153 lines
19 KiB
XML
153 lines
19 KiB
XML
<document id="E52F69DBD22B1C8A0FEA51E63E45F0C5" ID-CLB-Dataset="27497" ID-DOI="10.11646/zootaxa.4550.1.6" ID-GBIF-Dataset="264af183-e8ff-4d6f-9b42-b18f800af618" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625193" ID-ZooBank="7086C72D-9C8C-48D8-8228-F990D964D839" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554276780989" checkinUser="plazi" docAuthor="Bechly, Günter & Rasmussen, Jan Audun" docDate="2019" docId="03B9D118FFFDC02629E8FA58FBCE011B" docLanguage="en" docName="zootaxa.4550.1.6.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parabaissaeshna ejerslevense Bechly & Rasmussen 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="125" masterDocId="FF80A960FFFCC024297FFF87FFE60543" masterDocTitle="A new genus of hawker dragonfly (Odonata: Anisoptera: Aeshnidae) from the Early Eocene Fur Formation of Denmark" masterLastPageNumber="128" masterPageNumber="123" pageNumber="124" updateTime="1698716869983" updateUser="ExternalLinkService">
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<mods:title id="5ECCAB54B5E775B2A37E2273D248E72D">A new genus of hawker dragonfly (Odonata: Anisoptera: Aeshnidae) from the Early Eocene Fur Formation of Denmark</mods:title>
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<mods:namePart id="F9D8F87958E7A037342700111B7F9A05">Bechly, Günter</mods:namePart>
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<mods:name id="53BD3E3034D6F8D9FE07C9C4FF347B0A" type="personal">
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<mods:roleTerm id="D93324FD1C6F61D046B1A786FB5BB4ED">Author</mods:roleTerm>
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<mods:namePart id="17C997314682980AF18860C945D25D6C">Rasmussen, Jan Audun</mods:namePart>
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<mods:typeOfResource id="2CF0C77E70929FB6DDE5D68EA1708ABF">text</mods:typeOfResource>
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<mods:title id="E581CB499848E7F205F2E092B7F0B5D0">Zootaxa</mods:title>
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<mods:part id="9F8B4DE0846A407AF51856F88F0F8FF6">
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<mods:date id="18DF3C88761F7CD13ED51A6BA51D1AFF">2019</mods:date>
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<mods:number id="5433071DB85C6A0BD4999E71905D0D44">2019-01-23</mods:number>
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<mods:number id="B189C386BAEB687E000EAB06B73D3276">4550</mods:number>
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<mods:identifier id="E77C190D48998D7395ED93AE6B81C44B" type="DOI">10.11646/zootaxa.4550.1.6</mods:identifier>
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<mods:identifier id="41B205E81F029F9BE03ABBD5DBF72533" type="GBIF-Dataset">264af183-e8ff-4d6f-9b42-b18f800af618</mods:identifier>
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<treatment id="03B9D118FFFDC02629E8FA58FBCE011B" ID-DOI="http://doi.org/10.5281/zenodo.5927220" ID-GBIF-Taxon="154534327" ID-Zenodo-Dep="5927220" LSID="urn:lsid:plazi:treatment:03B9D118FFFDC02629E8FA58FBCE011B" httpUri="http://treatment.plazi.org/id/03B9D118FFFDC02629E8FA58FBCE011B" lastPageId="2" lastPageNumber="125" pageId="1" pageNumber="124">
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<subSubSection id="C30A3385FFFDC02529E8FA58FD8700B9" box="[151,609,1503,1530]" pageId="1" pageNumber="124" type="nomenclature">
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<paragraph id="8BAF600EFFFDC02529E8FA58FD8700B9" blockId="1.[151,609,1503,1563]" box="[151,609,1503,1530]" pageId="1" pageNumber="124">
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<heading id="D0E7D762FFFDC02529E8FA58FD8700B9" bold="true" box="[151,609,1503,1530]" fontSize="11" level="1" pageId="1" pageNumber="124" reason="1">
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<emphasis id="B964BC1CFFFDC02529E8FA58FD8700B9" bold="true" box="[151,609,1503,1530]" pageId="1" pageNumber="124">
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<taxonomicName id="4C101B8DFFFDC02529E8FA58FE1A00BA" authority="Bechly & Rasmussen, 2019" authorityName="Bechly & Rasmussen" authorityYear="2019" box="[151,508,1503,1529]" class="Insecta" family="Aeshnidae" genus="Parabaissaeshna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="ejerslevense" status="sp. nov.">
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<emphasis id="B964BC1CFFFDC02529E8FA58FE1A00BA" bold="true" box="[151,508,1503,1529]" italics="true" pageId="1" pageNumber="124">Parabaissaeshna ejerslevense</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2570167FFFDC0252B7CFA67FD8700B9" box="[515,609,1504,1530]" pageId="1" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A3385FFFDC02529E8F985FEFA0358" box="[151,284,1538,1563]" pageId="1" pageNumber="124" type="description">
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<paragraph id="8BAF600EFFFDC02529E8F985FEFA0358" blockId="1.[151,609,1503,1563]" box="[151,284,1538,1563]" pageId="1" pageNumber="124">
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<figureCitation id="132B7C8BFFFDC02529E8F985FEFA0358" box="[151,284,1538,1563]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1621,1643]" captionStartId-1="3.[151,250,616,638]" captionTargetBox-0="[151,1437,1167,1596]" captionTargetBox-1="[205,1392,229,546]" captionTargetId-0="figure@2.[151,1437,1167,1596]" captionTargetId-1="figure@3.[151,1436,193,595]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Photograph of holotype no. MGUH 33076 of Parabaissaeshna ejerslevense gen. et sp. nov. Scale bar = 10 mm." captionText-1="FIGURE 2. Drawing of holotype no. MGUH 33076 of Parabaissaeshna ejerslevense gen. et sp. nov. Scale bar = 10 mm." httpUri-0="https://zenodo.org/record/2625195/files/figure.png" httpUri-1="https://zenodo.org/record/2625197/files/figure.png" pageId="1" pageNumber="124">Figures 1–2</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30A3385FFFDC02529E8F9CDFC7802AC" pageId="1" pageNumber="124" type="materials_examined">
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<paragraph id="8BAF600EFFFDC02529E8F9CDFDE0038C" blockId="1.[151,1437,1610,2031]" pageId="1" pageNumber="124">
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<materialsCitation id="3B786A53FFFDC02529E8F9CDFDE0038C" ID-GBIF-Occurrence="2234226774" collectionCode="MGUH" collectorName="Frank Pedersen" country="Denmark" location="University of Copenhagen" municipality="Copenhagen in Copenhagen" pageId="1" pageNumber="124" specimenCode="MGUH 33076" specimenCount="1" typeStatus="holotype">
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<emphasis id="B964BC1CFFFDC02529E8F9CDFEED0320" bold="true" box="[151,267,1610,1635]" pageId="1" pageNumber="124">
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<typeStatus id="54ABDEACFFFDC02529E8F9CDFEE00320" box="[151,262,1610,1635]" pageId="1" pageNumber="124" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Isolated complete forewing of a dragonfly with collection no.
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<specimenCode id="DBB6C875FFFDC0252A8FF9CCFB7F0320" box="[1008,1177,1610,1635]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="1" pageNumber="124">MGUH 33076</specimenCode>
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, deposited in the type collection of the Natural History Museum of
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<collectingCountry id="F307209EFFFDC0252BEDF9E9FD1B03C4" box="[658,765,1646,1671]" name="Denmark" pageId="1" pageNumber="124">Denmark</collectingCountry>
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,
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<location id="8ECF36D5FFFDC0252A77F9E9FBD603C4" LSID="urn:lsid:plazi:treatment:03B9D118FFFDC02629E8FA58FBCE011B:8ECF36D5FFFDC0252A77F9E9FBD603C4" box="[776,1072,1646,1671]" country="Denmark" municipality="Copenhagen in Copenhagen" name="University of Copenhagen" pageId="1" pageNumber="124">
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University of
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<collectingMunicipality id="6BCBFA74FFFDC0252ADBF9E9FBD603C4" box="[932,1072,1646,1671]" pageId="1" pageNumber="124">Copenhagen</collectingMunicipality>
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</location>
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in Copenhagen (
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<collectingCountry id="F307209EFFFDC0252D90F9E9FAB803C4" box="[1263,1374,1646,1671]" name="Denmark" pageId="1" pageNumber="124">Denmark</collectingCountry>
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). The specimen was discovered by the amateur fossil collectors
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<collectorName id="26E505D8FFFDC0252A5EF915FC2803E8" box="[801,974,1682,1707]" pageId="1" pageNumber="124">Frank Pedersen</collectorName>
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and Viggo Pedersen, Mors, and declared as Danekrae fossil trove DK 792.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BAF600EFFFDC02529B8F95DFC1A0278" blockId="1.[151,1437,1610,2031]" pageId="1" pageNumber="124">
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<emphasis id="B964BC1CFFFDC02529B8F95DFDE503B0" bold="true" box="[199,515,1754,1779]" pageId="1" pageNumber="124">Type locality and horizon.</emphasis>
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Mo-clay quarry near the village Ejerslev (
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<geoCoordinate id="EE2406C9FFFDC0252A95F95DFBB003B1" box="[1002,1110,1754,1779]" direction="north" orientation="latitude" pageId="1" pageNumber="124" precision="555" value="56.93">56.93° N</geoCoordinate>
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,
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<geoCoordinate id="EE2406C9FFFDC0252D1DF95CFB5D03B0" box="[1122,1211,1755,1779]" direction="east" orientation="longitude" pageId="1" pageNumber="124" precision="555" value="8.91">8.91° E</geoCoordinate>
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), northern island of Mors, western Limfjord area, N. Jutland,
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<collectingCountry id="F307209EFFFDC0252B17F979FD320254" box="[616,724,1790,1815]" name="Denmark" pageId="1" pageNumber="124">Denmark</collectingCountry>
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. Calcareous concretion approximately
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<quantity id="4CE8CDEBFFFDC0252DF6F978FB520255" box="[1161,1204,1791,1815]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="124" unit="m" value="2.0">2 m</quantity>
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above Ash layer -
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<quantity id="4CE8CDEBFFFDC0252CFEF978FF4B0278" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="124" unit="in" value="11.0">11 in</quantity>
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the diatomites of the upper part of the Knudeklint Member, Fur Formation.
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</paragraph>
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<paragraph id="8BAF600EFFFDC02529B8F8C1FC7802AC" blockId="1.[151,1437,1610,2031]" pageId="1" pageNumber="124">
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<emphasis id="B964BC1CFFFDC02529B8F8C1FE83021C" bold="true" box="[199,357,1862,1887]" pageId="1" pageNumber="124">Stratigraphy.</emphasis>
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Dinoflagellate Zone 7 (corresponding to the acme of
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<taxonomicName id="4C101B8DFFFDC0252AB9F8C1FB12021D" box="[966,1268,1862,1886]" class="Dinophyceae" family="Peridiniaceae" genus="Deflandrea" higherTaxonomySource="GBIF" kingdom="Chromista" order="Peridiniales" pageId="1" pageNumber="124" phylum="Myzozoa" rank="species" species="oesbisfeldensis">
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<emphasis id="B964BC1CFFFDC0252AB9F8C1FB12021D" box="[966,1268,1862,1886]" italics="true" pageId="1" pageNumber="124">Deflandrea oesbisfeldensis</emphasis>
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</taxonomicName>
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) of Heilmann-
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<bibRefCitation id="EF811DFFFFFDC02529E8F8EDFEA002C0" author="Heilmann-Clausen, C." box="[151,326,1898,1923]" pageId="1" pageNumber="124" refId="ref3300" refString="Heilmann-Clausen, C. (1985) Dinoflagellate stratigraphy of the uppermost Danian to Ypresian in the Viborg 1 borehole, central Jylland, Denmark. Danmarks Geologiske Undersogelse, Serie A 7, 69 pp." type="book" year="1985">Clausen (1985)</bibRefCitation>
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, earliest Ypresian, Early Eocene, Paleogene, ca. 56.0-54.5 million years. The Early Eocene age of the Fur Formation was recently established due to the presence of the Paleocene-Eocene Thermal Maximum (PETM) in the Stolleklint clay of the Ølst Formation, which is placed stratigraphically immediately below the Fur Formation (
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<bibRefCitation id="EF811DFFFFFDC0252865F851FD5602AC" author="Heilmann-Clausen, C. & Schmitz, B." box="[282,688,2006,2031]" pageId="1" pageNumber="124" pagination="70" refId="ref3337" refString="Heilmann-Clausen, C. & Schmitz, B. (2000) The late Paleocene thermal maximum δ 13 C excursion in Denmark? GFF, 122 (1), 70. https: // doi. org / 10.1080 / 11035890001221070" type="journal article" year="2000">Heilmann-Clausen & Schmitz 2000</bibRefCitation>
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;
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<bibRefCitation id="EF811DFFFFFDC0252BC4F851FC7402AC" author="Schoon, P. L. & Heilmann-Clausen, C. & Schultz B. P. & Sluijs, A. & Sinninghe Damste, J. S. & Schouten, S." box="[699,914,2006,2031]" pageId="1" pageNumber="124" pagination="160 - 168" refId="ref4153" refString="Schoon, P. L., Heilmann-Clausen, C., Schultz B. P., Sluijs, A., Sinninghe Damste, J. S. & Schouten, S. (2013) Recognition of Early Eocene global carbon isotope excursions using lipids of marine Thaumarchaeota. Earth and Planetary Science Letters, 373, 160 - 168. https: // doi. org / 10.1016 / j. epsl. 2013.04.037" type="journal article" year="2013">
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Schoon
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<emphasis id="B964BC1CFFFDC0252A69F850FCA802AD" box="[790,846,2006,2030]" italics="true" pageId="1" pageNumber="124">et al.</emphasis>
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2013
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A3385FFFEC02629B8FF10FC1205F3" box="[199,1012,151,176]" pageId="2" pageNumber="125" type="etymology">
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<paragraph id="8BAF600EFFFEC02629B8FF10FC1205F3" blockId="2.[151,1437,151,1112]" box="[199,1012,151,176]" pageId="2" pageNumber="125">
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<emphasis id="B964BC1CFFFEC02629B8FF10FEA805F3" bold="true" box="[199,334,151,176]" pageId="2" pageNumber="125">Etymology.</emphasis>
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A noun in the genitive case, named after the place of origin.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A3385FFFEC02629B8FF3BFD520596" box="[199,692,188,213]" pageId="2" pageNumber="125" type="diagnosis">
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<paragraph id="8BAF600EFFFEC02629B8FF3BFD520596" blockId="2.[151,1437,151,1112]" box="[199,692,188,213]" pageId="2" pageNumber="125">
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<emphasis id="B964BC1CFFFEC02629B8FF3BFEA70596" bold="true" box="[199,321,188,213]" pageId="2" pageNumber="125">Diagnosis.</emphasis>
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Same as genus since monotypic.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A3385FFFEC02629B8FF58FBCE011B" pageId="2" pageNumber="125" type="description">
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||
<paragraph id="8BAF600EFFFEC02629B8FF58FBCE011B" blockId="2.[151,1437,151,1112]" pageId="2" pageNumber="125">
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||
<emphasis id="B964BC1CFFFEC02629B8FF58FD6E05BB" bold="true" box="[199,648,223,248]" pageId="2" pageNumber="125">
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||
Description of
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<typeStatus id="54ABDEACFFFEC0262809FF58FE3B05BB" box="[374,477,223,248]" pageId="2" pageNumber="125" type="holotype">holotype</typeStatus>
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(
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<figureCitation id="132B7C8BFFFEC0262894FF58FD9F05BB" box="[491,633,223,248]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1621,1643]" captionStartId-1="3.[151,250,616,638]" captionTargetBox-0="[151,1437,1167,1596]" captionTargetBox-1="[205,1392,229,546]" captionTargetId-0="figure@2.[151,1437,1167,1596]" captionTargetId-1="figure@3.[151,1436,193,595]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Photograph of holotype no. MGUH 33076 of Parabaissaeshna ejerslevense gen. et sp. nov. Scale bar = 10 mm." captionText-1="FIGURE 2. Drawing of holotype no. MGUH 33076 of Parabaissaeshna ejerslevense gen. et sp. nov. Scale bar = 10 mm." httpUri-0="https://zenodo.org/record/2625195/files/figure.png" httpUri-1="https://zenodo.org/record/2625197/files/figure.png" pageId="2" pageNumber="125">Figures 1–2</figureCitation>
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).
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</emphasis>
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Well-preserved and complete isolated forewing of a hawker dragonfly of unknown sex. Total length 45.0 mm, max. width
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<quantity id="4CE8CDEBFFFEC0262A79FE83FC94045F" box="[774,882,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="125" unit="mm" value="10.4">10.4 mm</quantity>
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||
. 4 secondary antenodal crossveins between the primary antenodal crossveins Ax1 and Ax2; 22 and 18 non-aligned secondary antenodal crossveins present distal of Ax
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<quantity id="4CE8CDEBFFFEC026299FFECBFEF70426" box="[224,273,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="125" unit="in" value="2.0">2 in</quantity>
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||
both rows respectively. Space between RA and RP basal of subnodus regularly crossed by 18 antesubnodal crossveins (no cordulegastrid gap of antesubnodals near subnodus). Nodus at 48.6% of wing length; nodal veinlet short and perpendicular; subnodal veinlet oblique. About 14 postnodal crossveins non-aligned with the 19 postsubnodal crossveins beneath them; no ‘libellulid gap’ of postsubnodal crossveins near subnodus. Pterostigma elongate, elongate (covering 6 cells), with oblique sides and broadened margins; oblique and strong stigmal brace vein aligned with anterior margin of pterostigma; about six crossveins between C and RA distal of pterostigma. Primary IR1 not developed; secondary IR1 originating beneath middle of pterostigma with 3 rows of cells between it and RP1, and three rows of cells between it and RP2; RP1 and RP2 basally parallel (2 rows of cells between them) and distally divergent with 3-9 rows of cells between them; RP2 originates at subnodus; RP2 undulating; only one lestine oblique vein between RP2 and IR2, a single cell distal of subnodus. IR2 originating on RP1/2, and not forked; space between RP2 and IR2 distally widened with 2-7 (mostly four) rows of cells between them. Rspl well defined, parallel to IR2 with two rows of cells between them; about
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<specimenCount id="9D16AB87FFFEC0262D34FD50FB3E07B3" box="[1099,1240,727,752]" pageId="2" pageNumber="125" type="intercalary">5 intercalary</specimenCount>
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veins originating from Rspl. Midfork six cells distal of triangle; 13 antefurcal crossveins between RP and MA. RP3/4 and MA weakly undulate and parallel, with two rows of cells between them distally; no bulged ‘aeshnid bulla’ between MA and RP3/4. Posttrigonal area between MA and MP with two rows of cells near the triangle; trigonal planate well defined, originating at a slight kink on the distal side (MAb) of triangle; Mspl well defined, parallel to MA, with two rows of cells between them. Space between MP and CuA narrow; MP and CuA reaching far beyond level of nodus. CuA with 7-8 posterior branches; cubital field max. 5 cells in width. Anal field with two rows of cells. Arculus complete (closed) and broken (kinked), situated midway between Ax1 and Ax2. Hypertriangle long and narrow, divided by 4 crossveins. Discoidal triangle elongate and divided into seven cells. Subtriangle divided into 2 cells. Between CuP-crossing (= ‘anal crossing’) and basal subdiscoidal vein there are two crossveins. Basal space free. Wing hyaline without trace of color pattern, except for the dark pterostigma.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |