765 lines
129 KiB
XML
765 lines
129 KiB
XML
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<mods:title id="C8C3F056F77F9B98324A2E44F4924695">A new species of the genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura: Paguridae) from a semi-submerged marine cave in Okinawa Island, southwestern Japan</mods:title>
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<mods:namePart id="40201475898397470F9499A2AADA2930">Nakajima, Hiroki</mods:namePart>
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<mods:affiliation id="50C52690212ADC2AB740A4D7866DB3BD">Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903 - 0213, Japan.</mods:affiliation>
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<mods:nameIdentifier id="A65E20F7377933D811DAA208E3383CB7" type="email">sawagani.is.kani@gmail.com</mods:nameIdentifier>
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<mods:namePart id="636A2329B154C20F3267E1B10D1821E8">Fujita, Yoshihisa</mods:namePart>
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<mods:affiliation id="94A452685D2C53089A9AB478DC6DB0F0">Okinawa Prefectual University of Arts, 1 - 4 Shuri-Tounokura, Naha, Okinawa 903 - 8602, Japan.</mods:affiliation>
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<mods:namePart id="E911BFC1310A7B8FDDFB6994693EE12D">Osawa, Masayuki</mods:namePart>
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<mods:affiliation id="C8376F2F38414D097A3D1F73FE41E089">Estuary Research Center, Shimane University, 1060 Nishikawatsu-cho, Matsue, Shimane, 690 - 8504 Japan.</mods:affiliation>
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<mods:title id="6AAB4763F33402D590CC4E89B87EE4FF">Zootaxa</mods:title>
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<mods:date id="C4AF825498492BA978FB7CEB13F9F533">2024</mods:date>
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<treatment id="03A4047A8634FFA348EAFDC7FB97FA49" ID-DOI="http://doi.org/10.5281/zenodo.10787808" ID-Zenodo-Dep="10787808" LSID="urn:lsid:plazi:treatment:03A4047A8634FFA348EAFDC7FB97FA49" httpUri="http://treatment.plazi.org/id/03A4047A8634FFA348EAFDC7FB97FA49" lastPageId="7" lastPageNumber="128" pageId="1" pageNumber="122">
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<heading id="D0FA02008634FFA548EAFDC7FD93FDB5" bold="true" box="[151,588,546,573]" fontSize="11" level="1" pageId="1" pageNumber="122" reason="1">
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<taxonomicName id="4C0DCEEF8634FFA548EAFDC7FDDDFDB5" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[151,514,546,573]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
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<emphasis id="B979697E8634FFA548EAFDC7FDDDFDB5" bold="true" box="[151,514,546,573]" italics="true" pageId="1" pageNumber="122">Trichopagurus spinibrachium</emphasis>
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<taxonomicNameLabel id="A24AD4058634FFA54A77FDC6FD93FDB5" box="[522,588,547,573]" pageId="1" pageNumber="122" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BB2B56C8634FFA548EAFDA2FD30FDE9" blockId="1.[151,751,546,645]" box="[151,751,583,609]" pageId="1" pageNumber="122">[New Japanese name: Doukutsu-yawakuda-yadokari]</paragraph>
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<subSubSection id="C317E6E78634FFA548EAFD8EFECDFD0D" box="[151,274,619,645]" pageId="1" pageNumber="122" type="description">
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<paragraph id="8BB2B56C8634FFA548EAFD8EFECDFD0D" blockId="1.[151,751,546,645]" box="[151,274,619,645]" pageId="1" pageNumber="122">
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(
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<figureCitation id="1336A9E98634FFA548E2FD8EFED5FD0D" box="[159,266,619,645]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1869,1893]" captionStartId-1="2.[151,250,1938,1962]" captionStartId-2="4.[151,250,1873,1897]" captionStartId-3="5.[151,250,1922,1946]" captionTargetBox-0="[159,1428,821,1840]" captionTargetBox-1="[172,1413,389,1907]" captionTargetBox-2="[159,1428,317,1837]" captionTargetBox-3="[151,1436,273,1898]" captionTargetId-0="figure-121@1.[151,1436,813,1845]" captionTargetId-1="figure-102@2.[164,1423,381,1915]" captionTargetId-2="figure-92@4.[151,1436,309,1849]" captionTargetId-3="figure-57@5.[151,1436,273,1898]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." captionText-1="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." captionText-2="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." captionText-3="FIGURE 4. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664, entire animal.A, anterior view, size not measured; B, right lateral view; C, dorsal view; D, ventral view. A, photographed while alive; B–D, photographed after freezing. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10782046" figureDoi-1="http://doi.org/10.5281/zenodo.10782048" figureDoi-2="http://doi.org/10.5281/zenodo.10782050" figureDoi-3="http://doi.org/10.5281/zenodo.10782054" httpUri-0="https://zenodo.org/record/10782046/files/figure.png" httpUri-1="https://zenodo.org/record/10782048/files/figure.png" httpUri-2="https://zenodo.org/record/10782050/files/figure.png" httpUri-3="https://zenodo.org/record/10782054/files/figure.png" pageId="1" pageNumber="122">Figs. 1–4</figureCitation>
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)
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</paragraph>
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<subSubSection id="C317E6E78634FFA548EAFD56FEFEFC9D" pageId="1" pageNumber="122" type="materials_examined">
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<paragraph id="8BB2B56C8634FFA548EAFD56FEFEFC9D" blockId="1.[151,1436,690,789]" pageId="1" pageNumber="122">
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<emphasis id="B979697E8634FFA548EAFD56FE98FD44" bold="true" box="[151,327,690,717]" pageId="1" pageNumber="122">Type material.</emphasis>
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<materialsCitation id="3B65BF318634FFA5492DFD56FEFEFC9D" collectingDate="2023-07-02" collectionCode="T" collectorName="H. Nakajima & Sato, M & Y. Fujita" county="Onna Village" elevation="1" location="Ryukyu Islands" municipality="Island" pageId="1" pageNumber="122" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
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<typeStatus id="54B60BCE8634FFA5492DFD56FE62FD45" box="[336,445,691,717]" pageId="1" pageNumber="122" type="holotype">Holotype</typeStatus>
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: RUMF-ZC-7664, male (sl
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<quantity id="4CF518898634FFA54A80FD56FC86FD45" box="[765,857,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="122" unit="mm" value="1.4">1.4 mm</quantity>
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), a semi-submerged marine cave on
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<collectingCounty id="62D3CDE08634FFA54C82FD56FA47FD45" box="[1279,1432,691,717]" pageId="1" pageNumber="122">Onna Village</collectingCounty>
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,
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<collectingRegion id="49C97B8E8634FFA548EAFD32FF23FD79" box="[151,252,727,753]" country="Japan" name="Okinawa" pageId="1" pageNumber="122">Okinawa</collectingRegion>
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<collectingMunicipality id="6BD62F168634FFA5497EFD32FE95FD79" box="[259,330,727,753]" pageId="1" pageNumber="122">Island</collectingMunicipality>
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,
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<location id="8ED2E3B78634FFA5492BFD32FDD8FD79" LSID="urn:lsid:plazi:treatment:03A4047A8634FFA348EAFDC7FB97FA49:8ED2E3B78634FFA5492BFD32FDD8FD79" box="[342,519,727,753]" county="Onna Village" municipality="Island" name="Ryukyu Islands" pageId="1" pageNumber="122" stateProvince="Okinawa">Ryukyu Islands</location>
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, about
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<elevation id="0020525F8634FFA54A25FD32FD14FD79" box="[600,715,727,753]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="122" unit="m" value="1.0">
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<quantity id="4CF518898634FFA54A25FD32FD5CFD79" box="[600,643,727,753]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="122" unit="m" value="1.0">1 m</quantity>
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depth
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</elevation>
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, collected by
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<collectorName id="26F8D0BA8634FFA54B1BFD32FC26FD79" box="[870,1017,727,753]" pageId="1" pageNumber="122">H. Nakajima</collectorName>
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,
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<collectionCode id="ED1C2DA98634FFA54C7EFD32FBCAFD79" box="[1027,1045,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="122">T</collectionCode>
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.
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<collectorName id="26F8D0BA8634FFA54C5DFD32FBA3FD79" box="[1056,1148,727,753]" pageId="1" pageNumber="122">Sato, M</collectorName>
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. Mizuyama and
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<collectorName id="26F8D0BA8634FFA54D48FD32FA47FD79" box="[1333,1432,727,753]" pageId="1" pageNumber="122">Y. Fujita</collectorName>
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,
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<date id="FFB393AC8634FFA548EAFD1EFEC1FC9D" box="[151,286,763,789]" pageId="1" pageNumber="122" value="2023-07-02">
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<collectingDate id="EFF76A448634FFA548EAFD1EFEC1FC9D" box="[151,286,763,789]" pageId="1" pageNumber="122" value="2023-07-02">2 July 2023</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<caption id="DF72E5E48634FFA548EAF8A8FE75F87D" ID-DOI="http://doi.org/10.5281/zenodo.10782046" ID-Zenodo-Dep="10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="1" pageNumber="122" startId="1.[151,250,1869,1893]" targetBox="[159,1428,821,1840]" targetPageId="1" targetType="figure">
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<paragraph id="8BB2B56C8634FFA548EAF8A8FE75F87D" blockId="1.[151,1437,1869,2037]" pageId="1" pageNumber="122">
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<emphasis id="B979697E8634FFA548EAF8A8FEC6F8ED" bold="true" box="[151,281,1869,1893]" pageId="1" pageNumber="122">FIGURE 1.</emphasis>
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<taxonomicName id="4C0DCEEF8634FFA54958F8A8FD86F8ED" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[293,601,1869,1893]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
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<emphasis id="B979697E8634FFA54958F8A8FD86F8ED" box="[293,601,1869,1893]" italics="true" pageId="1" pageNumber="122">Trichopagurus spinibrachium</emphasis>
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</taxonomicName>
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<emphasis id="B979697E8634FFA54A18F8A8FD7BF8ED" bold="true" box="[613,676,1869,1893]" pageId="1" pageNumber="122">
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<taxonomicNameLabel id="A24AD4058634FFA54A18F8A8FD7BF8ED" box="[613,676,1869,1893]" pageId="1" pageNumber="122" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
, holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm.
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||
</paragraph>
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</caption>
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<subSubSection id="C317E6E78637FFA248BAFF73FF07FD45" lastPageId="6" lastPageNumber="127" pageId="2" pageNumber="123" type="description">
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<paragraph id="8BB2B56C8637FFA648BAFF73FD15FF5D" blockId="2.[151,1437,150,357]" pageId="2" pageNumber="123">
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<emphasis id="B979697E8637FFA648BAFF73FE86FF38" bold="true" box="[199,345,150,176]" pageId="2" pageNumber="123">Description.</emphasis>
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Eleven pairs of biserial gills, including 1 pleurobranch above pereopod 4 (thoracomere 7) and 2 arthrobranchs on each maxilliped 3 to pereopod 4.
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</paragraph>
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||
<paragraph id="8BB2B56C8637FFA648BAFF3AFD1EFEED" blockId="2.[151,1437,150,357]" pageId="2" pageNumber="123">
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Shield (
|
||
<figureCitation id="1336A9E98637FFA64963FF3AFEAAFF71" box="[286,373,223,249]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="2" pageNumber="123">Fig. 1A</figureCitation>
|
||
) 1.06 times longer than wide; anterior margin between rostrum and lateral projections slightly concave; anterolateral margins sloping; posterior margin rounded; dorsal surface polished, convex, with some short setae anteriorly. Rostrum triangular, slightly falls short of proximal half of ocular acicles, terminating in spinule. Lateral projections with small submarginal spine.
|
||
</paragraph>
|
||
<caption id="DF72E5E48637FFA648EAF877FDE4F879" ID-DOI="http://doi.org/10.5281/zenodo.10782048" ID-Zenodo-Dep="10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="2" pageNumber="123" startId="2.[151,250,1938,1962]" targetBox="[172,1413,389,1907]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BB2B56C8637FFA648EAF877FDE4F879" blockId="2.[151,1437,1938,2034]" pageId="2" pageNumber="123">
|
||
<emphasis id="B979697E8637FFA648EAF877FECDF822" bold="true" box="[151,274,1938,1962]" pageId="2" pageNumber="123">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0DCEEF8637FFA64965F877FD9AF822" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[280,581,1938,1962]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
|
||
<emphasis id="B979697E8637FFA64965F877FD9AF822" box="[280,581,1938,1962]" italics="true" pageId="2" pageNumber="123">Trichopagurus spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8637FFA64A37F877FD5CF822" bold="true" box="[586,643,1938,1962]" pageId="2" pageNumber="123">
|
||
<taxonomicNameLabel id="A24AD4058637FFA64A37F877FD5CF822" box="[586,643,1938,1962]" pageId="2" pageNumber="123" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped. A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2B56C8636FFA748BAFF72FE3CFE95" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Ocular peduncles (including cornea) (
|
||
<figureCitation id="1336A9E98636FFA74A10FF72FD1CFF39" box="[621,707,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="3" pageNumber="124">Fig. 1A</figureCitation>
|
||
) stout, cylindrical, 0.47 length of shield; dorsal surface with tufts of stiff setae near base of cornea; lateral margin somewhat inflated medially; cornea not dilated, rounded, corneal width less than half length of peduncle. Ocular acicles subtriangular, moderately widely separated, each with small submarginal spine terminally.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA748BAFEC2FC32FE25" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Antennular peduncles (
|
||
<figureCitation id="1336A9E98636FFA749B6FEC2FDFDFEC9" box="[459,546,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="3" pageNumber="124">Fig. 1A</figureCitation>
|
||
) moderately long, when fully extended, overreaching distal corneal margins by half length of article 2. Article 3 2.2 times as long as high, widened distally in lateral view, with tuft of long setae distally and few short setae on dorsal surface. Article 2 without setae. Article 1 with lateral margin distally produced into small spine; statocyst lobe slightly inflated, with spinule on lateral face.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA748BAFE52FBA1FD0D" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Antennal peduncles (
|
||
<figureCitation id="1336A9E98636FFA749C5FE52FDCEFE59" box="[440,529,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="3" pageNumber="124">Fig. 1A</figureCitation>
|
||
) overreaching distal corneal margins by nearly full length of article 5. Articles 4 and 5 subcylindrical, with few setae. Article 3 with spinule at mesiodistal margin. Article 2 dorsolateral angle produced, nearly reaching distal margin of article 4, terminating in simple spine, bearing few setae; dorsomesial distal angle with acute spine. Article 1 with small distal spine on lateral surface. Antennal acicle reaching midlength of article 5, overreaching anterior margin of cornea by half length, terminating in acutely or bluntly. Antennal flagellum 3.7 times length of shield; each article with few short to moderately long setae.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA748BAFD6AFD50FCB1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Mouthparts not dissected. Maxilliped 3 (
|
||
<figureCitation id="1336A9E98636FFA74AD6FD6AFCF1FD21" box="[683,814,655,681]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="3" pageNumber="124">Fig. 1D, E</figureCitation>
|
||
) moderately slender. Dactylus, propodus and carpus unarmed, with numerous setae on ventral margins. Merus with strong dorsodistal spine and small ventromedian spine; surfaces with spares setae. Ischium with well-developed crista dentata composed of very small corneous teeth and 2 accessory teeth; ventral surface with sparse setae. Basis with single spinule distally on mesial margin. Exopod slightly overreaching distal margin of merus.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA748BAFCA6FADCFAF1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Chelipeds strongly unequal and dissimilar. Right cheliped (
|
||
<figureCitation id="1336A9E98636FFA74B08FCA6FC2FFCD5" box="[885,1008,835,861]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="3" pageNumber="124">Fig. 2A–E</figureCitation>
|
||
) elongate, moderately slender. Chela suboval in dorsal view, 2.45 times as long as wide (widest on proximal part of palm). Dactylus 0.53 length of palm, only slightly curved ventrally, terminating in small corneous claw; dorsal surface smooth, convex; dorsomesial margin not delimited; all surfaces with scattered short setae; occlusal margin with 2 large, triangular calcareous teeth medially. Palm slightly shorter than carpus, entirely smooth, with sparse setae (longer on ventral surface); lateral margin weakly convex; dorsal surface gently convex from side to side, dorsomesial and dorsolateral margins not delimited; ventral surface moderately convex. Fixed finger with scattered short setae on dorsal and ventral surfaces, terminating in small, blunt calcareous claw; occlusal margin with 2 distinct obtuse teeth, median tooth much larger. Carpus flattened dorsoventrally, medially widened in dorsal view, distinctly longer than merus; dorsomesial margin distinctly convex and delimited with 6 irregular-sized spines on proximal two-thirds; dorsolateral margin delimited with 3 spines on median one-third, spines increasing in size distally and larger than dorsomesial spines; ventrolateral margin with short distal spine; dorsal surface gently convex, with scattered short setae; ventral surface weakly convex in lateral and mesial views, with sparse short and long setae. Merus smooth, unarmed on dorsal surface; ventromesial margin with 4 spines, distal 3 spines subequal in size and much larger than proximalmost spine; ventrolateral margin somewhat sinuous, with 7 spines subequal in size; ventral surface concave, with sparse long setae. Ischium unarmed, dorsodistally with tuft of plumose setae. Coxa with small ventrodistal spine.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA748BAFA66FD6AF8A1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="124">
|
||
Left cheliped (
|
||
<figureCitation id="1336A9E98636FFA74915FA66FE3EFA15" box="[360,481,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="3" pageNumber="124">Fig. 3A–D</figureCitation>
|
||
) slender, reaching half length of right chela. Chela 3.52 times longer than wide (widest on bases of fingers). Dactylus 1.20 times longer than palm, terminating in small corneous claw; dorsomesial margin rounded; surfaces nearly smooth, with scattered short setae; occlusal margin with row of small corneous teeth. Palm about half as long as carpus; dorsal surface slightly convex, dorsomesial margin not delimited, dorsolateral margin weakly delimited on proximal one-fourth; mesial and lateral surfaces smooth, only with sparse setae. Fixed finger with sparse setae on surfaces, terminating in small corneous claw; occlusal margin with row of small corneous teeth. Carpus longer than merus; dorsal surface flattish, dorsomesial margin with 5 small spines on proximal two-thirds, dorsolateral margin with 4 spines including distal spine, spines stronger than dorsomesial spines; ventromesial margin with blunt, small distal spine; surfaces with sparse setae, ventral setae much longer. Merus with 3 and 4 slender spines on ventromesial and ventrolateral margins, respectively; mesial and lateral surfaces with sparse setae distally and ventrally; ventral surface with few long setae. Ischium with small distal spine on mesial surface and plumose setae on dorsal surface. Coxa unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8636FFA048BAF8D6FAE2FE95" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124">
|
||
Ambulatory legs (
|
||
<figureCitation id="1336A9E98636FFA749EEF8D6FDD5F8C5" box="[403,522,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="3" pageNumber="124">Fig. 3E–H</figureCitation>
|
||
) moderately slender, sparsely setose. Dactyli 8.3 (pereopod 2) and 9.0 (pereopod 3) times longer than high, 0.72 (pereopod 2) and 0.79 (pereopod 3) length of respective propodus, in dorsal view straight, in lateral view nearly straight with slightly curved distal claw; dorsal and ventral margins each with irregular row of setae; lateral and mesial faces non-sulcate; ventral margins each with 9 (pereopod 2) and 10 (pereopod 3) slender corneous spines. Propodi with sparse, short and moderately long setae on surfaces; dorsal surfaces unarmed; ventral surface with 4 (pereopod 2) and 3 (pereopod 3) slender corneous spines, distal margin with pair of slender corneous spines. Carpi with 2 tiny proximal spines (pereopod 2) and unarmed (pereopod 3) on dorsal surfaces. Meri somewhat compressed laterally; dorsal margins with sparse setae including some bristle-like setae; ventrolateral margin with 3 small spines on median one-third (pereopod 2) and unarmed (pereopod 3). Ischia unarmed.
|
||
</paragraph>
|
||
<caption id="DF72E5E48631FFA048EAF8B4FB17F838" ID-DOI="http://doi.org/10.5281/zenodo.10782050" ID-Zenodo-Dep="10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="4" pageNumber="125" startId="4.[151,250,1873,1897]" targetBox="[159,1428,317,1837]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BB2B56C8631FFA048EAF8B4FB17F838" blockId="4.[151,1436,1873,1969]" pageId="4" pageNumber="125">
|
||
<emphasis id="B979697E8631FFA048EAF8B4FECBF8E1" bold="true" box="[151,276,1873,1897]" pageId="4" pageNumber="125">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C0DCEEF8631FFA04966F8B4FD95F8E1" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[283,586,1873,1897]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
|
||
<emphasis id="B979697E8631FFA04966F8B4FD95F8E1" box="[283,586,1873,1897]" italics="true" pageId="4" pageNumber="125">Trichopagurus spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8631FFA04A2CF8B4FD54F8E1" bold="true" box="[593,651,1873,1897]" pageId="4" pageNumber="125">
|
||
<taxonomicNameLabel id="A24AD4058631FFA04A2CF8B4FD54F8E1" box="[593,651,1873,1897]" pageId="4" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2B56C8630FFA148BAFF72FE83FF71" blockId="5.[151,1437,151,249]" pageId="5" pageNumber="126">
|
||
Pereopod 4 (
|
||
<figureCitation id="1336A9E98630FFA1492CFF72FE7BFF39" box="[337,420,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="5" pageNumber="126">Fig. 1B</figureCitation>
|
||
) semichelate. Dactylus nearly straight, with tufts of setae, distal setae much longer. Propodal rasp consisting of single row of small corneous scales. Carpus with distal tuft of long setae; lateral surface ventrally with sparse setae.
|
||
</paragraph>
|
||
<caption id="DF72E5E48630FFA148EAF867FE6EF869" ID-DOI="http://doi.org/10.5281/zenodo.10782054" ID-Zenodo-Dep="10782054" httpUri="https://zenodo.org/record/10782054/files/figure.png" pageId="5" pageNumber="126" startId="5.[151,250,1922,1946]" targetBox="[151,1436,273,1898]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BB2B56C8630FFA148EAF867FE6EF869" blockId="5.[151,1436,1922,2018]" pageId="5" pageNumber="126">
|
||
<emphasis id="B979697E8630FFA148EAF867FECCF812" bold="true" box="[151,275,1922,1946]" pageId="5" pageNumber="126">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C0DCEEF8630FFA14964F867FD99F812" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[281,582,1922,1946]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
|
||
<emphasis id="B979697E8630FFA14964F867FD99F812" box="[281,582,1922,1946]" italics="true" pageId="5" pageNumber="126">Trichopagurus spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8630FFA14A30F867FD59F812" bold="true" box="[589,646,1922,1946]" pageId="5" pageNumber="126">
|
||
<taxonomicNameLabel id="A24AD4058630FFA14A30F867FD59F812" box="[589,646,1922,1946]" pageId="5" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (sl 1.4 mm), RUMF-ZC-7664, entire animal. A, anterior view, size not measured; B, right lateral view; C, dorsal view; D, ventral view. A, photographed while alive; B–D, photographed after freezing. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2B56C8633FFA248BAFF72FD90FF71" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
Pereopod 5 chelate. Male coxae (
|
||
<figureCitation id="1336A9E98633FFA24A3DFF72FD6BFF39" box="[576,692,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1F, G</figureCitation>
|
||
) slightly asymmetrical; right coxa with short sexual tube directed posterolaterally (about 1.4 times longer than width of right coxa), with distal part filamentous, recurved mesially; left sexual tube very short, papilla-like.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFEE6FD2EFEED" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
Thoracic sternite 3 with anterior margin slightly produced medially, unarmed. Anterior lobe of thoracic sternite 6 (
|
||
<figureCitation id="1336A9E98633FFA248C9FEC2FED6FEC9" box="[180,265,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1C</figureCitation>
|
||
) weakly marked, oblong, with row of sparse short setae on anterior margin. Thoracic sternite 8 (
|
||
<figureCitation id="1336A9E98633FFA24D40FEC2FA4CFEC9" box="[1341,1427,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1G</figureCitation>
|
||
) with pair of obsolete lobes, each with few short setae.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFE8AFDC1FE25" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">Pleon dextrally twisted. Male with 3 unpaired, unequally biramous left pleopods. Uropods markedly asymmetrical; protopods unarmed.</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFE52FD91FD91" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
Telson (
|
||
<figureCitation id="1336A9E98633FFA2495FFE52FEA6FE59" box="[290,377,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1H</figureCitation>
|
||
) with distinct lateral indentations; posterior lobes somewhat asymmetrical, rounded; terminal margins oblique, each with 4 tiny spinules posteriorly; lateral indentations and posterolateral margins with tufts of setae, posterolateral setae much longer.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFDC6FE4EFDB5" blockId="6.[151,1437,150,2013]" box="[199,401,547,573]" pageId="6" pageNumber="127">Female unknown.</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFDA3FF07FD45" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
<emphasis id="B979697E8633FFA248BAFDA3FE8BFDE8" bold="true" box="[199,340,582,608]" pageId="6" pageNumber="127">Color in life</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA2491FFDA2FE7AFDE9" box="[354,421,583,609]" captionStart="FIGURE 4" captionStartId="5.[151,250,1922,1946]" captionTargetBox="[151,1436,273,1898]" captionTargetId="figure-57@5.[151,1436,273,1898]" captionTargetPageId="5" captionText="FIGURE 4. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664, entire animal.A, anterior view, size not measured; B, right lateral view; C, dorsal view; D, ventral view. A, photographed while alive; B–D, photographed after freezing. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782054" httpUri="https://zenodo.org/record/10782054/files/figure.png" pageId="6" pageNumber="127">Fig. 4</figureCitation>
|
||
). Shield and posterior carapace generally white, with scattered pale red blotches (numerous on posterolateral parts). Ocular, antennule and antennal peduncles, chelipeds and ambulatory legs entirely orangish brown in general, paler or yellowish orange on distal parts. Corneas black, with scattered small whitish yellow spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317E6E78633FFA248BAFD33FE7AFC41" pageId="6" pageNumber="127" type="distribution">
|
||
<paragraph id="8BB2B56C8633FFA248BAFD33FA46FD79" blockId="6.[151,1437,150,2013]" box="[199,1433,726,753]" pageId="6" pageNumber="127">
|
||
<emphasis id="B979697E8633FFA248BAFD33FEBEFD78" bold="true" box="[199,353,726,752]" pageId="6" pageNumber="127">Distribution.</emphasis>
|
||
So far known only from the
|
||
<typeStatus id="54B60BCE8633FFA24AD8FD32FD0AFD79" box="[677,725,727,753]" pageId="6" pageNumber="127">type</typeStatus>
|
||
locality,
|
||
<collectingRegion id="49C97B8E8633FFA24B41FD32FC7EFD79" box="[828,929,727,753]" country="Japan" name="Okinawa" pageId="6" pageNumber="127">Okinawa</collectingRegion>
|
||
Island, Ryukyu Islands, southwestern
|
||
<collectingCountry id="F31AF5FC8633FFA24D2CFD32FA4BFD79" box="[1361,1428,727,753]" name="Japan" pageId="6" pageNumber="127">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA248BAFD1FFE7AFC41" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
<emphasis id="B979697E8633FFA248BAFD1FFEF5FC9C" bold="true" box="[199,298,762,788]" pageId="6" pageNumber="127">Habitat.</emphasis>
|
||
The
|
||
<typeStatus id="54B60BCE8633FFA2491BFD1EFE17FC9D" box="[358,456,763,789]" pageId="6" pageNumber="127" type="holotype">holotype</typeStatus>
|
||
was obtained from a semi-submerged marine cave, at
|
||
<quantity id="4CF518898633FFA24C4AFD1FFB44FC9D" box="[1079,1179,762,789]" metricMagnitude="1" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="6" pageNumber="127" unit="m" value="32.5" valueMax="35.0" valueMin="30.0">30–35 m</quantity>
|
||
from the entrance and about
|
||
<quantity id="4CF518898633FFA248A3FCFAFED3FCB1" box="[222,268,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="127" unit="m" value="1.0">1 m</quantity>
|
||
deep. The collection environment is anchialine (slightly lower salinity) and completely darkness. The hermit crab species co-occurring with the present new species include
|
||
<taxonomicName id="4C0DCEEF8633FFA24BD6FCA6FF10FC09" authority="Osawa, Fujita & Okuno, 2006" authorityName="Osawa, Fujita & Okuno" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B979697E8633FFA24BD6FCA6FB5BFCD5" box="[939,1156,835,861]" italics="true" pageId="6" pageNumber="127">Pagurixus longipes</emphasis>
|
||
<bibRefCitation id="EF9CC89D8633FFA24CF7FCA6FF10FC09" author="Osawa, M. & Fujita, Y. & Okuno, J." pageId="6" pageNumber="127" pagination="27 - 45" refId="ref5285" refString="Osawa, M., Fujita, Y. & Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148 (1), 27 - 45. https: // doi. org / 10.11646 / zootaxa. 1148.1.2" type="journal article" year="2006">Osawa, Fujita & Okuno, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C0DCEEF8633FFA2489CFC82FE8AFC09" authorityName="Latreille" authorityYear="1802" box="[225,341,871,897]" class="Malacostraca" family="Paguridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C0DCEEF8633FFA249E4FC82FD21FC09" authority="Morgan, 1992" authorityName="Morgan" authorityYear="1992" box="[409,766,871,897]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="jalur">
|
||
<emphasis id="B979697E8633FFA249E4FC82FD8DFC09" box="[409,594,871,897]" italics="true" pageId="6" pageNumber="127">Paguristes jalur</emphasis>
|
||
<bibRefCitation id="EF9CC89D8633FFA24A21FC82FD21FC09" author="Morgan, G. J." box="[604,766,871,897]" pageId="6" pageNumber="127" pagination="163 - 174" refId="ref5101" refString="Morgan, G. J. (1992). The hermit crabs (Crustacea: Decapoda: Coenobitidae, Diogenidae, Paguridae) of Christmas and Cocos (Keeling) Islands, Indian Ocean, with description of a new species of Paguristes. Raffles Bulletin of Zoology, 49 (2), 163 - 174." type="journal article" year="1992">Morgan, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C0DCEEF8633FFA24B6DFC82FC47FC09" authorityName="Ortmann" authorityYear="1892" box="[784,920,871,897]" class="Malacostraca" family="Diogenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
|
||
). Additionally, a new plagusiid crab species,
|
||
<emphasis id="B979697E8633FFA248EAFC6EFE75FC2D" box="[151,426,907,933]" italics="true" pageId="6" pageNumber="127">Caligoplagusia okinawa</emphasis>
|
||
<bibRefCitation id="EF9CC89D8633FFA249CFFC6EFD6EFC2D" author="Fujita, Y. & Naruse, T." box="[434,689,907,933]" pageId="6" pageNumber="127" pagination="408 - 418" refId="ref4683" refString="Fujita, Y. & Naruse, T. (2024) A new genus and species of a submarine cave crab of the family Plagusiidae Dana, 1851 (Crustacea: Brachyura: Grapsoidea) from Okinawa Island, Ryukyu Islands, southwestern Japan. Zootaxa, 5410 (3), 408 - 418. https: // doi. org / 10.11646 / zootaxa. 5410.3.8" type="journal article" year="2024">Fujita & Naruse, 2024</bibRefCitation>
|
||
, was recently described from the inner most part of the same cave (
|
||
<bibRefCitation id="EF9CC89D8633FFA248E3FC4AFE46FC41" author="Fujita, Y. & Naruse, T." box="[158,409,943,969]" pageId="6" pageNumber="127" pagination="408 - 418" refId="ref4683" refString="Fujita, Y. & Naruse, T. (2024) A new genus and species of a submarine cave crab of the family Plagusiidae Dana, 1851 (Crustacea: Brachyura: Grapsoidea) from Okinawa Island, Ryukyu Islands, southwestern Japan. Zootaxa, 5410 (3), 408 - 418. https: // doi. org / 10.11646 / zootaxa. 5410.3.8" type="journal article" year="2024">Fujita & Naruse 2024</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317E6E78633FFA248BAFC36FE5CFBBD" pageId="6" pageNumber="127" type="etymology">
|
||
<paragraph id="8BB2B56C8633FFA248BAFC36FE5CFBBD" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
<emphasis id="B979697E8633FFA248BAFC36FE91FC65" bold="true" box="[199,334,979,1005]" pageId="6" pageNumber="127">Etymology.</emphasis>
|
||
The specific name is derived from a combination of the Latin,
|
||
<emphasis id="B979697E8633FFA24C54FC36FBB9FC64" box="[1065,1126,979,1004]" italics="true" pageId="6" pageNumber="127">spina</emphasis>
|
||
(= spine) and
|
||
<emphasis id="B979697E8633FFA24D72FC36FAA5FC65" box="[1295,1402,979,1005]" italics="true" pageId="6" pageNumber="127">brachium</emphasis>
|
||
(= arm), referring the possession of strong spines on the carpi and meri of both chelipeds in the new species. Used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317E6E78633FFA348BAFBDAFB97FA49" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127" type="discussion">
|
||
<paragraph id="8BB2B56C8633FFA248BAFBDAFA54FB29" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="127">
|
||
<emphasis id="B979697E8633FFA248BAFBDAFEE4FBD1" bold="true" box="[199,315,1087,1113]" pageId="6" pageNumber="127">Remarks.</emphasis>
|
||
Although only a single male specimen is available for study, the present new species closely matches the generic diagnosis of
|
||
<taxonomicName id="4C0DCEEF8633FFA249DFFB86FD98FBF5" authorityName="de Saint Laurent" authorityYear="1968" box="[418,583,1123,1149]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979697E8633FFA249DFFB86FD98FBF5" box="[418,583,1123,1149]" italics="true" pageId="6" pageNumber="127">Trichopagurus</emphasis>
|
||
</taxonomicName>
|
||
revised by
|
||
<bibRefCitation id="EF9CC89D8633FFA24ABEFB86FC10FBF5" author="Komai, T. & Osawa, M." box="[707,975,1122,1149]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa (2005)</bibRefCitation>
|
||
. Within the genus,
|
||
<taxonomicName id="4C0DCEEF8633FFA24CE3FB86FA85FBF5" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1182,1370,1123,1149]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24CE3FB86FA85FBF5" box="[1182,1370,1123,1149]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24D22FB86FA43FBF5" bold="true" box="[1375,1436,1123,1149]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24D22FB86FA43FBF5" box="[1375,1436,1123,1149]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may be distinguished from all known congeners by the stronger armature of the carpi and meri of both chelipeds.
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8633FFA348BAFB4EFC29FE25" blockId="6.[151,1437,150,2013]" lastBlockId="7.[151,1437,150,1473]" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127">
|
||
<taxonomicName id="4C0DCEEF8633FFA248BAFB4EFDC7FB4D" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[199,536,1195,1221]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="sp. nov.">
|
||
<emphasis id="B979697E8633FFA248BAFB4EFDC7FB4D" box="[199,536,1195,1221]" italics="true" pageId="6" pageNumber="127">Trichopagurus spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24A5FFB4EFDBCFB4D" bold="true" box="[546,611,1195,1221]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24A5FFB4EFDBCFB4D" box="[546,611,1195,1221]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely resembles two congeners,
|
||
<taxonomicName id="4C0DCEEF8633FFA24C45FB4EFB06FB4D" authorityName="Komai & Osawa" authorityYear="2005" box="[1080,1241,1195,1221]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24C45FB4EFB06FB4D" box="[1080,1241,1195,1221]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
known from the south
|
||
<collectingCountry id="F31AF5FC8633FFA248A6FB2AFEC1FB61" box="[219,286,1231,1257]" name="Japan" pageId="6" pageNumber="127">Japan</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C97B8E8633FFA24955FB2AFE8BFB61" box="[296,340,1231,1257]" country="Micronesia" name="Yap" pageId="6" pageNumber="127">Yap</collectingRegion>
|
||
Islands in
|
||
<collectingCountry id="F31AF5FC8633FFA249B2FB2AFD8FFB61" box="[463,592,1231,1257]" name="Micronesia" pageId="6" pageNumber="127">Micronesia</collectingCountry>
|
||
, and the
|
||
<collectingCountry id="F31AF5FC8633FFA24AC8FB2AFCE9FB61" box="[693,822,1231,1257]" name="Philippines" pageId="6" pageNumber="127">Philippines</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C0DCEEF8633FFA24B0CFB2AFBFDFB61" authorityName="Komai" authorityYear="2013" box="[881,1058,1231,1257]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24B0CFB2AFBFDFB61" box="[881,1058,1231,1257]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
known only from the
|
||
<collectingCountry id="F31AF5FC8633FFA24D63FB2AFA43FB61" box="[1310,1436,1231,1257]" name="Philippines" pageId="6" pageNumber="127">Philippines</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA248E2FB16FEEAFA85" author="Komai, T." box="[159,309,1267,1293]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
). Shared characters of the three species are: chelipeds and ambulatory legs only sparsely setose; carpus of right cheliped with spines on dorsomesial margin; palm of right cheliped almost smooth on dorsal surface. However, the new species is distinguished from
|
||
<taxonomicName id="4C0DCEEF8633FFA24AD0FADEFC94FADD" authorityName="Komai & Osawa" authorityYear="2005" box="[685,843,1339,1365]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24AD0FADEFC94FADD" box="[685,843,1339,1365]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24B02FADEFBF0FADD" authorityName="Komai" authorityYear="2013" box="[895,1071,1339,1365]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24B02FADEFBF0FADD" box="[895,1071,1339,1365]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
by the shape and armature of the ocular, antennular and antennal peduncles, chelipeds, and ambulatory legs. The cornea is narrower than the basal width of ocular peduncle in
|
||
<taxonomicName id="4C0DCEEF8633FFA249A9FA66FD4DFA15" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[468,658,1411,1437]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA249A9FA66FD4DFA15" box="[468,658,1411,1437]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24AE4FA66FD07FA15" bold="true" box="[665,728,1411,1437]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24AE4FA66FD07FA15" box="[665,728,1411,1437]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA24A95FA66FC9EFA15" box="[744,833,1411,1437]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1A</figureCitation>
|
||
), rather being slightly wider than or equal in width to the ocular peduncle in
|
||
<taxonomicName id="4C0DCEEF8633FFA249E7FA42FDE6FA49" authorityName="Komai & Osawa" authorityYear="2005" box="[410,569,1447,1473]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA249E7FA42FDE6FA49" box="[410,569,1447,1473]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24A0CFA42FCFCFA49" authorityName="Komai" authorityYear="2013" box="[625,803,1447,1473]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24A0CFA42FCFCFA49" box="[625,803,1447,1473]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA24B4FFA42FBE9FA49" author="Komai, T. & Osawa, M." box="[818,1078,1446,1473]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 1B;
|
||
<bibRefCitation id="EF9CC89D8633FFA24CDDFA42FAEFFA49" author="Komai, T." box="[1184,1328,1447,1473]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 1A). The antennular and antennal peduncles are proportionally longer in
|
||
<taxonomicName id="4C0DCEEF8633FFA24BE5FA2EFB88FA6D" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[920,1111,1483,1509]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24BE5FA2EFB88FA6D" box="[920,1111,1483,1509]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24C23FA2EFB41FA6D" bold="true" box="[1118,1182,1483,1509]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24C23FA2EFB41FA6D" box="[1118,1182,1483,1509]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName id="4C0DCEEF8633FFA24C80FA2EFA43FA6D" authorityName="Komai & Osawa" authorityYear="2005" box="[1277,1436,1483,1509]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24C80FA2EFA43FA6D" box="[1277,1436,1483,1509]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA248B5FA0AFEA6F981" authorityName="Komai" authorityYear="2013" box="[200,377,1519,1545]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA248B5FA0AFEA6F981" box="[200,377,1519,1545]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
. The antennular peduncle is comparatively longer and overreaches the distal corneal margins by half the length of article
|
||
<quantity id="4CF518898633FFA249B7F9F6FE2BF9A5" box="[458,500,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="127" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA24987F9F6FD68F9A5" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[506,695,1555,1581]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24987F9F6FD68F9A5" box="[506,695,1555,1581]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24AC0F9F6FD24F9A5" bold="true" box="[701,763,1555,1581]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24AC0F9F6FD24F9A5" box="[701,763,1555,1581]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA24B77F9F6FCBDF9A5" box="[778,866,1555,1581]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1A</figureCitation>
|
||
), instead by 0.5–0.7 length and 0.5 length of article
|
||
<quantity id="4CF518898633FFA248EAF9D2FF1CF9D9" box="[151,195,1591,1617]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="127" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA248B6F9D2FEB4F9D9" authorityName="Komai & Osawa" authorityYear="2005" box="[203,363,1591,1617]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA248B6F9D2FEB4F9D9" box="[203,363,1591,1617]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA249D9F9D2FD89F9D9" authorityName="Komai" authorityYear="2013" box="[420,598,1591,1617]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA249D9F9D2FD89F9D9" box="[420,598,1591,1617]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
, respectively (
|
||
<bibRefCitation id="EF9CC89D8633FFA24A81F9D2FBDEF9D9" author="Komai, T. & Osawa, M." box="[764,1025,1590,1617]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 1B;
|
||
<bibRefCitation id="EF9CC89D8633FFA24C11F9D2FB23F9D9" author="Komai, T." box="[1132,1276,1591,1617]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 1A). The antennal peduncle overreaches the distal corneal margins by nearly the full length of article
|
||
<quantity id="4CF518898633FFA24CD7F9BFFB09F9FD" box="[1194,1238,1626,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="127" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA24CA3F9BEFA43F9FD" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1246,1436,1627,1653]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24CA3F9BEFA43F9FD" box="[1246,1436,1627,1653]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA248EAF99AFF08F911" bold="true" box="[151,215,1663,1689]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA248EAF99AFF08F911" box="[151,215,1663,1689]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA24895F99AFE9DF911" box="[232,322,1663,1689]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1A</figureCitation>
|
||
), instead by at most only 0.2–0.3 length of article
|
||
<quantity id="4CF518898633FFA24BFDF99BFC73F911" box="[896,940,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="127" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA24BC9F99AFB8BF911" authorityName="Komai & Osawa" authorityYear="2005" box="[948,1108,1663,1689]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24BC9F99AFB8BF911" box="[948,1108,1663,1689]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24CF0F99AFAE1F911" authorityName="Komai" authorityYear="2013" box="[1165,1342,1663,1689]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24CF0F99AFAE1F911" box="[1165,1342,1663,1689]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA24D33F99AFE96F935" author="Komai, T. & Osawa, M." pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 1B;
|
||
<bibRefCitation id="EF9CC89D8633FFA249C7F946FD93F935" author="Komai, T." box="[442,588,1699,1725]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 1A). The dorsolateral distal angle of the antennal peduncle article 2 nearly reaches the distal margin of article
|
||
<quantity id="4CF518898633FFA24A15F922FD4CF969" box="[616,659,1735,1761]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="127" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA24AE4F922FC89F969" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[665,854,1735,1761]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24AE4F922FC89F969" box="[665,854,1735,1761]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24B21F922FC45F969" bold="true" box="[860,922,1735,1761]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24B21F922FC45F969" box="[860,922,1735,1761]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA24BD4F922FBDEF969" box="[937,1025,1735,1761]" captionStart="FIGURE 1" captionStartId="1.[151,250,1869,1893]" captionTargetBox="[159,1428,821,1840]" captionTargetId="figure-121@1.[151,1436,813,1845]" captionTargetPageId="1" captionText="FIGURE 1. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, shield and cephalic appendages, dorsal view (setae omitted from left ocular and antennal peduncle); B, left pereopod 4, lateral view; C, thoracic sternites 5–6, ventral view; D, left maxilliped 3, lateral view; E, same, basis, ischium, merus, ventral view; F, coxa of right pereopod 5 and right sexual tube, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10782046" httpUri="https://zenodo.org/record/10782046/files/figure.png" pageId="6" pageNumber="127">Fig. 1A</figureCitation>
|
||
), whereas it slightly falls short of the midlength of article
|
||
<quantity id="4CF518898633FFA24907F90EFE7AF88D" box="[378,421,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="127" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C0DCEEF8633FFA249D6F90EFD96F88D" authorityName="Komai & Osawa" authorityYear="2005" box="[427,585,1771,1797]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA249D6F90EFD96F88D" box="[427,585,1771,1797]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24A02F90EFCF0F88D" authorityName="Komai" authorityYear="2013" box="[639,815,1771,1797]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24A02F90EFCF0F88D" box="[639,815,1771,1797]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA24B43F90EFBE2F88D" author="Komai, T. & Osawa, M." box="[830,1085,1770,1797]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 1B;
|
||
<bibRefCitation id="EF9CC89D8633FFA24CD9F90EFAECF88D" author="Komai, T." box="[1188,1331,1771,1797]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 1A). The carpi of both chelipeds are much more weakly convex on the ventral surfaces in
|
||
<taxonomicName id="4C0DCEEF8633FFA24C2CF8EAFAD0F8A1" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1105,1295,1807,1833]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24C2CF8EAFAD0F8A1" box="[1105,1295,1807,1833]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24D68F8EAFA8BF8A1" bold="true" box="[1301,1364,1807,1833]" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24D68F8EAFA8BF8A1" box="[1301,1364,1807,1833]" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA24D19F8EAFF03F8C5" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="6" pageNumber="127">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="1336A9E98633FFA2489AF8D6FEF5F8C5" box="[231,298,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="6" pageNumber="127">3A, B</figureCitation>
|
||
) than in
|
||
<taxonomicName id="4C0DCEEF8633FFA249F1F8D6FDF5F8C5" authorityName="Komai & Osawa" authorityYear="2005" box="[396,554,1843,1869]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA249F1F8D6FDF5F8C5" box="[396,554,1843,1869]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24A22F8D6FCD0F8C5" authorityName="Komai" authorityYear="2013" box="[607,783,1843,1869]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24A22F8D6FCD0F8C5" box="[607,783,1843,1869]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA24B60F8D6FBC3F8C5" author="Komai, T. & Osawa, M." box="[797,1052,1842,1869]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3B, C, E, F;
|
||
<bibRefCitation id="EF9CC89D8633FFA24CA0F8D6FAB3F8C5" author="Komai, T." box="[1245,1388,1843,1869]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 2A–D). The carpus of the right cheliped is convex on the median part of the dorsomesial margin in the new species (
|
||
<figureCitation id="1336A9E98633FFA248E2F89EFF27F81D" box="[159,248,1915,1941]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="6" pageNumber="127">Fig. 2D</figureCitation>
|
||
), but it is widened toward the distal in
|
||
<taxonomicName id="4C0DCEEF8633FFA24AD4F89EFC98F81D" authorityName="Komai & Osawa" authorityYear="2005" box="[681,839,1915,1941]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24AD4F89EFC98F81D" box="[681,839,1915,1941]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24B00F89EFBF2F81D" authorityName="Komai" authorityYear="2013" box="[893,1069,1915,1941]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24B00F89EFBF2F81D" box="[893,1069,1915,1941]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8633FFA24C46F89EFAE4F81D" author="Komai, T. & Osawa, M." box="[1083,1339,1914,1941]" pageId="6" pageNumber="127" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3A;
|
||
<bibRefCitation id="EF9CC89D8633FFA248EAF87AFEF7F831" author="Komai, T." box="[151,296,1951,1977]" pageId="6" pageNumber="127" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 1B). The dorsolateral margin of the same article has three distinct spines in
|
||
<taxonomicName id="4C0DCEEF8633FFA24CC2F87AFAA1F831" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1215,1406,1951,1977]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8633FFA24CC2F87AFAA1F831" box="[1215,1406,1951,1977]" italics="true" pageId="6" pageNumber="127">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8633FFA24DFBF87AFF67F855" bold="true" pageId="6" pageNumber="127">
|
||
<taxonomicNameLabel id="A24AD4058633FFA24DFBF87AFF67F855" pageId="6" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98633FFA248BAF826FE9EF855" box="[199,321,1987,2013]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="6" pageNumber="127">Fig. 2A, D</figureCitation>
|
||
), whereas it possesses a row of low tubercles or protuberances in
|
||
<taxonomicName id="4C0DCEEF8633FFA24C64F826FB68F855" authorityName="Komai & Osawa" authorityYear="2005" box="[1049,1207,1987,2013]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8633FFA24C64F826FB68F855" box="[1049,1207,1987,2013]" italics="true" pageId="6" pageNumber="127">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8633FFA24C91F826FA43F855" authorityName="Komai" authorityYear="2013" box="[1260,1436,1987,2013]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8633FFA24C91F826FA43F855" box="[1260,1436,1987,2013]" italics="true" pageId="6" pageNumber="127">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8632FFA348E2FF72FE42FF39" author="Komai, T. & Osawa, M." box="[159,413,150,177]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3A, C;
|
||
<bibRefCitation id="EF9CC89D8632FFA34A5FFF72FD6FFF39" author="Komai, T." box="[546,688,151,177]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: figs. 1B, 2B). The ventromesial margin of the right cheliped merus is armed with three strong spines and one small spine in
|
||
<taxonomicName id="4C0DCEEF8632FFA34B67FF5EFC06FF5D" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[794,985,187,213]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34B67FF5EFC06FF5D" box="[794,985,187,213]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34B9DFF5EFBFFFF5D" bold="true" box="[992,1056,187,213]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34B9DFF5EFBFFFF5D" box="[992,1056,187,213]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98632FFA34C4DFF5EFB56FF5D" box="[1072,1161,187,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="7" pageNumber="128">Fig. 2B</figureCitation>
|
||
), instead of only a blunt spine proximally and no spines in
|
||
<taxonomicName id="4C0DCEEF8632FFA34A5AFF3AFD17FF71" authorityName="Komai & Osawa" authorityYear="2005" box="[551,712,223,249]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34A5AFF3AFD17FF71" box="[551,712,223,249]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8632FFA34B79FF3AFC68FF71" authorityName="Komai" authorityYear="2013" box="[772,951,223,249]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8632FFA34B79FF3AFC68FF71" box="[772,951,223,249]" italics="true" pageId="7" pageNumber="128">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
, respectively (
|
||
<bibRefCitation id="EF9CC89D8632FFA34C1DFF3AFAB6FF71" author="Komai, T. & Osawa, M." box="[1120,1385,222,249]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3B;
|
||
<bibRefCitation id="EF9CC89D8632FFA348BAFEE6FE88FE95" author="Komai, T." box="[199,343,259,285]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 2A). The carpus of the left cheliped is longer than the merus in
|
||
<taxonomicName id="4C0DCEEF8632FFA34C26FEE6FAC6FE95" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1115,1305,259,285]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34C26FEE6FAC6FE95" box="[1115,1305,259,285]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34D5DFEE6FA80FE95" bold="true" box="[1312,1375,259,285]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34D5DFEE6FA80FE95" box="[1312,1375,259,285]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98632FFA34D12FEE6FF02FEC9" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">Fig. 3A, B</figureCitation>
|
||
), rather than shorter in
|
||
<taxonomicName id="4C0DCEEF8632FFA3499AFEC2FD59FEC9" authorityName="Komai & Osawa" authorityYear="2005" box="[487,646,295,321]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA3499AFEC2FD59FEC9" box="[487,646,295,321]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8632FFA34AC3FEC2FCB0FEC9" authorityName="Komai" authorityYear="2013" box="[702,879,295,321]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8632FFA34AC3FEC2FCB0FEC9" box="[702,879,295,321]" italics="true" pageId="7" pageNumber="128">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8632FFA34B02FEC2FB5DFEC9" author="Komai, T. & Osawa, M." box="[895,1154,294,321]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3E, F;
|
||
<bibRefCitation id="EF9CC89D8632FFA34D75FEC2FA47FEC9" author="Komai, T." box="[1288,1432,295,321]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 2C, D). The dactylus of the ambulatory legs is shorter than the propodus (about 0.7–0.8 times longer than the propodus) in the new species (
|
||
<figureCitation id="1336A9E98632FFA34992FE8AFDB6FE01" box="[495,617,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">Fig. 3E, G</figureCitation>
|
||
), instead of longer in
|
||
<taxonomicName id="4C0DCEEF8632FFA34B1CFE8AFBDFFE01" authorityName="Komai & Osawa" authorityYear="2005" box="[865,1024,367,393]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34B1CFE8AFBDFFE01" box="[865,1024,367,393]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
(1.1–1.3 times) and
|
||
<taxonomicName id="4C0DCEEF8632FFA34C96FE8AFA43FE01" authorityName="Komai" authorityYear="2013" box="[1259,1436,367,393]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8632FFA34C96FE8AFA43FE01" box="[1259,1436,367,393]" italics="true" pageId="7" pageNumber="128">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(1.1–1.2 times) (
|
||
<bibRefCitation id="EF9CC89D8632FFA3492DFE76FD8EFE25" author="Komai, T. & Osawa, M." box="[336,593,402,429]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 2H, G;
|
||
<bibRefCitation id="EF9CC89D8632FFA34AA1FE76FCB4FE25" author="Komai, T." box="[732,875,403,429]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 3A, C).
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8632FFA348BAFE52FDF7FD21" blockId="7.[151,1437,150,1473]" pageId="7" pageNumber="128">
|
||
In addition to the distinctions mentioned above,
|
||
<taxonomicName id="4C0DCEEF8632FFA34A99FE52FC7DFE59" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[740,930,439,465]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34A99FE52FC7DFE59" box="[740,930,439,465]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34BD4FE52FC38FE59" bold="true" box="[937,999,439,465]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34BD4FE52FC38FE59" box="[937,999,439,465]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C0DCEEF8632FFA34C01FE52FAC5FE59" authorityName="Komai & Osawa" authorityYear="2005" box="[1148,1306,439,465]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34C01FE52FAC5FE59" box="[1148,1306,439,465]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
in the right cheliped palm being entirely smooth on all the surfaces, the right cheliped carpus having three distinct spines on the median one-third, and the carpus of the left cheliped with a row of distinct spines on both the dorsomesial and dorsolateral margins (
|
||
<figureCitation id="1336A9E98632FFA349F0FDC6FDCCFDB5" box="[397,531,547,573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="7" pageNumber="128">Figs. 2A–C</figureCitation>
|
||
,
|
||
<figureCitation id="1336A9E98632FFA34A62FDC6FD55FDB5" box="[543,650,547,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">3A, B, D</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C0DCEEF8632FFA34AC3FDC6FC82FDB5" authorityName="Komai & Osawa" authorityYear="2005" box="[702,861,547,573]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34AC3FDC6FC82FDB5" box="[702,861,547,573]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
, the right palm is minutely granular at least on the mesial and lateral surfaces, the dorsolateral margin of the right cheliped carpus is weakly delimited by a row of low tubercles or protuberances, and the carpus of the left cheliped has no conspicuous spines on the dorsal surface (
|
||
<bibRefCitation id="EF9CC89D8632FFA348E2FD6AFE7FFD21" author="Komai, T. & Osawa, M." box="[159,416,654,681]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: fig. 3A–F).
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8632FFA348BAFD56FBFCFCB1" blockId="7.[151,1437,150,1473]" pageId="7" pageNumber="128">
|
||
The proportionally shorter dactylus of the ambulatory legs further distinguishes the present new species from
|
||
<taxonomicName id="4C0DCEEF8632FFA34DF5FD56FEF3FD79" authorityName="Komai" authorityYear="2013" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8632FFA34DF5FD56FEF3FD79" italics="true" pageId="7" pageNumber="128">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
; the dactyli are 8.3–9.0 versus 11–14 times longer than the height (
|
||
<figureCitation id="1336A9E98632FFA34C59FD32FB41FD79" box="[1060,1182,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">Fig. 3E, G</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF9CC89D8632FFA34CD7FD32FAE5FD79" author="Komai, T." box="[1194,1338,727,753]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 3A, C). The ventral margins of the same articles also have nine or ten corneous spines in
|
||
<taxonomicName id="4C0DCEEF8632FFA34C24FD1EFAC8FC9D" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1113,1303,763,789]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34C24FD1EFAC8FC9D" box="[1113,1303,763,789]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34D62FD1EFA81FC9D" bold="true" box="[1311,1374,763,789]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34D62FD1EFA81FC9D" box="[1311,1374,763,789]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1336A9E98632FFA34D12FD1EFF03FCB1" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">Fig. 3E–H</figureCitation>
|
||
), instead of only eight spines in
|
||
<taxonomicName id="4C0DCEEF8632FFA34A3BFCFAFD28FCB1" authorityName="Komai" authorityYear="2013" box="[582,759,799,825]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="tenuidactylus">
|
||
<emphasis id="B979697E8632FFA34A3BFCFAFD28FCB1" box="[582,759,799,825]" italics="true" pageId="7" pageNumber="128">T. tenuidactylus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8632FFA34B78FCFAFC49FCB1" author="Komai, T." box="[773,918,799,825]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
: fig. 3A–D).
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8632FFA348BAFCA6FD43FC41" blockId="7.[151,1437,150,1473]" pageId="7" pageNumber="128">
|
||
The new species can be immediately distinguished from
|
||
<taxonomicName id="4C0DCEEF8632FFA34B3BFCA6FC7FFCD4" authorityName="Komai & Poupin" authorityYear="2012" box="[838,928,835,860]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="asper">
|
||
<emphasis id="B979697E8632FFA34B3BFCA6FC7FFCD4" box="[838,928,835,860]" italics="true" pageId="7" pageNumber="128">T. asper</emphasis>
|
||
</taxonomicName>
|
||
by the dorsal surface of the palm of the right cheliped being smooth (
|
||
<figureCitation id="1336A9E98632FFA349D8FC82FE21FC09" box="[421,510,871,897]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="7" pageNumber="128">Fig. 2C</figureCitation>
|
||
), instead of distinctly tuberculate (
|
||
<bibRefCitation id="EF9CC89D8632FFA34BFFFC82FB58FC09" author="Komai, T. & Poupin, J." box="[898,1159,871,897]" pageId="7" pageNumber="128" pagination="56 - 68" refId="ref4881" refString="Komai, T. & Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai & Poupin 2012</bibRefCitation>
|
||
: fig. 6A–C), and from
|
||
<taxonomicName id="4C0DCEEF8632FFA34DF5FC82FE90FC2D" baseAuthorityName="Forest" baseAuthorityYear="1954" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34DF5FC82FE90FC2D" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
by both the chelipeds and ambulatory legs being much less setose (
|
||
<figureCitation id="1336A9E98632FFA34C3DFC6EFB50FC2D" box="[1088,1167,907,933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[172,1413,389,1907]" captionTargetId="figure-102@2.[164,1423,381,1915]" captionTargetPageId="2" captionText="FIGURE 2. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. Right cheliped.A, entire, lateral view; B, same, mesial view; C, chela, dorsal view; D, carpus, dorsal view; E, same, mesial view (setae omitted). Plumose setae on ischium simplified. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782048" httpUri="https://zenodo.org/record/10782048/files/figure.png" pageId="7" pageNumber="128">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="1336A9E98632FFA34CE6FC6EFB75FC2D" box="[1179,1194,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1873,1897]" captionTargetBox="[159,1428,317,1837]" captionTargetId="figure-92@4.[151,1436,309,1849]" captionTargetPageId="4" captionText="FIGURE 3. Trichopagurus spinibrachium n. sp., holotype male (sl 1.4 mm), RUMF-ZC-7664. A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view; D, same, carpus, dorsal view; E, right pereopod 2, lateral view; F, same, dactylus, mesial view; G, left pereopod 3, lateral view; H, same, dactylus, mesial view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10782050" httpUri="https://zenodo.org/record/10782050/files/figure.png" pageId="7" pageNumber="128">3</figureCitation>
|
||
), instead of distinctly setose (
|
||
<bibRefCitation id="EF9CC89D8632FFA34897FC4AFE34FC41" author="Komai, T. & Osawa, M." box="[234,491,942,969]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
: figs.
|
||
<geoCoordinate id="EE39D3AB8632FFA34A53FC4BFDB4FC41" box="[558,619,942,969]" degrees="5, 6" direction="east" orientation="longitude" pageId="7" pageNumber="128" precision="5555" value="5.6">5, 6E</geoCoordinate>
|
||
–G).
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8632FFA348BAFC36FC05FBBD" blockId="7.[151,1437,150,1473]" pageId="7" pageNumber="128">
|
||
The fresh coloration of the entire body is generally similar in all of
|
||
<taxonomicName id="4C0DCEEF8632FFA34BA1FC36FBA2FC65" authorityName="Komai & Osawa" authorityYear="2005" box="[988,1149,979,1005]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34BA1FC36FBA2FC65" box="[988,1149,979,1005]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DCEEF8632FFA34CF3FC36FABBFC65" baseAuthorityName="Forest" baseAuthorityYear="1954" box="[1166,1380,979,1005]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34CF3FC36FABBFC65" box="[1166,1380,979,1005]" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0DCEEF8632FFA348EAFC12FE86FB99" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[151,345,1015,1041]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA348EAFC12FE86FB99" box="[151,345,1015,1041]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA3491EFC12FE79FB99" bold="true" box="[355,422,1015,1041]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA3491EFC12FE79FB99" box="[355,422,1015,1041]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but the pereopods are pale brown in
|
||
<taxonomicName id="4C0DCEEF8632FFA34B10FC12FBD0FB99" authorityName="Komai & Osawa" authorityYear="2005" box="[877,1039,1015,1041]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34B10FC12FBD0FB99" box="[877,1039,1015,1041]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
, rather than orangish brown in
|
||
<taxonomicName id="4C0DCEEF8632FFA34DF5FC12FE90FBBD" baseAuthorityName="Forest" baseAuthorityYear="1954" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34DF5FC12FE90FBBD" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0DCEEF8632FFA349F1FBFEFD95FBBD" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[396,586,1051,1077]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA349F1FBFEFD95FBBD" box="[396,586,1051,1077]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34A2DFBFEFD50FBBD" bold="true" box="[592,655,1051,1077]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34A2DFBFEFD50FBBD" box="[592,655,1051,1077]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9CC89D8632FFA34AE3FBFEFCF5FBBD" author="Arima, H." box="[670,810,1051,1077]" pageId="7" pageNumber="128" refId="ref4560" refString="Arima, H. (2014) Hermit Crabs Illustrated. Seibundo-Shinkosha Co. Ltd, Tokyo, 223 pp. [in Japanese]" type="book" year="2014">Arima 2014</bibRefCitation>
|
||
; present study).
|
||
</paragraph>
|
||
<paragraph id="8BB2B56C8632FFA348BAFBDAFB97FA49" blockId="7.[151,1437,150,1473]" pageId="7" pageNumber="128">
|
||
Although the sole present specimen of
|
||
<taxonomicName id="4C0DCEEF8632FFA34AFBFBDAFC9AFBD1" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[646,837,1087,1113]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34AFBFBDAFC9AFBD1" box="[646,837,1087,1113]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34B33FBDAFC51FBD1" bold="true" box="[846,910,1087,1113]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34B33FBDAFC51FBD1" box="[846,910,1087,1113]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was collected from a semi-submerged marine cave, the new species may also occur on exposed shallow reef walls and nearby environments, as in three known congeners,
|
||
<taxonomicName id="4C0DCEEF8632FFA34965FB62FEACFB28" authorityName="Komai & Poupin" authorityYear="2012" box="[280,371,1159,1184]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="asper">
|
||
<emphasis id="B979697E8632FFA34965FB62FEACFB28" box="[280,371,1159,1184]" italics="true" pageId="7" pageNumber="128">T. asper</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DCEEF8632FFA349FFFB62FDFDFB29" authorityName="Komai & Osawa" authorityYear="2005" box="[386,546,1159,1185]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA349FFFB62FDFDFB29" box="[386,546,1159,1185]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0DCEEF8632FFA34A1FFB62FCE8FB29" baseAuthorityName="Forest" baseAuthorityYear="1954" box="[610,823,1159,1185]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34A1FFB62FCE8FB29" box="[610,823,1159,1185]" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
. Actually,
|
||
<taxonomicName id="4C0DCEEF8632FFA34BCFFB62FB8DFB29" authorityName="Komai & Osawa" authorityYear="2005" box="[946,1106,1159,1185]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34BCFFB62FB8DFB29" box="[946,1106,1159,1185]" italics="true" pageId="7" pageNumber="128">T. macrochela</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8632FFA34CF6FB62FABFFB29" baseAuthorityName="Forest" baseAuthorityYear="1954" box="[1163,1376,1159,1185]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34CF6FB62FABFFB29" box="[1163,1376,1159,1185]" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
have also been collected from coral reefs and lagoons as well as submarine caves in the
|
||
<collectingCountry id="F31AF5FC8632FFA34C22FB4EFB02FB4D" box="[1119,1245,1195,1221]" name="Philippines" pageId="7" pageNumber="128">Philippines</collectingCountry>
|
||
and the Ryukyu Islands, respectively (
|
||
<bibRefCitation id="EF9CC89D8632FFA349F2FB2AFDFCFB61" author="Komai, T." box="[399,547,1231,1257]" pageId="7" pageNumber="128" pagination="274 - 282" refId="ref4752" refString="Komai, T. (2013) Additional records of the pagurid hermit crab genus Trichopagurus de Saint Laurent, 1968 (Crustacea: Decapoda: Anomura), with description of a new species from the Philippines. Zootaxa, 3670 (2), 274 - 282. https: // doi. org / 10.11646 / zootaxa. 3670.2.10" type="journal article" year="2013">Komai 2013</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C0DCEEF8632FFA34A45FB2AFCB5FB61" authorityName="Komai & Osawa" authorityYear="2005" box="[568,874,1231,1257]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="macrochela">
|
||
<emphasis id="B979697E8632FFA34A45FB2AFCB5FB61" box="[568,874,1231,1257]" italics="true" pageId="7" pageNumber="128">Trichopagurus macrochela</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DCEEF8632FFA34B07FB2AFB8FFB61" baseAuthorityName="Forest" baseAuthorityYear="1954" box="[890,1104,1231,1257]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="trichophthalmus">
|
||
<emphasis id="B979697E8632FFA34B07FB2AFB8FFB61" box="[890,1104,1231,1257]" italics="true" pageId="7" pageNumber="128">T. trichophthalmus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0DCEEF8632FFA34CEFFB2AFA8DFB61" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[1170,1362,1231,1257]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34CEFFB2AFA8DFB61" box="[1170,1362,1231,1257]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34D26FB2AFA43FB61" bold="true" box="[1371,1436,1231,1257]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34D26FB2AFA43FB61" box="[1371,1436,1231,1257]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have been recorded from Japanese waters (
|
||
<bibRefCitation id="EF9CC89D8632FFA34A04FB16FCA6FA85" author="Komai, T. & Osawa, M." box="[633,889,1266,1293]" pageId="7" pageNumber="128" pagination="1 - 20" refId="ref4814" refString="Komai, T. & Osawa, M. (2005) A new species of Trichopagurus de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 801.1.1" type="journal article" year="2005">Komai & Osawa 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9CC89D8632FFA34BF9FB16FBD1FA85" author="Arima, H." box="[900,1038,1267,1293]" pageId="7" pageNumber="128" refId="ref4560" refString="Arima, H. (2014) Hermit Crabs Illustrated. Seibundo-Shinkosha Co. Ltd, Tokyo, 223 pp. [in Japanese]" type="book" year="2014">Arima 2014</bibRefCitation>
|
||
; present study), whereas
|
||
<taxonomicName id="4C0DCEEF8632FFA34D55FB16FA5DFA84" authorityName="Komai & Poupin" authorityYear="2012" box="[1320,1410,1267,1292]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="asper">
|
||
<emphasis id="B979697E8632FFA34D55FB16FA5DFA84" box="[1320,1410,1267,1292]" italics="true" pageId="7" pageNumber="128">T. asper</emphasis>
|
||
</taxonomicName>
|
||
is known only from
|
||
<collectingCountry id="F31AF5FC8632FFA3491EFAF2FDD3FAB9" box="[355,524,1303,1329]" name="Mayotte Island" pageId="7" pageNumber="128">Mayotte Island</collectingCountry>
|
||
in the southwestern Indian Ocean (
|
||
<bibRefCitation id="EF9CC89D8632FFA34BE3FAF2FB79FAB9" author="Komai, T. & Poupin, J." box="[926,1190,1303,1329]" pageId="7" pageNumber="128" pagination="56 - 68" refId="ref4881" refString="Komai, T. & Poupin, J. (2012) Two new species of shallow-water hermit crabs (Crustacea: Decapoda: Paguridae) from Mayotte Island, southwestern Indian Ocean. Zootaxa, 3277 (1), 56 - 68. https: // doi. org / 10.11646 / zootaxa. 3277.1.4" type="journal article" year="2012">Komai & Poupin 2012</bibRefCitation>
|
||
). As mentioned in the
|
||
<emphasis id="B979697E8632FFA348EAFADFFF2CFADC" bold="true" box="[151,243,1338,1364]" pageId="7" pageNumber="128">Habitat</emphasis>
|
||
section above, two hermit crabs,
|
||
<taxonomicName id="4C0DCEEF8632FFA34A1AFADEFCC4FADD" authorityName="Morgan" authorityYear="1992" box="[615,795,1339,1365]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="jalur">
|
||
<emphasis id="B979697E8632FFA34A1AFADEFCC4FADD" box="[615,795,1339,1365]" italics="true" pageId="7" pageNumber="128">Paguristes jalur</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8632FFA34B2CFADEFBF6FADD" authorityName="Osawa, Fujita & Okuno" authorityYear="2006" box="[849,1065,1339,1365]" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B979697E8632FFA34B2CFADEFBF6FADD" box="[849,1065,1339,1365]" italics="true" pageId="7" pageNumber="128">Pagurixus longipes</emphasis>
|
||
</taxonomicName>
|
||
, were observed in the nearby site in the same semi-submerged marine cave where
|
||
<taxonomicName id="4C0DCEEF8632FFA34AC8FABAFCADFAF1" authority="Nakajima & Fujita & Osawa, 2024" authorityName="Nakajima & Fujita & Osawa" authorityYear="2024" box="[693,882,1375,1401]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="spinibrachium" status="n. sp.">
|
||
<emphasis id="B979697E8632FFA34AC8FABAFCADFAF1" box="[693,882,1375,1401]" italics="true" pageId="7" pageNumber="128">T. spinibrachium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979697E8632FFA34B05FABAFC68FAF1" bold="true" box="[888,951,1375,1401]" pageId="7" pageNumber="128">
|
||
<taxonomicNameLabel id="A24AD4058632FFA34B05FABAFC68FAF1" box="[888,951,1375,1401]" pageId="7" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was found.
|
||
<taxonomicName id="4C0DCEEF8632FFA34C42FABAFB2CFAF1" authorityName="Morgan" authorityYear="1992" box="[1087,1267,1375,1401]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="jalur">
|
||
<emphasis id="B979697E8632FFA34C42FABAFB2CFAF1" box="[1087,1267,1375,1401]" italics="true" pageId="7" pageNumber="128">Paguristes jalur</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DCEEF8632FFA34D54FABAFF29FA15" authorityName="Osawa, Fujita & Okuno" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B979697E8632FFA34D54FABAFF29FA15" italics="true" pageId="7" pageNumber="128">Pagurixus longipes</emphasis>
|
||
</taxonomicName>
|
||
have been also recorded from non-cave but dark environments (cf.
|
||
<bibRefCitation id="EF9CC89D8632FFA34B91FA66FB3FFA15" author="Osawa, M. & Fujita, Y." box="[1004,1248,1411,1437]" pageId="7" pageNumber="128" pagination="463 - 482" refId="ref5160" refString="Osawa, M. & Fujita, Y. (2019) Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan. Zootaxa, 4560 (3), 463 - 482. https: // doi. org / 10.11646 / zootaxa. 4560.3.3" type="journal article" year="2019">Osawa & Fujita 2019</bibRefCitation>
|
||
), suggesting that the new species may have a relatively wide habitat preference, as in the two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |