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<document id="D5FAE432847EBC0CC1773B40642D5340" ID-CLB-Dataset="199799" ID-DOI="10.11646/zootaxa.5214.3.4" ID-GBIF-Dataset="9c8b2ed9-a132-4df0-afa9-b153547b9027" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389267" ID-ZooBank="395A3FB5-DA5E-4145-B9EA-7801C67D6CE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970496187" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2022" docId="038987F7BC0008789495FA840F359ACA" docLanguage="en" docName="zootaxa.5214.3.4.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis thatsonensis Huynh 2022, new species" docType="treatment" docVersion="7" lastPageNumber="413" masterDocId="FFB0FF8FBC12086C9402FF890C769D01" masterDocTitle="Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam" masterLastPageNumber="420" masterPageNumber="393" pageNumber="411" updateTime="1699412757114" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9CBD44FC21F526475F8C6DD816F5DA39" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="71D89F65ABC014464A5F951CBBD760B7">
<mods:title id="18A0C1B0CF4E58F3D7C44BADAC15F2F9">Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="AE648BAD8BA88E7965246AF8AF2A2C31" type="personal">
<mods:role id="D9F5BFE57C0BBF16A1E05B36E1EFA309">
<mods:roleTerm id="E5A98808B04B9DC0F0ACBC4A2334FB88">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB4DAB063B54258A9D6DB15C2133B9E0">Huynh, Cuong</mods:namePart>
<mods:affiliation id="E7CE246E9861A9F5F7869E885F8F18FB">Centre for Cellular and Molecular Biology (CCMB), Deakin University, 221 Burwood Hwy, Burwood, Melbourne, Australia 3125</mods:affiliation>
</mods:name>
<mods:typeOfResource id="429065E2C7A5BD93BABA150511DEF6C0">text</mods:typeOfResource>
<mods:relatedItem id="12CE0CBE8B99D7AA92F3FF8A72E25F6B" type="host">
<mods:titleInfo id="73603F5C801F4CB8F1AFDDE554E46073">
<mods:title id="03D2366EDD280117DDCF040103F192D1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8BA8B06E96993808E10D5DD836D63BB4">
<mods:date id="93302E1D339904FCB8612F00B825F9E0">2022</mods:date>
<mods:detail id="4B4CE08227782FD6F70BA1DBA497B551" type="pubDate">
<mods:number id="0C0A21F4AEEBD0775E0EBF9998A0EDE1">2022-12-02</mods:number>
</mods:detail>
<mods:detail id="27FB2D16344780E6445D978FEB0A9500" type="volume">
<mods:number id="D09785D605EBC8CA0DF1F1911ED355C4">5214</mods:number>
</mods:detail>
<mods:detail id="4E2931C4BA1950155A9EE4590ACE8DE5" type="issue">
<mods:number id="DC0B6E6FF3D67A1B47540E23D197415E">3</mods:number>
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<mods:extent id="102C0D8C5CA8456CD71CFB0DA8F901CF" unit="page">
<mods:start id="D329000D8F3BB0275B1179DD9BCA19E0">393</mods:start>
<mods:end id="9479E2815F416BAD7B0A3D39E5D54C2B">420</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="946E8E5263275DE152D32C8A456F1C85">journal article</mods:classification>
<mods:identifier id="D40ACCC2E4674953F21334F5A2A7E9A4" type="CLB-Dataset">199799</mods:identifier>
<mods:identifier id="78EE49B9638EF9126F414037B858146D" type="DOI">10.11646/zootaxa.5214.3.4</mods:identifier>
<mods:identifier id="39E6CDC7AD477803A8543453FF74117E" type="GBIF-Dataset">9c8b2ed9-a132-4df0-afa9-b153547b9027</mods:identifier>
<mods:identifier id="B6CB97ED15FE2EE4BE038449F94D88AF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="74CC6907D1EA3F7650BCE75F19CC95A2" type="Zenodo-Dep">7389267</mods:identifier>
<mods:identifier id="965273F88793166188B841A4FEB9D3A2" type="ZooBank">395A3FB5-DA5E-4145-B9EA-7801C67D6CE3</mods:identifier>
</mods:mods>
<treatment id="038987F7BC0008789495FA840F359ACA" ID-DOI="http://doi.org/10.5281/zenodo.7389488" ID-GBIF-Taxon="205697562" ID-Zenodo-Dep="7389488" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA" httpUri="http://treatment.plazi.org/id/038987F7BC0008789495FA840F359ACA" lastPageId="20" lastPageNumber="413" pageId="18" pageNumber="411">
<subSubSection id="C33A656ABC00087E9495FA840E199829" box="[151,623,1293,1320]" pageId="18" pageNumber="411" type="nomenclature">
<paragraph id="8B9F36E1BC00087E9495FA840E199829" blockId="18.[151,623,1293,1356]" box="[151,623,1293,1320]" pageId="18" pageNumber="411">
<heading id="D0D7818DBC00087E9495FA840E199829" bold="true" box="[151,623,1293,1320]" fontSize="11" level="1" pageId="18" pageNumber="411" reason="1">
<emphasis id="B954EAF3BC00087E9495FA840E199829" bold="true" box="[151,623,1293,1320]" pageId="18" pageNumber="411">
<taxonomicName id="4C204D62BC00087E9495FA840DAE9829" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[151,472,1293,1320]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="18" pageNumber="411" phylum="Arthropoda" rank="species" species="thatsonensis" status="sp. nov.">
<emphasis id="B954EAF3BC00087E9495FA840DAE9829" bold="true" box="[151,472,1293,1320]" italics="true" pageId="18" pageNumber="411">Monographis thatsonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2675788BC00087E95DDFA870E199829" box="[479,623,1294,1320]" pageId="18" pageNumber="411" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC00087E9495FABB0D22984D" box="[151,340,1330,1356]" pageId="18" pageNumber="411" type="description">
<paragraph id="8B9F36E1BC00087E9495FABB0D22984D" blockId="18.[151,623,1293,1356]" box="[151,340,1330,1356]" pageId="18" pageNumber="411">
<figureCitation id="131B2A64BC00087E9495FABB0D74984D" box="[151,258,1330,1356]" captionStart="FIGURE 2" captionStartId="4.[152,255,1543,1567]" captionTargetBox="[178,1409,597,1520]" captionTargetId="figure-244@4.[178,1409,597,1520]" captionTargetPageId="4" captionText="FIGURE 2. Live specimen of Monographis thatsonensis sp. n., its appearance is typical of Monographis species (Diplopoda, Polyxenida)—commonly yellowish brown in colour, darker on the latero-posterior rosette trichomes, forming two dark lateral bands contrasting with a light-yellow medial band." figureDoi="http://doi.org/10.5281/zenodo.7389277" httpUri="https://zenodo.org/record/7389277/files/figure.png" pageId="18" pageNumber="411">Figures 2</figureCitation>
,
<figureCitation id="131B2A64BC00087E950CFABB0D22984D" box="[270,340,1330,1356]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStart-3="FIGURE 21" captionStart-4="FIGURE 22" captionStart-5="FIGURE 23" captionStartId-0="19.[152,255,1782,1806]" captionStartId-1="20.[152,255,1430,1454]" captionStartId-2="21.[152,255,1896,1920]" captionStartId-3="22.[152,255,1079,1103]" captionStartId-4="22.[152,255,1902,1926]" captionStartId-5="23.[152,255,1170,1194]" captionTargetBox-0="[163,1435,317,1758]" captionTargetBox-1="[159,1419,190,1401]" captionTargetBox-2="[163,1413,191,1870]" captionTargetBox-3="[280,1307,181,1055]" captionTargetBox-4="[297,1291,1236,1879]" captionTargetBox-5="[152,1436,181,1146]" captionTargetId-0="figure-79@19.[151,1436,308,1758]" captionTargetId-1="figure-29@20.[151,1436,181,1406]" captionTargetId-2="figure-19@21.[158,1430,181,1872]" captionTargetId-3="figure-29@22.[280,1307,181,1055]" captionTargetId-4="figure-94@22.[297,1291,1236,1879]" captionTargetId-5="figure-19@23.[151,1436,181,1146]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="22" captionTargetPageId-5="23" captionText-0="FIGURE 18. Holotype male Monographis thatsonensis sp. n.. A. A head capsule showing eye patterns with eight ommatidia and trichobothria (a, b, and c) each side, vertex region (v), anterior vertex (av), posterior vertex trichome sockets (pv); B. Enlarged images of posterior vertex (pv) trichome sockets on left (pv (L)) and right (pv (R)). C. Clypeo-labrum showing labrum (l) with asetose surface, lateral labrum (Ll), anterior setae (sa), backward-pointing setae (sb), and posterior setae (Se, 4 setae and 6 sockets of missing setae)." captionText-1="FIGURE 19. Holotype male Monographis thatsonensis sp. n.. A. Antennal articles V, VI, VII and VIII with sensory cones (sc), and patterns of sensilla on article VI and VII; B. Article VII showing two thick bacilliform sensilla (Ta: anterior &amp; Tp: posterior), setiform (S), and a conical sensillum (C); C. Article VI showing three thick bacilliform sensilla (T), three thin bacilliform sensilla (t), three medium-length thin bacilliform sensilla (tm), and six short thin bacilliform sensilla (ts), a conical sensillum (C)." captionText-2="FIGURE 20. Holotype male Monographis thatsonensis sp. n.. A. Second leg showing seven articles: coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2); claw (c) and penis (p). B. Telotarsus showing claw (c) and all processes: anterior (a), posterior (p), setiform (s) and the lamella (la). C. Ornamental trichome sockets showing the arrangements on both sides: trichome sockets a, b, c, and indentation d." captionText-3="FIGURE 21. SEM images of Monographis thatsonensis sp. n.. A. Head capsule showing left antenna with eight articles, eye pattern with eight ommatidia, and the trichobothria. B. Antennal articles VI–VIII, showing the arrangement of sensilla. C. Antennal articles VII showing the typical pattern of sensilla and VIII with sensory cones (sc). D. Arrangement of sensilla on article VI." captionText-4="FIGURE 22. SEM images of Monographis thatsonensis sp. n.. A. The head—right side showing the posterior vertex (pv) trichome sockets and trichobothria (a, b, and c). B. Coxal glands on legs VIII and IX (pointed arrows). C. Clypeo-labrum showing structure of labrum (l) with asetose surface, lateral labrum (Ll), medial cleft (mc), anterior setae (sa), posterior setae (Se), backward-pointing setae (sb). D. Gnathochilaria (g): lateral palps (Lp) and median palps (Mp), the frons (f), labrum (l)." captionText-5="FIGURE 23. SEM images of Monographis thatsonensis sp. n.. A. Male’s caudal bundle with ornamental trichome sockets (ots) a, b, c and d; and telson (Te), and the bundle with caudal bundle trichome sockets (cbts), barbate trichome sockets (bts); the last pleural projection (pp). B. Female’s caudal bundle structure showing the same dorsally, except the presence of two bundles of nest trichome sockets ventrally (nts). C. Presence of two spines—Tarsus 1 and 2, typical of Monographis species. D. Telotarsus showing a claw (c) and all processes: anterior (a), posterior (p), anterior setiform process (s) and lamella (la)." figureDoi-0="http://doi.org/10.5281/zenodo.7389346" figureDoi-1="http://doi.org/10.5281/zenodo.7389348" figureDoi-2="http://doi.org/10.5281/zenodo.7389352" figureDoi-3="http://doi.org/10.5281/zenodo.7389354" figureDoi-4="http://doi.org/10.5281/zenodo.7389358" figureDoi-5="http://doi.org/10.5281/zenodo.7389361" httpUri-0="https://zenodo.org/record/7389346/files/figure.png" httpUri-1="https://zenodo.org/record/7389348/files/figure.png" httpUri-2="https://zenodo.org/record/7389352/files/figure.png" httpUri-3="https://zenodo.org/record/7389354/files/figure.png" httpUri-4="https://zenodo.org/record/7389358/files/figure.png" httpUri-5="https://zenodo.org/record/7389361/files/figure.png" pageId="18" pageNumber="411">18–23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC00087E9495FAF308C59B25" pageId="18" pageNumber="411" type="materials_examined">
<paragraph id="8B9F36E1BC00087E9495FAF308C59B25" blockId="18.[151,1437,1402,2041]" pageId="18" pageNumber="411">
<emphasis id="B954EAF3BC00087E9495FAF30D0F9895" bold="true" box="[151,377,1402,1428]" pageId="18" pageNumber="411">Material examined</emphasis>
:
<materialsCitation id="3B483CBCBC00087E958AFAF308C59B25" ID-GBIF-Occurrence="3981599310" collectingDate="2017-07-08" country="Vietnam" county="Specimens" elevation="662" latitude="10.496944" location="Cam Mountain" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" pageId="18" pageNumber="411" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="An Giang" typeStatus="holotype">
<typeStatus id="549B8843BC00087E958AFAF30D859895" box="[392,499,1402,1428]" pageId="18" pageNumber="411" type="holotype">Holotype</typeStatus>
, adult male (IEBR-Myr 993) and
<typeStatus id="549B8843BC00087E976CFAF20FA89894" box="[878,990,1403,1429]" pageId="18" pageNumber="411" type="paratype">paratypes</typeStatus>
: Adult,
<specimenCount id="9D26FD68BC00087E9037FAF208D19895" box="[1077,1191,1402,1429]" pageId="18" pageNumber="411" type="male">two males</specimenCount>
(IEBR-Myr 994)
<specimenCount id="9D26FD68BC00087E9170FAF20C9898B9" pageId="18" pageNumber="411" type="female">two females</specimenCount>
(IEBR-Myr 995) and subadults,
<specimenCount id="9D26FD68BC00087E9667FA170E8A98B9" box="[613,764,1438,1464]" pageId="18" pageNumber="411" type="female">three females</specimenCount>
with 12 pairs of legs (IEBR-Myr 996) were collected from
<location id="8EFF603ABC00087E9495FA4B0D3598DD" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9495FA4B0D3598DD" box="[151,323,1474,1500]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Cam Mountain" pageId="18" pageNumber="411" stateProvince="An Giang">Cam Mountain</location>
,
<geoCoordinate id="EE145026BC00087E954FFA4B0DAA98DD" box="[333,476,1474,1500]" degrees="10.496944" direction="north" orientation="latitude" pageId="18" pageNumber="411" precision="1" value="10.496944">10.496944N</geoCoordinate>
,
<geoCoordinate id="EE145026BC00087E95E4FA4B0EF698DD" box="[486,640,1474,1500]" degrees="104.981389" direction="east" orientation="longitude" pageId="18" pageNumber="411" precision="1" value="104.98139">104.981389E</geoCoordinate>
;
<elevation id="000DD1D2BC00087E9689FA4B0F3798DC" box="[651,833,1474,1501]" metricMagnitude="2" metricUnit="m" metricValue="6.62" pageId="18" pageNumber="411" unit="m" value="662.0">
elevation
<quantity id="4CD89B04BC00087E96FAFA4B0F3798DC" box="[760,833,1474,1501]" metricMagnitude="2" metricUnit="m" metricValue="6.62" pageId="18" pageNumber="411" unit="m" value="662.0">662 m</quantity>
</elevation>
, in the
<location id="8EFF603ABC00087E9793FA4B08D498DC" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9793FA4B08D498DC" box="[913,1186,1474,1501]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Seven Mountain Ranges" pageId="18" pageNumber="411" stateProvince="An Giang">
<location id="8EFF603ABC00087E9793FA4B083F98DD" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9793FA4B083F98DD" box="[913,1097,1474,1500]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Seven Mountain" pageId="18" pageNumber="411" stateProvince="An Giang">
Seven
<location id="8EFF603ABC00087E97DEFA4B083F98DD" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E97DEFA4B083F98DD" box="[988,1097,1474,1500]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Mountain Ranges" pageId="18" pageNumber="411" stateProvince="An Giang">Mountain</location>
</location>
Ranges
</location>
(That Son), located in
<location id="8EFF603ABC00087E9495FA6F0C9D9B00" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9495FA6F0C9D9B00" box="[151,235,1510,1537]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Tri Ton" pageId="18" pageNumber="411" stateProvince="An Giang">Tri Ton</location>
and
<location id="8EFF603ABC00087E9520FA6F0DE59B01" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9520FA6F0DE59B01" box="[290,403,1510,1536]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Tinh Bien" pageId="18" pageNumber="411" stateProvince="An Giang">Tinh Bien</location>
districts,
<collectingRegion id="49E4F803BC00087E9603FA6F0EAB9B01" box="[513,733,1510,1536]" country="Vietnam" name="An Giang" pageId="18" pageNumber="411">An Giang Province</collectingRegion>
,
<collectingCountry id="F3377671BC00087E96EBFA6F0F3A9B01" box="[745,844,1510,1536]" name="Vietnam" pageId="18" pageNumber="411">Vietnam</collectingCountry>
.
<location id="8EFF603ABC00087E9757FA6F0F0F9B01" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E9757FA6F0F0F9B01" box="[853,889,1510,1536]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="All" pageId="18" pageNumber="411" stateProvince="An Giang">All</location>
types were deposited in the
<location id="8EFF603ABC00087E90BEFA6F09EA9B01" LSID="urn:lsid:plazi:treatment:038987F7BC0008789495FA840F359ACA:8EFF603ABC00087E90BEFA6F09EA9B01" box="[1212,1436,1510,1536]" country="Vietnam" county="Specimens" latitude="10.496944" longLatPrecision="1" longitude="104.98139" municipality="Biological Resources" name="Institute of Ecology" pageId="18" pageNumber="411" stateProvince="An Giang">Institute of Ecology</location>
and
<collectingMunicipality id="6BFBAC9BBC00087E94C5F9830DCE9B24" box="[199,440,1546,1573]" pageId="18" pageNumber="411">Biological Resources</collectingMunicipality>
,
<collectingCountry id="F3377671BC00087E95C1F9830E509B25" box="[451,550,1546,1572]" name="Vietnam" pageId="18" pageNumber="411">Vietnam</collectingCountry>
.
<collectingCounty id="62FE4E6DBC00087E9632F9830EDF9B25" box="[560,681,1546,1572]" pageId="18" pageNumber="411">Specimens</collectingCounty>
were collected by author CH on
<date id="FF9E1021BC00087E901EF98308D99B25" box="[1052,1199,1546,1572]" pageId="18" pageNumber="411" value="2017-07-08">
<collectingDate id="EFDAE9C9BC00087E901EF98308D99B25" box="[1052,1199,1546,1572]" pageId="18" pageNumber="411" value="2017-07-08">
8
<superScript id="7C559BA9BC00087E9028F98308419B19" attach="left" box="[1066,1079,1546,1560]" fontSize="6" pageId="18" pageNumber="411">th</superScript>
July 2017
</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC00087E94C5F9A709FF9BB5" pageId="18" pageNumber="411" type="diagnosis">
<paragraph id="8B9F36E1BC00087E94C5F9A709FF9BB5" blockId="18.[151,1437,1402,2041]" pageId="18" pageNumber="411">
<emphasis id="B954EAF3BC00087E94C5F9A70D4C9B49" bold="true" box="[199,314,1582,1608]" pageId="18" pageNumber="411">Diagnosis</emphasis>
: Body length 2.0–3.0 mm. Antennal article VI with 15–16 bacilliform sensilla forming crescent shape, setiform sensillum located in anterior position, conical sensillum present near centre of crescent. Labrum asetose, long slender anterior setae on anterior margin. Telotarsus with anterior and posterior lateral processes equal in length and shorter than half-length of claw, lamella processes present, anterior setiform process longer than claw.
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC00087894C5F9370D5F9A16" lastPageId="20" lastPageNumber="413" pageId="18" pageNumber="411" type="description">
<paragraph id="8B9F36E1BC00087E94C5F9370D2C9BD8" blockId="18.[151,1437,1402,2041]" box="[199,346,1726,1753]" pageId="18" pageNumber="411">
<emphasis id="B954EAF3BC00087E94C5F9370D279BD9" bold="true" box="[199,337,1726,1752]" pageId="18" pageNumber="411">Description</emphasis>
:
</paragraph>
<paragraph id="8B9F36E1BC00087E94C5F96B0D1C9A20" blockId="18.[151,1437,1402,2041]" pageId="18" pageNumber="411">
<typeStatus id="549B8843BC00087E94C5F96B0D599BFD" box="[199,303,1762,1788]" pageId="18" pageNumber="411" type="holotype">Holotype</typeStatus>
male body 2.0 mm;
<typeStatus id="549B8843BC00087E961BF96A0EFF9BFC" box="[537,649,1763,1789]" pageId="18" pageNumber="411" type="paratype">paratypes</typeStatus>
: males 2.0–
<quantity id="4CD89B04BC00087E970CF96B0F109BFC" box="[782,870,1762,1789]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="411" unit="mm" value="2.2">2.2 mm</quantity>
, females 2.4–3.0 mm; caudal bundle
<quantity id="4CD89B04BC00087E9109F96B09159BFC" box="[1291,1379,1762,1789]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="411" unit="mm" value="0.5">0.5 mm</quantity>
, nest trichomes
<quantity id="4CD89B04BC00087E950FF88F0D109A20" box="[269,358,1798,1825]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="411" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
<paragraph id="8B9F36E1BC00087E94C5F8A309349AF9" blockId="18.[151,1437,1402,2041]" pageId="18" pageNumber="411">
<emphasis id="B954EAF3BC00087E94C5F8A30D709A45" bold="true" box="[199,262,1834,1860]" pageId="18" pageNumber="411">Head</emphasis>
:
<materialsCitation id="3B483CBCBC00087E9511F8A309489AF9" ID-GBIF-Occurrence="3981599311" collectionCode="R" pageId="18" pageNumber="411" specimenCount="4" typeStatus="holotype">
Arrangement of post vertex trichomes as for
<taxonomicName id="4C204D62BC00087E9702F8A308699A45" authorityName="Huynh" authorityYear="2022" box="[768,1055,1834,1860]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="18" pageNumber="411" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B954EAF3BC00087E9702F8A308699A45" box="[768,1055,1834,1860]" italics="true" pageId="18" pageNumber="411">Monographis cattienensis</emphasis>
</taxonomicName>
with the
<typeStatus id="549B8843BC00087E9087F8A308909A45" box="[1157,1254,1834,1860]" pageId="18" pageNumber="411" type="holotype">holotype</typeStatus>
having 12 sockets on both sides in anterior rows, posterior rows with 4 sockets (L) and 3 sockets (
<collectionCode id="ED31AE24BC00087E9040F8C708219A69" box="[1090,1111,1870,1896]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="411">R</collectionCode>
) (
<figureCitation id="131B2A64BC00087E906CF8C709269A69" box="[1134,1360,1870,1896]" captionStart="FIGURE 18" captionStartId="19.[152,255,1782,1806]" captionTargetBox="[163,1435,317,1758]" captionTargetId="figure-79@19.[151,1436,308,1758]" captionTargetPageId="19" captionText="FIGURE 18. Holotype male Monographis thatsonensis sp. n.. A. A head capsule showing eye patterns with eight ommatidia and trichobothria (a, b, and c) each side, vertex region (v), anterior vertex (av), posterior vertex trichome sockets (pv); B. Enlarged images of posterior vertex (pv) trichome sockets on left (pv (L)) and right (pv (R)). C. Clypeo-labrum showing labrum (l) with asetose surface, lateral labrum (Ll), anterior setae (sa), backward-pointing setae (sb), and posterior setae (Se, 4 setae and 6 sockets of missing setae)." figureDoi="http://doi.org/10.5281/zenodo.7389346" httpUri="https://zenodo.org/record/7389346/files/figure.png" pageId="18" pageNumber="411">Figures 18A &amp; 18B</figureCitation>
,
<figureCitation id="131B2A64BC00087E915EF8C709E79A69" box="[1372,1425,1870,1896]" captionStart="FIGURE 22" captionStartId="22.[152,255,1902,1926]" captionTargetBox="[297,1291,1236,1879]" captionTargetId="figure-94@22.[297,1291,1236,1879]" captionTargetPageId="22" captionText="FIGURE 22. SEM images of Monographis thatsonensis sp. n.. A. The head—right side showing the posterior vertex (pv) trichome sockets and trichobothria (a, b, and c). B. Coxal glands on legs VIII and IX (pointed arrows). C. Clypeo-labrum showing structure of labrum (l) with asetose surface, lateral labrum (Ll), medial cleft (mc), anterior setae (sa), posterior setae (Se), backward-pointing setae (sb). D. Gnathochilaria (g): lateral palps (Lp) and median palps (Mp), the frons (f), labrum (l)." figureDoi="http://doi.org/10.5281/zenodo.7389358" httpUri="https://zenodo.org/record/7389358/files/figure.png" pageId="18" pageNumber="411">22A</figureCitation>
).
<typeStatus id="549B8843BC00087E9495F8FA0C8E9A8C" box="[151,248,1907,1933]" pageId="18" pageNumber="411" type="paratype">Paratype</typeStatus>
showed anterior rows with 12–16 sockets, posterior rows with 3–4, these setae shorter in length than labrum width (
<typeStatus id="549B8843BC00087E94E1F81F0D309AB1" box="[227,326,1942,1968]" pageId="18" pageNumber="411" type="paratype">Paratype</typeStatus>
with 8–10 setae).
<emphasis id="B954EAF3BC00087E960FF81F0EB79AB1" bold="true" box="[525,705,1942,1968]" pageId="18" pageNumber="411">Clypeo-labrum</emphasis>
:
<typeStatus id="549B8843BC00087E96CCF81F0F4E9AB1" box="[718,824,1942,1968]" pageId="18" pageNumber="411" type="holotype">Holotype</typeStatus>
, labrum surface asetose (l), labrum posterior margin has a row of minute backward-pointing setae (sb). Lateral lamella (Ll) and single broad lamella each side median cleft (mc), 12 (L) and 8 (
<collectionCode id="ED31AE24BC00087E9570F8570DF19AF9" box="[370,391,2014,2040]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="411">R</collectionCode>
) slender setae (sa) present on anterior margin of labrum (
<figureCitation id="131B2A64BC00087E9000F85708FA9AF9" box="[1026,1164,2014,2040]" captionStart="FIGURE 18" captionStartId="19.[152,255,1782,1806]" captionTargetBox="[163,1435,317,1758]" captionTargetId="figure-79@19.[151,1436,308,1758]" captionTargetPageId="19" captionText="FIGURE 18. Holotype male Monographis thatsonensis sp. n.. A. A head capsule showing eye patterns with eight ommatidia and trichobothria (a, b, and c) each side, vertex region (v), anterior vertex (av), posterior vertex trichome sockets (pv); B. Enlarged images of posterior vertex (pv) trichome sockets on left (pv (L)) and right (pv (R)). C. Clypeo-labrum showing labrum (l) with asetose surface, lateral labrum (Ll), anterior setae (sa), backward-pointing setae (sb), and posterior setae (Se, 4 setae and 6 sockets of missing setae)." figureDoi="http://doi.org/10.5281/zenodo.7389346" httpUri="https://zenodo.org/record/7389346/files/figure.png" pageId="18" pageNumber="411">Figures 18C</figureCitation>
&amp;
<figureCitation id="131B2A64BC00087E90B2F85709409AF9" box="[1200,1334,2014,2040]" captionStart="FIGURE 22" captionStartId="22.[152,255,1902,1926]" captionTargetBox="[297,1291,1236,1879]" captionTargetId="figure-94@22.[297,1291,1236,1879]" captionTargetPageId="22" captionText="FIGURE 22. SEM images of Monographis thatsonensis sp. n.. A. The head—right side showing the posterior vertex (pv) trichome sockets and trichobothria (a, b, and c). B. Coxal glands on legs VIII and IX (pointed arrows). C. Clypeo-labrum showing structure of labrum (l) with asetose surface, lateral labrum (Ll), medial cleft (mc), anterior setae (sa), posterior setae (Se), backward-pointing setae (sb). D. Gnathochilaria (g): lateral palps (Lp) and median palps (Mp), the frons (f), labrum (l)." figureDoi="http://doi.org/10.5281/zenodo.7389358" httpUri="https://zenodo.org/record/7389358/files/figure.png" pageId="18" pageNumber="411">22C &amp; 22D</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F36E1BC01087F94C5FF1F0EB99C1D" blockId="19.[151,1436,150,285]" pageId="19" pageNumber="412">
<emphasis id="B954EAF3BC01087F94C5FF1F0D4C9DB1" bold="true" box="[199,314,150,176]" pageId="19" pageNumber="412">Antennae</emphasis>
:
<materialsCitation id="3B483CBCBC01087F9545FF1F0EB99C1D" ID-GBIF-Occurrence="3981599301" collectionCode="VI, T" pageId="19" pageNumber="412" specimenCount="1" typeStatus="holotype">
Antennal article
<collectionCode id="ED31AE24BC01087F95FDFF1E0E6B9DB0" box="[511,541,151,177]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="19" pageNumber="412">VI</collectionCode>
with 15–16 bacilliform sensilla.
<typeStatus id="549B8843BC01087F9788FF1F0F849DB1" box="[906,1010,150,176]" pageId="19" pageNumber="412" type="holotype">Holotype</typeStatus>
with 15 bacilliform sensilla forming a crescent shape with 3 thick (
<collectionCode id="ED31AE24BC01087F95D7FF330D9F9DD5" box="[469,489,186,212]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="412">T</collectionCode>
), 3 thin bacilliform sensilla (t), 3 medium-length thin bacilliform sensilla (tm), and 6 short thin bacilliform sensilla (ts), setiform sensillum (S) located in anterior position, conical sensillum (C) present near centre of crescent (
<figureCitation id="131B2A64BC01087F95A0FE8B0E469C1D" box="[418,560,258,284]" captionStart="FIGURE 19" captionStartId="20.[152,255,1430,1454]" captionTargetBox="[159,1419,190,1401]" captionTargetId="figure-29@20.[151,1436,181,1406]" captionTargetPageId="20" captionText="FIGURE 19. Holotype male Monographis thatsonensis sp. n.. A. Antennal articles V, VI, VII and VIII with sensory cones (sc), and patterns of sensilla on article VI and VII; B. Article VII showing two thick bacilliform sensilla (Ta: anterior &amp; Tp: posterior), setiform (S), and a conical sensillum (C); C. Article VI showing three thick bacilliform sensilla (T), three thin bacilliform sensilla (t), three medium-length thin bacilliform sensilla (tm), and six short thin bacilliform sensilla (ts), a conical sensillum (C)." figureDoi="http://doi.org/10.5281/zenodo.7389348" httpUri="https://zenodo.org/record/7389348/files/figure.png" pageId="19" pageNumber="412">Figures 19C</figureCitation>
,
<figureCitation id="131B2A64BC01087F963EFE8B0EB59C1D" box="[572,707,258,284]" captionStart="FIGURE 21" captionStartId="22.[152,255,1079,1103]" captionTargetBox="[280,1307,181,1055]" captionTargetId="figure-29@22.[280,1307,181,1055]" captionTargetPageId="22" captionText="FIGURE 21. SEM images of Monographis thatsonensis sp. n.. A. Head capsule showing left antenna with eight articles, eye pattern with eight ommatidia, and the trichobothria. B. Antennal articles VI–VIII, showing the arrangement of sensilla. C. Antennal articles VII showing the typical pattern of sensilla and VIII with sensory cones (sc). D. Arrangement of sensilla on article VI." figureDoi="http://doi.org/10.5281/zenodo.7389354" httpUri="https://zenodo.org/record/7389354/files/figure.png" pageId="19" pageNumber="412">21B &amp; 21D</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="DF5F6669BC01087F949AF97F0DD89A9F" ID-DOI="http://doi.org/10.5281/zenodo.7389346" ID-Zenodo-Dep="7389346" httpUri="https://zenodo.org/record/7389346/files/figure.png" pageId="19" pageNumber="412" startId="19.[152,255,1782,1806]" targetBox="[163,1435,317,1758]" targetPageId="19">
<paragraph id="8B9F36E1BC01087F949AF97F0DD89A9F" blockId="19.[151,1437,1782,1950]" pageId="19" pageNumber="412">
<emphasis id="B954EAF3BC01087F949AF97F0D509A0F" bold="true" box="[152,294,1782,1806]" pageId="19" pageNumber="412">FIGURE 18.</emphasis>
Holotype male
<taxonomicName id="4C204D62BC01087F95C9F97F0EA19A0F" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[459,727,1782,1806]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="19" pageNumber="412" phylum="Arthropoda" rank="species" species="thatsonensis" status="sp. nov.">
<emphasis id="B954EAF3BC01087F95C9F97F0EA19A0F" box="[459,727,1782,1806]" italics="true" pageId="19" pageNumber="412">Monographis thatsonensis</emphasis>
</taxonomicName>
<emphasis id="B954EAF3BC01087F96DDF97F0F6F9A0F" bold="true" box="[735,793,1782,1806]" pageId="19" pageNumber="412">
<taxonomicNameLabel id="A2675788BC01087F96DDF97F0F6F9A0F" box="[735,793,1782,1806]" pageId="19" pageNumber="412" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B954EAF3BC01087F9724F97F0F499A0F" bold="true" box="[806,831,1782,1806]" pageId="19" pageNumber="412">A.</emphasis>
A head capsule showing eye patterns with eight ommatidia and trichobothria (
<emphasis id="B954EAF3BC01087F9556F8920D179A33" bold="true" box="[340,353,1819,1842]" italics="true" pageId="19" pageNumber="412">a</emphasis>
,
<emphasis id="B954EAF3BC01087F956FF8930D0C9A33" bold="true" box="[365,378,1818,1842]" italics="true" pageId="19" pageNumber="412">b</emphasis>
, and
<emphasis id="B954EAF3BC01087F95B3F8920DCA9A33" bold="true" box="[433,444,1819,1842]" italics="true" pageId="19" pageNumber="412">c</emphasis>
) each side, vertex region (v), anterior vertex (av), posterior vertex trichome sockets (pv);
<emphasis id="B954EAF3BC01087F915BF89309199A33" bold="true" box="[1369,1391,1818,1842]" pageId="19" pageNumber="412">B.</emphasis>
Enlarged images of posterior vertex (pv) trichome sockets on left (pv (L)) and right (pv (R)).
<emphasis id="B954EAF3BC01087F9038F8B708249A57" bold="true" box="[1082,1106,1854,1878]" pageId="19" pageNumber="412">C.</emphasis>
Clypeo-labrum showing labrum (l) with asetose surface, lateral labrum (Ll), anterior setae (sa), backward-pointing setae (sb), and posterior setae (Se, 4 setae and 6 sockets of missing setae).
</paragraph>
</caption>
<caption id="DF5F6669BC060878949AFA1F09E49B1A" ID-DOI="http://doi.org/10.5281/zenodo.7389348" ID-Zenodo-Dep="7389348" httpUri="https://zenodo.org/record/7389348/files/figure.png" pageId="20" pageNumber="413" startId="20.[152,255,1430,1454]" targetBox="[159,1419,190,1401]" targetPageId="20">
<paragraph id="8B9F36E1BC060878949AFA1F09E49B1A" blockId="20.[151,1437,1430,1563]" pageId="20" pageNumber="413">
<emphasis id="B954EAF3BC060878949AFA1F0D5398AF" bold="true" box="[152,293,1430,1454]" pageId="20" pageNumber="413">FIGURE 19.</emphasis>
Holotype male
<taxonomicName id="4C204D62BC06087895C4FA1F0EA798AF" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[454,721,1430,1454]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="413" phylum="Arthropoda" rank="species" species="thatsonensis" status="sp. nov.">
<emphasis id="B954EAF3BC06087895C4FA1F0EA798AF" box="[454,721,1430,1454]" italics="true" pageId="20" pageNumber="413">Monographis thatsonensis</emphasis>
</taxonomicName>
<emphasis id="B954EAF3BC06087896D5FA1E0F6698AE" bold="true" box="[727,784,1431,1455]" pageId="20" pageNumber="413">
<taxonomicNameLabel id="A2675788BC06087896D5FA1E0F6698AE" box="[727,784,1431,1455]" pageId="20" pageNumber="413" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B954EAF3BC060878971EFA1F0F4398AF" bold="true" box="[796,821,1430,1454]" pageId="20" pageNumber="413">A.</emphasis>
Antennal articles V, VI, VII and VIII with sensory cones (sc), and patterns of sensilla on article VI and VII;
<emphasis id="B954EAF3BC060878965AFA320E1F98D2" bold="true" box="[600,617,1467,1491]" pageId="20" pageNumber="413">B</emphasis>
. Article VII showing two thick bacilliform sensilla (Ta: anterior &amp; Tp: posterior), setiform (S), and a conical sensillum (C);
<emphasis id="B954EAF3BC0608789640FA570E2D98F7" bold="true" box="[578,603,1502,1526]" pageId="20" pageNumber="413">C.</emphasis>
Article VI showing three thick bacilliform sensilla (T), three thin bacilliform sensilla (t), three medium-length thin bacilliform sensilla (tm), and six short thin bacilliform sensilla (ts), a conical sensillum (C).
</paragraph>
</caption>
<paragraph id="8B9F36E1BC06087894C5F9C00CAE9BAA" blockId="20.[151,1437,1609,1995]" pageId="20" pageNumber="413">
<emphasis id="B954EAF3BC06087894C5F9C00D349B62" bold="true" box="[199,322,1609,1635]" pageId="20" pageNumber="413">Telotarsus</emphasis>
: Claw slender, anterior (a) and posterior lateral (p) processes equal in length and shorter than halflength of claw (c), lamella processes (la) present, anterior setiform process (s) longer than claw (
<figureCitation id="131B2A64BC06087890F2F9E409089B86" box="[1264,1406,1645,1671]" captionStart="FIGURE 20" captionStartId="21.[152,255,1896,1920]" captionTargetBox="[163,1413,191,1870]" captionTargetId="figure-19@21.[158,1430,181,1872]" captionTargetPageId="21" captionText="FIGURE 20. Holotype male Monographis thatsonensis sp. n.. A. Second leg showing seven articles: coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2); claw (c) and penis (p). B. Telotarsus showing claw (c) and all processes: anterior (a), posterior (p), setiform (s) and the lamella (la). C. Ornamental trichome sockets showing the arrangements on both sides: trichome sockets a, b, c, and indentation d." figureDoi="http://doi.org/10.5281/zenodo.7389352" httpUri="https://zenodo.org/record/7389352/files/figure.png" pageId="20" pageNumber="413">Figures 20B</figureCitation>
&amp;
<figureCitation id="131B2A64BC0608789495F9180CBD9BAA" box="[151,203,1681,1707]" captionStart="FIGURE 23" captionStartId="23.[152,255,1170,1194]" captionTargetBox="[152,1436,181,1146]" captionTargetId="figure-19@23.[151,1436,181,1146]" captionTargetPageId="23" captionText="FIGURE 23. SEM images of Monographis thatsonensis sp. n.. A. Male’s caudal bundle with ornamental trichome sockets (ots) a, b, c and d; and telson (Te), and the bundle with caudal bundle trichome sockets (cbts), barbate trichome sockets (bts); the last pleural projection (pp). B. Female’s caudal bundle structure showing the same dorsally, except the presence of two bundles of nest trichome sockets ventrally (nts). C. Presence of two spines—Tarsus 1 and 2, typical of Monographis species. D. Telotarsus showing a claw (c) and all processes: anterior (a), posterior (p), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389361" httpUri="https://zenodo.org/record/7389361/files/figure.png" pageId="20" pageNumber="413">23D</figureCitation>
).
</paragraph>
<paragraph id="8B9F36E1BC06087894C5F93C0D5F9A16" blockId="20.[151,1437,1609,1995]" pageId="20" pageNumber="413">
<emphasis id="B954EAF3BC06087894C5F93C0D629BCE" bold="true" box="[199,276,1717,1743]" pageId="20" pageNumber="413">Telson</emphasis>
: Arrangement of dorsal ornamental trichome sockets similar to
<taxonomicName id="4C204D62BC06087897DAF93C08F69BCF" authorityName="Huynh" authorityYear="2022" box="[984,1152,1717,1742]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="413" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B954EAF3BC06087897DAF93C08F69BCF" box="[984,1152,1717,1742]" italics="true" pageId="20" pageNumber="413">M. cattienensis</emphasis>
</taxonomicName>
with 14 trichome sockets
<emphasis id="B954EAF3BC0608789495F9500CD39BF2" bold="true" box="[151,165,1753,1779]" italics="true" pageId="20" pageNumber="413">a</emphasis>
left side and 15 sockets on right side of the telson in the
<typeStatus id="549B8843BC0608789730F9500FE29BF2" box="[818,916,1753,1779]" pageId="20" pageNumber="413" type="holotype">holotype</typeStatus>
(
<typeStatus id="549B8843BC06087897A6F95008709BF2" box="[932,1030,1753,1779]" pageId="20" pageNumber="413" type="paratype">paratype</typeStatus>
with 14–18 sockets). (
<figureCitation id="131B2A64BC060878910BF95009EE9BF2" box="[1289,1432,1753,1779]" captionStart="FIGURE 20" captionStartId="21.[152,255,1896,1920]" captionTargetBox="[163,1413,191,1870]" captionTargetId="figure-19@21.[158,1430,181,1872]" captionTargetPageId="21" captionText="FIGURE 20. Holotype male Monographis thatsonensis sp. n.. A. Second leg showing seven articles: coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2); claw (c) and penis (p). B. Telotarsus showing claw (c) and all processes: anterior (a), posterior (p), setiform (s) and the lamella (la). C. Ornamental trichome sockets showing the arrangements on both sides: trichome sockets a, b, c, and indentation d." figureDoi="http://doi.org/10.5281/zenodo.7389352" httpUri="https://zenodo.org/record/7389352/files/figure.png" pageId="20" pageNumber="413">Figures 20C</figureCitation>
,
<figureCitation id="131B2A64BC0608789495F9740D6A9A16" box="[151,284,1789,1815]" captionStart="FIGURE 23" captionStartId="23.[152,255,1170,1194]" captionTargetBox="[152,1436,181,1146]" captionTargetId="figure-19@23.[151,1436,181,1146]" captionTargetPageId="23" captionText="FIGURE 23. SEM images of Monographis thatsonensis sp. n.. A. Male’s caudal bundle with ornamental trichome sockets (ots) a, b, c and d; and telson (Te), and the bundle with caudal bundle trichome sockets (cbts), barbate trichome sockets (bts); the last pleural projection (pp). B. Female’s caudal bundle structure showing the same dorsally, except the presence of two bundles of nest trichome sockets ventrally (nts). C. Presence of two spines—Tarsus 1 and 2, typical of Monographis species. D. Telotarsus showing a claw (c) and all processes: anterior (a), posterior (p), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389361" httpUri="https://zenodo.org/record/7389361/files/figure.png" pageId="20" pageNumber="413">23A &amp; 23B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC06087894C5F8A80E619A82" pageId="20" pageNumber="413" type="discussion">
<paragraph id="8B9F36E1BC06087894C5F8A80E619A82" blockId="20.[151,1437,1609,1995]" pageId="20" pageNumber="413">
<emphasis id="B954EAF3BC06087894C5F8A80D429A3A" bold="true" box="[199,308,1825,1851]" pageId="20" pageNumber="413">Remarks</emphasis>
:
<taxonomicName id="4C204D62BC0608789546F8A80E1A9A3A" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[324,620,1825,1851]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="413" phylum="Arthropoda" rank="species" species="thatsonensis" status="sp. nov.">
<emphasis id="B954EAF3BC0608789546F8A80E1A9A3A" box="[324,620,1825,1851]" italics="true" pageId="20" pageNumber="413">Monographis thatsonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2675788BC0608789677F8A80EC29A3A" box="[629,692,1825,1851]" pageId="20" pageNumber="413" rank="species">
<emphasis id="B954EAF3BC0608789677F8A80ED89A3A" bold="true" box="[629,686,1825,1851]" pageId="20" pageNumber="413">sp. n</emphasis>
.
</taxonomicNameLabel>
showed a distinctive pattern of sensilla on antennal article VI, labrum structures, anterior and posterior lateral processes equal in length and shorter than half-length of claw, anterior setiform process longer than claw.
</paragraph>
</subSubSection>
<subSubSection id="C33A656ABC06087894C5F8040F359ACA" pageId="20" pageNumber="413" type="etymology">
<paragraph id="8B9F36E1BC06087894C5F8040F359ACA" blockId="20.[151,1437,1609,1995]" pageId="20" pageNumber="413">
<emphasis id="B954EAF3BC06087894C5F8040D3E9AA6" bold="true" box="[199,328,1933,1959]" pageId="20" pageNumber="413">Etymology</emphasis>
:
<taxonomicName id="4C204D62BC060878955BF8040EF59AA6" authorityName="Huynh" authorityYear="2022" box="[345,643,1933,1959]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="20" pageNumber="413" phylum="Arthropoda" rank="species" species="thatsonensis">
<emphasis id="B954EAF3BC060878955BF8040EF59AA6" box="[345,643,1933,1959]" italics="true" pageId="20" pageNumber="413">Monographis thatsonensis</emphasis>
</taxonomicName>
is named after the Seven Mountain Ranges, derived from Sino-Vietnamese, meaning: That Son, in
<collectingRegion id="49E4F803BC06087895F5F8380EA49ACA" box="[503,722,1969,1995]" country="Vietnam" name="An Giang" pageId="20" pageNumber="413">An Giang Province</collectingRegion>
,
<collectingCountry id="F3377671BC06087896DFF8380F369ACA" box="[733,832,1969,1995]" name="Vietnam" pageId="20" pageNumber="413">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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