407 lines
71 KiB
XML
407 lines
71 KiB
XML
<document id="A06AF1F5632C410A274D7EAA27D61A45" ID-CLB-Dataset="53542" ID-DOI="10.11646/zootaxa.5297.1.5" ID-GBIF-Dataset="326fb044-60e8-49a6-b043-5a2f9a91da92" ID-ISSN="1175-5326" ID-Zenodo-Dep="7989367" ID-ZooBank="7C3B9BE1-C23A-4416-893E-B5E68A20BD75" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685520503012" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Lin, Chia-Wei" docDate="2023" docId="038987C9300FFF8CFF0CFAACFAC7ECCE" docLanguage="en" docName="zootaxa.5297.1.5.pdf" docOrigin="Zootaxa 5297 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteropilumnus planus Ng & Lin 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="106" masterDocId="FFB0FFB1300EFF89FF9BFFF3FFADEE4B" masterDocTitle="Two new species of hairy crabs (Pilumnidae) and the first record of Crinitocinus alcocki (Borradaile, 1900) (Acidopsidae) (Crustacea: Brachyura) from Taiwan" masterLastPageNumber="114" masterPageNumber="101" pageNumber="102" updateTime="1699512094808" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DA526B4C8541829D412CF278F7BC9F0F">Two new species of hairy crabs (Pilumnidae) and the first record of Crinitocinus alcocki (Borradaile, 1900) (Acidopsidae) (Crustacea: Brachyura) from Taiwan</mods:title>
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<mods:name id="11F495BB7E5A4A4EBF341C680B7A4E24" type="personal">
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<mods:roleTerm id="1B4F8DE52E3CC85F058DB816E3C0A8A8">Author</mods:roleTerm>
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<mods:namePart id="620BC367420562BAD659BDDE20EE459C">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="00817FE9C3D4DC1A88C56F2107EC9344">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:roleTerm id="3C8DB09605631B98E21DD8B329B75690">Author</mods:roleTerm>
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<mods:namePart id="ACA37E43A9BCB8D446EB327D9FA9A13A">Lin, Chia-Wei</mods:namePart>
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<mods:affiliation id="46685E416573DA0AEBBDB720333A7C0E">National Museum of Marine Biology and Aquarium, 2 Houwan Road, Checheng, Pingtung, Taiwan 944, Taiwan. y 9300923 @ ms 8. hinet. net; https: // orcid. org / 0000 - 0002 - 8142 - 4491</mods:affiliation>
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<mods:title id="14690C0F3236DA46B5C3F221D065E7E8">Zootaxa</mods:title>
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<mods:date id="F29C552851C6F4AAF1FD15984CAB2969">2023</mods:date>
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<mods:number id="D3F2CCAE65039B53FCF7CCDB8A39F85F">2023-05-31</mods:number>
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<treatment id="038987C9300FFF8CFF0CFAACFAC7ECCE" ID-DOI="http://doi.org/10.5281/zenodo.7989345" ID-GBIF-Taxon="209664024" ID-Zenodo-Dep="7989345" LSID="urn:lsid:plazi:treatment:038987C9300FFF8CFF0CFAACFAC7ECCE" httpUri="http://treatment.plazi.org/id/038987C9300FFF8CFF0CFAACFAC7ECCE" lastPageId="5" lastPageNumber="106" pageId="1" pageNumber="102">
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<subSubSection id="C33A6554300FFF88FF0CFAACFE52EB32" box="[151,511,1375,1402]" pageId="1" pageNumber="102" type="nomenclature">
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<paragraph id="8B9F36DF300FFF88FF0CFAACFE52EB32" blockId="1.[151,511,1375,1437]" box="[151,511,1375,1402]" pageId="1" pageNumber="102">
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<heading id="D0D781B3300FFF88FF0CFAACFE52EB32" bold="true" box="[151,511,1375,1402]" fontSize="11" level="1" pageId="1" pageNumber="102" reason="1">
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<emphasis id="B954EACD300FFF88FF0CFAACFE52EB32" bold="true" box="[151,511,1375,1402]" pageId="1" pageNumber="102">
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<taxonomicName id="4C204D5C300FFF88FF0CFAACFE1BEB31" ID-CoL="f7719545-8c93-4107-b3a5-201893c40f32" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[151,438,1375,1402]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
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<emphasis id="B954EACD300FFF88FF0CFAACFE1BEB31" bold="true" box="[151,438,1375,1402]" italics="true" pageId="1" pageNumber="102">Heteropilumnus planus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26757B6300FFF88FE26FAACFE52EB32" box="[445,511,1375,1401]" pageId="1" pageNumber="102" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A6554300FFF88FF0CFA70FE28EBD6" box="[151,389,1411,1437]" pageId="1" pageNumber="102" type="description">
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<paragraph id="8B9F36DF300FFF88FF0CFA70FE28EBD6" blockId="1.[151,511,1375,1437]" box="[151,389,1411,1437]" pageId="1" pageNumber="102">
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(
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<figureCitation id="131B2A5A300FFF88FF04FA70FEEAEBD6" box="[159,327,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1887]" captionTargetBox="[151,1436,273,1840]" captionTargetId="figure-68@2.[151,1436,273,1840]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life. A, G, Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625); B, H. planus n. sp., paratype female (11.4 × 8.2 mm) (NMMBCD5626); C, H. planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044); D, Pseudolitochira taiwang n. sp., holotype female (11.2 × 8.1 mm) (NMMBCD5627); E, Crinitocinus alcocki (Borradaile, 1902), male (9.3 × 7.9 mm) (ZRC 2022.0825); F, H, C. alcocki (Borradaile, 1902), male (6.8 × 6.2 mm) (NMMBCD5628). G, H, in situ in habitat; H at night with filter. All specimens from Taiwan." figureDoi="http://doi.org/10.5281/zenodo.7989377" httpUri="https://zenodo.org/record/7989377/files/figure.png" pageId="1" pageNumber="102">Figs. 1A–C, G</figureCitation>
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,
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<figureCitation id="131B2A5A300FFF88FEC9FA70FED0EBD6" box="[338,381,1411,1437]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1874,1898]" captionStartId-1="4.[151,250,877,901]" captionStartId-2="6.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1850]" captionTargetBox-1="[153,1434,183,851]" captionTargetBox-2="[183,1405,181,1919]" captionTargetId-0="figure-17@3.[151,1436,181,1850]" captionTargetId-1="figure-18@4.[151,1436,181,853]" captionTargetId-2="figure-18@6.[183,1405,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." captionText-1="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." captionText-2="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi-0="http://doi.org/10.5281/zenodo.7989383" figureDoi-1="http://doi.org/10.5281/zenodo.7989387" figureDoi-2="http://doi.org/10.5281/zenodo.7989393" httpUri-0="https://zenodo.org/record/7989383/files/figure.png" httpUri-1="https://zenodo.org/record/7989387/files/figure.png" httpUri-2="https://zenodo.org/record/7989393/files/figure.png" pageId="1" pageNumber="102">2–4</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C33A6554300FFF88FF0CFA38FE00E81A" pageId="1" pageNumber="102" type="materials_examined">
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<paragraph id="8B9F36DF300FFF88FF0CFA38FE00E81A" blockId="1.[151,1437,1483,2014]" pageId="1" pageNumber="102">
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||
<emphasis id="B954EACD300FFF88FF0CFA38FEE9EBAE" bold="true" box="[151,324,1483,1509]" pageId="1" pageNumber="102">Type material.</emphasis>
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||
<materialsCitation id="3B483C82300FFF88FED2FA38FE46E842" ID-GBIF-Occurrence="4121719304" collectingDate="2018-09-01" collectionCode="SCUBA" collectorName="C. - W. Lin" country="Taiwan" location="Toucheng" pageId="1" pageNumber="102" specimenCode="NMMBCD5625" specimenCount="1" stateProvince="Yilan" typeStatus="holotype">
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<typeStatus id="549B887D300FFF88FED2FA38FE1BEBAE" box="[329,438,1483,1509]" pageId="1" pageNumber="102" type="holotype">Holotype</typeStatus>
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: male (10.7 × 8.0 mm) (
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<specimenCode id="DB869EA4300FFF88FD20FA38FCD4EBAD" box="[699,889,1483,1510]" collectionCode="NMMBCD" pageId="1" pageNumber="102">NMMBCD5625</specimenCode>
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),
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||
<location id="8EFF6004300FFF88FC11FA38FC56EBAE" LSID="urn:lsid:plazi:treatment:038987C9300FFF8CFF0CFAACFAC7ECCE:8EFF6004300FFF88FC11FA38FC56EBAE" box="[906,1019,1483,1509]" country="Taiwan" name="Toucheng" pageId="1" pageNumber="102" stateProvince="Yilan">Toucheng</location>
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||
,
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<collectingRegion id="49E4F83D300FFF88FB98FA38FB37EBAE" box="[1027,1178,1483,1509]" country="Taiwan" name="Yilan" pageId="1" pageNumber="102">Yilan County</collectingRegion>
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,
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<collectingCountry id="F337764F300FFF88FB38FA38FB55EBAE" box="[1187,1272,1483,1509]" name="Taiwan" pageId="1" pageNumber="102">Taiwan</collectingCountry>
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||
, coll.
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<collectionCode id="ED31AE1A300FFF88FAA3FA38FA35EBAE" box="[1336,1432,1483,1509]" pageId="1" pageNumber="102">SCUBA</collectionCode>
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,
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<collectorName id="26D55309300FFF88FF0CFA1CFEA6E842" box="[151,267,1519,1545]" pageId="1" pageNumber="102">C.-W. Lin</collectorName>
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,
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<date id="FF9E101F300FFF88FE83FA1CFE46E842" box="[280,491,1519,1545]" pageId="1" pageNumber="102" value="2018-09-01">
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||
<collectingDate id="EFDAE9F7300FFF88FE83FA1CFE46E842" box="[280,491,1519,1545]" pageId="1" pageNumber="102" value="2018-09-01">1 September 2018</collectingDate>
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</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B483C82300FFF88FE6DFA03FA2FE842" ID-GBIF-Occurrence="4121719306" box="[502,1410,1519,1546]" collectingDate="2018-09-01" collectionCode="NMMBCD" collectorName="C. - W. Lin" country="Taiwan" location="Toucheng" pageId="1" pageNumber="102" specimenCode="NMMBCD 5626" specimenCount="1" specimenCount-female="1" stateProvince="Yilan" typeStatus="paratype">
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<typeStatus id="549B887D300FFF88FE6DFA03FDCAE841" box="[502,615,1520,1546]" pageId="1" pageNumber="102" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D26FD56300FFF88FDE8FA1CFD7BE842" box="[627,726,1519,1545]" pageId="1" pageNumber="102" type="female">1 female</specimenCount>
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||
(11.4 ×
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<quantity id="4CD89B3A300FFF88FCA3FA1CFC3CE842" box="[824,913,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="102" unit="mm" value="8.2">8.2 mm</quantity>
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||
) (
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||
<specimenCode id="DB869EA4300FFF88FC33FA1CFBC3E841" box="[936,1134,1519,1546]" collectionCode="NMMBCD" pageId="1" pageNumber="102">NMMBCD 5626</specimenCode>
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||
), same data as holotype
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B483C82300FFF88FA15FA1CFE07E81A" ID-GBIF-Occurrence="4121719308" collectingDate="2014-07-18" collectionCode="SCUBA" collectorName="C. - W. Lin" country="Taiwan" location="Northeast Cape" pageId="1" pageNumber="102" specimenCode="ZRC 2017.1044" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<specimenCount id="9D26FD56300FFF88FA15FA1CFF4EE866" pageId="1" pageNumber="102" type="female">1 female</specimenCount>
|
||
(10.5 ×
|
||
<quantity id="4CD89B3A300FFF88FEDCF9E0FE0DE866" box="[327,416,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="102" unit="mm" value="7.9">7.9 mm</quantity>
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||
) (
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||
<specimenCode id="DB869EA4300FFF88FE2CF9E0FDD9E866" box="[439,628,1555,1581]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="102">ZRC 2017.1044</specimenCode>
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||
), muddy substrate under rocks,
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||
<location id="8EFF6004300FFF88FC44F9E0FB3DE866" LSID="urn:lsid:plazi:treatment:038987C9300FFF8CFF0CFAACFAC7ECCE:8EFF6004300FFF88FC44F9E0FB3DE866" box="[991,1168,1555,1581]" country="Taiwan" name="Northeast Cape" pageId="1" pageNumber="102">Northeast Cape</location>
|
||
,
|
||
<collectingCountry id="F337764F300FFF88FB06F9E0FB5FE866" box="[1181,1266,1555,1581]" name="Taiwan" pageId="1" pageNumber="102">Taiwan</collectingCountry>
|
||
, coll.
|
||
<collectionCode id="ED31AE1A300FFF88FAA3F9E0FA35E866" box="[1336,1432,1555,1581]" pageId="1" pageNumber="102">SCUBA</collectionCode>
|
||
,
|
||
<collectorName id="26D55309300FFF88FF0CF9C4FEA7E81A" box="[151,266,1591,1617]" pageId="1" pageNumber="102">C.-W. Lin</collectorName>
|
||
,
|
||
<date id="FF9E101F300FFF88FE8EF9C4FE07E81A" box="[277,426,1591,1617]" pageId="1" pageNumber="102" value="2014-07-18">
|
||
<collectingDate id="EFDAE9F7300FFF88FE8EF9C4FE07E81A" box="[277,426,1591,1617]" pageId="1" pageNumber="102" value="2014-07-18">18 July 2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6554300FFF8DFF5CF9A8FB47EA16" lastPageId="4" lastPageNumber="105" pageId="1" pageNumber="102" type="diagnosis">
|
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<paragraph id="8B9F36DF300FFF8BFF5CF9A8FB25EEB2" blockId="1.[151,1437,1483,2014]" lastBlockId="2.[151,1437,151,249]" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102">
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<emphasis id="B954EACD300FFF88FF5CF9A8FEEFE83E" bold="true" box="[199,322,1627,1653]" pageId="1" pageNumber="102">Diagnosis.</emphasis>
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Carapace subhexagonal, wider than long; dorsal surface of carapace, chelipeds and ambulatory legs with low, covered with felt-like tomentum; anterior part of carapace and anterolateral margins with relatively dense, long, plumose setae, obscuring surface, setae lining postfrontal ridge relatively shorter, more plumose; carapace not high, with dorsal surface gently convex to almost flat in frontal view (
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<figureCitation id="131B2A5A300FFF88FC32F934FBA6E8AA" box="[937,1035,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">Figs. 2C</figureCitation>
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,
|
||
<figureCitation id="131B2A5A300FFF88FB8CF934FB91E8AA" box="[1047,1084,1735,1761]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="1" pageNumber="102">4C</figureCitation>
|
||
); anterolateral margin with low external orbital tooth and 3 low anterolateral teeth separated by clear concavities; first anterolateral tooth relatively low, triangular in shape (
|
||
<figureCitation id="131B2A5A300FFF88FE35F8FCFDBCE962" box="[430,529,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300FFF88FD86F8FCFDECE962" box="[541,577,1807,1833]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="1" pageNumber="102">4B</figureCitation>
|
||
); median lobes of posterior margin of epistome projecting downwards, truncate in shape with each margin concave, separated from lateral part by deep V-shaped cleft (
|
||
<figureCitation id="131B2A5A300FFF88FB05F8C0FAB2E905" box="[1182,1311,1843,1870]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">Figs. 2C, E</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300FFF88FAB0F8C0FAE0E906" box="[1323,1357,1843,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="1" pageNumber="102">4C</figureCitation>
|
||
).Adult male chelipeds heterochelous, outer surface of major chela smooth and glabrous on ventral half (
|
||
<figureCitation id="131B2A5A300FFF88FB46F8A4FAEEE93A" box="[1245,1347,1879,1905]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">Figs. 2A</figureCitation>
|
||
); dorsal surface of cheliped carpus and chela with mixture of long and short plumose setae obscuring surface. Extensor and flexor margins of ambulatory legs with mixture of long and short plumose setae obscuring margins (
|
||
<figureCitation id="131B2A5A300FFF88FA89F86CFA3AE9F2" box="[1298,1431,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1887]" captionTargetBox="[151,1436,273,1840]" captionTargetId="figure-68@2.[151,1436,273,1840]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life. A, G, Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625); B, H. planus n. sp., paratype female (11.4 × 8.2 mm) (NMMBCD5626); C, H. planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044); D, Pseudolitochira taiwang n. sp., holotype female (11.2 × 8.1 mm) (NMMBCD5627); E, Crinitocinus alcocki (Borradaile, 1902), male (9.3 × 7.9 mm) (ZRC 2022.0825); F, H, C. alcocki (Borradaile, 1902), male (6.8 × 6.2 mm) (NMMBCD5628). G, H, in situ in habitat; H at night with filter. All specimens from Taiwan." figureDoi="http://doi.org/10.5281/zenodo.7989377" httpUri="https://zenodo.org/record/7989377/files/figure.png" pageId="1" pageNumber="102">Figs. 1A–C</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300FFF88FF0CF830FEEEE996" box="[151,323,1987,2014]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">2A, C, E, G, H</figureCitation>
|
||
); meri relatively long, slender (
|
||
<figureCitation id="131B2A5A300FFF88FD3FF830FC81E995" box="[676,812,1987,2014]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="1" pageNumber="102">Figs. 2A, H</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300FFF88FCACF830FCD6E996" box="[823,891,1987,2013]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="1" pageNumber="102">4A, F</figureCitation>
|
||
).
|
||
<collectingRegion id="49E4F83D300FFF88FC14F830FC64E996" box="[911,969,1987,2013]" country="Maldives" name="Maale" pageId="1" pageNumber="102">Male</collectingRegion>
|
||
pleon triangular, somite 1 subrectangular reaching to coxae of last ambulatory legs, somite 3 trapezoidal with rounded lateral margins, as wide as somite 1 (
|
||
<figureCitation id="131B2A5A300CFF8BFF04FF48FF54EE9E" box="[159,249,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,877,901]" captionTargetBox="[153,1434,183,851]" captionTargetId="figure-18@4.[151,1436,181,853]" captionTargetPageId="4" captionText="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="2" pageNumber="103">Fig. 3A</figureCitation>
|
||
); telson relatively short, semicircular in shape (
|
||
<figureCitation id="131B2A5A300CFF8BFC9DFF48FCCFEE9E" box="[774,866,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="2" pageNumber="103">Figs. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300CFF8BFCF7FF48FC3FEE9E" box="[876,914,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,877,901]" captionTargetBox="[153,1434,183,851]" captionTargetId="figure-18@4.[151,1436,181,853]" captionTargetPageId="4" captionText="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="2" pageNumber="103">3A</figureCitation>
|
||
). G1 gently sinuous, with proximal half almost straight with only distal half curved, distal part distinctly hooked downwards (
|
||
<figureCitation id="131B2A5A300CFF8BFB98FF2CFBD6EEB2" box="[1027,1147,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,877,901]" captionTargetBox="[153,1434,183,851]" captionTargetId="figure-18@4.[151,1436,181,853]" captionTargetPageId="4" captionText="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="2" pageNumber="103">Fig. 3B–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5F6657300CFF8BFF0CF8B4FBEEE9A4" ID-DOI="http://doi.org/10.5281/zenodo.7989377" ID-Zenodo-Dep="7989377" httpUri="https://zenodo.org/record/7989377/files/figure.png" pageId="2" pageNumber="103" startId="2.[151,250,1863,1887]" targetBox="[151,1436,273,1840]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B9F36DF300CFF8BFF0CF8B4FBEEE9A4" blockId="2.[151,1437,1863,2031]" pageId="2" pageNumber="103">
|
||
<emphasis id="B954EACD300CFF8BFF0CF8B4FEBDE914" bold="true" box="[151,272,1863,1887]" pageId="2" pageNumber="103">FIGURE 1</emphasis>
|
||
. Colour in life. A, G,
|
||
<taxonomicName id="4C204D5C300CFF8BFD9AF8B4FD5CE914" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[513,753,1863,1887]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300CFF8BFD9AF8B4FD5CE914" box="[513,753,1863,1887]" italics="true" pageId="2" pageNumber="103">Heteropilumnus planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300CFF8BFD61F8B4FC9BE914" bold="true" box="[762,822,1863,1887]" pageId="2" pageNumber="103">
|
||
<taxonomicNameLabel id="A26757B6300CFF8BFD61F8B4FC9BE914" box="[762,822,1863,1887]" pageId="2" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (10.7 × 8.0 mm) (NMMBCD5625); B,
|
||
<taxonomicName id="4C204D5C300CFF8BFA1FF8BBFF76E9C8" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300CFF8BFA1FF8BBFF76E9C8" italics="true" pageId="2" pageNumber="103">H. planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300CFF8BFF7FF898FE8CE9C8" bold="true" box="[228,289,1899,1923]" pageId="2" pageNumber="103">
|
||
<taxonomicNameLabel id="A26757B6300CFF8BFF7FF898FE8CE9C8" box="[228,289,1899,1923]" pageId="2" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype female (11.4 × 8.2 mm) (NMMBCD5626); C,
|
||
<taxonomicName id="4C204D5C300CFF8BFC1AF89FFC45E9C8" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[897,1000,1899,1923]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300CFF8BFC1AF89FFC45E9C8" box="[897,1000,1899,1923]" italics="true" pageId="2" pageNumber="103">H. planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300CFF8BFC6AF898FB80E9C8" bold="true" box="[1009,1069,1899,1923]" pageId="2" pageNumber="103">
|
||
<taxonomicNameLabel id="A26757B6300CFF8BFC6AF898FB80E9C8" box="[1009,1069,1899,1923]" pageId="2" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype female (10.5 × 7.9 mm) (ZRC 2017.1044); D,
|
||
<taxonomicName id="4C204D5C300CFF8BFE1BF87CFDD0E9EC" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[384,637,1935,1959]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="taiwang" status="sp. nov.">
|
||
<emphasis id="B954EACD300CFF8BFE1BF87CFDD0E9EC" box="[384,637,1935,1959]" italics="true" pageId="2" pageNumber="103">Pseudolitochira taiwang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300CFF8BFD1DF87CFD6FE9EC" bold="true" box="[646,706,1935,1959]" pageId="2" pageNumber="103">
|
||
<taxonomicNameLabel id="A26757B6300CFF8BFD1DF87CFD6FE9EC" box="[646,706,1935,1959]" pageId="2" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (11.2 × 8.1 mm) (NMMBCD5627); E,
|
||
<taxonomicName id="4C204D5C300CFF8BFA85F87CFE0BE980" authority="(Borradaile, 1902)" baseAuthorityName="Borradaile" baseAuthorityYear="1902" class="Malacostraca" family="Acidopsidae" genus="Crinitocinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="alcocki">
|
||
<emphasis id="B954EACD300CFF8BFA85F87CFF4DE980" italics="true" pageId="2" pageNumber="103">Crinitocinus alcocki</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB14B2E300CFF8BFF74F840FE32E980" author="Borradaile, L. A." box="[239,415,1971,1995]" pageId="2" pageNumber="103" pagination="237 - 271" refId="ref5787" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, I (III), pp. 237 - 271, figs. 41 - 60." type="book chapter" year="1902">Borradaile, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (9.3 × 7.9 mm) (ZRC 2022.0825); F, H,
|
||
<taxonomicName id="4C204D5C300CFF8BFC16F840FB16E980" authority="(Borradaile, 1902)" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[909,1211,1971,1995]" class="Malacostraca" family="Acidopsidae" genus="Crinitocinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="alcocki">
|
||
<emphasis id="B954EACD300CFF8BFC16F840FC58E980" box="[909,1013,1971,1995]" italics="true" pageId="2" pageNumber="103">C. alcocki</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB14B2E300CFF8BFB9FF840FB19E980" author="Borradaile, L. A." box="[1028,1204,1971,1995]" pageId="2" pageNumber="103" pagination="237 - 271" refId="ref5787" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, I (III), pp. 237 - 271, figs. 41 - 60." type="book chapter" year="1902">Borradaile, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (6.8 × 6.2 mm) (NMMBCD5628). G, H, in situ in habitat; H at night with filter. All specimens from Taiwan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6657300DFF8AFF0CF8A1FAF0E9F9" ID-DOI="http://doi.org/10.5281/zenodo.7989383" ID-Zenodo-Dep="7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="3" pageNumber="104" startId="3.[151,250,1874,1898]" targetBox="[151,1436,181,1850]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B9F36DF300DFF8AFF0CF8A1FAF0E9F9" blockId="3.[151,1436,1874,1970]" pageId="3" pageNumber="104">
|
||
<emphasis id="B954EACD300DFF8AFF0CF8A1FEBFE921" bold="true" box="[151,274,1874,1898]" pageId="3" pageNumber="104">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C204D5C300DFF8AFE8CF8A1FDAEE921" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[279,515,1874,1898]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300DFF8AFE8CF8A1FDAEE921" box="[279,515,1874,1898]" italics="true" pageId="3" pageNumber="104">Heteropilumnus planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300DFF8AFD93F8A1FDEDE921" bold="true" box="[520,576,1874,1898]" pageId="3" pageNumber="104">
|
||
<taxonomicNameLabel id="A26757B6300DFF8AFD93F8A1FDEDE921" box="[520,576,1874,1898]" pageId="3" pageNumber="104" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6657300AFF8DFF0CFC9EFD65ED86" ID-DOI="http://doi.org/10.5281/zenodo.7989387" ID-Zenodo-Dep="7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,877,901]" targetBox="[153,1434,183,851]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B9F36DF300AFF8DFF0CFC9EFD65ED86" blockId="4.[151,1437,877,973]" pageId="4" pageNumber="105">
|
||
<emphasis id="B954EACD300AFF8DFF0CFC9EFEB8EDCE" bold="true" box="[151,277,877,901]" pageId="4" pageNumber="105">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C204D5C300AFF8DFE85FC9EFDA1EDCE" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[286,524,877,901]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300AFF8DFE85FC9EFDA1EDCE" box="[286,524,877,901]" italics="true" pageId="4" pageNumber="105">Heteropilumnus planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300AFF8DFD8FFC9EFDFDEDCE" bold="true" box="[532,592,877,901]" pageId="4" pageNumber="105">
|
||
<taxonomicNameLabel id="A26757B6300AFF8DFD8FFC9EFDFDEDCE" box="[532,592,877,901]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36DF300AFF8DFF5CFBECFB47EA16" blockId="4.[151,1437,1055,2018]" pageId="4" pageNumber="105">
|
||
<emphasis id="B954EACD300AFF8DFF5CFBECFEC3EA72" bold="true" box="[199,366,1055,1081]" pageId="4" pageNumber="105">Colour in life.</emphasis>
|
||
Overall orange-brown setae covering carapace, with surface white; surface of ambulatory legs with patches and streaks of red and orange; distal half of fingers pigmented brown (
|
||
<figureCitation id="131B2A5A300AFF8DFBDBFBB0FB70EA16" box="[1088,1245,1091,1117]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1887]" captionTargetBox="[151,1436,273,1840]" captionTargetId="figure-68@2.[151,1436,273,1840]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life. A, G, Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625); B, H. planus n. sp., paratype female (11.4 × 8.2 mm) (NMMBCD5626); C, H. planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044); D, Pseudolitochira taiwang n. sp., holotype female (11.2 × 8.1 mm) (NMMBCD5627); E, Crinitocinus alcocki (Borradaile, 1902), male (9.3 × 7.9 mm) (ZRC 2022.0825); F, H, C. alcocki (Borradaile, 1902), male (6.8 × 6.2 mm) (NMMBCD5628). G, H, in situ in habitat; H at night with filter. All specimens from Taiwan." figureDoi="http://doi.org/10.5281/zenodo.7989377" httpUri="https://zenodo.org/record/7989377/files/figure.png" pageId="4" pageNumber="105">Fig. 1A–C, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6554300AFF8DFF5CFB94FA3FEACA" box="[199,1426,1127,1154]" pageId="4" pageNumber="105" type="etymology">
|
||
<paragraph id="8B9F36DF300AFF8DFF5CFB94FA3FEACA" blockId="4.[151,1437,1055,2018]" box="[199,1426,1127,1154]" pageId="4" pageNumber="105">
|
||
<emphasis id="B954EACD300AFF8DFF5CFB94FEE3EACA" bold="true" box="[199,334,1127,1153]" pageId="4" pageNumber="105">Etymology.</emphasis>
|
||
The name is derived from the Latin for flat, alluding to the relatively flat carapace of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6554300AFF8CFF5CFB78FB2FEC2A" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105" type="discussion">
|
||
<paragraph id="8B9F36DF300AFF8DFF5CFB78FBCEE942" blockId="4.[151,1437,1055,2018]" pageId="4" pageNumber="105">
|
||
<emphasis id="B954EACD300AFF8DFF5CFB78FE96EAEE" bold="true" box="[199,315,1163,1189]" pageId="4" pageNumber="105">Remarks.</emphasis>
|
||
<taxonomicName id="4C204D5C300AFF8DFEDEFB78FDE0EAEE" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[325,589,1163,1189]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300AFF8DFEDEFB78FDE0EAEE" box="[325,589,1163,1189]" italics="true" pageId="4" pageNumber="105">Heteropilumnus planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300AFF8DFDCCFB78FD34EAEE" bold="true" box="[599,665,1163,1189]" pageId="4" pageNumber="105">
|
||
<taxonomicNameLabel id="A26757B6300AFF8DFDCCFB78FD34EAEE" box="[599,665,1163,1189]" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinctive in that the carapace has a felt-like tomentum overall with the long plumose setae protruding from the anterior surfaces. In aspect and overall appearance,
|
||
<taxonomicName id="4C204D5C300AFF8DFA94FB43FAD3EA82" authority="Ng & Lin, 2023" authorityName="Ng & Lin" authorityYear="2023" box="[1295,1406,1199,1225]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
|
||
<emphasis id="B954EACD300AFF8DFA94FB43FAD3EA82" box="[1295,1406,1199,1225]" italics="true" pageId="4" pageNumber="105">H. planus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EACD300AFF8DFA1DFB5CFF15EAA6" bold="true" pageId="4" pageNumber="105">
|
||
<taxonomicNameLabel id="A26757B6300AFF8DFA1DFB5CFF15EAA6" pageId="4" pageNumber="105" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely resembles
|
||
<emphasis id="B954EACD300AFF8DFE52FB27FDFCEAA6" box="[457,593,1235,1261]" italics="true" pageId="4" pageNumber="105">
|
||
<taxonomicName id="4C204D5C300AFF8DFE52FB27FDE0EAA6" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[457,589,1235,1261]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">H. hirsutior</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but the carapace is lower, with the dorsal surface distinctly flatter in frontal view (
|
||
<figureCitation id="131B2A5A300AFF8DFF46FB04FE92EB5A" box="[221,319,1271,1297]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="4" pageNumber="105">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300AFF8DFED1FB04FEC1EB5A" box="[330,364,1271,1297]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="4" pageNumber="105">4C</figureCitation>
|
||
) (carapace higher with the dorsal surface gently convex in frontal view in
|
||
<taxonomicName id="4C204D5C300AFF8DFB36FB0BFA9CEB5A" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[1197,1329,1271,1297]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFB36FB0BFA9CEB5A" box="[1197,1329,1271,1297]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="131B2A5A300AFF8DFADBFB04FF62EB7D" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5A, B, D</figureCitation>
|
||
; Ng & Tan 1988; fig. 3A, B); the first anterolateral tooth is relatively lower and more triangular in shape, with the teeth separated by concavities (
|
||
<figureCitation id="131B2A5A300AFF8DFDBCFACCFD21EB12" box="[551,652,1343,1369]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="4" pageNumber="105">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300AFF8DFD02FACCFD16EB12" box="[665,699,1343,1369]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="4" pageNumber="105">4B</figureCitation>
|
||
) (first anterolateral tooth more truncate, with the teeth separated by deeper clefts in
|
||
<taxonomicName id="4C204D5C300AFF8DFEF7FA97FE5DEB36" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[364,496,1379,1405]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFEF7FA97FE5DEB36" box="[364,496,1379,1405]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="131B2A5A300AFF8DFE65FA90FDDAEB35" box="[510,631,1379,1406]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5A, B</figureCitation>
|
||
; Ng & Tan 1988: fig. 1A); the two median lobes of the posterior margin of the epistome project downwards, are truncate with each margin concave and separated from the lateral part by a prominent V-shaped cleft (
|
||
<figureCitation id="131B2A5A300AFF8DFE59FA58FDEEEB8D" box="[450,579,1451,1478]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="4" pageNumber="105">Figs. 2C, E</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300AFF8DFDCBFA58FDDCEB8E" box="[592,625,1451,1477]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="4" pageNumber="105">4C</figureCitation>
|
||
) (median lobes forming one triangular structure, joining lateral part as a concavity and separated by a narrow fissure in
|
||
<taxonomicName id="4C204D5C300AFF8DFD3BFA23FC89EBA2" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[672,804,1487,1513]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFD3BFA23FC89EBA2" box="[672,804,1487,1513]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="131B2A5A300AFF8DFCA9FA3CFC27EBA1" box="[818,906,1487,1514]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5D</figureCitation>
|
||
); the chelipeds are heterochelous in adult males with the outer surface of the major chela smooth on the ventral half (
|
||
<figureCitation id="131B2A5A300AFF8DFC14FA00FC5FE846" box="[911,1010,1523,1549]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="4" pageNumber="105">Figs. 2A</figureCitation>
|
||
) (chelipeds more or less homochelous with the outer surfaces of both distinctly and evenly granulated in adult male
|
||
<taxonomicName id="4C204D5C300AFF8DFB83F9EBFB32E87A" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[1048,1183,1559,1585]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFB83F9EBFB32E87A" box="[1048,1183,1559,1585]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
; Ng & Tan 1988; fig. 3C, G); the ambulatory meri are relatively longer and more slender (
|
||
<figureCitation id="131B2A5A300AFF8DFC07F9C8FB89E81D" box="[924,1060,1595,1622]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-17@3.[151,1436,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan.A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, antennules, antennae, orbits and epistome; F, anterior thoracic sternum and pleon; G, outer view of chelae; H, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.7989383" httpUri="https://zenodo.org/record/7989383/files/figure.png" pageId="4" pageNumber="105">Figs. 2A, H</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A5A300AFF8DFBABF9C8FBDCE81E" box="[1072,1137,1595,1621]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[183,1405,181,1919]" captionTargetId="figure-18@6.[183,1405,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Heteropilumnus planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044), Taiwan. A, overall dorsal habitus; B, dorsal view of carapace (right side denuded); C, frontal view of cephalothorax; D, left third maxilliped; E, outer view of left chela; F, right fourth ambulatory leg; G, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7989393" httpUri="https://zenodo.org/record/7989393/files/figure.png" pageId="4" pageNumber="105">4A, F</figureCitation>
|
||
) (shorter and stouter in
|
||
<taxonomicName id="4C204D5C300AFF8DFA1AF9CFFF54E832" baseAuthorityName="Lanchester" baseAuthorityYear="1900" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFA1AF9CFFF54E832" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="131B2A5A300AFF8DFE93F9ACFECDE831" box="[264,352,1631,1658]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5A</figureCitation>
|
||
; Ng & Tan 1988; fig. 3E); the male telson in similarly sized specimens is more rounded and proportionately shorter (
|
||
<figureCitation id="131B2A5A300AFF8DFE1AF970FE75E8D6" box="[385,472,1667,1693]" captionStart="FIGURE 3" captionStartId="4.[151,250,877,901]" captionTargetBox="[153,1434,183,851]" captionTargetId="figure-18@4.[151,1436,181,853]" captionTargetPageId="4" captionText="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="4" pageNumber="105">Fig. 3A</figureCitation>
|
||
) (distinctly more elongate and linguiform in
|
||
<taxonomicName id="4C204D5C300AFF8DFC50F977FBE3E8D6" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[971,1102,1667,1693]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFC50F977FBE3E8D6" box="[971,1102,1667,1693]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="131B2A5A300AFF8DFBC7F970FB1CE8D5" box="[1116,1201,1667,1694]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5C</figureCitation>
|
||
; Ng & Tan 1988; fig. 3D; see later for discussion of this character); and the G1 is less sinuous, with the proximal half almost straight and only the distal half curved, with the distal part distinctly hooked downwards (
|
||
<figureCitation id="131B2A5A300AFF8DFC65F938FBD9E8AE" box="[1022,1140,1739,1765]" captionStart="FIGURE 3" captionStartId="4.[151,250,877,901]" captionTargetBox="[153,1434,183,851]" captionTargetId="figure-18@4.[151,1436,181,853]" captionTargetPageId="4" captionText="FIGURE 3. Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625), Taiwan. A, pleon; B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (dorsal view); F, left G2. Scales: A = 1.0 mm; B, C, F = 0.5 mm; D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7989387" httpUri="https://zenodo.org/record/7989387/files/figure.png" pageId="4" pageNumber="105">Fig. 3B–E</figureCitation>
|
||
) (G1 is uniformly sinuous with the distal part directed more laterally in
|
||
<taxonomicName id="4C204D5C300AFF8DFD0AF903FCB8E942" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[657,789,1775,1801]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFD0AF903FCB8E942" box="[657,789,1775,1801]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
; Ng & Tan 1988; fig. 4A–D).
|
||
</paragraph>
|
||
<paragraph id="8B9F36DF300AFF8CFF5CF8E7FB2FEC2A" blockId="4.[151,1437,1055,2018]" lastBlockId="5.[151,1437,150,645]" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105">
|
||
<bibRefCitation id="EFB14B2E300AFF8DFF5CF8E7FE36E966" author="Maenosono T." box="[199,411,1811,1838]" pageId="4" pageNumber="105" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C204D5C300AFF8DFD9EF8E7FD24E966" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[517,649,1811,1837]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300AFF8DFD9EF8E7FD24E966" box="[517,649,1811,1837]" italics="true" pageId="4" pageNumber="105">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
from the Ryukyus,
|
||
<collectingCountry id="F337764F300AFF8DFCFDF8E7FC04E965" box="[870,937,1812,1838]" name="Japan" pageId="4" pageNumber="105">Japan</collectingCountry>
|
||
. We compared one of his specimens with the material from
|
||
<collectingCountry id="F337764F300AFF8DFEA3F8C4FE07E91A" box="[312,426,1847,1873]" name="Singapore" pageId="4" pageNumber="105">Singapore</collectingCountry>
|
||
and they are very similar. The male chela structure requires comment. The large
|
||
<typeStatus id="549B887D300AFF8DFAA1F8C4FA31E91A" box="[1338,1436,1847,1873]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
|
||
male has only one chela, which appears to be the minor one (
|
||
<figureCitation id="131B2A5A300AFF8DFCCAF8A8FC61E93D" box="[849,972,1883,1910]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="4" pageNumber="105">Fig. 5A; E</figureCitation>
|
||
; Ng & Tan 1988: fig. 3C, G), and all the other male specimens from
|
||
<collectingCountry id="F337764F300AFF8DFE57F88CFD93E9D2" box="[460,574,1919,1945]" name="Singapore" pageId="4" pageNumber="105">Singapore</collectingCountry>
|
||
we have examined so far have homochelous chelipeds with both chelae similarly structured. Even in the largest male examined (11.7 ×
|
||
<quantity id="4CD89B3A300AFF8DFCABF850FC24E9F5" box="[816,905,1955,1982]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="4" pageNumber="105" unit="mm" value="8.9">8.9 mm</quantity>
|
||
, ZRC 1992.10569), both chelae are similar and closely resemble that of the
|
||
<typeStatus id="549B887D300AFF8DFE4AF834FD9EE9AA" box="[465,563,1991,2017]" pageId="4" pageNumber="105" type="holotype">holotype</typeStatus>
|
||
male. For the Japanese material, however,
|
||
<bibRefCitation id="EFB14B2E300AFF8DFB8DF83BFB47E9AA" author="Maenosono T." box="[1046,1258,1991,2018]" pageId="4" pageNumber="105" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
|
||
figured a larger male (8.6 ×
|
||
<quantity id="4CD89B3A300BFF8CFEBAFF65FED5EEFA" box="[289,376,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="106" unit="mm" value="5.9">5.9 mm</quantity>
|
||
) as having distinctly heterochelous chelipeds, with the major one having the lower part of the outer surface mostly glabrous and smooth (
|
||
<bibRefCitation id="EFB14B2E300BFF8CFDE0FF48FCEFEE9E" author="Maenosono T." box="[635,834,187,213]" pageId="5" pageNumber="106" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono 2019</bibRefCitation>
|
||
: fig. 4D), whereas a smaller male (7.1 ×
|
||
<quantity id="4CD89B3A300BFF8CFA90FF48FACFEE9E" box="[1291,1378,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="106" unit="mm" value="4.8">4.8 mm</quantity>
|
||
) was homochelous, with chelae like those from
|
||
<collectingCountry id="F337764F300BFF8CFDE5FF2CFD5DEEB2" box="[638,752,223,249]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
specimens. The ambulatory legs of the Japanese specimens are slightly longer (
|
||
<bibRefCitation id="EFB14B2E300BFF8CFEEDFEF0FDEDEF56" author="Maenosono T." box="[374,576,259,285]" pageId="5" pageNumber="106" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono 2019</bibRefCitation>
|
||
: fig. 1C) than those from
|
||
<collectingCountry id="F337764F300BFF8CFCF1FEF0FC71EF56" box="[874,988,259,285]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
(
|
||
<figureCitation id="131B2A5A300BFF8CFC77FEF0FBEAEF56" box="[1004,1095,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="5" pageNumber="106">Fig. 5A</figureCitation>
|
||
; Ng & Tan 1988: fig. 3E) but this does not seem significant. The male pleon figured by
|
||
<bibRefCitation id="EFB14B2E300BFF8CFCBFFED4FC5EEF0A" author="Maenosono T." box="[804,1011,295,321]" pageId="5" pageNumber="106" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono (2019</bibRefCitation>
|
||
: fig. 6C) is comparatively shorter and more rounded than that of the type (
|
||
<figureCitation id="131B2A5A300BFF8CFDB7FEB8FD29EF2E" box="[556,644,330,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-17@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 5. Heteropilumnus hirsutior (Lanchester, 1900), holotype male (10.8 × 8.0 mm) (NHM 1900.10.22.181), Singapore. A, overall dorsal habitus; B, dorsal view of carapace; C, anterior thoracic sternum and pleon; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7989395" httpUri="https://zenodo.org/record/7989395/files/figure.png" pageId="5" pageNumber="106">Fig. 5C</figureCitation>
|
||
; Ng & Tan 1988: fig. 3D) but there appears to be some variation, with smaller males possessing a more rounded telson. It must be noted that the
|
||
<typeStatus id="549B887D300BFF8CFC52FE9CFB86EFC2" box="[969,1067,367,393]" pageId="5" pageNumber="106" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="4C204D5C300BFF8CFB10FE9CFAA3EFC2" baseAuthorityName="Lanchester" baseAuthorityYear="1900" box="[1163,1294,367,393]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="hirsutior">
|
||
<emphasis id="B954EACD300BFF8CFB10FE9CFAA3EFC2" box="[1163,1294,367,393]" italics="true" pageId="5" pageNumber="106">H. hirsutior</emphasis>
|
||
</taxonomicName>
|
||
is very large (10.8 × 8.0 mm) and a similarly sized male in
|
||
<collectingCountry id="F337764F300BFF8CFD3FFE60FCBBEFE6" box="[676,790,403,429]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
(ZRC 1992.10569) also has the same telson shape.
|
||
<bibRefCitation id="EFB14B2E300BFF8CFAFAFE60FE98EF9A" author="Maenosono T." pageId="5" pageNumber="106" pagination="19 - 44" refId="ref6259" refString="Maenosono T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44." type="journal article" year="2019">Maenosono (2019</bibRefCitation>
|
||
: fig. 14E, F) figured the G1 of the species and his figure gives the appearance that the proximal half is straighter with the distalmost part more hooked compared to that of the
|
||
<collectingCountry id="F337764F300BFF8CFC4EFE28FBEAEFBE" box="[981,1095,475,501]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
material (Ng & Tan 1988: fig. 4A–D). Examination of the Japanese male on hand shows its G1 to be almost identical to the
|
||
<collectingCountry id="F337764F300BFF8CFB33FE0CFAB7EC52" box="[1192,1306,511,537]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
specimens. As such, we treat all the
|
||
<collectingCountry id="F337764F300BFF8CFE30FDD0FDB0EC76" box="[427,541,547,573]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
and Japanese material as conspecific for now. It would be useful to compare all of the Japanese and
|
||
<collectingCountry id="F337764F300BFF8CFEE2FDB4FE46EC2A" box="[377,491,583,609]" name="Singapore" pageId="5" pageNumber="106">Singapore</collectingCountry>
|
||
specimens to determine if they belong to the same species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6554300BFF8CFF5CFD99FAC7ECCE" box="[199,1386,618,645]" pageId="5" pageNumber="106" type="biology_ecology">
|
||
<paragraph id="8B9F36DF300BFF8CFF5CFD99FAC7ECCE" blockId="5.[151,1437,150,645]" box="[199,1386,618,645]" pageId="5" pageNumber="106">
|
||
<emphasis id="B954EACD300BFF8CFF5CFD99FE8AECCF" bold="true" box="[199,295,618,644]" pageId="5" pageNumber="106">Biology.</emphasis>
|
||
The specimens were all collected from a coral reef flat covered with muddy sediments (
|
||
<figureCitation id="131B2A5A300BFF8CFA98FD98FAF1ECCE" box="[1283,1372,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1887]" captionTargetBox="[151,1436,273,1840]" captionTargetId="figure-68@2.[151,1436,273,1840]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life. A, G, Heteropilumnus planus n. sp., holotype male (10.7 × 8.0 mm) (NMMBCD5625); B, H. planus n. sp., paratype female (11.4 × 8.2 mm) (NMMBCD5626); C, H. planus n. sp., paratype female (10.5 × 7.9 mm) (ZRC 2017.1044); D, Pseudolitochira taiwang n. sp., holotype female (11.2 × 8.1 mm) (NMMBCD5627); E, Crinitocinus alcocki (Borradaile, 1902), male (9.3 × 7.9 mm) (ZRC 2022.0825); F, H, C. alcocki (Borradaile, 1902), male (6.8 × 6.2 mm) (NMMBCD5628). G, H, in situ in habitat; H at night with filter. All specimens from Taiwan." figureDoi="http://doi.org/10.5281/zenodo.7989377" httpUri="https://zenodo.org/record/7989377/files/figure.png" pageId="5" pageNumber="106">Fig. 1G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |