389 lines
43 KiB
XML
389 lines
43 KiB
XML
<document id="9D750D3F7D24B4E4222CE6034064087C" ID-CLB-Dataset="20325" ID-DOI="10.11646/zootaxa.5104.3.4" ID-GBIF-Dataset="17de5cf1-e498-4883-b0e7-8bcd5e976d47" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332094" ID-ZooBank="9A9BA43F-3B32-4595-A95A-853B8F72FD90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594675974" checkinUser="plazi" docAuthor="Kaneko, Naoki & Wada, Kaoru" docDate="2022" docId="03EBE556FFFEFFA63795928CA835FCEB" docLanguage="en" docName="zootaxa.5104.3.4.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaia (M) medea Heller 1892, new species" docType="treatment" docVersion="8" lastPageNumber="429" masterDocId="FFD29D2EFFFFFFA53702931EAB7AFFB6" masterDocTitle="A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia" masterLastPageNumber="432" masterPageNumber="426" pageNumber="427" updateTime="1699316905168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4475354CC6ED0759DDF9FD5059407111">
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<mods:title id="8DF6DBD2A8F2F682A7123436D05C0800">A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia</mods:title>
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<mods:name id="048ECAD769836C5000C2AA8E0C3B6B5B" type="personal">
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<mods:namePart id="78B510BFF3C36D21774EAF088926A3B1">Kaneko, Naoki</mods:namePart>
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<mods:roleTerm id="D25B9474641FC4644AEE2DA98C04D122">Author</mods:roleTerm>
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<mods:namePart id="C3B0404AF02DE0450D2B811EAF47E13A">Wada, Kaoru</mods:namePart>
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<mods:typeOfResource id="07126D507AB5295E1A18E6409345927B">text</mods:typeOfResource>
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<mods:title id="7C1C4414311EF00EC8B80D597E9BA74F">Zootaxa</mods:title>
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<mods:part id="9590C661742B2D62C932A3BD55C8DE06">
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<mods:date id="3EB9F3C877210977EA1F88143583DC2E">2022</mods:date>
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<mods:number id="77DF6C530D633CF39171B80077C1CFAA">2022-02-28</mods:number>
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<mods:number id="9ED8B1CC8DE2A2941CE3F23632DBF6D2">5104</mods:number>
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<mods:identifier id="4DC08C3077DF4646FEA1224AC5C8F913" type="CLB-Dataset">20325</mods:identifier>
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<mods:identifier id="99F7182327074AC3FCF99E3F13F6443B" type="DOI">10.11646/zootaxa.5104.3.4</mods:identifier>
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<mods:identifier id="40897FEC8D9954AA4644972D20590E52" type="GBIF-Dataset">17de5cf1-e498-4883-b0e7-8bcd5e976d47</mods:identifier>
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<treatment id="03EBE556FFFEFFA63795928CA835FCEB" ID-DOI="http://doi.org/10.5281/zenodo.6343750" ID-GBIF-Taxon="193892592" ID-Zenodo-Dep="6343750" LSID="urn:lsid:plazi:treatment:03EBE556FFFEFFA63795928CA835FCEB" httpUri="http://treatment.plazi.org/id/03EBE556FFFEFFA63795928CA835FCEB" lastPageId="3" lastPageNumber="429" pageId="1" pageNumber="427">
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<subSubSection id="C35807CBFFFEFFA43795928CA8A6FE1A" box="[151,988,402,429]" pageId="1" pageNumber="427" type="nomenclature">
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<paragraph id="8BFD5440FFFEFFA43795928CA8A6FE1A" blockId="1.[151,988,402,429]" box="[151,988,402,429]" pageId="1" pageNumber="427">
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<heading id="D0B5E32CFFFEFFA43795928CA8A6FE1A" bold="true" box="[151,988,402,429]" fontSize="11" level="1" pageId="1" pageNumber="427" reason="1">
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<emphasis id="B9368852FFFEFFA43795928CA8A6FE1A" bold="true" box="[151,988,402,429]" pageId="1" pageNumber="427">
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<taxonomicName id="4C422FC3FFFEFFA43795928CA920FE1B" authority="Kaneko & Wada" authorityName="Heller" authorityYear="1892" box="[151,602,402,429]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B9368852FFFEFFA43795928CAB97FE1B" bold="true" box="[151,237,402,429]" italics="true" pageId="1" pageNumber="427">Malaia</emphasis>
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||
(
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<emphasis id="B9368852FFFEFFA437FD928DAA63FE1B" bold="true" box="[255,281,403,429]" italics="true" pageId="1" pageNumber="427">M</emphasis>
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||
.)
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<emphasis id="B9368852FFFEFFA43633928CAA05FE1B" bold="true" box="[305,383,402,429]" italics="true" pageId="1" pageNumber="427">medea</emphasis>
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Kaneko & Wada
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2053529FFFEFFA43565928DA98DFE1A" box="[615,759,402,429]" pageId="1" pageNumber="427" rank="species">new species</taxonomicNameLabel>
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||
(
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<figureCitation id="137948C5FFFEFFA43405928CA800FE1A" box="[775,890,402,428]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1678,1702]" captionTargetBox="[162,1427,189,1638]" captionTargetId="figure-17@2.[151,1436,179,1652]" captionTargetPageId="2" captionText="FIGURES 1–9. Habitus of Malaia (M.) medea Kaneko & Wada, new species (1–6) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (7–9). 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, paratype, dorsal view (color variability); 5, allotype, dorsal view; 6, allotype, ventral view; 7, specimen from RIEB, dorsal view; 8, the same, ventral view; 9, the same, lateral view. Scale: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332096" httpUri="https://zenodo.org/record/6332096/files/figure.png" pageId="1" pageNumber="427">Figs. 1−6</figureCitation>
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,
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<figureCitation id="137948C5FFFEFFA4348A928CA8A8FE1A" box="[904,978,402,428]" captionStart="FIGURES 10–13" captionStartId="4.[151,264,1911,1935]" captionTargetBox="[234,1358,191,1880]" captionTargetId="figure-17@4.[215,1371,181,1887]" captionTargetPageId="4" captionText="FIGURES 10–13. Aedeagus of Malaia (M.) medea Kaneko & Wada, new species (10–11) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="1" pageNumber="427">10−11</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35807CBFFFEFFA4379592C5AF82FD33" pageId="1" pageNumber="427" type="materials_examined">
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<paragraph id="8BFD5440FFFEFFA4379592C5A849FE43" blockId="1.[151,1437,474,2013]" box="[151,819,474,501]" pageId="1" pageNumber="427">
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<materialsCitation id="3B2A5E1DFFFEFFA4379592C5A855FE43" ID-GBIF-Occurrence="3698900303" box="[151,815,474,501]" country="Indonesia" location="Puncak Palopo" pageId="1" pageNumber="427" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9368852FFFEFFA4379592C5AA4CFE42" bold="true" box="[151,310,474,501]" pageId="1" pageNumber="427">
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||
<typeStatus id="54F9EAE2FFFEFFA4379592C5ABA8FE43" box="[151,210,475,501]" pageId="1" pageNumber="427">Type</typeStatus>
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locality.
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</emphasis>
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||
<location id="8E9D029BFFFEFFA4363E92C5AA9DFE43" LSID="urn:lsid:plazi:treatment:03EBE556FFFEFFA63795928CA835FCEB:8E9D029BFFFEFFA4363E92C5AA9DFE43" box="[316,487,475,501]" country="Indonesia" name="Puncak Palopo" pageId="1" pageNumber="427">Puncak Palopo</location>
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,
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<taxonomicName id="4C422FC3FFFEFFA436F092C5A9C9FE43" authority="Selatan" authorityName="Selatan" box="[498,691,475,501]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi Selatan</taxonomicName>
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,
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<collectingCountry id="F35514D0FFFEFFA435BC92C5A855FE43" box="[702,815,475,501]" name="Indonesia" pageId="1" pageNumber="427">Indonesia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BFD5440FFFEFFA437C592E1AEFDFDD7" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
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||
<emphasis id="B9368852FFFEFFA437C592E1AA08FDAE" bold="true" box="[199,370,510,537]" pageId="1" pageNumber="427">Type material</emphasis>
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||
(
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<specimenCount id="9D449FC9FFFEFFA4368592E1A961FDAC" box="[391,539,511,538]" pageId="1" pageNumber="427" type="generic">8 specimens</specimenCount>
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||
).
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<typeStatus id="54F9EAE2FFFEFFA4353392E1A9E6FDAF" box="[561,668,511,537]" pageId="1" pageNumber="427" type="holotype">Holotype</typeStatus>
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,
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<materialsCitation id="3B2A5E1DFFFEFFA435AE92E1A8B6FDAF" ID-GBIF-Occurrence="3698900302" box="[684,972,511,537]" collectionCode="NMNS" pageId="1" pageNumber="427" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<specimenCount id="9D449FC9FFFEFFA435AE92E1A9BBFDAF" box="[684,705,511,537]" pageId="1" pageNumber="427" type="male">♂</specimenCount>
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(deposited in
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<collectionCode id="ED53CC85FFFEFFA4346F92E1A8BEFDAF" box="[877,964,511,537]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="427">NMNS</collectionCode>
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||
)
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||
</materialsCitation>
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||
:
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<materialsCitation id="3B2A5E1DFFFEFFA434D992E0A980FD8B" ID-GBIF-Occurrence="3698900309" collectingDate="2012-06-23" collectingDateMax="2012-06-25" collectingDateMin="2012-06-23" country="Indonesia" location="Sulawesi" pageId="1" pageNumber="427" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
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“[
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||
<collectingCountry id="F35514D0FFFEFFA434E992E0AFF6FDAF" box="[1003,1164,510,537]" name="Indonesia" pageId="1" pageNumber="427">INDONESIA</collectingCountry>
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] / Puncak Palopo, / C.
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<collectingRegion id="49869AA2FFFEFFA43795913DAA7AFD8B" box="[151,256,547,573]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="427">Sulawesi</collectingRegion>
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||
. /
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<date id="FFFC7280FFFEFFA43614913CAAA7FD8B" box="[278,477,546,573]" pageId="1" pageNumber="427" value="2012-06-23" valueMax="2012-06-25" valueMin="2012-06-23">
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||
<collectingDate id="EFB88B68FFFEFFA43614913CAAA7FD8B" box="[278,477,546,573]" pageId="1" pageNumber="427" value="2012-06-23" valueMax="2012-06-25" valueMin="2012-06-23">23—25. VI. 2012</collectingDate>
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</date>
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. / KIYOSHI ANDO leg.”
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</materialsCitation>
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.
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<typeStatus id="54F9EAE2FFFEFFA43401913DA80AFD8B" box="[771,880,547,573]" pageId="1" pageNumber="427" type="paratype">Paratypes</typeStatus>
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||
(
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<specimenCount id="9D449FC9FFFEFFA4347D913CA8C5FD8B" box="[895,959,546,573]" pageId="1" pageNumber="427" type="male">5 ♂♂</specimenCount>
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,
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<specimenCount id="9D449FC9FFFEFFA434CB913DAF7DFD8B" box="[969,1031,547,573]" pageId="1" pageNumber="427" type="female">2 ♀♀</specimenCount>
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||
):
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<specimenCount id="9D449FC9FFFEFFA4331C913DAF25FD8B" box="[1054,1119,547,573]" pageId="1" pageNumber="427" type="male">2 ♂♂</specimenCount>
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||
:
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<materialsCitation id="3B2A5E1DFFFEFFA4336B913DA954FDD7" ID-GBIF-Occurrence="3698900311" country="Indonesia" location="Central SULAWESI" pageId="1" pageNumber="427" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi Tengah" typeStatus="paratype">
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“Puncak, Palopo / VI—VII, 1989 /
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<collectingRegion id="49869AA2FFFEFFA437E59159AAAEFDD7" box="[231,468,583,609]" country="Indonesia" name="Sulawesi Tengah" pageId="1" pageNumber="427">Central SULAWESI</collectingRegion>
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”;
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<specimenCount id="9D449FC9FFFEFFA436E89159A954FDD7" box="[490,558,583,609]" pageId="1" pageNumber="427" type="male">2 ♂♂</specimenCount>
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||
</materialsCitation>
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||
: “
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<materialsCitation id="3B2A5E1DFFFEFFA435419159AEF8FDD7" ID-GBIF-Occurrence="3698900304" box="[579,1410,583,609]" collectorName="Puncak Palopo" country="Indonesia" location="INDONESIA" pageId="1" pageNumber="427" specimenCount="1" typeStatus="paratype">
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||
<collectingCountry id="F35514D0FFFEFFA435419159A9A7FDD7" box="[579,733,583,609]" name="Indonesia" pageId="1" pageNumber="427">INDONESIA</collectingCountry>
|
||
/
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||
<collectorName id="26B73196FFFEFFA435F79159A8E4FDD7" box="[757,926,583,609]" pageId="1" pageNumber="427">Puncak Palopo</collectorName>
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||
/
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||
<taxonomicName id="4C422FC3FFFEFFA434B49159AF0FFDD7" authority="Selatan" authorityName="Selatan" box="[950,1141,583,609]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="427" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi Selatan</taxonomicName>
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||
/ 30 31. XII, 1999 / N
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||
</materialsCitation>
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||
.
|
||
</paragraph>
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||
<paragraph id="8BFD5440FFFEFFA437959175AF82FD33" blockId="1.[151,1437,474,2013]" box="[151,1272,619,645]" pageId="1" pageNumber="427">
|
||
Ohbayashi leg.”;
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||
<materialsCitation id="3B2A5E1DFFFEFFA436589175AF8EFD33" ID-GBIF-Occurrence="3698900306" box="[346,1268,619,645]" collectingDate="2012-06-23" collectingDateMax="2012-06-25" collectingDateMin="2012-06-23" country="Indonesia" location="Sulawesi" pageId="1" pageNumber="427" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
|
||
<specimenCount id="9D449FC9FFFEFFA436589175AAFCFD33" box="[346,390,619,645]" pageId="1" pageNumber="427" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D449FC9FFFEFFA436909175AAA9FD33" box="[402,467,619,645]" pageId="1" pageNumber="427" type="female">2 ♀♀</specimenCount>
|
||
: same data as holotype.
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||
<typeStatus id="54F9EAE2FFFEFFA435E79175A828FD33" box="[741,850,619,645]" pageId="1" pageNumber="427" type="paratype">Paratypes</typeStatus>
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||
are preserved in the RIEB and EUM
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFEFFA637C59190AB86FF63" lastPageId="3" lastPageNumber="429" pageId="1" pageNumber="427" type="description">
|
||
<paragraph id="8BFD5440FFFEFFA437C59190AF69FD1F" blockId="1.[151,1437,474,2013]" box="[199,1043,654,683]" pageId="1" pageNumber="427">
|
||
<emphasis id="B9368852FFFEFFA437C59190A945FD1D" bold="true" box="[199,575,654,683]" pageId="1" pageNumber="427">
|
||
Description of the
|
||
<typeStatus id="54F9EAE2FFFEFFA436A19191A970FD1F" box="[419,522,655,681]" pageId="1" pageNumber="427" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="9D449FC9FFFEFFA43519918EA94AFD1D" box="[539,560,656,683]" pageId="1" pageNumber="427" type="male">♂</specimenCount>
|
||
).
|
||
</emphasis>
|
||
Body shape ovoid, slightly bent laterally.
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||
</paragraph>
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||
<paragraph id="8BFD5440FFFEFFA437C591ADAFF7FC8F" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">Color almost blackish-brown with a sparkly green gloss; legs black with green vitreous luster; antennomeres 1−5 dark orange, antennal club, maxillary palpi, and labial palpi black; elytra black with four reddish-brown maculae, including a pair of wide, elongated maculae in the anterior 1/4 and encircling scutellar shield, and remaining two maculae in the posterior 1/4 at a near apico-sutural angle; marginal membrane pale brown.</paragraph>
|
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<paragraph id="8BFD5440FFFEFFA437C5905DAFC4FC7F" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">Head distinctly convex, densely and transversely rugoso punctured throughout; clypeus rectangular with rounded angles, weakly reflexed anterior margin, weakly concave at the center; ambiguous front-clypeal suture, depressed frons, and a shallow median groove extending to a vertex that is narrowly concave along the midline; vertex coarsely rugoso punctured, transforming into a concentric arrangement at side and base.</paragraph>
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||
<paragraph id="8BFD5440FFFEFFA437C590CDA849FBCB" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Pronotum approximately 1.4 times as wide as long, sides weakly constricted in the posterior half, then strongly curved and strongly converging in the anterior half; anterior angles acute and protrude strongly; posterior angles rectangular; disc with dense granulose texture and a pair of oblique depressions in the middle to anterior part of the lateral sides, furnished with sparse short appressed yellowish-white scaly hairs (
|
||
<quantity id="4CBAF9A5FFFEFFA433149720AFC1FBEF" box="[1046,1211,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" metricValueMax="1.7" metricValueMin="0.5" pageId="1" pageNumber="427" unit="mm" value="0.11" valueMax="0.17" valueMin="0.05">0.05–0.17 mm</quantity>
|
||
in length) along the anterior to posterior angles; all marginal lines are complete.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C59799AA81FB70" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Scutellar shield rounded triangle, densely rugoso punctured, covered with appressed yellowish-white scaly hairs (
|
||
<quantity id="4CBAF9A5FFFEFFA437DE97B5AAFBFB73" box="[220,385,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="1.85" metricValueMax="2.3" metricValueMin="1.4" pageId="1" pageNumber="427" unit="mm" value="0.185" valueMax="0.23" valueMin="0.14">0.14–0.23 mm</quantity>
|
||
in length).
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C597D1A8A2FAE3" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">Elytra with very ambiguous rows; intervals almost flattened; humeral calli and posterior portion of the scutellar shield slightly depressed; disc with dense granulose texture; sides weakly rounded from anterior to posterior; lateral marginal lines rimmed completely and continuing to elytral apices; posterior margin evenly rounded; marginal membrane narrow, starting from basal 1/4, and extending to elytral apices.</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C59641A87BFA2B" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Propygidium roughly rugoso punctured; disc furnished with dense fringe of appressed yellowish-white scaly hairs (
|
||
<quantity id="4CBAF9A5FFFEFFA437DE969DAAFBFA2B" box="[220,385,1411,1437]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="427" unit="mm" value="0.135" valueMax="0.17" valueMin="0.1">0.10–0.17 mm</quantity>
|
||
in length) along posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C596B9AA62F9BF" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Pygidium weakly convex and apex broadly rounded; disc densely rugoso-reticulately punctured throughout, with dense appressed yellowish-white scaly hairs (
|
||
<quantity id="4CBAF9A5FFFEFFA4359196D4A842FA53" box="[659,824,1482,1509]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="1" pageNumber="427" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.15–0.17 mm</quantity>
|
||
in length), transforming into a concentric arrangement at the apex.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C5950DA90BF9C3" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Mesoventrite roughly rugoso punctured and covered with sparse appressed white setae (
|
||
<quantity id="4CBAF9A5FFFEFFA433D0950DAE01F99B" box="[1234,1403,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="427" unit="mm" value="0.135" valueMax="0.17" valueMin="0.1">0.10–0.17 mm</quantity>
|
||
in length). Mesoventral process short, compressed between mesocoxae, slightly projecting upward in lateral view; apex rounded and rimmed in ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C59561AF8AF957" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Abdominal ventrites covered with rugoso-reticulate punctures, with a sparse transverse band of yellowish-white scaly hair (
|
||
<quantity id="4CBAF9A5FFFEFFA4361395BDAACFF90B" box="[273,437,1699,1725]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="427" unit="mm" value="0.135" valueMax="0.17" valueMin="0.1">0.10–0.17 mm</quantity>
|
||
in length) in the middle, transforming laterally into dense long yellowish-white scaly hair (
|
||
<quantity id="4CBAF9A5FFFEFFA4379D95D9AA3EF957" box="[159,324,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="1" pageNumber="427" unit="mm" value="0.245" valueMax="0.26" valueMin="0.23">0.23–0.26 mm</quantity>
|
||
in length); ventrite 5 approximately as long as combined width of ventrites 3 and 4.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C595F5AF50F823" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Legs with bidentate protibia, proximal tooth very weak and situated close to the apical tooth; pro-, meso-, and metafemur with sparse yellowish-white scaly hair (
|
||
<quantity id="4CBAF9A5FFFEFFA435E79411A8F7F89F" box="[741,909,1807,1833]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="427" unit="mm" value="0.135" valueMax="0.17" valueMin="0.1">0.10–0.17 mm</quantity>
|
||
in length); protarsomere 1−4 almost equal in length; inner protarsal claw strongly curved and incised apically, upper branch spiniform, lower branch broad and obliquely truncated; outer mesotarsal claw incised at apex, upper branch spiniform; metatarsal claws very unequal, with outer claws approximately 1.5 times thicker and 1/3 longer than inner claws.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFEFFA437C59481AFFFF86B" blockId="1.[151,1437,474,2013]" pageId="1" pageNumber="427">
|
||
Aedeagus curved in lateral view, shape simple as shown in
|
||
<figureCitation id="137948C5FFFEFFA4346F9481A88EF80F" box="[877,1012,1951,1977]" captionStart="FIGURES 10–13" captionStartId="4.[151,264,1911,1935]" captionTargetBox="[234,1358,191,1880]" captionTargetId="figure-17@4.[215,1371,181,1887]" captionTargetPageId="4" captionText="FIGURES 10–13. Aedeagus of Malaia (M.) medea Kaneko & Wada, new species (10–11) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="1" pageNumber="427">Figs. 10–11</figureCitation>
|
||
. Parameres almost symmetrical, with each lobe narrowly sinuate at the apex, and multiple sparse long reddish setae at the apex.
|
||
</paragraph>
|
||
<caption id="DF3D04C8FFFDFFA737959590A9C7F8A4" ID-DOI="http://doi.org/10.5281/zenodo.6332096" ID-Zenodo-Dep="6332096" httpUri="https://zenodo.org/record/6332096/files/figure.png" pageId="2" pageNumber="428" startId="2.[151,264,1678,1702]" targetBox="[162,1427,189,1638]" targetPageId="2">
|
||
<paragraph id="8BFD5440FFFDFFA737959590A9C7F8A4" blockId="2.[151,1436,1678,1811]" pageId="2" pageNumber="428">
|
||
<emphasis id="B9368852FFFDFFA737959590AA40F910" bold="true" box="[151,314,1678,1702]" pageId="2" pageNumber="428">FIGURES 1–9.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C422FC3FFFDFFA736AD9590AA82F910" authorityName="Heller" authorityYear="1892" box="[431,504,1678,1702]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="428" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFDFFA736AD9590AA82F910" box="[431,504,1678,1702]" italics="true" pageId="2" pageNumber="428">Malaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C422FC3FFFDFFA736FC9591A85BF910" authority="Kaneko & Wada" authorityName="Kaneko & Wada" authorityYear="2022" box="[510,801,1678,1703]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="428" phylum="Arthropoda" rank="species" species="medea" status="sp. nov." subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFDFFA735049591A961F910" box="[518,539,1679,1702]" italics="true" pageId="2" pageNumber="428">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFDFFA735329590A908F910" box="[560,626,1678,1702]" italics="true" pageId="2" pageNumber="428">medea</emphasis>
|
||
Kaneko & Wada
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2053529FFFDFFA734289591A8D9F910" box="[810,931,1678,1702]" pageId="2" pageNumber="428" rank="species">new species</taxonomicNameLabel>
|
||
(1–6) and
|
||
<emphasis id="B9368852FFFDFFA733129591AF56F910" box="[1040,1068,1679,1702]" italics="true" pageId="2" pageNumber="428">M.</emphasis>
|
||
<taxonomicName id="4C422FC3FFFDFFA733339591AE37F911" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1073,1357,1678,1703]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="428" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFDFFA7333B9591AF34F910" box="[1081,1102,1679,1702]" italics="true" pageId="2" pageNumber="428">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFDFFA733619590AFB0F910" box="[1123,1226,1678,1702]" italics="true" pageId="2" pageNumber="428">simulatrix</emphasis>
|
||
<bibRefCitation id="EFD329B1FFFDFFA733D29590AE37F911" author="Heller, K. M." box="[1232,1357,1678,1703]" pageId="2" pageNumber="428" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
♂ from Pulau Peleng, Sulawesi (7–9). 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, paratype, dorsal view (color variability); 5, allotype, dorsal view; 6, allotype, ventral view; 7, specimen from RIEB, dorsal view; 8, the same, ventral view; 9, the same, lateral view. Scale: 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD5440FFFDFFA737C5945FA811F8C9" blockId="2.[151,1437,1856,2027]" pageId="2" pageNumber="428">
|
||
<emphasis id="B9368852FFFDFFA737C5945FAA5FF8ED" bold="true" box="[199,293,1857,1883]" pageId="2" pageNumber="428">Female.</emphasis>
|
||
Pygidium strongly convex. Abdominal ventrite 5 shorter than combined width of ventrites 3 and 4. Apical tooth of protibia long and spatulate; proximal tooth stout.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFDFFA737C59497AF05F871" blockId="2.[151,1437,1856,2027]" pageId="2" pageNumber="428">
|
||
<emphasis id="B9368852FFFDFFA737C59497AA04F815" bold="true" box="[199,382,1929,1955]" pageId="2" pageNumber="428">Measurements.</emphasis>
|
||
Males (n = 6): total body length
|
||
<quantity id="4CBAF9A5FFFDFFA735E89497A80AF815" box="[746,880,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.9" metricValueMin="6.4" pageId="2" pageNumber="428" unit="mm" value="6.65" valueMax="6.9" valueMin="6.4">6.4–6.9 mm</quantity>
|
||
(
|
||
<typeStatus id="54F9EAE2FFFDFFA7347F9497A89FF815" box="[893,997,1929,1955]" pageId="2" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="4CBAF9A5FFFDFFA734ED9497AF32F815" box="[1007,1096,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="428" unit="mm" value="6.4">6.4 mm</quantity>
|
||
), total body width 3.2–4.0 mm (
|
||
<typeStatus id="54F9EAE2FFFDFFA7379D94B3AA7DF871" box="[159,263,1965,1991]" pageId="2" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="4CBAF9A5FFFDFFA7361394B3AA11F871" box="[273,363,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="428" unit="mm" value="3.2">3.2 mm</quantity>
|
||
). Females (n = 2): total body length 7.3–7.4, total body width 3.8–4.0.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFDFFA637C594CEAB86FF63" blockId="2.[151,1437,1856,2027]" lastBlockId="3.[151,1437,151,861]" lastPageId="3" lastPageNumber="429" pageId="2" pageNumber="428">
|
||
<emphasis id="B9368852FFFDFFA737C594CEAA88F85C" bold="true" box="[199,498,2000,2026]" pageId="2" pageNumber="428">Morphological variation.</emphasis>
|
||
Elytra color varies from dark blue to spotted reddish-brown on black. The reddishbrown maculae present as four maculae or two elongated maculae formed by connecting anterior and posterior maculae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA637C593C0A9EEFDAF" pageId="3" pageNumber="429" type="diagnosis">
|
||
<paragraph id="8BFD5440FFFCFFA637C593C0A9EEFDAF" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C593C0AAB5FF4E" bold="true" box="[199,463,222,248]" pageId="3" pageNumber="429">Differential diagnosis.</emphasis>
|
||
The newly described species is similar to
|
||
<taxonomicName id="4C422FC3FFFCFFA634B193C1AFFEFF4F" authorityName="Heller" authorityYear="1892" box="[947,1156,223,249]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA634B193C1A8B0FF4E" box="[947,970,223,248]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA634E193C1A880FF4E" box="[995,1018,223,248]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA6331093C1AFFEFF4F" box="[1042,1156,223,249]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137948C5FFFCFFA6339793C1AE7BFF4F" box="[1173,1281,223,249]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1678,1702]" captionTargetBox="[162,1427,189,1638]" captionTargetId="figure-17@2.[151,1436,179,1652]" captionTargetPageId="2" captionText="FIGURES 1–9. Habitus of Malaia (M.) medea Kaneko & Wada, new species (1–6) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (7–9). 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, paratype, dorsal view (color variability); 5, allotype, dorsal view; 6, allotype, ventral view; 7, specimen from RIEB, dorsal view; 8, the same, ventral view; 9, the same, lateral view. Scale: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332096" httpUri="https://zenodo.org/record/6332096/files/figure.png" pageId="3" pageNumber="429">Figs. 7–9</figureCitation>
|
||
,
|
||
<figureCitation id="137948C5FFFCFFA6320C93C1AE2CFF4F" box="[1294,1366,223,249]" captionStart="FIGURES 10–13" captionStartId="4.[151,264,1911,1935]" captionTargetBox="[234,1358,191,1880]" captionTargetId="figure-17@4.[215,1371,181,1887]" captionTargetPageId="4" captionText="FIGURES 10–13. Aedeagus of Malaia (M.) medea Kaneko & Wada, new species (10–11) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="3" pageNumber="429">12–13</figureCitation>
|
||
) from Bangkai,
|
||
<collectingCountry id="F35514D0FFFCFFA63605921DAA02FEAB" box="[263,376,259,285]" name="Indonesia" pageId="3" pageNumber="429">Indonesia</collectingCountry>
|
||
. However, this species is readily distinguishable from
|
||
<taxonomicName id="4C422FC3FFFCFFA634FB921DAFAAFEAB" authorityName="Heller" authorityYear="1892" box="[1017,1232,259,285]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA634FB921DAF6AFEAA" box="[1017,1040,259,284]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA6332E921DAF39FEAA" box="[1068,1091,259,284]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA6335C921DAFAAFEAB" box="[1118,1232,259,285]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: 1) sparkly luster of pronotum, with strong oblique impression on each side.
|
||
<taxonomicName id="4C422FC3FFFCFFA633989239AF91FEF7" authorityName="Heller" authorityYear="1892" box="[1178,1259,295,321]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA633989239AF91FEF7" box="[1178,1259,295,321]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C422FC3FFFCFFA633F09239AEE7FEF7" authorityName="Heller" authorityYear="1892" box="[1266,1437,295,321]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFCFFA633FE9239AE69FEF6" box="[1276,1299,295,320]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA632299239AEE7FEF7" box="[1323,1437,295,321]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
displays less sparkly luster, with weaker oblique impression; 2) protibial proximal tooth very weak, whereas, in
|
||
<taxonomicName id="4C422FC3FFFCFFA637959271AA10FE3F" authorityName="Heller" authorityYear="1892" box="[151,362,367,393]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA637959271ABD4FE3E" box="[151,174,367,392]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA637CA9271ABA5FE3E" box="[200,223,367,392]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA637FA9271AA10FE3F" box="[248,362,367,393]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
, it is well developed; 3) mesotibia are slender (length: width = 4.5:1) as compared to
|
||
<taxonomicName id="4C422FC3FFFCFFA632469271AA73FE1B" authorityName="Heller" authorityYear="1892" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA632469271AE21FE3E" box="[1348,1371,367,392]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA632779271AEF6FE3E" box="[1397,1420,367,392]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA63795928DAA73FE1B" box="[151,265,403,429]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
(length: width = 3.5:1); 4) metatibia are slender (length: width = 3.8:1) as compared to
|
||
<taxonomicName id="4C422FC3FFFCFFA633CD928DAEE7FE1B" authorityName="Heller" authorityYear="1892" box="[1231,1437,403,429]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA633CD928DAF9CFE1A" box="[1231,1254,403,428]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA633FF928DAE6EFE1A" box="[1277,1300,403,428]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA63229928DAEE7FE1B" box="[1323,1437,403,429]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
(length: width = 3:1); 5) width of abdominal ventrites
|
||
<quantity id="4CBAF9A5FFFCFFA635F092A9A843FE67" box="[754,825,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="3" pageNumber="429" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
|
||
the male is shorter than those of
|
||
<taxonomicName id="4C422FC3FFFCFFA633AB92A9AE0DFE67" authorityName="Heller" authorityYear="1892" box="[1193,1399,439,465]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA633AB92A9AFBAFE66" box="[1193,1216,439,464]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA633D592A9AF94FE66" box="[1239,1262,439,464]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA6320792A9AE0DFE67" box="[1285,1399,439,465]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
; 6) abdominal ventrite hairs are very sparse; 7) development of the mesoventral process is weak compared to
|
||
<taxonomicName id="4C422FC3FFFCFFA6324792C5AA73FDAF" authorityName="Heller" authorityYear="1892" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA6324792C5AE26FE42" box="[1349,1372,475,500]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA6327792C5AEF6FE42" box="[1397,1420,475,500]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA6379592E1AA73FDAF" box="[151,265,511,537]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
; 8) aedeagus is distinctly different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA637C5913DAE31FD33" pageId="3" pageNumber="429" type="etymology">
|
||
<paragraph id="8BFD5440FFFCFFA637C5913DAE31FD33" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C5913DAA34FD8B" bold="true" box="[199,334,547,573]" pageId="3" pageNumber="429">Etymology.</emphasis>
|
||
The new species is named after “Medea”, a character in Greek mythology. In Latin, “Simulatrix” is synonymous with “Circe”, who is also a character in Greek mythology, suggested to be a sister or aunt to Medea. The present species and
|
||
<taxonomicName id="4C422FC3FFFCFFA636AB9175A902FD33" authorityName="Heller" authorityYear="1892" box="[425,632,619,645]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA636AB9175AABAFD32" box="[425,448,619,644]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA636D59175AA8CFD32" box="[471,502,619,644]" italics="true" pageId="3" pageNumber="429">M.</emphasis>
|
||
)
|
||
<emphasis id="B9368852FFFCFFA635049175A902FD33" box="[518,632,619,645]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
are closely similar, so we named it
|
||
<emphasis id="B9368852FFFCFFA6330B9175AF5AFD32" box="[1033,1056,619,644]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.
|
||
<taxonomicName id="4C422FC3FFFCFFA6332C9175AFD5FD33" authority="Kaneko & Wada, 2022" authorityName="Kaneko & Wada" authorityYear="2022" box="[1070,1199,619,645]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="medea" status="sp. nov." subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFCFFA633359175AF34FD32" box="[1079,1102,619,644]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA633649175AFD5FD33" box="[1126,1199,619,645]" italics="true" pageId="3" pageNumber="429">medea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2053529FFFCFFA633BF9175AE3DFD33" box="[1213,1351,619,645]" pageId="3" pageNumber="429" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA637C59190A852FD1C" box="[199,808,654,682]" pageId="3" pageNumber="429" type="distribution">
|
||
<paragraph id="8BFD5440FFFCFFA637C59190A852FD1C" blockId="3.[151,1437,151,861]" box="[199,808,654,682]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C59190AA1BFD1E" bold="true" box="[199,353,654,680]" pageId="3" pageNumber="429">Distribution.</emphasis>
|
||
<collectingCountry id="F35514D0FFFCFFA636659191AAA2FD1F" box="[359,472,655,681]" name="Indonesia" pageId="3" pageNumber="429">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49869AA2FFFCFFA636E19191A932FD1F" box="[483,584,655,681]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">Sulawesi</collectingRegion>
|
||
(
|
||
<taxonomicName id="4C422FC3FFFCFFA635559191A866FD1C" authority="Selatan" authorityName="Selatan" box="[599,796,655,682]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi Selatan</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA637C591ADA835FCEB" pageId="3" pageNumber="429" type="discussion">
|
||
<paragraph id="8BFD5440FFFCFFA637C591ADA835FCEB" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C591ADAA41FD7B" bold="true" box="[199,315,691,717]" pageId="3" pageNumber="429">Remarks.</emphasis>
|
||
The present species has the following characteristics of the subgenus
|
||
<taxonomicName id="4C422FC3FFFCFFA6334291ADAFEBFD7B" authorityName="Heller" authorityYear="1892" box="[1088,1169,691,717]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA6334291ADAFEBFD7B" box="[1088,1169,691,717]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
: 1) the posterior margin of the pronotum is rounded in an arch or truncated anterior to the scutellar shield; 2) in lateral view, the semicylindrical mesoventral process protrudes from the mesocoxa forward, but not downward. Thus, we distinguished this species as belonging to the subgenus
|
||
<taxonomicName id="4C422FC3FFFCFFA635649001A9CDFC8F" authorityName="Heller" authorityYear="1892" box="[614,695,799,825]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA635649001A9CDFC8F" box="[614,695,799,825]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
. However, we consider the above characteristics to be unstable. In the future, the definition of subgenera may need to be revised.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |