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<document id="4FB44209D8516E2B438B6935D84FC9C1" ID-DOI="10.5281/zenodo.13244881" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244881" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970250919" checkinUser="felipe" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2004" docId="03D687B7A73AFFB039AAFDD8BD5F4D41" docLanguage="en" docName="RafflesBZool.52.1.97-145.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Copelatus minutissimus Balfour-Browne 1939" docType="treatment" docVersion="2" lastPageNumber="118" masterDocId="FFEFFFCFA72FFFA53903FF87BF1E4871" masterDocTitle="Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #" masterLastPageNumber="145" masterPageNumber="97" pageNumber="118" updateTime="1722975545229" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6CFFD26F3A80B2789B8F6E0D7326F668" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8BF73ED30FF8ACB89FFE029D1F8EF0B6">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="6B9CB939208101FF731A437B253DDDA1">Hendrich, Lars</mods:namePart>
</mods:name>
<mods:name id="347E8B75637377535116367DF23D2ABC" type="personal">
<mods:role id="D4815232EB4BD3BEE70246AA3D7EFA43">
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<mods:namePart id="92C00F2FD6818CC03B7143F06BB7B314">Balke, Michael</mods:namePart>
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<mods:title id="0D0EA41B74585F232CE18F9C7B2443CA">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="BEB354FBB85BAFA20B7D3C86938B549A">
<mods:date id="D6BAC45BE7C8CE9628A920EB76EF2758">2004</mods:date>
<mods:detail id="88EE9BCBFD44CB4F28B86C0AB14E1494" type="volume">
<mods:number id="42502EF5B61031F49C4FEC6201737F85">52</mods:number>
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<mods:detail id="810B2191A7CD4B2EB39D306B76C97233" type="issue">
<mods:number id="9AFA820B871292F9658ED9F88BF098FD">1</mods:number>
</mods:detail>
<mods:extent id="B4C9A6936C45E5561C49628E3F9A0DDA" unit="page">
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<mods:classification id="FAAB5A33FEF62A0A65B6FF4F0203593C">journal article</mods:classification>
<mods:identifier id="8A0A11095B5E3828D970C81D8CA78EBB" type="DOI">10.5281/zenodo.13244881</mods:identifier>
<mods:identifier id="5BDC84443BFCB3983AEC77E4467B883D" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="69057DC70F26A737E83221D7C3E124B2" type="Zenodo-Dep">13244881</mods:identifier>
</mods:mods>
<treatment id="03D687B7A73AFFB039AAFDD8BD5F4D41" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB039AAFDD8BD5F4D41" httpUri="http://treatment.plazi.org/id/03D687B7A73AFFB039AAFDD8BD5F4D41" lastPageNumber="118" pageId="21" pageNumber="118">
<subSubSection id="C365652AA73AFFB039AAFDD8BDA74A06" box="[169,697,607,631]" pageId="21" pageNumber="118" type="nomenclature">
<paragraph id="8BC036A1A73AFFB039AAFDD8BDA74A06" blockId="21.[169,697,607,663]" box="[169,697,607,631]" pageId="21" pageNumber="118">
<heading id="D08881CDA73AFFB039AAFDD8BDA74A06" bold="true" box="[169,697,607,631]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="118" reason="2">
<taxonomicName id="4C7F4D22A73AFFB039AAFDD8BDA74A06" ID-CoL="5ZWWL" authority="Balfour-Browne, 1939" authorityName="Balfour-Browne" authorityYear="1939" box="[169,697,607,631]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B90BEAB3A73AFFB039AAFDD8BDA74A06" bold="true" box="[169,697,607,631]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB039AAFDD8BEAD4A06" bold="true" box="[169,435,607,631]" italics="true" pageId="21" pageNumber="118">Copelatus minutissimus</emphasis>
Balfour-Browne, 1939
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03882FD07BEFF4AE6" box="[385,481,640,663]" pageId="21" pageNumber="118" type="description">
<paragraph id="8BC036A1A73AFFB03882FD07BEFF4AE6" blockId="21.[169,697,607,663]" box="[385,481,640,663]" pageId="21" pageNumber="118">
(
<figureCitation id="13442A24A73AFFB0388AFD07BEC74AE6" box="[393,473,640,663]" captionStart="Figs" captionStartId="15.[113,158,2006,2027]" captionTargetBox="[253,1312,191,1946]" captionTargetPageId="15" captionText="Figs. 18-23. 18.Hydaticus fabricii (MacLeay, 1825) (Fam. Dytiscidae, 10.5 mm); 19. Leiodytes nicobaricus (Redtenbacher, 1867) (Fam. Dytiscidae, 2.1 mm); 20. Hydrovatus rufoniger rufoniger (Clark, 1863) (Fam. Dytiscidae, 3.6 mm); 21. Laccophilus medialis Sharp, 1882 (Fam. Dytiscidae, 3.3 mm); 22. Copelatus minutissimus Balfour-Browne, 1939 (Fam. Dytiscidae, 4.1 mm); 23. Eretes griseus (Fabricius, 1781) (Fam. Dytiscidae, 13.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13244912" httpUri="https://zenodo.org/record/13244912/files/figure.png" pageId="21" pageNumber="118">Fig. 22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03972FD3BBEE54B7B" pageId="21" pageNumber="118" type="reference_group">
<paragraph id="8BC036A1A73AFFB03972FD3BBEE54B7B" blockId="21.[113,753,700,778]" pageId="21" pageNumber="118">
<treatmentCitationGroup id="AB6F118FA73AFFB03972FD3BBEE54B7B" pageId="21" pageNumber="118">
<treatmentCitation id="0ADE10B0A73AFFB03972FD3BBD734AA0" author="Balfour-Browne, J" box="[113,621,700,721]" page="79" pageId="21" pageNumber="118" year="1939">
<taxonomicName id="4C7F4D22A73AFFB03972FD3BBD734AA0" ID-CoL="5ZWWL" authority="Balfour-Browne, 1939 a: 79" authorityName="Balfour-Browne" authorityPageNumber="79" authorityYear="1939" box="[113,621,700,721]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B90BEAB3A73AFFB03972FD3BBE474AA0" box="[113,345,700,721]" italics="true" pageId="21" pageNumber="118">Copelatus minutissimus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB0385CFD3BBD734AA0" author="Balfour-Browne, J" box="[351,621,700,721]" pageId="21" pageNumber="118" pagination="57 - 88" refId="ref35364" refString="Balfour-Browne, J., 1939 a. On Copelatus Erichson and Leiopterus Stephens (Col. Dytiscidae) with description of new species. Transactions of the Royal Entomological Society of London, 88 (2): 57 - 88." type="journal article" year="1939">Balfour-Browne, 1939a: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03B74FD3BBFF34A9F" author="Gueorguiev, V" page="22" pageId="21" pageNumber="118" year="1968">
<bibRefCitation id="EFEE4B50A73AFFB03B74FD3BBFF34A9F" author="Gueorguiev, V" pageId="21" pageNumber="118" pagination="5 - 45" refId="ref37343" refString="Gueorguiev, V., 1968. Essai de classification des Coleopteres Dytiscidae, Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia, 28: 5 - 45." type="journal article" year="1968">Guéorguiev, 1968: 22</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB039F6FD5EBEAD4A9F" author="Vazirani, T" box="[245,435,729,750]" page="315" pageId="21" pageNumber="118" year="1970">
<bibRefCitation id="EFEE4B50A73AFFB039F6FD5EBEAD4A9F" author="Vazirani, T" box="[245,435,729,750]" pageId="21" pageNumber="118" pagination="93 - 129" refId="ref41247" refString="Vazirani, T. G., 1970. Contribution to the study of aquatic beetles (Coleoptera) V. A review of Hydroporinae: Dytiscidae in part, from India. Oriental Insects, 4 (1): 93 - 129." type="journal article" year="1970">Vazirani, 1970: 315</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB038B8FD5EBD0F4A9F" author="Vazirani, T" box="[443,529,729,750]" page="56" pageId="21" pageNumber="118" year="1977">
<bibRefCitation id="EFEE4B50A73AFFB038B8FD5EBD0F4A9F" author="Vazirani, T" box="[443,529,729,750]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">1977: 56</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03B1AFD5EBDAF4A9F" author="Balke, M" box="[537,689,729,750]" page="32" pageId="21" pageNumber="118" year="1994">
<bibRefCitation id="EFEE4B50A73AFFB03B1AFD5EBDAF4A9F" author="Balke, M" box="[537,689,729,750]" pageId="21" pageNumber="118" pagination="32 - 34" refId="ref35508" refString="Balke, M., 1994. Regenwald in der Stadt. Okowerkmagazin, Berlin 8 (12): 32 - 34." type="journal article" year="1994">Balke, 1994: 32</bibRefCitation>
</treatmentCitation>
; Balke et al., 1999: 325;
<treatmentCitation id="0ADE10B0A73AFFB03846FD72BEE84B7B" author="Nilsson, A" box="[325,502,757,778]" page="70" pageId="21" pageNumber="118" year="2001">
<bibRefCitation id="EFEE4B50A73AFFB03846FD72BEE84B7B" author="Nilsson, A" box="[325,502,757,778]" pageId="21" pageNumber="118" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 70</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03972FCA8BD124BA5" pageId="21" pageNumber="118" type="materials_examined">
<paragraph id="8BC036A1A73AFFB03972FCA8BD124BA5" blockId="21.[113,753,815,980]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972FCA8BE214B35" bold="true" box="[113,319,815,836]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972FCA8BE374B35" bold="true" box="[113,297,815,836]" italics="true" pageId="21" pageNumber="118">Material examined</emphasis>
. –
</emphasis>
<typeStatus id="54C48803A73AFFB03840FCA8BE6A4B35" box="[323,372,815,836]" pageId="21" pageNumber="118">Type</typeStatus>
material:
<materialsCitation id="3B173CFCA73AFFB038D2FCA8BDF04B35" box="[465,750,815,836]" collectionCode="BMNH" country="Singapore" location="Singapore" pageId="21" pageNumber="118" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D79FD28A73AFFB038D2FCA8BD0D4B35" box="[465,531,815,836]" count="1" pageId="21" pageNumber="118" type="male">1 male</specimenCount>
: “
<collectingCountry id="F3687631A73AFFB03B20FCA8BD994B35" box="[547,647,815,836]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
” (
<collectionCode id="ED6EAE64A73AFFB03B99FCA8BDF94B35" box="[666,743,815,836]" pageId="21" pageNumber="118">BMNH</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54C48803A73AFFB03972FCCBBE284B10" box="[113,310,844,865]" isOtherTypeStatus="true" pageId="21" pageNumber="118">Additional material</typeStatus>
:
<specimenCount id="9D79FD28A73AFFB03842FCCBBEA54B10" box="[321,443,844,865]" count="2" pageId="21" pageNumber="118" type="generic">2 specimens</specimenCount>
-
<emphasis id="B90BEAB3A73AFFB038D1FCCBBDD24B10" bold="true" box="[466,716,844,865]" pageId="21" pageNumber="118">Central catchment area</emphasis>
-
<materialsCitation id="3B173CFCA73AFFB03BE7FCCBBFAA4BC6" collectingDate="1992-05-22" collectionCode="ZRC" collectorName="Range II Training Shed &amp; H. K. Lua &amp; K. Lim" country="Singapore" location="Nee Soon Swamp Forest" municipality="Transect" pageId="21" pageNumber="118" specimenCount="1" typeStatus="Additional material">
<specimenCount id="9D79FD28A73AFFB03BE7FCCBBF8E4B0F" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB0399BFCEEBE9C4B0F" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB039AAFDD8BD5F4D41:8EA0607AA73AFFB0399BFCEEBE9C4B0F" box="[152,386,873,894]" country="Singapore" municipality="Transect" name="Nee Soon Swamp Forest" pageId="21" pageNumber="118">Nee Soon Swamp Forest</location>
,
<collectorName id="268A5377A73AFFB03889FCEEBD764B0F" box="[394,616,873,894]" pageId="21" pageNumber="118">Range II Training Shed</collectorName>
,
<collectingMunicipality id="6BA4ACDBA73AFFB03B73FCEEBDDC4B0F" box="[624,706,873,894]" pageId="21" pageNumber="118">Transect</collectingMunicipality>
“E”,
<collectingMethod id="523E4EB6A73AFFB03972FC02BE184BEB" box="[113,262,901,922]" pageId="21" pageNumber="118">UV Light Trap</collectingMethod>
,
<date id="FFC11061A73AFFB03812FC02BE884BEB" box="[273,406,901,922]" pageId="21" pageNumber="118" value="1992-05-22">
<collectingDate id="EF85E989A73AFFB03812FC02BE884BEB" box="[273,406,901,922]" pageId="21" pageNumber="118" value="1992-05-22">22 May.1992</collectingDate>
</date>
, NS 10d, coll.
<collectorName id="268A5377A73AFFB03B2BFC02BD944BEB" box="[552,650,901,922]" pageId="21" pageNumber="118">H. K. Lua</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03BA9FC02BDEE4BEB" box="[682,752,901,922]" pageId="21" pageNumber="118">K. Lim</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB0397BFC25BFB24BC6" box="[120,172,930,951]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB039BDFC25BD164BA5" collectingDate="1993-08-26" collectionCode="ZRC" collectorName="M. Balke &amp; Murphy &amp; C. M. Yang" country="Singapore" location="Nee Soon Swamp Forest" pageId="21" pageNumber="118" specimenCount="1" typeStatus="Additional material">
<specimenCount id="9D79FD28A73AFFB039BDFC25BFF14BC6" box="[190,239,930,951]" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB039FAFC25BEF04BC6" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB039AAFDD8BD5F4D41:8EA0607AA73AFFB039FAFC25BEF04BC6" box="[249,494,930,951]" country="Singapore" name="Nee Soon Swamp Forest" pageId="21" pageNumber="118">Nee Soon Swamp Forest</location>
,
<date id="FFC11061A73AFFB038F4FC25BD664BC6" box="[503,632,930,951]" pageId="21" pageNumber="118" value="1993-08-26">
<collectingDate id="EF85E989A73AFFB038F4FC25BD664BC6" box="[503,632,930,951]" pageId="21" pageNumber="118" value="1993-08-26">26 Aug.1993</collectingDate>
</date>
, y 840, coll.
<collectorName id="268A5377A73AFFB03972FC38BFD14BA5" box="[113,207,959,980]" pageId="21" pageNumber="118">M. Balke</collectorName>
,
<collectorName id="268A5377A73AFFB039D9FC38BE364BA5" box="[218,296,959,980]" pageId="21" pageNumber="118">Murphy</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03849FC38BEA14BA5" box="[330,447,959,980]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB038CFFC38BD1F4BA5" box="[460,513,959,980]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03972FC7EBFA94CE1" pageId="21" pageNumber="118" type="distribution">
<paragraph id="8BC036A1A73AFFB03972FC7EBFA94CE1" blockId="21.[113,753,1017,1168]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972FC7EBE154C60" bold="true" box="[113,267,1017,1041]" italics="true" pageId="21" pageNumber="118">Distribution. –</emphasis>
<collectingCountry id="F3687631A73AFFB03812FC7EBE594C61" box="[273,327,1017,1040]" name="India" pageId="21" pageNumber="118">India</collectingCountry>
[
<collectingRegion id="49BBF843A73AFFB03852FC7EBE874C61" box="[337,409,1017,1040]" country="India" name="Odisha" pageId="21" pageNumber="118">Orissa</collectingRegion>
] needs to be confirmed,
<collectingCountry id="F3687631A73AFFB03B92FC7EBDEF4C61" box="[657,753,1017,1040]" name="Malaysia" pageId="21" pageNumber="118">Malaysia</collectingCountry>
(
<bibRefCitation id="EFEE4B50A73AFFB0397AFB9EBE074C41" author="Vazirani, T" box="[121,281,1049,1072]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977</bibRefCitation>
),
<collectingCountry id="F3687631A73AFFB03828FB9EBE884C41" box="[299,406,1049,1072]" name="Indonesia" pageId="21" pageNumber="118">Indonesia</collectingCountry>
:
<collectingRegion id="49BBF843A73AFFB0389CFB9EBD004C41" box="[415,542,1049,1072]" country="Indonesia" name="Kalimantan Utara" pageId="21" pageNumber="118">Kalimantan</collectingRegion>
,
<collectingRegion id="49BBF843A73AFFB03B24FB9EBD9E4C41" box="[551,640,1049,1072]" country="Indonesia" name="Sumatera Utara" pageId="21" pageNumber="118">Sumatra</collectingRegion>
(Hendrich vid.),
<collectingCountry id="F3687631A73AFFB039B2FBBEBE0F4C21" box="[177,273,1081,1104]" name="Thailand" pageId="21" pageNumber="118">Thailand</collectingCountry>
(
<bibRefCitation id="EFEE4B50A73AFFB0381CFBBEBD3A4C21" author="Balfour-Browne, J" box="[287,548,1081,1104]" pageId="21" pageNumber="118" pagination="57 - 88" refId="ref35364" refString="Balfour-Browne, J., 1939 a. On Copelatus Erichson and Leiopterus Stephens (Col. Dytiscidae) with description of new species. Transactions of the Royal Entomological Society of London, 88 (2): 57 - 88." type="journal article" year="1939">Balfour-Browne, 1939a</bibRefCitation>
),
<collectingCountry id="F3687631A73AFFB03B3BFBBEBD8B4C21" box="[568,661,1081,1104]" name="Vietnam" pageId="21" pageNumber="118">Vietnam</collectingCountry>
(in coll. Balke &amp; Hendrich) and
<collectingCountry id="F3687631A73AFFB0386FFBDEBEC64C01" box="[364,472,1113,1136]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
(
<bibRefCitation id="EFEE4B50A73AFFB038E6FBDEBD744C01" author="Balke, M" box="[485,618,1113,1136]" pageId="21" pageNumber="118" pagination="32 - 34" refId="ref35508" refString="Balke, M., 1994. Regenwald in der Stadt. Okowerkmagazin, Berlin 8 (12): 32 - 34." type="journal article" year="1994">Balke, 1994</bibRefCitation>
; Balke et al., 1999).
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03972FB3EBD5F4D41" pageId="21" pageNumber="118" type="biology_ecology">
<paragraph id="8BC036A1A73AFFB03972FB3EBD5F4D41" blockId="21.[113,752,1209,1328]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972FB3EBFF84CA0" bold="true" box="[113,230,1209,1233]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972FB3EBFD54CA0" bold="true" box="[113,203,1209,1233]" italics="true" pageId="21" pageNumber="118">Ecology</emphasis>
. –
</emphasis>
A forest species which occurs among leaf litter, roots and submerged plants of very small, temporary, shallow puddles in or near forested areas. This species is confined to the forests of the Central Catchment Area.
</paragraph>
</subSubSection>
</treatment>
</document>
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