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<mods:title id="DC195BB0F51FD48A5D9BC352B97A3B89">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<paragraph id="8B9F36B6FFEA4B09B38AA5B81026FA4A" blockId="16.[924,1340,1469,1493]" box="[924,1340,1469,1493]" pageId="16" pageNumber="241">
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<taxonomicName id="4C204D35FFEA4B09B38AA5B81026FA4A" ID-CoL="78PXW" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[924,1340,1469,1493]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEA4B09B38AA5B81026FA4A" bold="true" box="[924,1340,1469,1493]" pageId="16" pageNumber="241">
<emphasis id="B954EAA4FFEA4B09B38AA5B81160FA4A" bold="true" box="[924,1146,1469,1493]" italics="true" pageId="16" pageNumber="241">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFEA4B09B497A5BB1026FA4A" author="BERNHAUER M." box="[1153,1340,1470,1493]" pageId="16" pageNumber="241" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
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<paragraph id="8B9F36B6FFEA4B09B3F0A5D911E9FA70" blockId="16.[828,1437,1500,1633]" box="[998,1267,1500,1519]" pageId="16" pageNumber="241">
(
<figureCitation id="131B2A33FFEA4B09B3FAA5D91138FA70" box="[1004,1058,1500,1519]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="16" pageNumber="241">Figs 1</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B43AA5D91122FA70" box="[1068,1080,1500,1519]" captionStart="Fig" captionStartId="11.[151,182,2008,2027]" captionTargetBox="[151,1437,191,1879]" captionTargetId="figure-13@11.[151,1437,191,1968]" captionTargetPageId="11" captionText="Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399708" httpUri="https://zenodo.org/record/7399708/files/figure.png" pageId="16" pageNumber="241">3</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B457A5D91146FA70" box="[1089,1116,1500,1519]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="16" pageNumber="241">6B</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B473A5D91165FA70" box="[1125,1151,1500,1519]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="16" pageNumber="241">7B</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B49FA5D911B7FA70" box="[1161,1197,1500,1519]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="16" pageNumber="241">11C</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B4A0A5D911D7FA70" box="[1206,1229,1500,1519]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="16" pageNumber="241">12</figureCitation>
, 
<figureCitation id="131B2A33FFEA4B09B4C0A5D911F6FA70" box="[1238,1260,1500,1519]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="16" pageNumber="241">18</figureCitation>
)
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<taxonomicName id="4C204D35FFEA4B09B32AA5FB11AEF98E" ID-CoL="78PXW" authority="Bernhauer, 1908 e: 335" authorityName="Bernhauer" authorityPageNumber="335" authorityYear="1908" box="[828,1204,1534,1553]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEA4B09B32AA5FB16F3F98E" box="[828,1001,1534,1553]" italics="true" pageId="16" pageNumber="241">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFEA4B09B3E6A5FB119BF98E" author="BERNHAUER M." box="[1008,1153,1534,1553]" pageId="16" pageNumber="241" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
e: 335
</taxonomicName>
[
<typeStatus id="549B8814FFEA4B09B4D6A5FB11F7F98E" box="[1216,1261,1534,1553]" pageId="16" pageNumber="241">Type</typeStatus>
locality: Faroe; 
<collectingCountry id="F3377626FFEA4B09B56AA5FB168BF9B3" name="Romania" pageId="16" pageNumber="241">Romania</collectingCountry>
; Buchara; 
<collectingCountry id="F3377626FFEA4B09B3FAA61C112DF9B3" box="[1004,1079,1561,1580]" name="Czech Republic" pageId="16" pageNumber="241">Böhmen</collectingCountry>
, Wran a. Moldau].
</paragraph>
<paragraph id="8B9F36B6FFEA4B09B32AA6311087F9FE" blockId="16.[828,1437,1500,1633]" pageId="16" pageNumber="241">
<taxonomicName id="4C204D35FFEA4B09B32AA6311163F9D8" ID-CoL="4R3NG" authority="Stephens, 1832: 215" authorityName="Stephens" authorityPageNumber="215" authorityYear="1832" box="[828,1145,1588,1607]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B954EAA4FFEA4B09B32AA63116D2F9D8" box="[828,968,1588,1607]" italics="true" pageId="16" pageNumber="241">Quedius gracilis</emphasis>
<bibRefCitation id="EFB14B47FFEA4B09B3DBA6311163F9D8" author="STEPHENS J. F." box="[973,1145,1588,1607]" pageId="16" pageNumber="241" refId="ref92742" refString="STEPHENS J. F. 1832: Illustrations of British entomology; or a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin &amp; Cradock, London, 240 pp." type="book" year="1832">Stephens, 1832: 215</bibRefCitation>
</taxonomicName>
[
<typeStatus id="549B8814FFEA4B09B495A63111AAF9D8" box="[1155,1200,1588,1607]" pageId="16" pageNumber="241">Type</typeStatus>
locality: London], 
<emphasis id="B954EAA4FFEA4B09B544A6311086F9D8" bold="true" box="[1362,1436,1588,1607]" pageId="16" pageNumber="241">syn. rev.</emphasis>
<taxonomicName id="4C204D35FFEA4B09B32AA64B1174F9FE" authority="Hatch, 1957: 216" authorityName="Hatch" authorityPageNumber="216" authorityYear="1957" box="[828,1134,1614,1633]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEA4B09B32AA64B16C1F9FE" box="[828,987,1614,1633]" italics="true" pageId="16" pageNumber="241">Quedius parallelus</emphasis>
<bibRefCitation id="EFB14B47FFEA4B09B3C9A64B1174F9FE" author="HATCH M. H." box="[991,1134,1614,1633]" pageId="16" pageNumber="241" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch,1957:216</bibRefCitation>
</taxonomicName>
[
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locality:Washington, Seattle]
</paragraph>
<paragraph id="8B9F36B6FFEA4B0CB32AA67D1797FE14" blockId="16.[828,1437,1656,2075]" lastBlockId="21.[151,761,190,394]" lastPageId="21" lastPageNumber="246" pageId="16" pageNumber="241">
<emphasis id="B954EAA4FFEA4B09B32AA67D16BCF914" bold="true" box="[828,934,1656,1675]" pageId="16" pageNumber="241">References.</emphasis>
Gඋංൽൾඅඅං (1924): 79 (characters), SർHൾൾඋඉൾඅඍඓ (1933): 1441 (characters), Hൺඇඌൾඇ (1952): 141 (characters); Sආൾඍൺඇൺ (1958): 362 (characters and ecology); (1962): 133, (1965b): 38 (characters); (1971a): 129 (characters, synonymy of 
<taxonomicName id="4C204D35FFEA4B09B4B5A6CD1003F944" authorityName="Hatch" authorityYear="1957" box="[1187,1305,1736,1755]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEA4B09B4B5A6CD1003F944" box="[1187,1305,1736,1755]" italics="true" pageId="16" pageNumber="241">Q. parallelus</emphasis>
</taxonomicName>
); (1965a): 12, (1970): 62, (1990): 98, (1993): 50 (distribution); (1978a): 826 (biology); KඈඋGൾ (1962b): 332 (suspected synonymy of 
<taxonomicName id="4C204D35FFEA4B09B4C3A6F81053F88F" authorityName="Hatch" authorityYear="1957" box="[1237,1353,1789,1808]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEA4B09B4C3A6F81053F88F" box="[1237,1353,1789,1808]" italics="true" pageId="16" pageNumber="241">Q. parallelus</emphasis>
</taxonomicName>
); (1964): 119 (distribution); Pൺඅආ (1963): 141 (characters); LඈHඌൾ (1964): 211 (characters); Dඏඈෞගĸ (1965): 89 (distribution); Hඈඋංඈඇ (1965): 272 (distribution); Sඓඎඃൾർĸං (1968): 735 (distribution); Fඋൺඇĸ (1969): 267 (biology); Bඈඋൽඈඇං (1974b): 10, (1976b): 92 (characters); Pඈඉൾ (1977): 31 (distribution); Cඈංൿൿൺංඍ (1978): 191 (characters); Oඎඍൾඋൾඅඈ (1978): 281 (pupal characters); Bඎඋൺĸඈඐඌĸං et al. (1980): 134 (distribution); TඬඍH (1984): 118 (characters); DඋඎGආൺඇൽ (1987): 312 (distribution); Nඈඐඈඌൺൽ (1990): 145 (biology); BඈඋGൾඌ (1990): Table II (distribution); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 32 (distribution); Oඐൾඇ (1997): 149 (distribution); Oඎඍൾඋൾඅඈ et al. (1998): 129 (biology); AඌඌංඇG (2001): 75 (biology); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඃĸൺ &amp; Sආൾඍൺඇൺ (2007): 429 (characters and distribution); ÖඓGൾඇ (2011):204 (distribution); Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ (2011): 59 (characters and distribution); Wൾൻඌඍൾඋ et al. (2012): 313 (distribution); MൺGඎඋൺ et al. (2013): 719 (biology); TඬඍHආඣඋඣඌඓ et al. (2014): 688 (biology); Sൾආൾඇඈඏ et al. (2015): 127 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 330 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 125 (notes on types and distribution); (2019): 48 (distribution).
</paragraph>
</subSubSection>
<caption id="DF5F663EFFEB4B08B081A7E216EAF78B" ID-DOI="http://doi.org/10.5281/zenodo.7399720" ID-Zenodo-Dep="7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="17" pageNumber="242" startId="17.[151,182,2023,2042]" targetBox="[180,1386,189,1955]" targetPageId="17">
<paragraph id="8B9F36B6FFEB4B08B081A7E216EAF78B" blockId="17.[151,1437,2023,2068]" pageId="17" pageNumber="242">
Fig. 7. Habitus of 
<taxonomicName id="4C204D35FFEB4B08B120A7E21467F865" authorityName="Stephens" authorityYear="1829" box="[310,381,2023,2042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFEB4B08B120A7E21467F865" box="[310,381,2023,2042]" italics="true" pageId="17" pageNumber="242">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFEB4B08B195A7E214ABF865" box="[387,433,2023,2042]" pageId="17" pageNumber="242" sensu="stricto">s. str.</taxonomicNameLabel>
A – 
<taxonomicName id="4C204D35FFEB4B08B1CCA7E217E1F865" authority="Gusarov, 1991" authorityName="Gusarov" authorityYear="1991" box="[474,763,2023,2042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="afrofuliginosus">
<emphasis id="B954EAA4FFEB4B08B1CCA7E2176FF865" box="[474,629,2023,2042]" italics="true" pageId="17" pageNumber="242">Q. afrofuliginosus</emphasis>
<bibRefCitation id="EFB14B47FFEB4B08B26DA7E217E1F865" author="GUSAROV V. I." box="[635,763,2023,2042]" pageId="17" pageNumber="242" pagination="3 - 12" refId="ref83976" refString="GUSAROV V. I. 1991: Novye i maloizvestnye palearkticheskie stafilinidy (Coleoptera: Staphylinidae) [New and little-known Palearctic rove beetles (Coleoptera: Staphylinidae)] Soobshchenie 3: 3 - 12 (in Russian)." type="journal article" year="1991">Gusarov, 1991</bibRefCitation>
</taxonomicName>
; B – 
<taxonomicName id="4C204D35FFEB4B08B33CA7E21121F865" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[810,1083,2023,2042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEB4B08B33CA7E216BFF865" box="[810,933,2023,2042]" italics="true" pageId="17" pageNumber="242">Q. curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFEB4B08B3BDA7E21121F865" author="BERNHAUER M." box="[939,1083,2023,2042]" pageId="17" pageNumber="242" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
</taxonomicName>
; C – 
<taxonomicName id="4C204D35FFEB4B08B47CA7E21082F865" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[1130,1432,2023,2042]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEB4B08B47CA7E211F9F865" box="[1130,1251,2023,2042]" italics="true" pageId="17" pageNumber="242">Q. fuliginosus</emphasis>
(
<bibRefCitation id="EFB14B47FFEB4B08B4F8A7E21089F865" author="GRAVENHORST J. L. C." box="[1262,1427,2023,2042]" pageId="17" pageNumber="242" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
; D – 
<taxonomicName id="4C204D35FFEB4B08B0AAA804148FF78B" authority="Brulle, 1832" authorityName="Brulle" authorityYear="1832" box="[188,405,2049,2068]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFEB4B08B0AAA8041439F78B" box="[188,291,2049,2068]" italics="true" pageId="17" pageNumber="242">Q. levicollis</emphasis>
<bibRefCitation id="EFB14B47FFEB4B08B13EA804148FF78B" author="BRULLE G. A." box="[296,405,2049,2068]" pageId="17" pageNumber="242" pagination="289 - 400" refId="ref80797" refString="BRULLE G. A. 1832 - 1833: Expedition scientifique de Moree. Section des Sciences Physiques. Tome III. 1. re Partie. Zoologie. Deuxieme Section. Des animales articules. F. G. Levrault, Paris, 400 pp. [Pp. 1 - 288 published in 1832, pp. 289 - 400 in 1833.]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
</taxonomicName>
; E – 
<taxonomicName id="4C204D35FFEB4B08B1D7A80417CAF78B" authority="Kiesenwetter, 1847" authorityName="Kiesenwetter" authorityYear="1847" box="[449,720,2049,2068]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B954EAA4FFEB4B08B1D7A8041739F78B" box="[449,547,2049,2068]" italics="true" pageId="17" pageNumber="242">Q. unicolor</emphasis>
<bibRefCitation id="EFB14B47FFEB4B08B23EA80417CAF78B" author="KIESENWETTER H. &amp; MARKEL F." box="[552,720,2049,2068]" pageId="17" pageNumber="242" pagination="73 - 87" refId="ref85769" refString="KIESENWETTER H. ඏඈඇ &amp; MARKEL F. 1847: Eine entomologische Excursion im Riesengebirge im Juli 1846 (Fortsetzung). Entomologische Zeitung (Stettin) 8: 73 - 87." type="journal article" year="1847">Kiesenwetter, 1847</bibRefCitation>
</taxonomicName>
; F – 
<taxonomicName id="4C204D35FFEB4B08B2ECA80416F6F78B" authority="Korge, 1961" authorityName="Korge" authorityYear="1961" box="[762,1004,2049,2068]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="subunicolor">
<emphasis id="B954EAA4FFEB4B08B2ECA8041660F78B" box="[762,890,2049,2068]" italics="true" pageId="17" pageNumber="242">Q. subunicolor</emphasis>
<bibRefCitation id="EFB14B47FFEB4B08B369A80416F6F78B" author="KORGE H." box="[895,1004,2049,2068]" pageId="17" pageNumber="242" pagination="81 - 83" refId="ref86145" refString="KORGE H. 1961: Quedius subunicolor n. sp. aus Nordeuropa (Col. Staphyl.). Mitteilungen der Deutschen Entomologischen Gesellschaft 20: 81 - 83." type="journal article" year="1961">Korge, 1961</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF5F663EFFE84B0BB081A7EE11A4F787" ID-DOI="http://doi.org/10.5281/zenodo.7399722" ID-Zenodo-Dep="7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="18" pageNumber="243" startId="18.[151,182,2027,2046]" targetBox="[189,1393,195,2007]" targetPageId="18">
<paragraph id="8B9F36B6FFE84B0BB081A7EE11A4F787" blockId="18.[151,1436,2027,2072]" pageId="18" pageNumber="243">
Fig. 8. Habitus of 
<taxonomicName id="4C204D35FFE84B0BB125A7EE1460F861" authorityName="Stephens" authorityYear="1829" box="[307,378,2027,2046]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFE84B0BB125A7EE1460F861" box="[307,378,2027,2046]" italics="true" pageId="18" pageNumber="243">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFE84B0BB169A7EE14B7F861" box="[383,429,2027,2046]" pageId="18" pageNumber="243" sensu="stricto">s. str.</taxonomicNameLabel>
A – 
<taxonomicName id="4C204D35FFE84B0BB1C2A7EE17D1F861" authority="(Menetries, 1832)" baseAuthorityName="Menetries" baseAuthorityYear="1832" box="[468,715,2027,2046]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="B954EAA4FFE84B0BB1C2A7EE1730F861" box="[468,554,2027,2046]" italics="true" pageId="18" pageNumber="243">Q. vicinus</emphasis>
(
<bibRefCitation id="EFB14B47FFE84B0BB220A7EE17DFF861" author="MENETRIES E." box="[566,709,2027,2046]" pageId="18" pageNumber="243" refId="ref87999" refString="MENETRIES E. 1832: Catalogue raisonne des objets de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse entrepris par ordre de S. M. L'Empereur. L'Academie Imperiale des Sciences, St. Petersbourg, 271 + xxxii + iv pp." type="book" year="1832">Ménétriés, 1832</bibRefCitation>
)
</taxonomicName>
; B – 
<taxonomicName id="4C204D35FFE84B0BB2EEA7EE115AF861" authority="Smetana, 1958" authorityName="Smetana A. Solodovnikov" authorityYear="1958" box="[760,1088,2027,2046]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="meridiocarpathicus">
<emphasis id="B954EAA4FFE84B0BB2EEA7EE16A3F861" box="[760,953,2027,2046]" italics="true" pageId="18" pageNumber="243">Q. meridiocarpathicus</emphasis>
<bibRefCitation id="EFB14B47FFE84B0BB3A9A7EE115AF861" author="SMETANA A." box="[959,1088,2027,2046]" pageId="18" pageNumber="243" refId="ref91349" refString="SMETANA A. 1958: Fauna CSR. Svazek 12. Drabcikoviti - Staphylinidae I. Staphylinae (Coleoptera). [Fauna of the CSR. Volume 12. Rove beetles -- Staphylinidae I. Staphylinae (Coleoptera)]. Ceskoslovenska akademie ved, Praha, 435 pp (in Czech)." type="book" year="1958">Smetana, 1958</bibRefCitation>
</taxonomicName>
; C – 
<taxonomicName id="4C204D35FFE84B0BB47BA7EE1086F861" authority="(Gravenhorst, 1806)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[1133,1436,2027,2046]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFE84B0BB47BA7EE11F3F861" box="[1133,1257,2027,2046]" italics="true" pageId="18" pageNumber="243">Q. molochinus</emphasis>
(
<bibRefCitation id="EFB14B47FFE84B0BB4E3A7EE108CF861" author="GRAVENHORST J. L. C." box="[1269,1430,2027,2046]" pageId="18" pageNumber="243" refId="ref83841" refString="GRAVENHORST J. L. C. 1806: Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae, 248 pp." type="book" year="1806">Gravenhorst, 1806</bibRefCitation>
)
</taxonomicName>
(red elytra variant); D – 
<taxonomicName id="4C204D35FFE84B0BB171A80014F9F787" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[359,483,2053,2072]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFE84B0BB171A80014F9F787" box="[359,483,2053,2072]" italics="true" pageId="18" pageNumber="243">Q. molochinus</emphasis>
</taxonomicName>
(black elytra variant); E – 
<taxonomicName id="4C204D35FFE84B0BB2DFA8001683F787" authority="Korge, 1960" authorityName="Korge" authorityYear="1960" box="[713,921,2053,2072]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B954EAA4FFE84B0BB2DFA800163DF787" box="[713,807,2053,2072]" italics="true" pageId="18" pageNumber="243">Q. balticus</emphasis>
Korge, 1960
</taxonomicName>
; F – 
<taxonomicName id="4C204D35FFE84B0BB3D5A80011A0F787" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[963,1210,2053,2072]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="243" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFE84B0BB3D5A800112EF787" box="[963,1076,2053,2072]" italics="true" pageId="18" pageNumber="243">Q. sundukovi</emphasis>
<bibRefCitation id="EFB14B47FFE84B0BB42FA80011A0F787" author="SMETANA A." box="[1081,1210,2053,2072]" pageId="18" pageNumber="243" pagination="189 - 193" refId="ref92269" refString="SMETANA A. 2003: Quedius (Quedius) sundukovi (Coleoptera, Staphylinidae, Staphylinini, Quediina), an interesting new species from the Russian Far East. Elytra 31: 189 - 193." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF5F663EFFE94B0AB081A7E9119CF785" ID-DOI="http://doi.org/10.5281/zenodo.7399724" ID-Zenodo-Dep="7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="19" pageNumber="244" startId="19.[151,182,2028,2047]" targetBox="[203,1360,198,1958]" targetPageId="19">
<paragraph id="8B9F36B6FFE94B0AB081A7E9119CF785" blockId="19.[151,1437,2028,2074]" pageId="19" pageNumber="244">
Fig. 9. Habitus of 
<taxonomicName id="4C204D35FFE94B0AB121A7E91464F860" authorityName="Stephens" authorityYear="1829" box="[311,382,2028,2047]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFE94B0AB121A7E91464F860" box="[311,382,2028,2047]" italics="true" pageId="19" pageNumber="244">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFE94B0AB193A7E914AEF860" box="[389,436,2028,2047]" pageId="19" pageNumber="244" sensu="stricto">s. str.</taxonomicNameLabel>
A – 
<taxonomicName id="4C204D35FFE94B0AB1C9A7E917F7F860" authority="Bernhauer, 1898" authorityName="Bernhauer" authorityYear="1898" box="[479,749,2028,2047]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFE94B0AB1C9A7E9174FF860" box="[479,597,2028,2047]" italics="true" pageId="19" pageNumber="244">Q. hispanicus</emphasis>
<bibRefCitation id="EFB14B47FFE94B0AB24AA7E917F7F860" author="BERNHAUER M." box="[604,749,2028,2047]" pageId="19" pageNumber="244" pagination="338 - 341" refId="ref80314" refString="BERNHAUER M. 1898: Neue Staphyliniden aus Europa. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 48: 338 - 341." type="journal article" year="1898">Bernhauer, 1898</bibRefCitation>
</taxonomicName>
; B – 
<taxonomicName id="4C204D35FFE94B0AB30BA7E9168EF860" authorityName="Bernhauer" authorityYear="1898" box="[797,916,2028,2047]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFE94B0AB30BA7E9168EF860" box="[797,916,2028,2047]" italics="true" pageId="19" pageNumber="244">Q. hispanicus</emphasis>
</taxonomicName>
(pale sutures variant); C – 
<taxonomicName id="4C204D35FFE94B0AB492A7E9104CF860" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[1156,1366,2028,2047]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFE94B0AB492A7E911FEF860" box="[1156,1252,2028,2047]" italics="true" pageId="19" pageNumber="244">Q. pallipes</emphasis>
<bibRefCitation id="EFB14B47FFE94B0AB4FCA7E9104CF860" author="LUCAS P. H." box="[1258,1366,2028,2047]" pageId="19" pageNumber="244" refId="ref87139" refString="LUCAS P. H. 1846 - 1849: Histoire naturelle des animaux articules. Deuxieme partie. Insectes. In: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842 publiee par ordre du Gouvernement et avec le concours d'une Commission Academique. Sciences physiques. Zoologie II. A. Bertrand, Paris, 590 pp." type="book" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
; D – 
<taxonomicName id="4C204D35FFE94B0AB591A7E9148EF785" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFE94B0AB591A7E915F8F785" italics="true" pageId="19" pageNumber="244">Q. laticollis</emphasis>
(
<bibRefCitation id="EFB14B47FFE94B0AB0FBA8021494F785" author="GRAVENHORST J. L. C." box="[237,398,2055,2074]" pageId="19" pageNumber="244" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
(black elytra variant); E – 
<taxonomicName id="4C204D35FFE94B0AB26CA80217C5F785" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[634,735,2055,2074]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFE94B0AB26CA80217C5F785" box="[634,735,2055,2074]" italics="true" pageId="19" pageNumber="244">Q. laticollis</emphasis>
</taxonomicName>
(red elytra variant); F – 
<taxonomicName id="4C204D35FFE94B0AB3A6A802119BF785" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[944,1153,2055,2074]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B954EAA4FFE94B0AB3A6A8021108F785" box="[944,1042,2055,2074]" italics="true" pageId="19" pageNumber="244">Q. strenuus</emphasis>
<bibRefCitation id="EFB14B47FFE94B0AB401A802119BF785" author="CASEY T. L." box="[1047,1153,2055,2074]" pageId="19" pageNumber="244" pagination="395 - 450" refId="ref81365" refString="CASEY T. L. 1915: Studies in some staphylinid genera of North America. Memoirs on the Coleoptera 6: 395 - 450." type="journal article" year="1915">Casey, 1915</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF5F663EFFEE4B0DB081A7EB1165F783" ID-DOI="http://doi.org/10.5281/zenodo.7399726" ID-Zenodo-Dep="7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="20" pageNumber="245" startId="20.[151,182,2030,2049]" targetBox="[202,1343,201,1961]" targetPageId="20">
<paragraph id="8B9F36B6FFEE4B0DB081A7EB1165F783" blockId="20.[151,1436,2030,2076]" pageId="20" pageNumber="245">
Fig. 10. Habitus of 
<taxonomicName id="4C204D35FFEE4B0DB12DA7EB1498F79E" authorityName="Stephens" authorityYear="1829" box="[315,386,2030,2049]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFEE4B0DB12DA7EB1498F79E" box="[315,386,2030,2049]" italics="true" pageId="20" pageNumber="245">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFEE4B0DB191A7EB14AEF79E" box="[391,436,2030,2049]" pageId="20" pageNumber="245" sensu="stricto">s. str.</taxonomicNameLabel>
A – 
<taxonomicName id="4C204D35FFEE4B0DB1CFA7EB178AF79E" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[473,656,2030,2049]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="B954EAA4FFEE4B0DB1CFA7EB173BF79E" box="[473,545,2030,2049]" italics="true" pageId="20" pageNumber="245">Q. horni</emphasis>
<bibRefCitation id="EFB14B47FFEE4B0DB230A7EB178AF79E" author="HATCH M. H." box="[550,656,2030,2049]" pageId="20" pageNumber="245" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
; B – 
<taxonomicName id="4C204D35FFEE4B0DB2ADA7EB1674F79E" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[699,878,2030,2049]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B954EAA4FFEE4B0DB2ADA7EB17E5F79E" box="[699,767,2030,2049]" italics="true" pageId="20" pageNumber="245">Q. lanei</emphasis>
<bibRefCitation id="EFB14B47FFEE4B0DB312A7EB1674F79E" author="HATCH M. H." box="[772,878,2030,2049]" pageId="20" pageNumber="245" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
; C – 
<taxonomicName id="4C204D35FFEE4B0DB38FA7EB1195F79E" authority="Hansen &amp; Brunke" authorityName="Hansen &amp; Brunke &amp; Simonsen &amp; Solodovnikov" authorityYear="2022" box="[921,1167,2030,2049]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="altanai" status="sp. nov.">
<emphasis id="B954EAA4FFEE4B0DB38FA7EB16F4F79E" box="[921,1006,2030,2049]" italics="true" pageId="20" pageNumber="245">Q. altanai</emphasis>
Hansen &amp; Brunke
</taxonomicName>
, 
<taxonomicNameLabel id="A26757DFFFEE4B0DB481A7EB11C6F79E" box="[1175,1244,2030,2049]" pageId="20" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
; D – 
<taxonomicName id="4C204D35FFEE4B0DB510A7EB140CF783" authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="molochinoides">
<emphasis id="B954EAA4FFEE4B0DB510A7EB1086F79E" box="[1286,1436,2030,2049]" italics="true" pageId="20" pageNumber="245">Q. molochinoides</emphasis>
Smetana, 1965
</taxonomicName>
(red elytra variant); E – 
<taxonomicName id="4C204D35FFEE4B0DB1FFA80C1765F783" authorityName="Smetana" authorityYear="1965" box="[489,639,2057,2076]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="molochinoides">
<emphasis id="B954EAA4FFEE4B0DB1FFA80C1765F783" box="[489,639,2057,2076]" italics="true" pageId="20" pageNumber="245">Q. molochinoides</emphasis>
</taxonomicName>
(black elytra variant); F – 
<taxonomicName id="4C204D35FFEE4B0DB372A80C1160F783" authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" box="[868,1146,2057,2076]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="labradorensis">
<emphasis id="B954EAA4FFEE4B0DB372A80C16EEF783" box="[868,1012,2057,2076]" italics="true" pageId="20" pageNumber="245">Q. labradorensis</emphasis>
Smetana, 1965
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C33A653DFFEF4B0FB081A1A71702FF6E" lastPageId="22" lastPageNumber="247" pageId="21" pageNumber="246" type="materials_examined">
<paragraph id="8B9F36B6FFEF4B0CB081A1A7178EFD45" blockId="21.[151,761,416,917]" pageId="21" pageNumber="246">
<emphasis id="B954EAA4FFEF4B0CB081A1A71464FE2A" bold="true" box="[151,382,418,437]" pageId="21" pageNumber="246">Type material examined.</emphasis>
<taxonomicName id="4C204D35FFEF4B0CB193A1A7172EFE2A" authorityName="Bernhauer" authorityYear="1908" box="[389,564,418,437]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEF4B0CB193A1A7172EFE2A" box="[389,564,418,437]" italics="true" pageId="21" pageNumber="246">Quedius curtipennis</emphasis>
</taxonomicName>
: Lൾർඍඈඍඒඉൾ: (J, here designated): “Suderö 
<collectingCountry id="F3377626FFEF4B0CB142A1B81496FE4F" box="[340,396,445,464]" name="Faroe Islands" pageId="21" pageNumber="246">Faroer</collectingCountry>
Ins./ Dr. Cornu 
<date id="FF9E1076FFEF4B0CB208A1B8177CFE4F" box="[542,614,445,464]" bridgedPair="/" pageId="21" pageNumber="246" value="1907-05">
1907/ 
<taxonomicName id="4C204D35FFEF4B0CB240A1B8177CFE4F" authority="Brh." authorityName="Brh." box="[598,614,445,464]" pageId="21" pageNumber="246" rank="variety" variety="curtipennis">v.</taxonomicName>
</date>
curtipennis Brh. Typus / 
<taxonomicName id="4C204D35FFEF4B0CB0F4A1D214D6FE75" authority="Grav. Scheerp." authorityName="Grav. Scheerp." box="[226,460,471,490]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="fuliginosus">fuliginosus Grav. Scheerp.</taxonomicName>
/ Chicago NHMus M. Bernhauer Collection / D. Drugmand det. 1994 
<taxonomicName id="4C204D35FFEF4B0CB1C2A1F717E3FD9A" authority="Brnh." authorityName="Bernhauer" authorityYear="1908" box="[468,761,498,517]" class="Insecta" family="Staphylinidae" genus="Quedius" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="curtipennis">Quedius (s. str.) curtipennis Brnh.</taxonomicName>
/ 
<typeStatus id="549B8814FFEF4B0CB0B3A20815E1FDBF" box="[165,251,525,544]" pageId="21" pageNumber="246" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C204D35FFEF4B0CB115A2081415FDA5" authority="Bernhauer, 1908 A. K. Hansen &amp; A. Solodovnikov" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEF4B0CB115A20814A8FDBF" box="[259,434,525,544]" italics="true" pageId="21" pageNumber="246">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFEF4B0CB1ACA2081751FDBF" author="BERNHAUER M." box="[442,587,525,544]" pageId="21" pageNumber="246" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
A.K. Hansen &amp; A. Solodovnikov
</taxonomicName>
des. 2021” (FMNH). Pൺඋൺඅൾർඍඈඍඒඉൾඌ: “Nördl. 
<collectingCountry id="F3377626FFEF4B0CB2D6A22217E2FDA5" box="[704,760,551,570]" name="Faroe Islands" pageId="21" pageNumber="246">Faroer</collectingCountry>
Ins. / Dr. Cornu 
<date id="FF9E1076FFEF4B0CB132A247146AFDCA" box="[292,368,578,597]" bridgedPair="/" pageId="21" pageNumber="246" value="1907-05">
1907 / 
<taxonomicName id="4C204D35FFEF4B0CB176A247146AFDCA" authority="Brh." authorityName="Brh." box="[352,368,578,597]" pageId="21" pageNumber="246" rank="variety" variety="curtipennis">v.</taxonomicName>
</date>
curtipennis Brh. Typus / 
<taxonomicName id="4C204D35FFEF4B0CB25FA24717E3FDCA" authority="Scheerp." authorityName="Scheerp." box="[585,761,578,597]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="fuliginosus">fuliginosus Scheerp.</taxonomicName>
[sic!] det. [illegible] / Chicago NHMus M. Bernhauer Collection / D. Drugmand det. 1994 
<taxonomicName id="4C204D35FFEF4B0CB146A272176BFD15" authority="Brnh" authorityName="Bernhauer" authorityYear="1908" box="[336,625,631,650]" class="Insecta" family="Staphylinidae" genus="Quedius" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="curtipennis">Quedius (s. str.) curtipennis Brnh</taxonomicName>
/ 
<typeStatus id="549B8814FFEF4B0CB292A27217E2FD15" box="[644,760,631,650]" pageId="21" pageNumber="246" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C204D35FFEF4B0CB081A29717E2FD3A" authority="Bernhauer, 1908 A. K. Hansen &amp; A. Solodovnikov" authorityName="Bernhauer" authorityYear="1908" box="[151,760,658,677]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEF4B0CB081A297145EFD3A" box="[151,324,658,677]" italics="true" pageId="21" pageNumber="246">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFEF4B0CB15CA29714C2FD3A" author="BERNHAUER M." box="[330,472,658,677]" pageId="21" pageNumber="246" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
A.K. Hansen &amp; A. Solodovnikov
</taxonomicName>
des. 2021” (1 J FMNH); “ 
<taxonomicName id="4C204D35FFEF4B0CB16EA2A8178CFD5F" authority="Buchara Bang Haas" authorityName="Buchara Bang Haas" box="[376,662,685,704]" pageId="21" pageNumber="246" rank="variety" variety="curtipennis">v. curtipennis Buchara Bang Haas</taxonomicName>
det. Bernh. / Chicago NHMus M. Bernhauer Collection” (1 J FMNH).
</paragraph>
<paragraph id="8B9F36B6FFEF4B0CB0A3A2E71755FC0A" blockId="21.[151,761,416,917]" pageId="21" pageNumber="246">
<taxonomicName id="4C204D35FFEF4B0CB0A3A2E7144CFD6A" authorityName="Hatch" authorityYear="1957" box="[181,342,738,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEF4B0CB0A3A2E7144CFD6A" box="[181,342,738,757]" italics="true" pageId="21" pageNumber="246">Quedius parallelus</emphasis>
</taxonomicName>
: Hඈඅඈඍඒඉൾ: “Seattle, Wash. U.W.Campus 
<date id="FF9E1076FFEF4B0CB2D6A2E715D8FC8F" pageId="21" pageNumber="246" value="1949-04-12">IV-12- 1949</date>
/ Del. 1954 H. Houk / 
<typeStatus id="549B8814FFEF4B0CB196A2F814B0FC8F" box="[384,426,765,784]" pageId="21" pageNumber="246">Type</typeStatus>
J 
<taxonomicName id="4C204D35FFEF4B0CB1D5A2F817BAFC8F" authorityName="Hatch" authorityYear="1957" box="[451,672,765,784]" class="Insecta" family="Staphylinidae" genus="Quedius" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="parallelus">Quedius (s. str.) parallelus</taxonomicName>
- 1951 M. Hatch / 
<taxonomicName id="4C204D35FFEF4B0CB0C0A3121722FCB5" authority="Grav. M. Hatch" box="[214,568,791,810]" class="Insecta" family="Staphylinidae" genus="Quedius" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="fuliginosus">Quedius (s. str.) fuliginosus Grav.M. Hatch</taxonomicName>
– 1947” (1J UWBM). Pൺඋൺඍඒඉൾඌ: “Seattle,Wash. 
<date id="FF9E1076FFEF4B0CB16EA33714E8FCDA" box="[376,498,818,837]" pageId="21" pageNumber="246" value="1934-04-12">April 12, 1934</date>
” (1J UWBM); “Seattle,Wash. U.W.Campus, 
<date id="FF9E1076FFEF4B0CB101A3481460FCFF" box="[279,378,845,864]" pageId="21" pageNumber="246" value="1947-04-11">IV-11-1947</date>
” (1 J UWBM, N. Vowles coll.); “Vancouver, B.C., 5 March, 1949, G.B. Rich” (1 J CNC); “Wash. Seattle, 
<date id="FF9E1076FFEF4B0CB2A7A36215DEFC0A" pageId="21" pageNumber="246" value="1954-02-15">Feb. 15, 1954</date>
, B. Malkin” (
<specimenCount id="9D26FD3FFFEF4B0CB12CA3871443FC0A" box="[314,345,892,917]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
FMNH, ex coll. B. Malkin).
</paragraph>
<paragraph id="8B9F36B6FFEF4B0FB081A3A91702FF6E" blockId="21.[150,761,940,2079]" lastBlockId="22.[151,760,189,241]" lastPageId="22" lastPageNumber="247" pageId="21" pageNumber="246">
<emphasis id="B954EAA4FFEF4B0CB081A3A91729FC20" bold="true" box="[151,563,940,959]" pageId="21" pageNumber="246">
Additional material examined. 
<collectingCountry id="F3377626FFEF4B0CB1D4A3A9172AFC20" box="[450,560,940,959]" name="Armenia" pageId="21" pageNumber="246">ARMENIA</collectingCountry>
:
</emphasis>
Arzakanskaja dolina, Aghveran, [40.51, 44.56], 
<date id="FF9E1076FFEF4B0CB16EA3C214CDFC45" box="[376,471,967,986]" pageId="21" pageNumber="246" value="1978-07-03">3.VII.1978</date>
, leg. R. Rous (
<specimenCount id="9D26FD3FFFEF4B0CB240A3C2176FFC46" box="[598,629,960,986]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
MCZ); NE Dilizhan nr. Haghartsin Monastery, 
<geoCoordinate id="EE145071FFEF4B0CB1A4A3E414E3FC6B" box="[434,505,993,1012]" degrees="40.8013" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="40.8013">40.8013</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB217A3E41752FC6B" box="[513,584,993,1012]" degrees="44.8919" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="44.8919">44.8919</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB247A3E41789FC6B" box="[593,659,993,1012]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="246" unit="m" value="1450.0">1450 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB28DA3E417EFFC6B" box="[667,757,993,1012]" pageId="21" pageNumber="246" value="2001-05-21">21.V.2001</date>
, leg. Shaverdo &amp; Schillhammer (4 JJ NMW); 
<quantity id="4CD89B53FFEF4B0CB23CA3F9177AFB90" box="[554,608,1020,1039]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="246" unit="km" value="18.0">18 km</quantity>
E Dilizhan, 
<quantity id="4CD89B53FFEF4B0CB2DAA3F917E2FB90" box="[716,760,1020,1039]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="246" unit="km" value="3.0">3 km</quantity>
Gosh, 
<geoCoordinate id="EE145071FFEF4B0CB0DBA412140EFBB5" box="[205,276,1047,1066]" degrees="40.7405" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="40.7405">40.7405</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB10AA4121479FBB5" box="[284,355,1047,1066]" degrees="45.0191" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="45.0191">45.0191</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB17CA41214B6FBB5" box="[362,428,1047,1066]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="21" pageNumber="246" unit="m" value="1850.0">1850 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB1A5A4121716FBB5" box="[435,524,1047,1066]" pageId="21" pageNumber="246" value="2001-05-27">27.V.2001</date>
, leg. Shaverdo (1 J NMW); 
<quantity id="4CD89B53FFEF4B0CB081A43415CAFBDB" box="[151,208,1073,1092]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="246" unit="km" value="21.0">21 km</quantity>
N Hrazdan, Ankavan, 
<geoCoordinate id="EE145071FFEF4B0CB1B7A43414F3FBDB" box="[417,489,1073,1092]" degrees="40.6286" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="40.6286">40.6286</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB1E3A4341727FBDB" box="[501,573,1073,1092]" degrees="44.5533" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="44.5533">44.5533</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB25EA4341795FBDB" box="[584,655,1073,1092]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="246" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB28CA43417EEFBDB" box="[666,756,1073,1092]" pageId="21" pageNumber="246" value="2001-05-22">22.V.2001</date>
, leg. Shaverdo &amp; Schillhammer (1 J NMW); Teghut, Akhtala, 
<geoCoordinate id="EE145071FFEF4B0CB2BBA44917EEFBC0" box="[685,756,1100,1119]" degrees="41.0901" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="41.0901">41.0901</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB081A46215C4FBE5" box="[151,222,1127,1146]" degrees="44.8120" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="44.812">44.8120</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB0FEA4621431FBE5" box="[232,299,1127,1146]" metricMagnitude="3" metricUnit="m" metricValue="1.009" pageId="21" pageNumber="246" unit="m" value="1009.0">1009 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB122A462148DFBE5" box="[308,407,1127,1146]" pageId="21" pageNumber="246" value="2016-06-28">28.VI.2016</date>
, leg. J. Müller (1 J cSch); same locality, 
<geoCoordinate id="EE145071FFEF4B0CB081A48415C7FB0B" box="[151,221,1153,1172]" degrees="41.0911" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="41.0911">41.0911</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB0F3A4841436FB0B" box="[229,300,1153,1172]" degrees="44.8131" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="44.8131">44.8131</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB125A484146AFB0B" box="[307,368,1153,1172]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="21" pageNumber="246" unit="m" value="1020.0">1020m</quantity>
, 
<date id="FF9E1076FFEF4B0CB161A48414D5FB0B" box="[375,463,1153,1172]" pageId="21" pageNumber="246" value="2016-05-20">20.V.2016</date>
, leg.A. &amp; J. Müller (
<specimenCount id="9D26FD3FFFEF4B0CB26AA48417B2FB0B" box="[636,680,1147,1172]" pageId="21" pageNumber="246" type="female">2♀♀</specimenCount>
ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB081A49915E4FB30" bold="true" box="[151,254,1180,1199]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB081A49915E0FB30" box="[151,250,1180,1199]" name="Austria" pageId="21" pageNumber="246">AUSTRIA</collectingCountry>
:
</emphasis>
Bisamberg, [48.33, 16.36], leg. Luze (
<specimenCount id="9D26FD3FFFEF4B0CB253A4991778FB31" box="[581,610,1173,1199]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NMW); S 
<collectingRegion id="49E4F854FFEF4B0CB2A9A49915DFFB55" country="Austria" name="Steiermark" pageId="21" pageNumber="246">Steiermark</collectingRegion>
, [46.91, 15.60] (1 J NMW); 
<collectingRegion id="49E4F854FFEF4B0CB1DFA4B214EFFB55" box="[457,501,1207,1226]" country="Austria" name="Tirol" pageId="21" pageNumber="246">Tirol</collectingRegion>
, Musau, [47.53, 10.67], litter, moss, and grass, 28.C.1995, leg. M.Uhlig (1 J ZMHB); Wolfpassing, [48.08, 15.06], 
<date id="FF9E1076FFEF4B0CB132A4E91474FB60" box="[292,366,1260,1279]" pageId="21" pageNumber="246" value="1965-06">VI.1965</date>
, leg. Wewalka (
<specimenCount id="9D26FD3FFFEF4B0CB213A4E9173DFB61" box="[517,551,1253,1279]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NMW). 
<emphasis id="B954EAA4FFEF4B0CB26DA4E917E2FB60" bold="true" box="[635,760,1260,1279]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB26DA4E917EEFB60" box="[635,756,1260,1279]" name="Bulgaria" pageId="21" pageNumber="246">BULGARIA</collectingCountry>
:
</emphasis>
Rila Monastery, [42.13, 23.34], 
<quantity id="4CD89B53FFEF4B0CB1BBA50214EBFA85" box="[429,497,1287,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="21" pageNumber="246" unit="m" value="1150.0">1150 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB1ECA502177FFA85" box="[506,613,1287,1306]" pageId="21" pageNumber="246" value="1965-07-14">14.VII.1965</date>
, leg. P. Jørum (1 J NHMD). 
<emphasis id="B954EAA4FFEF4B0CB116A5241729FAAB" bold="true" box="[256,563,1313,1332]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB116A5241735FAAB" box="[256,559,1313,1332]" name="Bosnia and Herzegovina" pageId="21" pageNumber="246">BOSNIA AND HERZEGOVINA</collectingCountry>
:
</emphasis>
Zeleznica [Zeljeznica] n. Foinica, [43.89, 17.95], 
<quantity id="4CD89B53FFEF4B0CB195A53914C6FAD0" box="[387,476,1340,1359]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="21" pageNumber="246" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500-700m</quantity>
, 
<date id="FF9E1076FFEF4B0CB1F1A539175DFAD0" box="[487,583,1340,1359]" pageId="21" pageNumber="246" value="1969-04-13">13.IV.1969</date>
, leg. U. Heinz (1 J ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB0FDA5521715FAF5" bold="true" box="[235,527,1367,1386]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB0FDA5521456FAF5" box="[235,332,1367,1386]" name="Canada" pageId="21" pageNumber="246">CANADA</collectingCountry>
: BඋංඍංඌH Cඈඅඎආൻංൺ:
</emphasis>
Stanley Park, Vancouver, [49.30, -123.14], 
<date id="FF9E1076FFEF4B0CB13DA5741499FA1B" box="[299,387,1393,1412]" pageId="21" pageNumber="246" value="1954-06-09">9.VI.1954</date>
, leg. D. Lazorko (1 J 
<specimenCount id="9D26FD3FFFEF4B0CB256A5741745FA1B" box="[576,607,1387,1412]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NMW). 
<emphasis id="B954EAA4FFEF4B0CB2BAA5741416FA00" bold="true" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB2BAA5741412FA00" name="Czech Republic" pageId="21" pageNumber="246">CZECH REPUBLIC</collectingCountry>
:
</emphasis>
Brandeis, [50.18, 14.65], leg. Skalitzky (2 JJ NMW). 
<emphasis id="B954EAA4FFEF4B0CB081A5A21415FA25" bold="true" box="[151,271,1447,1466]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB081A5A21410FA25" box="[151,266,1447,1466]" name="Denmark" pageId="21" pageNumber="246">DENMARK</collectingCountry>
:
</emphasis>
Vigsø Bugt, 
<quantity id="4CD89B53FFEF4B0CB16CA5A214BEFA25" box="[378,420,1447,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="246" unit="km" value="1.0">1 km</quantity>
E of Vigsø, 
<geoCoordinate id="EE145071FFEF4B0CB21EA5A21755FA25" box="[520,591,1447,1466]" degrees="57.0950" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="57.095">57.0950</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB240A5A21788FA25" box="[598,658,1447,1466]" degrees="8.7394" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="8.7394">8.7394</geoCoordinate>
, pine forest, pitfall traps, 
<date id="FF9E1076FFEF4B0CB0E9A5C41498FA4B" box="[255,386,1473,1492]" pageId="21" pageNumber="246" value="2014-06-16" valueMax="2014-06-18" valueMin="2014-06-16">16.-18.VI.2014</date>
, leg.A.Hansen, M.Justesen,A.Solodovnikov &amp; L.Kraemer (1 J NHMD) 
<emphasis id="B954EAA4FFEF4B0CB16BA5D914A2FA70" bold="true" box="[381,440,1500,1519]" pageId="21" pageNumber="246">LඔඌØ:</emphasis>
Laesø Klitplantage, 
<geoCoordinate id="EE145071FFEF4B0CB276A5D917BDFA70" box="[608,679,1500,1519]" degrees="57.3082" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="57.3082">57.3082</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB2B8A5D917EEFA70" box="[686,756,1500,1519]" degrees="11.0081" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="11.0081">11.0081</geoCoordinate>
, mixed forest, sifting litter, 
<date id="FF9E1076FFEF4B0CB166A5F214D2F995" box="[368,456,1527,1546]" pageId="21" pageNumber="246" value="2013-05-28">28.V.2013</date>
, leg.A. Solodovnikov &amp; M.Justesen (1 NHMD). 
<emphasis id="B954EAA4FFEF4B0CB0EBA61414B6F9BB" bold="true" box="[253,428,1553,1572]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB0EBA61414B2F9BB" box="[253,424,1553,1572]" name="Faroe Islands" pageId="21" pageNumber="246">FAROE ISLANDS</collectingCountry>
:
</emphasis>
Suderö [Suðuroy], [61.51, -6.83], 1907, leg. Cornu (1 J FMNH). 
<emphasis id="B954EAA4FFEF4B0CB165A62914C9F9A0" bold="true" box="[371,467,1580,1599]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB165A62914D5F9A0" box="[371,463,1580,1599]" name="France" pageId="21" pageNumber="246">FRANCE</collectingCountry>
:
</emphasis>
Ain, Torcieu n. Amberieu, [45.91, 5.39] (1 J ZMHB); 
<collectingRegion id="49E4F854FFEF4B0CB15FA6421482F9C5" box="[329,408,1607,1626]" country="France" name="Bretagne" pageId="21" pageNumber="246">Bretagne</collectingRegion>
, Locronan, [48.09, -4.21], 
<date id="FF9E1076FFEF4B0CB269A64217D4F9C5" box="[639,718,1607,1626]" pageId="21" pageNumber="246" value="1983-07">VII.1983</date>
, leg. H.Korge (1 J ZMHB); Chigny n. Oise, [49.91, 3.76], leg. Gelchert (
<specimenCount id="9D26FD3FFFEF4B0CB2F8A66415B3F911" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZMHB); Morlaix, 
<collectingRegion id="49E4F854FFEF4B0CB15DA6791480F910" box="[331,410,1660,1679]" country="France" name="Bretagne" pageId="21" pageNumber="246">Bretagne</collectingRegion>
, [48.57, -3.82], 
<date id="FF9E1076FFEF4B0CB237A679179DF910" box="[545,647,1660,1679]" pageId="21" pageNumber="246" value="1950-08-01">1.VIII.1950</date>
, leg. A. Nilsson (2 JJ MZLU). 
<emphasis id="B954EAA4FFEF4B0CB159A69214A4F935" bold="true" box="[335,446,1687,1706]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB159A69214A0F935" box="[335,442,1687,1706]" name="Georgia" pageId="21" pageNumber="246">GEORGIA</collectingCountry>
:
</emphasis>
Borzhom Distr., Bakuriani, [41.75, 43.52], 
<date id="FF9E1076FFEF4B0CB0CDA6B4145FF95B" box="[219,325,1713,1732]" pageId="21" pageNumber="246" value="1947-07-27">27.VII.1947</date>
, leg. Bogachev (1 J 
<specimenCount id="9D26FD3FFFEF4B0CB213A6B4173DF95B" box="[517,551,1707,1732]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZIN); Kartliskij Chreb. [Kartalinskiy Khrebet], Sabaduris Tye [Sabaduri Pass] [41.91, 44.91], 
<quantity id="4CD89B53FFEF4B0CB081A6E215CFF965" box="[151,213,1767,1786]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="246" unit="m" value="1800.0">1800m</quantity>
, 
<date id="FF9E1076FFEF4B0CB0C9A6E2142DF965" box="[223,311,1767,1786]" pageId="21" pageNumber="246" value="1987-06-08">8.VI.1987</date>
, leg. Wrase (1 J cSch); same locality, 
<date id="FF9E1076FFEF4B0CB284A6E217EFF965" box="[658,757,1767,1786]" pageId="21" pageNumber="246" value="1988-06-25">25.VI.1988</date>
, leg. Wrase (1 J ZMHB); Trialetski, Khrebet Bakuriani, [41.74, 43.83], 
<quantity id="4CD89B53FFEF4B0CB081A719141FF8B0" box="[151,261,1820,1839]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="21" pageNumber="246" unit="m" value="2000.0" valueMax="2200.0" valueMin="1800.0">1800-2200m</quantity>
, 
<date id="FF9E1076FFEF4B0CB11BA7191499F8B0" box="[269,387,1820,1839]" pageId="21" pageNumber="246" value="1986-07-04" valueMax="1986-07-07" valueMin="1986-07-04">4.-7.VII.1986</date>
, leg.Wrase &amp; Schülke (8JJ 
<specimenCount id="9D26FD3FFFEF4B0CB26AA71917BDF8B1" box="[636,679,1813,1839]" pageId="21" pageNumber="246" type="female">4♀♀</specimenCount>
ZMHB); Zchneti [Tskneti], [41.69, 44.69], 
<quantity id="4CD89B53FFEF4B0CB1AAA73214E3F8D5" box="[444,505,1847,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="246" unit="m" value="1200.0">1200m</quantity>
, 
<date id="FF9E1076FFEF4B0CB214A732177CF8D5" box="[514,614,1847,1866]" pageId="21" pageNumber="246" value="1986-06-26">26.VI.1986</date>
, leg. Wrase (1 J cSch). 
<emphasis id="B954EAA4FFEF4B0CB0C4A754145DF8FB" bold="true" box="[210,327,1873,1892]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB0C4A754145EF8FB" box="[210,324,1873,1892]" name="Germany" pageId="21" pageNumber="246">GERMANY</collectingCountry>
:
</emphasis>
<collectingRegion id="49E4F854FFEF4B0CB15AA7541496F8FB" box="[332,396,1873,1892]" country="Germany" name="Bayern" pageId="21" pageNumber="246">Bayern</collectingRegion>
, Garmisch, [47.48, 11.08], 
<date id="FF9E1076FFEF4B0CB266A75417CBF8FB" box="[624,721,1873,1892]" pageId="21" pageNumber="246" value="1932-04-27">27.IV.1932</date>
, leg. Ihssen (1 J ZMHB); Bonn, [50.72, 7.09], X, leg. Verhoeff (
<specimenCount id="9D26FD3FFFEF4B0CB29DA76917B1F8E1" box="[651,683,1893,1919]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZMHB); 
<collectingRegion id="49E4F854FFEF4B0CB081A78215CFF805" box="[151,213,1927,1946]" country="Germany" name="Bayern" pageId="21" pageNumber="246">Bayern</collectingRegion>
, Eschenlohe, [47.58, 11.18], 
<date id="FF9E1076FFEF4B0CB1D7A782173FF805" box="[449,549,1927,1946]" pageId="21" pageNumber="246" value="1937-08-03">3.VIII.1937</date>
, leg.Ihssen (1J ZMHB); 
<collectingRegion id="49E4F854FFEF4B0CB081A7A41412F82B" box="[151,264,1953,1972]" country="Germany" name="Brandenburg" pageId="21" pageNumber="246">Brandenburg</collectingRegion>
, Strausberg, [52.52, 13.85], 
<date id="FF9E1076FFEF4B0CB1E3A7A4174FF82B" box="[501,597,1953,1972]" pageId="21" pageNumber="246" value="1987-04-17">17.IV.1987</date>
, leg. M. Schülke (1 J ZMHB); Dresden, Kötschenbroda,[51.11, 13.61], moss, 
<date id="FF9E1076FFEF4B0CB296A7B917C8F850" box="[640,722,1980,1999]" pageId="21" pageNumber="246" value="1920-01-23">23.I.1920</date>
, leg. Dahl (1 J ZMHB); Düsseldrof Unterbach, [51.20, 6.89], 
<date id="FF9E1076FFEF4B0CB29DA7D217EEF875" box="[651,756,2007,2026]" pageId="21" pageNumber="246" value="1974-07-27">27.VII.1974</date>
, leg. H.Bremer (
<specimenCount id="9D26FD3FFFEF4B0CB134A7F41458F79B" box="[290,322,2027,2052]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZMHB); Marburg, [50.80, 8.76], 
<date id="FF9E1076FFEF4B0CB278A7F417D7F79B" box="[622,717,2033,2052]" pageId="21" pageNumber="246" value="1904-04-16">16.IV.1904</date>
, leg. Strand (
<specimenCount id="9D26FD3FFFEF4B0CB0CAA80915E0F781" box="[220,250,2053,2079]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZMHB); Sarstadt, [52.24, 9.85], 
<date id="FF9E1076FFEF4B0CB202A809176DF780" box="[532,631,2060,2079]" pageId="21" pageNumber="246" value="1960-09-12">12.IX.1960</date>
, leg. Kuntze (1 J ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB389A0C616FFFF49" bold="true" box="[927,997,195,214]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB389A0C616FAFF49" box="[927,992,195,214]" name="Italy" pageId="21" pageNumber="246">ITALY</collectingCountry>
:
</emphasis>
Aniene, 
<collectingRegion id="49E4F854FFEF4B0CB426A0C61179FF49" box="[1072,1123,195,214]" country="Italy" name="Lazio" pageId="21" pageNumber="246">Lazio</collectingRegion>
, [41.95, 12.82], 
<quantity id="4CD89B53FFEF4B0CB4FBA0C61035FF49" box="[1261,1327,195,214]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="246" unit="m" value="1200.0">1200 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB521A0C61083FF49" box="[1335,1433,195,214]" pageId="21" pageNumber="246" value="1964-03-18">18.III.1964</date>
, leg. A. Vigna-Taglianti (1 J cBor); Barni, 
<collectingRegion id="49E4F854FFEF4B0CB4ACA0DB1000FF6E" box="[1210,1306,222,241]" country="Italy" name="Lombardia" pageId="21" pageNumber="246">Lombardia</collectingRegion>
, [45.91, 9.26], 
<date id="FF9E1076FFEF4B0CB32AA0FC168FFE93" box="[828,917,249,268]" pageId="21" pageNumber="246" value="2001-02-11">11.II.2001</date>
, leg. Diotti (1 J 
<specimenCount id="9D26FD3FFFEF4B0CB43EA0FC1153FE94" box="[1064,1097,242,268]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NHMD); Moliterno, 
<collectingRegion id="49E4F854FFEF4B0CB517A0FC104DFE93" box="[1281,1367,249,268]" country="Italy" name="Basilicata" pageId="21" pageNumber="246">Basilicata</collectingRegion>
, [40.24, 15.86], 
<date id="FF9E1076FFEF4B0CB368A11616FBFEB9" box="[894,993,275,294]" pageId="21" pageNumber="246" value="2000-09-30">30.IX.2000</date>
, leg. F. Angelini (
<specimenCount id="9D26FD3FFFEF4B0CB46BA1161184FEB9" box="[1149,1182,269,294]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NHMD); Pignola, 
<collectingRegion id="49E4F854FFEF4B0CB555A1161083FEB9" box="[1347,1433,275,294]" country="Italy" name="Basilicata" pageId="21" pageNumber="246">Basilicata</collectingRegion>
, [40.57, 15.78], 
<quantity id="4CD89B53FFEF4B0CB3A8A12B16ECFEDE" box="[958,1014,302,321]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="246" unit="m" value="700.0">700 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB3E8A12B1144FEDE" box="[1022,1118,302,321]" pageId="21" pageNumber="246" value="1998-04-19">19.IV.1998</date>
, leg. F. Angelini (1 J NHMD); Querciola, 
<collectingRegion id="49E4F854FFEF4B0CB365A14C16A6FEC3" box="[883,956,329,348]" country="Italy" name="Toscana" pageId="21" pageNumber="246">Toscana</collectingRegion>
, [43.84, 11.32], 
<date id="FF9E1076FFEF4B0CB446A14C11A8FEC3" box="[1104,1202,329,348]" pageId="21" pageNumber="246" value="2005-04-13">13.IV.2005</date>
, leg. Bordoni (1 J cBor). 
<emphasis id="B954EAA4FFEF4B0CB32AA166168EFEE9" bold="true" box="[828,916,355,374]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB32AA166168AFEE9" box="[828,912,355,374]" name="Latvia" pageId="21" pageNumber="246">LATVIA</collectingCountry>
:
</emphasis>
Curonia Libau [
<collectingRegion id="49E4F854FFEF4B0CB436A166117FFEE9" box="[1056,1125,355,374]" country="Latvia" name="Liepaja" pageId="21" pageNumber="246">Liepāja</collectingRegion>
], [56.49, 21.01], 
<date id="FF9E1076FFEF4B0CB4E1A1661051FEE9" box="[1271,1355,355,374]" pageId="21" pageNumber="246" value="1910-04-04">4.IV.1910</date>
(1 J 
<specimenCount id="9D26FD3FFFEF4B0CB56AA1661087FEE9" box="[1404,1437,349,374]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB39CA17B111FFE0E" bold="true" box="[906,1029,382,401]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB39CA17B111BFE0E" box="[906,1025,382,401]" name="Morocco" pageId="21" pageNumber="246">MOROCCO</collectingCountry>
:
</emphasis>
Cirque du Jaafar, [32.55, -4.91], 
<date id="FF9E1076FFEF4B0CB50BA17B106BFE0E" box="[1309,1393,382,401]" pageId="21" pageNumber="246" value="1964-03-04">4.III.1964</date>
(1 J NMW) [potentially mislabelled or introduced specimen clearly belonging to this species]. 
<emphasis id="B954EAA4FFEF4B0CB3D8A1B61153FE59" bold="true" box="[974,1097,435,454]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB3D8A1B6115FFE59" box="[974,1093,435,454]" name="Myanmar" pageId="21" pageNumber="246">MYANMAR</collectingCountry>
:
</emphasis>
Mountains, Tenasserim, Siam Border, [11.79, 99.62], 
<date id="FF9E1076FFEF4B0CB3A8A1CB110CFE7E" box="[958,1046,462,481]" pageId="21" pageNumber="246" value="1913-02" valueMax="1913-05-31" valueMin="1913-02-00">II.-V.1913</date>
, leg. K.G. Gairdner (1 J BMNH) [potentially mislabelled or introduced specimen clearly belonging to this species]. 
<emphasis id="B954EAA4FFEF4B0CB56EA1EC1665FD89" bold="true" pageId="21" pageNumber="246">PO-LAND:</emphasis>
Dunajec River n. Sromowce Nizne, [49.39, 20.39], 
<date id="FF9E1076FFEF4B0CB539A2061083FD89" box="[1327,1433,515,534]" pageId="21" pageNumber="246" value="1977-07-18">18.VII.1977</date>
, leg. F. Hieke (3 JJ 
<specimenCount id="9D26FD3FFFEF4B0CB3E6A21B1139FDAE" box="[1008,1059,536,561]" pageId="21" pageNumber="246" type="female">4 ♀♀</specimenCount>
ZMHB); Wyspowy, Krzozonowka River n. Poim, [49.69, 20.24] 
<quantity id="4CD89B53FFEF4B0CB3EBA23C112BFDD3" box="[1021,1073,569,588]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="246" unit="m" value="300.0">300m</quantity>
, 
<date id="FF9E1076FFEF4B0CB42AA23C11B5FDD3" box="[1084,1199,569,588]" pageId="21" pageNumber="246" value="1980-08-26">26.VIII.1980</date>
, leg. Hieke (1 J ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB32AA2561114FDF9" bold="true" box="[828,1038,595,614]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB32AA25616ADFDF9" box="[828,951,595,614]" name="Portugal" pageId="21" pageNumber="246">PORTUGAL</collectingCountry>
: Aඓඈඋൾඌ:
</emphasis>
Santa Maria, 
<quantity id="4CD89B53FFEF4B0CB492A25611B5FDF9" box="[1156,1199,595,614]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="246" unit="km" value="2.0">2 km</quantity>
ENE Almagreira, Miradoro dos Picos, [36.97, -25.07], 
<quantity id="4CD89B53FFEF4B0CB434A26B1140FD1E" box="[1058,1114,622,641]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="21" pageNumber="246" unit="m" value="440.0">440 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB474A26B11D9FD1E" box="[1122,1219,622,641]" pageId="21" pageNumber="246" value="1957-03-19">19.III.1957</date>
, leg. Brinck &amp; Dahl (1 J MZLU); São Miguel, 
<quantity id="4CD89B53FFEF4B0CB3EFA28C112AFD03" box="[1017,1072,649,668]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="246" unit="m" value="500.0">500 m</quantity>
E of Lagoa do Pau Pique, [37.83, -25.74], pond, 
<date id="FF9E1076FFEF4B0CB367A2A616DDFD29" box="[881,967,675,694]" pageId="21" pageNumber="246" value="1957-03-07">7.III.1957</date>
, leg. Brinck &amp; Dahl (1 J MZLU); São Miguel, Pico de Vara, 
<geoCoordinate id="EE145071FFEF4B0CB37AA2BB16A8FD4E" box="[876,946,702,721]" degrees="37.8119" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="37.8119">37.8119</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB3AEA2BB111CFD4E" box="[952,1030,702,721]" degrees="25.2105" direction="west" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="-25.2105">-25.2105</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB41BA2BB1151FD4E" box="[1037,1099,702,721]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="21" pageNumber="246" unit="m" value="1050.0">1050m</quantity>
, boggy slope 
<date id="FF9E1076FFEF4B0CB4ACA2BB1039FD4E" box="[1210,1315,702,721]" pageId="21" pageNumber="246" value="2013-07-24">24.VII.2013</date>
, leg.V.Assing (
<specimenCount id="9D26FD3FFFEF4B0CB354A2DC166EFD74" box="[834,884,722,748]" pageId="21" pageNumber="246" type="female">4 ♀♀</specimenCount>
ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB3DAA2DC113CFD73" bold="true" box="[972,1062,729,748]" pageId="21" pageNumber="246">Mൺൽൾංඋൺ:</emphasis>
Levada da Serra do Faial, [32.68, -16.85], creek, 
<quantity id="4CD89B53FFEF4B0CB361A2F616B0FC99" box="[887,938,755,774]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="21" pageNumber="246" unit="m" value="820.0">820m</quantity>
, 
<date id="FF9E1076FFEF4B0CB3A3A2F6110DFC99" box="[949,1047,755,774]" pageId="21" pageNumber="246" value="2014-09-11">11.IX.2014</date>
, leg. A. Kleeberg (1 J cSch). 
<emphasis id="B954EAA4FFEF4B0CB53DA2F61086FC99" bold="true" box="[1323,1436,755,774]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB53DA2F61083FC99" box="[1323,1433,755,774]" name="Romania" pageId="21" pageNumber="246">ROMANIA</collectingCountry>
:
</emphasis>
Klaussenburg [
<collectingRegion id="49E4F854FFEF4B0CB3AFA30B1136FCBE" box="[953,1068,782,801]" country="Romania" name="Cluj" pageId="21" pageNumber="246">Cluj-Napoca</collectingRegion>
], [46.76, 23.62], leg.Verhoeff (1 J ZMHB); 
<collectingRegion id="49E4F854FFEF4B0CB32AA32C16AFFCA3" box="[828,949,809,828]" country="Romania" name="Sibiu" pageId="21" pageNumber="246">Hermannstadt</collectingRegion>
, Siebenbürgen [
<collectingRegion id="49E4F854FFEF4B0CB42DA32C1177FCA3" box="[1083,1133,809,828]" country="Romania" name="Sibiu" pageId="21" pageNumber="246">Sibiu</collectingRegion>
], [45.79, 24.15], leg. Skalitzky (1 J NMW); Piatra, Virful Tiganului, Sighetu Marmației, 
<collectingRegion id="49E4F854FFEF4B0CB4E0A346104CFCC9" box="[1270,1366,835,854]" country="Romania" name="Maramures" pageId="21" pageNumber="246">Maramureș</collectingRegion>
, [47.99, 23.53], 
<quantity id="4CD89B53FFEF4B0CB396A35B16DCFCEE" box="[896,966,862,881]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="246" unit="m" value="1300.0">1300 m</quantity>
, pasture, in rotten logs of beech, 
<date id="FF9E1076FFEF4B0CB4E5A35B104CFCEE" box="[1267,1366,862,881]" pageId="21" pageNumber="246" value="2006-06-14">14.VI.2006</date>
, leg. A. Grabant, Z. György, O. Merkl &amp; A. Podlussany (1 J NHMD). 
<emphasis id="B954EAA4FFEF4B0CB551A37C110AFC39" bold="true" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB551A37C1082FC13" box="[1351,1432,889,908]" name="Russia" pageId="21" pageNumber="246">RUSSIA</collectingCountry>
: KൺඋൺർHൺඒ- 
<collectingCountry id="F3377626FFEF4B0CB388A39616ADFC39" box="[926,951,915,934]" name="Switzerland" pageId="21" pageNumber="246">CH</collectingCountry>
ൾඋKൾඌඌංൺ:
</emphasis>
Teberda,[43.44, 41.74], 
<date id="FF9E1076FFEF4B0CB4CAA3961038FC39" box="[1244,1314,915,934]" pageId="21" pageNumber="246" value="1912-06">VI.1912</date>
, leg.Roubal (
<specimenCount id="9D26FD3FFFEF4B0CB585A3961650FC5E" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
SNMC). 
<emphasis id="B954EAA4FFEF4B0CB38CA3AB1123FC5E" bold="true" box="[922,1081,942,961]" pageId="21" pageNumber="246">Mඈඌർඈඐ Oൻඅൺඌඍ:</emphasis>
Bykovo village, VNIIKR institute territory, [55.61, 38.05], 
<date id="FF9E1076FFEF4B0CB3AAA3CC110DFC43" box="[956,1047,969,988]" pageId="21" pageNumber="246" value="2013-10-16">16.X.2013</date>
, leg.S.A. Kurbatov (3 JJ ZIN); Odintsovsky Distr., near Zhavoronki vill., [55.64, 37.10], 
<date id="FF9E1076FFEF4B0CB4ADA3E61027FC69" box="[1211,1341,995,1014]" pageId="21" pageNumber="246" value="2017-08-06" valueMax="2017-08-13" valueMin="2017-08-06">6-13.VIII.2017</date>
, leg. E. M. Veselova &amp; A. B. Ryvkin (1 J ZIN). 
<emphasis id="B954EAA4FFEF4B0CB461A3FB1069FB8E" bold="true" box="[1143,1395,1022,1041]" pageId="21" pageNumber="246">NංඓHൾGඈඋඈൽඌKൺඃൺ Oൻඅൺඌඍ:</emphasis>
near Gorodets town, [56.64, 43.47], mixed forest, 
<date id="FF9E1076FFEF4B0CB4D7A41C1028FBB3" box="[1217,1330,1049,1068]" pageId="21" pageNumber="246" value="1989-08-17">17.VIII.1989</date>
, leg. A. Klimenko (1J ZIN). 
<emphasis id="B954EAA4FFEF4B0CB3C4A43611A2FBD9" bold="true" box="[978,1208,1075,1094]" pageId="21" pageNumber="246">NඈඏGඈඋඈൽඌKൺඃൺ Oൻඅൺඌඍ:</emphasis>
Torbino, Novgorod, [58.58, 32.87], 
<date id="FF9E1076FFEF4B0CB368A44B16C4FBFE" box="[894,990,1102,1121]" pageId="21" pageNumber="246" value="1914-04-10">10.IV.1914</date>
, leg. A. Filiniev (1 J cBor). 
<emphasis id="B954EAA4FFEF4B0CB4CFA44B1086FBFE" bold="true" box="[1241,1436,1102,1121]" pageId="21" pageNumber="246">NඈඋඍH Oඌඌൾඍංൺ Rൾඉ.:</emphasis>
Caucasus, Skalisty Khrebet, Fiagdon River, [42.87, 44.32], 
<quantity id="4CD89B53FFEF4B0CB556A46C1655FB09" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="21" pageNumber="246" unit="m" value="1950.0" valueMax="2200.0" valueMin="1700.0">1700-2200 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB34EA48616BAFB09" box="[856,928,1155,1174]" pageId="21" pageNumber="246" value="1976-06">VI.1976</date>
, leg. R. Rous (3 JJ 
<specimenCount id="9D26FD3FFFEF4B0CB44CA4861167FB09" box="[1114,1149,1149,1174]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
MCZ). 
<emphasis id="B954EAA4FFEF4B0CB4D5A4861023FB09" bold="true" box="[1219,1337,1155,1174]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB4D5A486102FFB09" box="[1219,1333,1155,1174]" name="Slovakia" pageId="21" pageNumber="246">SLOVAKIA</collectingCountry>
:
</emphasis>
Belá River n. Pribylina, [49.09, 19.81], 
<quantity id="4CD89B53FFEF4B0CB438A49B117AFB2E" box="[1070,1120,1182,1201]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="246" unit="m" value="600.0">600m</quantity>
, 
<date id="FF9E1076FFEF4B0CB47FA49B11C0FB2E" box="[1129,1242,1182,1201]" pageId="21" pageNumber="246" value="1981-08-18">18.VIII.1981</date>
, leg. Hieke (1 J 
<specimenCount id="9D26FD3FFFEF4B0CB57BA49B1086FB2E" box="[1389,1436,1176,1201]" pageId="21" pageNumber="246" type="female">3 ♀♀</specimenCount>
ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB39DA4BC16CAFB53" bold="true" box="[907,976,1209,1228]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB39DA4BC16D7FB53" box="[907,973,1209,1228]" name="Spain" pageId="21" pageNumber="246">SPAIN</collectingCountry>
:
</emphasis>
Bera de Bidasoa, 
<collectingRegion id="49E4F854FFEF4B0CB47DA4BC11A8FB53" box="[1131,1202,1209,1228]" country="Spain" name="Navarra" pageId="21" pageNumber="246">Navarra</collectingRegion>
, [43.27, -1.68], 
<date id="FF9E1076FFEF4B0CB52FA4BC1082FB53" box="[1337,1432,1209,1228]" pageId="21" pageNumber="246" value="2001-01-01" valueMax="2001-01-07" valueMin="2001-01-01">1.-7.I.2001</date>
, leg.A.Anichtchenko (1J NHMD); Caldetas n. Barcelona, [41.57, 2.52], leg. G. Heine (1 J ZMHB); La Molina, Pyrenees, [42.33, 1.93], 
<date id="FF9E1076FFEF4B0CB577A4EB1693FA83" pageId="21" pageNumber="246" value="1963-07-11" valueMax="1963-07-16" valueMin="1963-07-11">11.-16. VII.1963</date>
(1 J MZLU); O Rial, Serra do Mirador, [42.74, -7.83], 
<quantity id="4CD89B53FFEF4B0CB570A50C1083FA83" box="[1382,1433,1289,1308]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="246" unit="m" value="450.0">450m</quantity>
, 
<date id="FF9E1076FFEF4B0CB32AA526168EFAA9" box="[828,916,1315,1334]" pageId="21" pageNumber="246" value="2003-06-03">3.VI.2003</date>
, leg. J. P. Valcárcel (1 J cSch); Picos de Europa, Camping El Redondo [Fuente Dé], [43.14, 4.81], forest, 
<quantity id="4CD89B53FFEF4B0CB4B3A53B11FFFACE" box="[1189,1253,1342,1361]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="21" pageNumber="246" unit="m" value="1100.0">1100 m</quantity>
, 
<date id="FF9E1076FFEF4B0CB4FAA53B106CFACE" box="[1260,1398,1342,1361]" pageId="21" pageNumber="246" value="1996-07-14" valueMax="1996-07-17" valueMin="1996-07-14">14.-17.VII.1996</date>
, leg. Wrase (1 J cSch); Plaiaundi, Irun, Guipuzcoa, [43.34, -1.79], 
<date id="FF9E1076FFEF4B0CB54DA55C1083FAF3" box="[1371,1433,1369,1388]" pageId="21" pageNumber="246" value="2006-05">V.2006</date>
, A. Anichtchenko (1 J 
<specimenCount id="9D26FD3FFFEF4B0CB3E9A576113BFA19" box="[1023,1057,1389,1414]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NHMD); Pyrenees, Toses, 
<geoCoordinate id="EE145071FFEF4B0CB51AA5761049FA19" box="[1292,1363,1395,1414]" degrees="42.3502" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="42.3502">42.3502</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB54BA5761080FA19" box="[1373,1434,1395,1414]" degrees="1.9955" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="1.9955">1.9955</geoCoordinate>
, 
<quantity id="4CD89B53FFEF4B0CB32AA58B1663FA3E" box="[828,889,1422,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="21" pageNumber="246" unit="m" value="1660.0">1660m</quantity>
, fir-spruce forest with moss, 
<date id="FF9E1076FFEF4B0CB464A58B11D9FA3E" box="[1138,1219,1422,1441]" pageId="21" pageNumber="246" value="1997-10-09">9.X.1997</date>
, leg. Zerche (1 J SDEI); Sierra Ancares, [42.81, -6.86], 
<date id="FF9E1076FFEF4B0CB452A5AC11BAFA23" box="[1092,1184,1449,1468]" pageId="21" pageNumber="246" value="1984-10-20">20.X.1984</date>
, leg. J. C. Otero (6 JJ 
<specimenCount id="9D26FD3FFFEF4B0CB57BA5AC1087FA24" box="[1389,1437,1442,1468]" pageId="21" pageNumber="246" type="female">3 ♀♀</specimenCount>
ZMHB); Sierra Caurel, [42.59, -7.18], 
<date id="FF9E1076FFEF4B0CB49AA5C611ECFA49" box="[1164,1270,1475,1494]" pageId="21" pageNumber="246" value="1985-07-20">20.VII.1985</date>
, leg. J. C. Otero (5 JJ 
<specimenCount id="9D26FD3FFFEF4B0CB376A5DB1695FA6E" box="[864,911,1496,1521]" pageId="21" pageNumber="246" type="female">4 ♀♀</specimenCount>
ZMHB); Sierra de Gredos, Hoyos de Collado, [40.35, -5.21], 
<quantity id="4CD89B53FFEF4B0CB32AA5FC166DF993" box="[828,887,1529,1548]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="246" unit="m" value="1600.0">1600m</quantity>
, 
<date id="FF9E1076FFEF4B0CB368A5FC16C6F993" box="[894,988,1529,1548]" pageId="21" pageNumber="246" value="1991-04-14">14.IV.1991</date>
,leg.Heinz (
<specimenCount id="9D26FD3FFFEF4B0CB456A5FC1171F994" box="[1088,1131,1522,1548]" pageId="21" pageNumber="246" type="female">2♀♀</specimenCount>
ZMHB); Sierra Guadarrama, [40.85, -3.95], 
<date id="FF9E1076FFEF4B0CB36CA61616C9F9B9" box="[890,979,1555,1574]" pageId="21" pageNumber="246" value="1980-06-01">1.VI.1980</date>
, leg. C. Otero (2 JJ ZMHB); Sierra de la Demanda, Val de Zcaray [Ezcaray], [42.32, -3.02], forest, 
<quantity id="4CD89B53FFEF4B0CB4DBA62B1011F9DE" box="[1229,1291,1582,1601]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="21" pageNumber="246" unit="m" value="1650.0">1650m</quantity>
, 
<date id="FF9E1076FFEF4B0CB505A62B106FF9DE" box="[1299,1397,1582,1601]" pageId="21" pageNumber="246" value="1996-07-23">23.07.1996</date>
, leg. Zaballos &amp; Wrase (1 J cSch). 
<emphasis id="B954EAA4FFEF4B0CB42AA64C11BAF9C3" bold="true" box="[1084,1184,1609,1628]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB42AA64C1186F9C3" box="[1084,1180,1609,1628]" name="Sweden" pageId="21" pageNumber="246">SWEDEN</collectingCountry>
:
</emphasis>
Brönnestad, [56.08, 13.70] (1 MZLU); Omberg, [58.33, 14.64] (1 MZLU); Strömholm, [59.52, 16.23] (1 MZLU); Visseltofta, [56.42, 13.85] (1 MZLU); 
<emphasis id="B954EAA4FFEF4B0CB4E8A67B105EF90E" bold="true" box="[1278,1348,1662,1681]" pageId="21" pageNumber="246">Öඅൺඇൽ:</emphasis>
Vickleby, [56.57, 16.46] (1 MZLU). 
<emphasis id="B954EAA4FFEF4B0CB40BA69C11D9F933" bold="true" box="[1053,1219,1689,1708]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB40BA69C11DAF933" box="[1053,1216,1689,1708]" name="Switzerland" pageId="21" pageNumber="246">SWITZERLAND</collectingCountry>
:
</emphasis>
Wolfenschiessen, [46.91, 8.39], 
<date id="FF9E1076FFEF4B0CB367A6B616CCF959" box="[881,982,1715,1734]" pageId="21" pageNumber="246" value="1963-08-05">5.VIII.1963</date>
, moss (1 J ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB494A6B611FBF959" bold="true" box="[1154,1249,1715,1734]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB494A6B611C4F959" box="[1154,1246,1715,1734]" name="Turkey" pageId="21" pageNumber="246">TURKEY</collectingCountry>
:
</emphasis>
Abant n. 
<collectingRegion id="49E4F854FFEF4B0CB539A6B61040F959" box="[1327,1370,1715,1734]" country="Turkey" name="Bolu" pageId="21" pageNumber="246">Bolu</collectingRegion>
, [40.61, 31.27], 
<date id="FF9E1076FFEF4B0CB368A6CB16CDF97E" box="[894,983,1742,1761]" pageId="21" pageNumber="246" value="1964-05-30">30.V.1964</date>
, leg. H.Korge (1 J ZMHB); Akkus n. Ünye, [40.79, 37.01], 
<date id="FF9E1076FFEF4B0CB36CA6EC16CBF963" box="[890,977,1769,1788]" pageId="21" pageNumber="246" value="1964-05-28">28.V.1964</date>
,leg. H.Korge (1J 
<specimenCount id="9D26FD3FFFEF4B0CB467A6EC1186F964" box="[1137,1180,1762,1788]" pageId="21" pageNumber="246" type="female">2♀♀</specimenCount>
ZMHB); Ilgaz Dağları, [41.11, 33.90], 
<date id="FF9E1076FFEF4B0CB369A7061119F889" box="[895,1027,1795,1814]" pageId="21" pageNumber="246" value="1960-06-17" valueMax="1960-06-21" valueMin="1960-06-17">17.-21.VI.1960</date>
, leg. F. Schubert (
<specimenCount id="9D26FD3FFFEF4B0CB4B0A70611DDF889" box="[1190,1223,1789,1814]" pageId="21" pageNumber="246" type="female">1 ♀</specimenCount>
NMW); n. Ismetpasa &amp; Cerkes, [40.75, 32.82], 
<date id="FF9E1076FFEF4B0CB414A71B1178F8AE" box="[1026,1122,1822,1841]" pageId="21" pageNumber="246" value="1979-04-05">05.IV.1979</date>
, leg. Heinz (1 J NMEG); Kulakkaya n. 
<collectingRegion id="49E4F854FFEF4B0CB345A73C1682F8D3" box="[851,920,1849,1868]" country="Turkey" name="Giresun" pageId="21" pageNumber="246">Giresun</collectingRegion>
, [40.69, 38.33], 
<quantity id="4CD89B53FFEF4B0CB43CA73C117DF8D3" box="[1066,1127,1849,1868]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="246" unit="m" value="1450.0">1450m</quantity>
, 
<date id="FF9E1076FFEF4B0CB467A73C11C1F8D3" box="[1137,1243,1849,1868]" pageId="21" pageNumber="246" value="1963-07-27">27.VII.1963</date>
, leg. H. Korge (
<specimenCount id="9D26FD3FFFEF4B0CB57DA73C1086F8D4" box="[1387,1436,1842,1868]" pageId="21" pageNumber="246" type="female">2 ♀♀</specimenCount>
ZMHB). 
<emphasis id="B954EAA4FFEF4B0CB39BA75616E2F8F9" bold="true" box="[909,1016,1875,1894]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB39BA75616EFF8F9" box="[909,1013,1875,1894]" name="Ukraine" pageId="21" pageNumber="246">UKRAINE</collectingCountry>
:
</emphasis>
Tausban Bazar, [44.97, 34.59], 
<date id="FF9E1076FFEF4B0CB506A7561068F8F9" box="[1296,1394,1875,1894]" pageId="21" pageNumber="246" value="1907-06-20">20.VI.1907</date>
, leg. B. Grigoriev (1 J ZIN). 
<emphasis id="B954EAA4FFEF4B0CB41AA76B102DF81E" bold="true" box="[1036,1335,1902,1921]" pageId="21" pageNumber="246">
<collectingCountry id="F3377626FFEF4B0CB41AA76B11C9F81E" box="[1036,1235,1902,1921]" name="United Kingdom" pageId="21" pageNumber="246">UNITED KINGDOM</collectingCountry>
: EඇGඅൺඇൽ:
</emphasis>
Barnstaple, 
<collectingRegion id="49E4F854FFEF4B0CB32AA78C166CF803" box="[828,886,1929,1948]" country="United Kingdom" name="Devon" pageId="21" pageNumber="246">Devon</collectingRegion>
, [51.07, -4.06], 
<date id="FF9E1076FFEF4B0CB3EEA78C1168F803" box="[1016,1138,1929,1948]" pageId="21" pageNumber="246" value="1954-05-12" valueMax="1954-05-31" valueMin="1954-05-12">12.-31.V.1954</date>
, leg. Lindroth (1J MZLU); 
<collectingRegion id="49E4F854FFEF4B0CB549A78C1080F803" box="[1375,1434,1929,1948]" country="United Kingdom" name="Devon" pageId="21" pageNumber="246">Devon</collectingRegion>
, Bidefort [51.01, -4.21], leg. H. Korge (
<specimenCount id="9D26FD3FFFEF4B0CB493A7A611AEF829" box="[1157,1204,1949,1974]" pageId="21" pageNumber="246" type="female">2 ♀♀</specimenCount>
ZMHB); 
<collectingRegion id="49E4F854FFEF4B0CB51FA7A6102EF829" box="[1289,1332,1955,1974]" country="United Kingdom" name="Kent" pageId="21" pageNumber="246">Kent</collectingRegion>
, Knockholt, 
<geoCoordinate id="EE145071FFEF4B0CB32AA7BB1699F84E" box="[828,899,1982,2001]" degrees="51.3124" direction="north" orientation="latitude" pageId="21" pageNumber="246" precision="5" value="51.3124">51.3124</geoCoordinate>
, 
<geoCoordinate id="EE145071FFEF4B0CB39DA7BB16DDF84E" box="[907,967,1982,2001]" degrees="0.1191" direction="east" orientation="longitude" pageId="21" pageNumber="246" precision="5" value="0.1191">0.1191</geoCoordinate>
, 
<date id="FF9E1076FFEF4B0CB3D8A7BB1139F84E" box="[974,1059,1982,2001]" pageId="21" pageNumber="246" value="2011-04-02">2.IV.2011</date>
, leg. J.J. Shaw (1 J cJen); Shrops, Edgmond Harper Adams Uni. Campus, [52.77, -2.42], 
<date id="FF9E1076FFEF4B0CB4D4A7DC1003F873" box="[1218,1305,2009,2028]" pageId="21" pageNumber="246" value="2015-03-06">6.III.2015</date>
, leg. J.J. Shaw (1 J cJen); 
<emphasis id="B954EAA4FFEF4B0CB3BEA7F61111F799" bold="true" box="[936,1035,2035,2054]" pageId="21" pageNumber="246">Sർඈඍඅൺඇൽ:</emphasis>
Loch Hope, [58.44, -4.62], 
<date id="FF9E1076FFEF4B0CB511A7F6106BF799" box="[1287,1393,2035,2054]" pageId="21" pageNumber="246" value="1987-07-19">19.VII.1987</date>
, leg. G. Gillerfors (1 J MZLU). 
<collectingCountry id="F3377626FFEF4B0CB427A80B1145F7BE" box="[1073,1119,2062,2081]" name="United States of America" pageId="21" pageNumber="246">USA</collectingCountry>
: 
<collectingRegion id="49E4F854FFEF4B0CB47FA80B11B9F7BE" box="[1129,1187,2062,2081]" country="United States of America" name="Maine" pageId="21" pageNumber="246">Maine</collectingRegion>
: Booth Bay, [43.85, -69.62], 
<date id="FF9E1076FFEC4B0FB081A0C61410FF49" box="[151,266,195,214]" pageId="22" pageNumber="247" value="1982-08-15">15.VIII.1982</date>
, leg. Wewalka (
<specimenCount id="9D26FD3FFFEC4B0FB18AA0C614A4FF49" box="[412,446,189,214]" pageId="22" pageNumber="247" type="female">1 ♀</specimenCount>
NMW). 
<emphasis id="B954EAA4FFEC4B0FB207A0C617B9FF49" bold="true" box="[529,675,195,214]" pageId="22" pageNumber="247">
<collectingCountry id="F3377626FFEC4B0FB207A0C61785FF49" box="[529,671,195,214]" name="Uzbekistan" pageId="22" pageNumber="247">UZBEKISTAN</collectingCountry>
:
</emphasis>
Buchara, [39.77, 64.42], leg. Bang Haas (1 J FMNH).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0FB081A10F1784FADE" pageId="22" pageNumber="247" type="description">
<paragraph id="8B9F36B6FFEC4B0FB081A10F17E9FD1E" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">
<emphasis id="B954EAA4FFEC4B0FB081A10F1427FEBE" bold="true" box="[151,317,266,289]" pageId="22" pageNumber="247">Redescription.</emphasis>
Measurements JJ (n = 6): HW = 1.89– 1.96 (1.93); HL = 1.53–1.71 (1.61); HL/HW 0.81–0.88 (0.84); PW = 2.36–2.56 (2.49); PL = 2.16–2.36 (2.29); PL/PW 0.87–0.94 (0.92); EW = 2.51–2.67 (2.61); EL = 2.36–2.49 (2.44); EL/EW 0.92–0.96 (0.93); EL/PL 1.04– 1.10 (1.06); PW/HW 1.31–1.60 (1.49); forebody length 6.04–6.56 (6.34). J 
<specimenCount id="9D26FD3FFFEC4B0FB17CA1C61498FE7D" box="[362,386,451,482]" pageId="22" pageNumber="247" type="female">♀</specimenCount>
(n = 4): HW = 1.78–2.00 (1.89); HL = 1.51–1.64 (1.58); HL/HW 0.82–0.85 (0.84); PW = 2.22–2.56 (2.41); PL = 2.07–2.40 (2.18); PL/PW 0.86–0.94 (0.91); EW = 2.44–2.62 (2.52); EL = 2.20–2.49 (2.33); EL/EW 0.90–0.95 (0.92); EL/PL 1.03–1.15 (1.07); PW/ HW 1.47–1.55 (1.52); forebody length 5.78–6.53 (6.10).
</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB0A3A28F1770FD3E" blockId="22.[151,760,261,2081]" box="[181,618,650,673]" pageId="22" pageNumber="247">
Very large species; body black (
<figureCitation id="131B2A33FFEC4B0FB21FA28F1744FD3E" box="[521,606,650,673]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="22" pageNumber="247">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB0A3A2AF1797FCDE" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">
Head black, slightly transverse; eyes large (EyL/TL = 3.00–4.10 (3.55)) not protruding; microsculpture of transverse waves, four punctures between anterior frontal punctures (
<figureCitation id="131B2A33FFEC4B0FB118A30F1478FCBE" box="[270,354,778,801]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="22" pageNumber="247">Fig. 6B</figureCitation>
); antennae and palpi pale reddish, antennae long, all antennomeres clearly elongate.
</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB0A3A34F14EAFBDE" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">Thorax: pronotum black, slightly wider than long, clearly wider than head, microsculpture of transverse waves, three punctures in dorsal row and two in sublateral row with posteriormost puncture reaching level of middle puncture of dorsal row; scutellum smooth and glabrous; elytra black, uniformly pubescent, wider than long, of equal length as pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB0A3A44F141CFB1E" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">Abdomen black, tergites uniformly punctured, slightly iridescent.</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB0A3A48C1784FADE" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">
<emphasis id="B954EAA4FFEC4B0FB0A3A48C15EFFB3E" bold="true" box="[181,245,1161,1185]" italics="true" pageId="22" pageNumber="247">Male.</emphasis>
Aedeagus (
<figureCitation id="131B2A33FFEC4B0FB16BA48F14F9FB3E" box="[381,483,1162,1185]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="22" pageNumber="247">Fig. 11C</figureCitation>
): paramere broad, rather parallel-sided, reaching apex of median lobe, with sensory peg setae forming two single rows following parameral lateral margins for 2/3 of their length; median lobe with apical part dorsoventrally flattened to form a plate-like structure; internal sac with two pairs of sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0FB081A54F102BFDCA" pageId="22" pageNumber="247" type="diagnosis">
<paragraph id="8B9F36B6FFEC4B0FB081A54F171EF7BE" blockId="22.[151,760,261,2081]" pageId="22" pageNumber="247">
<emphasis id="B954EAA4FFEC4B0FB081A54F148FFAFE" bold="true" box="[151,405,1354,1377]" pageId="22" pageNumber="247">Differential diagnosis.</emphasis>
Within the 
<emphasis id="B954EAA4FFEC4B0FB200A54F178DFAFE" box="[534,663,1354,1377]" italics="true" pageId="22" pageNumber="247">fuliginosus-</emphasis>
group 
<taxonomicName id="4C204D35FFEC4B0FB2CBA54F1408FA1E" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB2CBA54F1408FA1E" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
is very close to 
<taxonomicName id="4C204D35FFEC4B0FB1D0A56F1745FA1E" box="[454,607,1386,1409]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB1D0A56F1745FA1E" box="[454,607,1386,1409]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
. Both species are very frequently encountered, often co-occurring, nearly non-distinguishable from each other based on external morphological characters alone. Our study of hundreds of specimens across their entire distribution ranges revealed that 
<taxonomicName id="4C204D35FFEC4B0FB191A60F1733F9BE" authorityName="Bernhauer" authorityYear="1908" box="[391,553,1546,1569]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB191A60F1733F9BE" box="[391,553,1546,1569]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
always have pale antennomeres 1–3, whereas 
<taxonomicName id="4C204D35FFEC4B0FB1DAA62F177CF9DE" box="[460,614,1578,1601]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB1DAA62F177CF9DE" box="[460,614,1578,1601]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
have them at least slightly darkened. A negligible fraction (ca. 2 %) of 
<taxonomicName id="4C204D35FFEC4B0FB081A66F142BF91E" box="[151,305,1642,1665]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB081A66F142BF91E" box="[151,305,1642,1665]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
with pale basal antennomeres are mostly teneral. Antennae are generally more elongate in 
<taxonomicName id="4C204D35FFEC4B0FB28BA68F15C7F95E" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB28BA68F15C7F95E" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
, as opposed to being stouter in 
<taxonomicName id="4C204D35FFEC4B0FB23FA6AF17A5F95E" box="[553,703,1706,1729]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB23FA6AF17A5F95E" box="[553,703,1706,1729]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A33FFEC4B0FB2DAA6AF15BDF97E" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="22" pageNumber="247">Fig. 7</figureCitation>
). Also 
<taxonomicName id="4C204D35FFEC4B0FB0E2A6CF148BF97E" authorityName="Bernhauer" authorityYear="1908" box="[244,401,1738,1761]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB0E2A6CF148BF97E" box="[244,401,1738,1761]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
has an almost smoothly rounded head with the eyes protruding while 
<taxonomicName id="4C204D35FFEC4B0FB226A6EF17D1F89E" box="[560,715,1770,1793]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB226A6EF17D1F89E" box="[560,715,1770,1793]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
has eyes clearly protruding and thus making its head seem more quadrate (
<figureCitation id="131B2A33FFEC4B0FB12AA72F1467F8DE" box="[316,381,1834,1857]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="22" pageNumber="247">Fig. 6</figureCitation>
). Only some rare specimens, maybe hybrids, faded and very old museum specimens, or actual rare intraspecific variants, do not fit these trends and their identity can be ascertained only through examination of the aedeagi. In 
<emphasis id="B954EAA4FFEC4B0FB12FA7AF14C0F85E" box="[313,474,1962,1985]" italics="true" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB12FA7AF14CCF85E" authorityName="Bernhauer" authorityYear="1908" box="[313,470,1962,1985]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">Q. curtipennis</taxonomicName>
,
</emphasis>
contrary to 
<taxonomicName id="4C204D35FFEC4B0FB24CA7AF17EAF85E" box="[602,752,1962,1985]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB24CA7AF17EAF85E" box="[602,752,1962,1985]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
, the paramere lacks clear medial attenuation, the apical part of median lobe dorso-ventrally flattened, and internal sac with 2 pairs of clear sclerites.
</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB34CA0BB16BEFE2A" blockId="22.[828,1437,190,2069]" pageId="22" pageNumber="247">
The examined material for both species clearly shows that 
<taxonomicName id="4C204D35FFEC4B0FB37FA0DB16E5FF6A" authorityName="Bernhauer" authorityYear="1908" box="[873,1023,222,245]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB37FA0DB16E5FF6A" box="[873,1023,222,245]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
is much more common in the south and west and becomes increasingly rare towards the north and east of its range, while 
<taxonomicName id="4C204D35FFEC4B0FB42EA11B11D5FEAA" box="[1080,1231,286,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB42EA11B11D5FEAA" box="[1080,1231,286,309]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
shows an opposite trend. 
<taxonomicName id="4C204D35FFEC4B0FB396A13B1143FECA" authorityName="Bernhauer" authorityYear="1908" box="[896,1113,318,341]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB396A13B1143FECA" box="[896,1113,318,341]" italics="true" pageId="22" pageNumber="247">Quedius curtipennis</emphasis>
</taxonomicName>
seems to be more common in warmer and drier areas with more sandy soils, where 
<taxonomicName id="4C204D35FFEC4B0FB594A15B16ABFE0A" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB594A15B16ABFE0A" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
seems to prefer wetter, colder and more humus rich soils.
</paragraph>
<paragraph id="8B9F36B6FFEC4B0FB34CA1BB102BFDCA" blockId="22.[828,1437,190,2069]" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB34CA1BB112BFE4A" authorityName="Bernhauer" authorityYear="1908" box="[858,1073,446,469]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB34CA1BB112BFE4A" box="[858,1073,446,469]" italics="true" pageId="22" pageNumber="247">Quedius curtipennis</emphasis>
</taxonomicName>
is also very similar to 
<taxonomicName id="4C204D35FFEC4B0FB532A1BB1697FE6A" authorityName="Gusarov" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="afrofuliginosus">
<emphasis id="B954EAA4FFEC4B0FB532A1BB1697FE6A" italics="true" pageId="22" pageNumber="247">Q. afrofuliginosus</emphasis>
</taxonomicName>
, from which it is easily recognized by having no additional punctures between anterior and posterior frontal punctures. Besides, distributions of 
<taxonomicName id="4C204D35FFEC4B0FB4AEA21B104AFDAA" authorityName="Bernhauer" authorityYear="1908" box="[1208,1360,542,565]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB4AEA21B104AFDAA" box="[1208,1360,542,565]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
and 
<emphasis id="B954EAA4FFEC4B0FB594A21B16F9FDCA" italics="true" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB594A21B16C5FDCA" authorityName="Gusarov" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="afrofuliginosus">Q. afrofuliginosus</taxonomicName>
,
</emphasis>
as far as known, are allopatric.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0FB34CA25B11F2FD0A" pageId="22" pageNumber="247" type="reference_group">
<paragraph id="8B9F36B6FFEC4B0FB34CA25B11F2FD0A" blockId="22.[828,1437,190,2069]" pageId="22" pageNumber="247">
From 
<emphasis id="B954EAA4FFEC4B0FB38DA25B11D8FDEA" box="[923,1218,606,629]" italics="true" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB38DA25B1105FDEA" authorityName="Brulle" authorityYear="1832" box="[923,1055,606,629]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="levicollis">Q. levicollis</taxonomicName>
, 
<taxonomicName id="4C204D35FFEC4B0FB43CA25B11D8FDEA" ID-CoL="78PXW" authorityName="Bernhauer" authorityYear="1908" box="[1066,1218,606,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">Q. curtipennis</taxonomicName>
</emphasis>
is easily recognized by unpunctured and glabrous scutellum.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0FB32AA29B105CFBEA" pageId="22" pageNumber="247" type="type_taxon">
<paragraph id="8B9F36B6FFEC4B0FB32AA29B105CFBEA" blockId="22.[828,1437,190,2069]" pageId="22" pageNumber="247">
<emphasis id="B954EAA4FFEC4B0FB32AA29B1126FD2A" bold="true" box="[828,1084,670,693]" pageId="22" pageNumber="247">
<typeStatus id="549B8814FFEC4B0FB32AA29B16B7FD2A" box="[828,941,670,693]" pageId="22" pageNumber="247" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
BൾඋඇHൺඎൾඋ (1908) described 
<taxonomicName id="4C204D35FFEC4B0FB594A29B16AEFD4A" ID-CoL="78PXW" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB594A29B16AEFD4A" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
as a variety of 
<taxonomicName id="4C204D35FFEC4B0FB449A2BB11ECFD4A" ID-CoL="4R2YF" box="[1119,1270,702,725]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB449A2BB11ECFD4A" box="[1119,1270,702,725]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
without giving any exact data on the type material. He had stressed that it is very common at 
<collectingCountry id="F3377626FFEC4B0FB41EA2FB118DFC8A" box="[1032,1175,766,789]" name="Faroe Islands" pageId="22" pageNumber="247">Faroe Islands</collectingCountry>
and mentioned its occurrence in Central Europe as well as in ‘Romania’ and even ‘Buchara’. Here (see above), we have listed specimens from 
<collectingCountry id="F3377626FFEC4B0FB363A35B1119FCEA" box="[885,1027,862,885]" name="Faroe Islands" pageId="22" pageNumber="247">Faroe Islands</collectingCountry>
and ‘Buchara’ at the FMNH which qualify as 
<typeStatus id="549B8814FFEC4B0FB39BA37B16F0FC0A" box="[909,1002,894,917]" pageId="22" pageNumber="247" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C204D35FFEC4B0FB41AA37B11B9FC0A" ID-CoL="78PXW" authorityName="Bernhauer" authorityYear="1908" box="[1036,1187,894,917]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB41AA37B11B9FC0A" box="[1036,1187,894,917]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
and which we recorded long ago but were unable to re-examine in detail recently. There is no doubt about the identity of the 
<typeStatus id="549B8814FFEC4B0FB512A3BB107BFC4A" box="[1284,1377,958,981]" pageId="22" pageNumber="247" type="syntype">syntypes</typeStatus>
from 
<collectingCountry id="F3377626FFEC4B0FB32AA3DB16CAFC6A" box="[828,976,990,1013]" name="Faroe Islands" pageId="22" pageNumber="247">Faroe Islands</collectingCountry>
. However, a 
<typeStatus id="549B8814FFEC4B0FB476A3DB11A9FC6A" box="[1120,1203,990,1013]" pageId="22" pageNumber="247" type="syntype">syntype</typeStatus>
from ‘Buchara’ must be carefully checked. Given, additionally, some ambiguity about the geographic origin of that Middle Asian 
<typeStatus id="549B8814FFEC4B0FB55FA41B1086FBAA" box="[1353,1436,1054,1077]" pageId="22" pageNumber="247" type="syntype">syntype</typeStatus>
(see Distribution), a 
<typeStatus id="549B8814FFEC4B0FB40BA43B116AFBCA" box="[1053,1136,1086,1109]" pageId="22" pageNumber="247" type="syntype">syntype</typeStatus>
male from 
<collectingCountry id="F3377626FFEC4B0FB4E7A43B1098FBCA" box="[1265,1410,1086,1109]" name="Faroe Islands" pageId="22" pageNumber="247">Faroe Islands</collectingCountry>
is here designated as a 
<typeStatus id="549B8814FFEC4B0FB40EA45B1160FBEA" box="[1048,1146,1118,1141]" pageId="22" pageNumber="247" type="lectotype">lectotype</typeStatus>
for 
<taxonomicName id="4C204D35FFEC4B0FB4B1A45B1025FBEA" ID-CoL="78PXW" authorityName="Bernhauer" authorityYear="1908" box="[1191,1343,1118,1141]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB4B1A45B1025FBEA" box="[1191,1343,1118,1141]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0FB32AA47B110BFA6A" pageId="22" pageNumber="247" type="synonymic_list">
<paragraph id="8B9F36B6FFEC4B0FB32AA47B110BFA6A" blockId="22.[828,1437,190,2069]" pageId="22" pageNumber="247">
<emphasis id="B954EAA4FFEC4B0FB32AA47B1114FB0A" bold="true" box="[828,1038,1150,1173]" pageId="22" pageNumber="247">Synonymic notes.</emphasis>
<taxonomicName id="4C204D35FFEC4B0FB40FA47B11EBFB0A" authorityName="Hatch" authorityYear="1957" box="[1049,1265,1150,1173]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEC4B0FB40FA47B11EBFB0A" box="[1049,1265,1150,1173]" italics="true" pageId="22" pageNumber="247">Quedius parallelus</emphasis>
</taxonomicName>
was described by HൺඍർH (1957) in his work on beetles of the Pacific Northwest of North America without reference to European species. Based on the description of 
<emphasis id="B954EAA4FFEC4B0FB4AFA4DB1051FB6A" box="[1209,1355,1246,1269]" italics="true" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB4AFA4DB105DFB6A" authorityName="Hatch" authorityYear="1957" box="[1209,1351,1246,1269]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="parallelus">Q. parallelus</taxonomicName>
,
</emphasis>
KඈඋGൾ (1962b) suspected it to be identical to the European 
<taxonomicName id="4C204D35FFEC4B0FB594A4FB16A9FAAA" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB594A4FB16A9FAAA" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
. This was confirmed by Sආൾඍൺඇൺ (1971a) who studied the type material of 
<taxonomicName id="4C204D35FFEC4B0FB472A53B11EAFACA" authorityName="Hatch" authorityYear="1957" box="[1124,1264,1342,1365]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFEC4B0FB472A53B11EAFACA" box="[1124,1264,1342,1365]" italics="true" pageId="22" pageNumber="247">Q. parallelus</emphasis>
</taxonomicName>
and found them conspecific with 
<specimenCount id="9D26FD3FFFEC4B0FB3E5A55B1126FAEA" box="[1011,1084,1374,1397]" pageId="22" pageNumber="247" type="male">2 male</specimenCount>
<typeStatus id="549B8814FFEC4B0FB452A55B11BBFAEA" box="[1092,1185,1374,1397]" pageId="22" pageNumber="247" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C204D35FFEC4B0FB4D1A55B107BFAEA" authorityName="Bernhauer" authorityYear="1908" box="[1223,1377,1374,1397]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB4D1A55B107BFAEA" box="[1223,1377,1374,1397]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3377626FFEC4B0FB32AA57B16D6FA0A" box="[828,972,1406,1429]" name="Faroe Islands" pageId="22" pageNumber="247">Faroe Islands</collectingCountry>
at the FMNH. It is now firmly known that 
<taxonomicName id="4C204D35FFEC4B0FB32AA59B110FFA2A" authorityName="Bernhauer" authorityYear="1908" box="[828,1045,1438,1461]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB32AA59B110FFA2A" box="[828,1045,1438,1461]" italics="true" pageId="22" pageNumber="247">Quedius curtipennis</emphasis>
</taxonomicName>
occurs in the North American West (Sආൾඍൺඇൺ 1971a) and East Coasts (Sආൾඍൺඇൺ 1990, Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFEC4B0EB34CA5FB17B5F9AA" lastPageId="23" lastPageNumber="248" pageId="22" pageNumber="247" type="description">
<paragraph id="8B9F36B6FFEC4B0EB34CA5FB1744FD8A" blockId="22.[828,1437,190,2069]" lastBlockId="23.[151,760,190,2069]" lastPageId="23" lastPageNumber="248" pageId="22" pageNumber="247">
<taxonomicName id="4C204D35FFEC4B0FB34CA5FB11A6F98A" authority="Stephens, 1832" authorityName="Stephens" authorityYear="1832" box="[858,1212,1534,1557]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B954EAA4FFEC4B0FB34CA5FB1117F98A" box="[858,1037,1534,1557]" italics="true" pageId="22" pageNumber="247">Quedius gracilis</emphasis>
<bibRefCitation id="EFB14B47FFEC4B0FB401A5FB11A6F98A" author="STEPHENS J. F." box="[1047,1212,1534,1557]" pageId="22" pageNumber="247" refId="ref92742" refString="STEPHENS J. F. 1832: Illustrations of British entomology; or a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin &amp; Cradock, London, 240 pp." type="book" year="1832">Stephens, 1832</bibRefCitation>
</taxonomicName>
was described from London, 
<collectingCountry id="F3377626FFEC4B0FB38DA61B16DFF9AA" box="[923,965,1566,1589]" name="United Kingdom" pageId="22" pageNumber="247">UK</collectingCountry>
. Later, SඍൾඉHൾඇඌ (1837) had it as a variety of 
<taxonomicName id="4C204D35FFEC4B0FB32AA63B1683F9CA" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[828,921,1598,1621]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B954EAA4FFEC4B0FB32AA63B1683F9CA" box="[828,921,1598,1621]" italics="true" pageId="22" pageNumber="247">Q. tristis</emphasis>
</taxonomicName>
(now 
<taxonomicName id="4C204D35FFEC4B0FB3CBA63B1147F9CA" authorityName="Brulle" authorityYear="1832" box="[989,1117,1598,1621]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFEC4B0FB3CBA63B1147F9CA" box="[989,1117,1598,1621]" italics="true" pageId="22" pageNumber="247">Q. levicollis</emphasis>
</taxonomicName>
). In more recent works it was cited as a synonym of 
<taxonomicName id="4C204D35FFEC4B0FB40BA65B11B7F9EA" box="[1053,1197,1630,1653]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFEC4B0FB40BA65B11B7F9EA" box="[1053,1197,1630,1653]" italics="true" pageId="22" pageNumber="247">Q. fuliginosus</emphasis>
</taxonomicName>
. Based on examination of its 
<specimenCount id="9D26FD3FFFEC4B0FB363A67B16E4F90A" box="[885,1022,1662,1685]" pageId="22" pageNumber="247" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
by Roger Booth (BMNH, pers. comm.) on our request, it was established that these specimens are conspecific with 
<taxonomicName id="4C204D35FFEC4B0FB3F9A6BB119CF94A" authorityName="Bernhauer" authorityYear="1908" box="[1007,1158,1726,1749]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB3F9A6BB119CF94A" box="[1007,1158,1726,1749]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
. As the name is older than 
<taxonomicName id="4C204D35FFEC4B0FB32AA6DB16CFF96A" authorityName="Bernhauer" authorityYear="1908" box="[828,981,1758,1781]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB32AA6DB16CFF96A" box="[828,981,1758,1781]" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
we here establish the prevailing use of 
<taxonomicName id="4C204D35FFEC4B0FB594A6DB16A9F88A" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB594A6DB16A9F88A" italics="true" pageId="22" pageNumber="247">Q. curtipennis</emphasis>
</taxonomicName>
as opposed to the rule of priority (ICZN Article 23.9). 
<taxonomicName id="4C204D35FFEC4B0FB396A71B114CF8AA" authorityName="Bernhauer" authorityYear="1908" box="[896,1110,1822,1845]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFEC4B0FB396A71B114CF8AA" box="[896,1110,1822,1845]" italics="true" pageId="22" pageNumber="247">Quedius curtipennis</emphasis>
</taxonomicName>
has been used at least 26 times in the last 50 years by more than ten authors (Bඈඋൽඈඇං 1974b, 1976b; Pඈඉൾ 1977; Cඈංൿൿൺංඍ 1978; Oඎඍൾඋൾඅඈ 1978; Bඎඋൺĸඈඐඌĸං et al. 1980; TඬඍH 1984; DඋඎGආൺඇൽ 1987; Nඈඐඈඌൺൽ 1990; BඈඋGൾඌ 1990; Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං 1995; Oඐൾඇ 1997; Oඎඍൾඋൾඅඈ et al. 1998; AඌඌංඇG 2001; UHඅංG et al. 2006; Tඋඈඇඊඎൾඍ 2006; Mൺඃĸൺ &amp; Sආൾඍൺඇൺ 2007; ÖඓGൾඇ 2011; Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011; Wൾൻඌඍൾඋ et al. 2012; MൺGඎඋൺ et al. 2013; TඬඍHආඣඋඣඌඓ et al. 2014; Sൾආൾඇඈඏ et al. 2015; Sൾආංඈඇൾඇĸඈඏ et al. 2015; Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ 2018a, 2019). On the other hand, 
<taxonomicName id="4C204D35FFED4B0EB2C8A0FB15F2FEAA" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B954EAA4FFED4B0EB2C8A0FB15F2FEAA" italics="true" pageId="23" pageNumber="248">Q. gracilis</emphasis>
</taxonomicName>
has only been used once after its description as a valid species which was more than 150 years ago (Cඎඋඍංඌ 1837). This deems the name 
<taxonomicName id="4C204D35FFED4B0EB1C3A15B15FAFE0A" authority="Stephens, 1832 a" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B954EAA4FFED4B0EB1C3A15B1792FEEA" box="[469,648,350,373]" italics="true" pageId="23" pageNumber="248">Quedius gracilis</emphasis>
<bibRefCitation id="EFB14B47FFED4B0EB287A15B15D6FE0A" author="STEPHENS J. F." pageId="23" pageNumber="248" refId="ref92742" refString="STEPHENS J. F. 1832: Illustrations of British entomology; or a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin &amp; Cradock, London, 240 pp." type="book" year="1832">Stephens, 1832</bibRefCitation>
a
</taxonomicName>
nomen oblitum in relation to 
<taxonomicName id="4C204D35FFED4B0EB236A17B1450FE2A" authority="Bernhauer 1908" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB236A17B17E2FE0A" box="[544,760,382,405]" italics="true" pageId="23" pageNumber="248">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFED4B0EB081A19B1450FE2A" author="BERNHAUER M." box="[151,330,414,437]" pageId="23" pageNumber="248" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer 1908</bibRefCitation>
</taxonomicName>
(nomen protectum). 
<taxonomicName id="4C204D35FFED4B0EB228A19B1426FE4A" authority="Stephens, 1832" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B954EAA4FFED4B0EB228A19B17E2FE2A" box="[574,760,414,437]" italics="true" pageId="23" pageNumber="248">Quedius gracilis</emphasis>
<bibRefCitation id="EFB14B47FFED4B0EB081A1BB1426FE4A" author="STEPHENS J. F." box="[151,316,446,469]" pageId="23" pageNumber="248" refId="ref92742" refString="STEPHENS J. F. 1832: Illustrations of British entomology; or a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. 5. Baldwin &amp; Cradock, London, 240 pp." type="book" year="1832">Stephens, 1832</bibRefCitation>
</taxonomicName>
syn. rev. is thus moved from synonymy with 
<taxonomicName id="4C204D35FFED4B0EB0DCA1DB1777FE6A" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[202,621,478,501]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFED4B0EB0DCA1DB1482FE6A" box="[202,408,478,501]" italics="true" pageId="23" pageNumber="248">Quedius fuliginosus</emphasis>
(
<bibRefCitation id="EFB14B47FFED4B0EB1B5A1DB177CFE6A" author="GRAVENHORST J. L. C." box="[419,614,478,501]" pageId="23" pageNumber="248" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
to synonymy with 
<taxonomicName id="4C204D35FFED4B0EB0DBA1FB1740FD8A" authority="Bernhauer 1908" authorityName="Bernhauer" authorityYear="1908" box="[205,602,510,533]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB0DBA1FB14BFFD8A" box="[205,421,510,533]" italics="true" pageId="23" pageNumber="248">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFED4B0EB1BDA1FB1740FD8A" author="BERNHAUER M." box="[427,602,510,533]" pageId="23" pageNumber="248" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer 1908</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F36B6FFED4B0EB081A21B178AFA8A" blockId="23.[151,760,190,2069]" pageId="23" pageNumber="248">
<emphasis id="B954EAA4FFED4B0EB081A21B1406FDAA" bold="true" box="[151,284,542,565]" pageId="23" pageNumber="248">Bionomics.</emphasis>
Based on published data as well as our observations, 
<taxonomicName id="4C204D35FFED4B0EB125A23B14D2FDCA" baseAuthorityName="Bernhauer" baseAuthorityYear="1908" box="[307,456,574,597]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipenni">
<emphasis id="B954EAA4FFED4B0EB125A23B14D2FDCA" box="[307,456,574,597]" italics="true" pageId="23" pageNumber="248">Q. curtipenni</emphasis>
</taxonomicName>
s is a polytopic, somewhat thermophilous species found in a large range of habitats and various ground-based microhabitats with a tendency to occur in open woodlands. For example, in Central and Eastern Europe it was found under stones in dry pastures in 
<collectingCountry id="F3377626FFED4B0EB0A2A2DB15E4FD6A" box="[180,254,734,757]" name="Poland" pageId="23" pageNumber="248">Poland</collectingCountry>
(Dඏඈෞගĸ 1965), in xerothermous grassland in 
<collectingCountry id="F3377626FFED4B0EB081A2FB15E0FC8A" box="[151,250,766,789]" name="Germany" pageId="23" pageNumber="248">Germany</collectingCountry>
(AඌඌංඇG 2001), and in closed oak forest with extensive ground vegetation and shrub cover in 
<collectingCountry id="F3377626FFED4B0EB28CA31B17EDFCAA" box="[666,759,798,821]" name="Hungary" pageId="23" pageNumber="248">Hungary</collectingCountry>
(TඬඍHආඣඋඣඌඓ et. al 2014). We have collected this species in a mixed forest on the sandy dry island of Laesø in 
<collectingCountry id="F3377626FFED4B0EB2D4A35B15CBFC0A" name="Denmark" pageId="23" pageNumber="248">Denmark</collectingCountry>
. There are also records from mole (
<taxonomicName id="4C204D35FFED4B0EB25FA37B17F5FC0A" authorityName="Linnaeus" authorityYear="1758" box="[585,751,894,917]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="23" pageNumber="248" phylum="Chordata" rank="species" species="europaea">
<emphasis id="B954EAA4FFED4B0EB25FA37B17F5FC0A" box="[585,751,894,917]" italics="true" pageId="23" pageNumber="248">Talpa europaea</emphasis>
</taxonomicName>
) nests built from grasses and reeds on wet meadow in 
<collectingCountry id="F3377626FFED4B0EB081A3BB15FBFC4A" box="[151,225,958,981]" name="Poland" pageId="23" pageNumber="248">Poland</collectingCountry>
(Nඈඐඈඌൺൽ 1990). In south-western Europe it was recorded from under rock in pine forest at an altitude of 
<quantity id="4CD89B53FFED4B0EB081A3FB15FCFB8A" box="[151,230,1022,1045]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="23" pageNumber="248" unit="m" value="1550.0">1550 m</quantity>
in Central Spain (Oඎඍൾඋൾඅඈ 1978) and from a cave in Northern Spain (Oඎඍൾඋൾඅඈ et al. 1998). On the Azores (Sආൾඍൺඇൺ 1970) it was found under rocks at the shore of a freshwater lake at an elevation of 
<quantity id="4CD89B53FFED4B0EB1E2A45B1774FBEA" box="[500,622,1118,1141]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="248" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
400– 
<specimenCount id="9D26FD3FFFED4B0EB23FA45B1774FBEA" box="[553,622,1118,1141]" pageId="23" pageNumber="248" type="generic">500 m</specimenCount>
</quantity>
. Presumably, the nest or cave-based subterranean records are accidental occurrences, as there is no evidence of the species preferring these habitats. The species is found within the wide range of elevations from sea level in the northern part of its range up to about 
<quantity id="4CD89B53FFED4B0EB16EA4FB14D5FA8A" box="[376,463,1278,1301]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="248" unit="m" value="2.0">2.000 m</quantity>
the southern part.
</paragraph>
<paragraph id="8B9F36B6FFED4B0EB0A3A51B176DFA4A" blockId="23.[151,760,190,2069]" pageId="23" pageNumber="248">
As an introduced species in North America, it was found especially around human settlements in various debris, under rocks, and in greenhouses. Some specimens were also collected by sifting barnyard litter on a pasture in 
<collectingRegion id="49E4F854FFED4B0EB0A5A59B1419FA2A" box="[179,259,1438,1461]" country="United States of America" name="Oregon" pageId="23" pageNumber="248">Oregon</collectingRegion>
(Sආൾඍൺඇൺ 1978a) and in natural habitats away from settlements (in moss, in leaf litter, etc.).
</paragraph>
<paragraph id="8B9F36B6FFED4B0EB0A3A5DB17B5F9AA" blockId="23.[151,760,190,2069]" pageId="23" pageNumber="248">
Pupae of the species are described in Oඎඍൾඋൾඅඈ (1978). Adult females from Wytham, 
<collectingCountry id="F3377626FFED4B0EB1DCA5FB14EAF98A" box="[458,496,1534,1557]" name="United Kingdom" pageId="23" pageNumber="248">UK</collectingCountry>
were observed to contain eggs between October and January (Fඋൺඇĸ 1969).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFED4B0EB081A63B16FBFC6A" pageId="23" pageNumber="248" type="distribution">
<paragraph id="8B9F36B6FFED4B0EB081A63B1702F78A" blockId="23.[151,760,190,2069]" pageId="23" pageNumber="248">
<emphasis id="B954EAA4FFED4B0EB081A63B1430F9CA" bold="true" box="[151,298,1598,1621]" pageId="23" pageNumber="248">Distribution.</emphasis>
<taxonomicName id="4C204D35FFED4B0EB139A63B171CF9CA" authorityName="Bernhauer" authorityYear="1908" box="[303,518,1598,1621]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB139A63B171CF9CA" box="[303,518,1598,1621]" italics="true" pageId="23" pageNumber="248">Quedius curtipennis</emphasis>
</taxonomicName>
is distributed from the Atlantic coast of Europe to 
<collectingCountry id="F3377626FFED4B0EB1D3A65B1759F9EA" box="[453,579,1630,1653]" name="Turkey" pageId="23" pageNumber="248">Asia Minor</collectingCountry>
, east to the Ural Mountains and Middle Asia (
<figureCitation id="131B2A33FFED4B0EB1DDA67B1703F90A" box="[459,537,1662,1685]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="23" pageNumber="248">Fig. 18</figureCitation>
). It is also present on the islands of Madeira, the Azores, and the 
<collectingCountry id="F3377626FFED4B0EB275A69B17EFF92A" box="[611,757,1694,1717]" name="Faroe Islands" pageId="23" pageNumber="248">Faroe Islands</collectingCountry>
. One specimen, which we studied was found in the central Morrocan mountains, and one old specimen from the 
<typeStatus id="549B8814FFED4B0EB2DCA6DB17E2F96A" box="[714,760,1758,1781]" pageId="23" pageNumber="248">type</typeStatus>
series is from ‘Buchara’, 
<collectingCountry id="F3377626FFED4B0EB1BCA6FB173CF88A" box="[426,550,1790,1813]" name="Uzbekistan" pageId="23" pageNumber="248">Uzbekistan</collectingCountry>
. These are the only specimens we know from North Africa and Middle Asia, respectively. The species’ occurrence in these areas needs confirmation to rule out possible mislabelling. Most previous records of 
<taxonomicName id="4C204D35FFED4B0EB15CA77B14F8F80A" authorityName="Bernhauer" authorityYear="1908" box="[330,482,1918,1941]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB15CA77B14F8F80A" box="[330,482,1918,1941]" italics="true" pageId="23" pageNumber="248">Q. curtipennis</emphasis>
</taxonomicName>
(and 
<taxonomicName id="4C204D35FFED4B0EB209A77B17AEF80A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[543,692,1918,1941]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFED4B0EB209A77B17AEF80A" box="[543,692,1918,1941]" italics="true" pageId="23" pageNumber="248">Q. fuliginosus</emphasis>
</taxonomicName>
) from North Africa could be assigned to 
<taxonomicName id="4C204D35FFED4B0EB1E4A79B17B6F82A" authorityName="Gusarov" authorityYear="1991" box="[498,684,1950,1973]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="afrofuliginosus">
<emphasis id="B954EAA4FFED4B0EB1E4A79B17B6F82A" box="[498,684,1950,1973]" italics="true" pageId="23" pageNumber="248">Q. afrofuliginosus</emphasis>
</taxonomicName>
, which is externally very similar. As 
<taxonomicName id="4C204D35FFED4B0EB1DEA7BB1745F84A" authorityName="Bernhauer" authorityYear="1908" box="[456,607,1982,2005]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB1DEA7BB1745F84A" box="[456,607,1982,2005]" italics="true" pageId="23" pageNumber="248">Q. curtipennis</emphasis>
</taxonomicName>
does not seem to occur in southern 
<collectingCountry id="F3377626FFED4B0EB163A7DB14AEF86A" box="[373,436,2014,2037]" name="Spain" pageId="23" pageNumber="248">Spain</collectingCountry>
, 
<collectingCountry id="F3377626FFED4B0EB1A9A7DB14E7F86A" box="[447,509,2014,2037]" name="France" pageId="23" pageNumber="248">Corse</collectingCountry>
and Sardinia, it is very likely absent from North Africa too.
</paragraph>
<paragraph id="8B9F36B6FFED4B0EB34CA0BB16FBFC6A" blockId="23.[828,1437,190,1013]" pageId="23" pageNumber="248">
<taxonomicName id="4C204D35FFED4B0EB34CA0BB1128FF4A" authorityName="Bernhauer" authorityYear="1908" box="[858,1074,190,213]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB34CA0BB1128FF4A" box="[858,1074,190,213]" italics="true" pageId="23" pageNumber="248">Quedius curtipennis</emphasis>
</taxonomicName>
is also known as introduced to at least four regions of North America (
<figureCitation id="131B2A33FFED4B0EB4A9A0DB1014FF6A" box="[1215,1294,222,245]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="23" pageNumber="248">Fig. 12</figureCitation>
). The presence of 
<taxonomicName id="4C204D35FFED4B0EB36CA0FB110EFE8A" authorityName="Bernhauer" authorityYear="1908" box="[890,1044,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB36CA0FB110EFE8A" box="[890,1044,254,277]" italics="true" pageId="23" pageNumber="248">Q. curtipennis</emphasis>
</taxonomicName>
in the Nearctic was first recognized by Sආൾඍൺඇൺ (1971a) when he reviewed specimens of 
<taxonomicName id="4C204D35FFED4B0EB594A11B16B0FECA" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFED4B0EB594A11B16B0FECA" italics="true" pageId="23" pageNumber="248">Q. parallelus</emphasis>
</taxonomicName>
, a newly described species from the Nearctic. 
<taxonomicName id="4C204D35FFED4B0EB370A15B112EFEEA" authorityName="Hatch" authorityYear="1957" box="[870,1076,350,373]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFED4B0EB370A15B112EFEEA" box="[870,1076,350,373]" italics="true" pageId="23" pageNumber="248">Quedius parallelus</emphasis>
</taxonomicName>
was found to be identical to the Palearctic 
<taxonomicName id="4C204D35FFED4B0EB3A6A17B1154FE0A" authorityName="Bernhauer" authorityYear="1908" box="[944,1102,382,405]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB3A6A17B1154FE0A" box="[944,1102,382,405]" italics="true" pageId="23" pageNumber="248">Q. curtipennis</emphasis>
</taxonomicName>
, something already hinted by KඈඋGൾ (1962b). One specimen from the 
<typeStatus id="549B8814FFED4B0EB4F8A19B1006FE2A" box="[1262,1308,414,437]" pageId="23" pageNumber="248">type</typeStatus>
series of 
<taxonomicName id="4C204D35FFED4B0EB594A19B16B2FE4A" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B954EAA4FFED4B0EB594A19B16B2FE4A" italics="true" pageId="23" pageNumber="248">Q. parallelus</emphasis>
</taxonomicName>
was collected in 1934 near the city of Seattle in 
<collectingRegion id="49E4F854FFED4B0EB32AA1DB16A7FE6A" box="[828,957,478,501]" country="United States of America" name="Washington" pageId="23" pageNumber="248">Washington</collectingRegion>
, which is currently the oldest known record of the species from North America. In 1971, the species was already found in a relatively large area along the Pacific coast of 
<collectingRegion id="49E4F854FFED4B0EB385A23B16FCFDCA" box="[915,998,574,597]" country="United States of America" name="Oregon" pageId="23" pageNumber="248">Oregon</collectingRegion>
, 
<collectingRegion id="49E4F854FFED4B0EB3F8A23B1175FDCA" box="[1006,1135,574,597]" country="United States of America" name="Washington" pageId="23" pageNumber="248">Washington</collectingRegion>
, and British Columbia (Sආൾ-ඍൺඇൺ 1971a), suggesting that it may have been established in North America even prior to 1934. Currently, 
<taxonomicName id="4C204D35FFED4B0EB556A27B1698FD2A" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFED4B0EB556A27B1698FD2A" italics="true" pageId="23" pageNumber="248">Q. curtipennis</emphasis>
</taxonomicName>
in western North America is still known only from these states and provinces, where it is found as far East as Cranbrook, British Columbia in the Canadian Rockies (from web www.boldsystems.org) (teal in 
<figureCitation id="131B2A33FFED4B0EB4E1A2FB105FFC8A" box="[1271,1349,766,789]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="23" pageNumber="248">Fig. 12</figureCitation>
). One to three additional introductions of 
<taxonomicName id="4C204D35FFED4B0EB480A31B1039FCAA" baseAuthorityName="Bernhauer" baseAuthorityYear="1908" box="[1174,1315,798,821]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="248" phylum="Arthropoda" rank="species" species="curtipenni">
<emphasis id="B954EAA4FFED4B0EB480A31B1039FCAA" box="[1174,1315,798,821]" italics="true" pageId="23" pageNumber="248">Q. curtipenni</emphasis>
</taxonomicName>
s were later reported from the East Coast, in 
<collectingRegion id="49E4F854FFED4B0EB49BA33B1026FCCA" box="[1165,1340,830,853]" country="United States of America" name="New Hampshire" pageId="23" pageNumber="248">New Hampshire</collectingRegion>
, 
<collectingCountry id="F3377626FFED4B0EB553A33B1067FCCA" box="[1349,1405,830,853]" name="United States of America" pageId="23" pageNumber="248">USA</collectingCountry>
, as well as 
<collectingRegion id="49E4F854FFED4B0EB39FA35B16C0FCEA" box="[905,986,862,885]" country="Canada" name="Ontario" pageId="23" pageNumber="248">Ontario</collectingRegion>
and 
<collectingRegion id="49E4F854FFED4B0EB41CA35B1194FCEA" box="[1034,1166,862,885]" country="Canada" name="Nova Scotia" pageId="23" pageNumber="248">Nova Scotia</collectingRegion>
, 
<collectingCountry id="F3377626FFED4B0EB480A35B11F3FCEA" box="[1174,1257,862,885]" name="Canada" pageId="23" pageNumber="248">Canada</collectingCountry>
. Specimens from near Guelph in 
<collectingRegion id="49E4F854FFED4B0EB3CAA37B1137FC0A" box="[988,1069,894,917]" country="Canada" name="Ontario" pageId="23" pageNumber="248">Ontario</collectingRegion>
were collected in 1976 (Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011), from 
<collectingRegion id="49E4F854FFED4B0EB424A39B11C6FC2A" box="[1074,1244,926,949]" inferredBySuffix="true" pageId="23" pageNumber="248">New Hampshire</collectingRegion>
in 1983 (Sආൾඍൺඇൺ 1990) and from 
<collectingRegion id="49E4F854FFED4B0EB3F9A3BB116FFC4A" box="[1007,1141,958,981]" country="Canada" name="Nova Scotia" pageId="23" pageNumber="248">Nova Scotia</collectingRegion>
in 1997 (Mൺඃĸൺ &amp; Kඅංආൺ-ඌඓൾඐඌĸං 2008).
</paragraph>
</subSubSection>
</treatment>
</document>