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<figureCitation id="131B2A33FFE14B02B3FFA1B91105FE50" box="[1001,1055,444,463]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="27" pageNumber="252">Figs 1</figureCitation>
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<figureCitation id="131B2A33FFE14B02B43FA1B9115EFE50" box="[1065,1092,444,463]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="27" pageNumber="252">2B</figureCitation>
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<figureCitation id="131B2A33FFE14B02B474A1B91164FE50" box="[1122,1150,444,463]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="27" pageNumber="252">6D</figureCitation>
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locality: Morée: plaine de Modon]
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locality: Gallia; Symperopoli]
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<emphasis id="B954EAA4FFE14B02B32AA24C16FAFDC3" box="[828,992,585,604]" italics="true" pageId="27" pageNumber="252">Quedius balthasari</emphasis>
<bibRefCitation id="EFB14B47FFE14B02B3F3A24C1172FDC3" author="MARAN J." box="[997,1128,585,604]" pageId="27" pageNumber="252" pagination="8 - 9" refId="ref87754" refString="MARAN J. 1935: Nove a zajimave formy rodu Quedius Steph. z ostrova Krety. De novis et raris formis generis Quedius Steph. ex insula Creta. Casopis Ceskoslovenske Spolecnosti Entomologicke 32: 8 - 9 (in Czech)." type="journal article" year="1935">Mařan, 1935: 9</bibRefCitation>
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<treatmentCitation id="0A8110A7FFE14B02B32AA26111A3FDE8" author="MENETRIES E." box="[828,1209,612,631]" page="144" pageId="27" pageNumber="252" year="1832">
<taxonomicName id="4C204D35FFE14B02B32AA26111A3FDE8" ID-CoL="4ZK82" authority="Menetries, 1832: 144" authorityName="Menetries" authorityPageNumber="144" authorityYear="1832" box="[828,1209,612,631]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B954EAA4FFE14B02B32AA26116E4FDE8" box="[828,1022,612,631]" italics="true" pageId="27" pageNumber="252">Staphylinus nigritarsis</emphasis>
<bibRefCitation id="EFB14B47FFE14B02B415A26111A3FDE8" author="MENETRIES E." box="[1027,1209,612,631]" pageId="27" pageNumber="252" refId="ref87999" refString="MENETRIES E. 1832: Catalogue raisonne des objets de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse entrepris par ordre de S. M. L'Empereur. L'Academie Imperiale des Sciences, St. Petersbourg, 271 + xxxii + iv pp." type="book" year="1832">Ménétriés, 1832: 144</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="549B8814FFE14B02B4D5A26111EBFDE8" box="[1219,1265,612,631]" pageId="27" pageNumber="252">Type</typeStatus>
locality: Caucase]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9F36B6FFE14B02B32AA28811BCF979" blockId="27.[828,1438,653,1766]" pageId="27" pageNumber="252">
<emphasis id="B954EAA4FFE14B02B32AA28816B8FD3F" bold="true" box="[828,930,653,672]" pageId="27" pageNumber="252">References.</emphasis>
GඋൺඏൾඇHඈඋඌඍ (1802):34,(1847):231 (characters); SඍൾඉHൾඇඌ (1832): 215, (1839): 389 (characters); Fൺඅൽൾඋආൺඇඇ (1835): 123 (characters); Lൺർඈඋൽൺංඋൾ (1835): 376 (characters); Nඈඋൽආൺඇඇ (1837): 75 (distribution); Hඈඅආൾ (1837): 63 (distribution); Wൾඌඍඐඈඈൽ (1838): 167 (larva and pupa); EඋංർHඌඈඇ (1840): 537 (characters); Kඈඅൾඇൺඍං (1846): 22 (characters); Kංൾඌൾඇඐൾඍඍൾඋ (1848): 54 (characters); HඈർHHඎඍH (1849): 154, (1862): 46 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 709, (1857): 201, (1874): 198 (characters); Fൺංඋආൺංඋൾ &amp; Lൺൻඈඎඅൻජඇൾ (1856): 536 (characters); RඈඌൾඇHൺඎൾඋ (1856): 71 (distribution); Kඋൺൺඍඓ (1857): 502 (characters); Fൺංඋආൺංඋൾ &amp; Cඈඊඎൾඋൾඅ (1860): 154 (distribution); Fൺඎඏൾඅ (1865): 293 (ecology); (1874): 289 (characters and distribution); (1878): 128, (1886): 58, (1902): 115 (distribution); Mඎඅඌൺඇඍ &amp; Rൾඒ (1876): 688 (characters); Fඈඐඅൾඋ (1888): 237 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Sൺංඇඍൾ- Cඅൺංඋൾ Dൾඏංඅඅൾ (1906): 102 (distribution); Pඈඋඍൺ (1907): 129 (characters); Rൾංඍඍൾඋ (1909): 111 (characters); JඈHൺඇඌൾඇ (1914): 363 (characters); Mඎඇඌඍൾඋ (1923): 196 (distribution); Gඋංൽൾඅඅං (1924):80 (characters and distribution); Pඈඋඍൾඏංඇ (1929):341 (characters); WඳඌඍHඈൿൿ (1938): fig. 22 (characters); TඈඍඍൾඇHൺආ (1949): 376 (characters); Hൺඋൽඒ (1851): 32 (distribution); Hൺඇඌൾඇ (1952): 142 (characters); SർHൾൾඋඉൾඅඍඓ (1956): 1102, (1965a): 407, (1965b): 511 (distribution); (1961): 133 (characters); Sආൾඍൺඇൺ (1958): 362 (characters and ecology); (1962): 134 (characters and distribution); (1963): 39, (1967): 558, (1970): 62, (1993):50 (distribution);(1978b):121 (distribution and ecology); Pൺඅආ (1963): 141 (characters and distribution); LඈHඌൾ (1964): 211 (characters); Fඈർൺඋංඅൾ (1964): 61 (distribution); Hඈඋංඈඇ (1965): 273 (distribution); Cඈංൿൿൺංඍ (1970):62 (distribution); (1978):192 (characters and distribution); Bඈඋൽඈඇං (1976a):93 (characters);(1982):140 (ecology); (1984):342 (distribution); Pඈඉൾ (1977): 31 (characters and distribution); Hංඇඍඈඇ (1981): 662 (characters of egg); TඬඍH (1984): 118 (distribution); Fඋൺඇඓ (1987): 70 (distribution); Gඎඌൺඋඈඏ (1993): 73 (synonymy); WൾඅർH (1993): 229 (ovary structure); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 33 (distribution); Nൾඐඍඈඇ (1995): 51 (requested to ICZN to place 
<taxonomicName id="4C204D35FFE14B02B494A5CD11FCFA44" authorityName="Brulle" authorityYear="1832" box="[1154,1254,1480,1499]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE14B02B494A5CD11FCFA44" box="[1154,1254,1480,1499]" italics="true" pageId="27" pageNumber="252">Q. levicollis</emphasis>
</taxonomicName>
on the Official List of Specific Names in Zoology); 
<bibRefCitation id="EFB14B47FFE14B02B420A5E611B2FA69" author="ICZN" box="[1078,1192,1507,1526]" pageId="27" pageNumber="252" pagination="215 - 217" refId="ref84977" refString="ICZN 1996: Opinion 1851. Xantholinini Erichson, 1839 and Quediini Kraatz, [1857] (Insecta, Coleoptera): given precedence over some senior synonyms; Quedius Stephens, 1829: Staphylinus levicollis Brulle, 1832 designated as the type species. Bulletin of Zoological Nomenclature 53: 215 - 217." type="journal article" year="1996">ICZN (1996)</bibRefCitation>
: 215 (
<taxonomicName id="4C204D35FFE14B02B4C8A5E6105EFA69" authorityName="Brulle" authorityYear="1832" box="[1246,1348,1507,1526]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE14B02B4C8A5E6105EFA69" box="[1246,1348,1507,1526]" italics="true" pageId="27" pageNumber="252">Q. levicollis</emphasis>
</taxonomicName>
placed on Official List of Specific Names in Zoology); Bඅൺඇൽ &amp; Sංඇർඅൺංඋ (1997): 204 (biology); MൺඋർHൺඅ &amp; 
<collectingCountry id="F3377626FFE14B02B408A61D112FF9B4" box="[1054,1077,1560,1579]" name="Switzerland" pageId="27" pageNumber="252">CH</collectingCountry>
ൺඋൽඈඇඇൾඍ (2000): 72 (distribution); Oඐൾඇ (2000): 252 (ecology); AඌඌංඇG &amp; Wඎඇൽൾඋඅൾ (2001): 37 (distribution); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Aඇඅൺ෡ (2009): 229 (distribution); Sඍൺඇ (2009): 242 (distribution); Vඈඋඌඍ et al. (2009): 16 (distribution); Mൺඃඓඅൺඇ &amp; BඈHගඹ (2012): 182 (distribution); Bൾൻൾඋආൺඇඌ et al. (2016): 135 (distribution); ÖඓGൾඇ et al. (2016): 620 (distribution); Rඎඍൺ et al. (2016): 37 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2019): 49 (distribution).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE14B05B32AA6F811A1FB94" lastPageId="28" lastPageNumber="253" pageId="27" pageNumber="252" type="materials_examined">
<paragraph id="8B9F36B6FFE14B02B32AA6F816F1F8FF" blockId="27.[828,1437,1789,2075]" pageId="27" pageNumber="252">
<emphasis id="B954EAA4FFE14B02B32AA6F81101F88F" bold="true" box="[828,1051,1789,1808]" pageId="27" pageNumber="252">
<typeStatus id="549B8814FFE14B02B32AA6F81673F88F" box="[828,873,1789,1808]" pageId="27" pageNumber="252">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="4C204D35FFE14B02B409A6F811FAF88F" authorityName="Menetries" authorityYear="1832" box="[1055,1248,1789,1808]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B954EAA4FFE14B02B409A6F811FAF88F" box="[1055,1248,1789,1808]" italics="true" pageId="27" pageNumber="252">Staphylinus nigritarsis</emphasis>
</taxonomicName>
: Lൾർඍඈඍඒඉൾ: designated by Gඎඌൺඋඈඏ (1993) (
<collectionCode id="ED31AE73FFE14B02B40CA712115BF8B5" box="[1050,1089,1815,1834]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="252" type="Museum">ZIN</collectionCode>
), labelled: “[golden square] / Caucas. / 
<taxonomicName id="4C204D35FFE14B02B32AA7371110F8DA" authority="Menet. Cauc" authorityName="Menet. Cauc" box="[828,1034,1842,1861]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="252" phylum="Arthropoda" rank="species" species="nigritarsis">nigritarsis Menet. Cauc</taxonomicName>
”. Pൺඋൺඅൾർඍඈඍඒඉൾ: labelled: “[golden square] / Caucas.” (1 J 
<collectionCode id="ED31AE73FFE14B02B3ACA74816FBF8FF" box="[954,993,1869,1888]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="252" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B9F36B6FFE14B05B32AA76211A1FB94" blockId="27.[828,1437,1789,2075]" lastBlockId="28.[826,1438,184,1035]" lastPageId="28" lastPageNumber="253" pageId="27" pageNumber="252">
<emphasis id="B954EAA4FFE14B02B32AA76211A7F8E5" bold="true" box="[828,1213,1895,1914]" pageId="27" pageNumber="252">
Additional material examined. 
<collectingCountry id="F3377626FFE14B02B443A76211A3F8E5" box="[1109,1209,1895,1914]" name="Albania" pageId="27" pageNumber="252">ALBANIA</collectingCountry>
:
</emphasis>
<quantity id="4CD89B53FFE14B02B4D2A76211E9F8E5" box="[1220,1267,1895,1914]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="252" unit="km" value="13.0">13km</quantity>
SE 
<collectingRegion id="49E4F854FFE14B02B50CA7621044F8E5" box="[1306,1374,1895,1914]" country="Albania" name="Elbasan" pageId="27" pageNumber="252">Elbasan</collectingRegion>
, Mali i Shpatit, Mt. Bukanikut, 
<geoCoordinate id="EE145071FFE14B02B41BA7871149F80A" box="[1037,1107,1922,1941]" degrees="41.0431" direction="north" orientation="latitude" pageId="27" pageNumber="252" precision="5" value="41.0431">41.0431</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE14B02B448A78711BEF80A" box="[1118,1188,1922,1941]" degrees="20.2172" direction="east" orientation="longitude" pageId="27" pageNumber="252" precision="5" value="20.2172">20.2172</geoCoordinate>
, brook bank in beech forest, 
<quantity id="4CD89B53FFE14B02B32AA7981662F82F" box="[828,888,1949,1968]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="252" unit="m" value="1500.0">1500m</quantity>
, 
<date id="FF9E1076FFE14B02B369A79816CDF82F" box="[895,983,1949,1968]" pageId="27" pageNumber="252" value="2010-05-22">22.V.2010</date>
, leg. D.W.Wrase (1J cSch). 
<emphasis id="B954EAA4FFE14B02B4DEA7981683F855" bold="true" pageId="27" pageNumber="252">BOSNIAAND HERZE-GOVINA:</emphasis>
Maklen-Pass,[43.85,17.65], 1902, leg.O.Leonhard (1 
<collectionCode id="ED31AE73FFE14B02B577A7B21089F855" box="[1377,1427,1975,1994]" pageId="27" pageNumber="252">SDEI</collectionCode>
); Prozor, [43.82, 17.61], 1902, leg. O. Leonhard (1 
<collectionCode id="ED31AE73FFE14B02B4F5A7D7100FF87A" box="[1251,1301,2002,2021]" pageId="27" pageNumber="252">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFE14B02B532A7D71086F87A" bold="true" box="[1316,1436,2002,2021]" pageId="27" pageNumber="252">
<collectingCountry id="F3377626FFE14B02B532A7D71083F87A" box="[1316,1433,2002,2021]" name="Bulgaria" pageId="27" pageNumber="252">BULGARIA</collectingCountry>
:
</emphasis>
Arkutino, S of 
<collectingRegion id="49E4F854FFE14B02B3D5A7E8111BF79F" box="[963,1025,2029,2048]" country="Bulgaria" name="Burgas" pageId="27" pageNumber="252">Burgas</collectingRegion>
, [42.41, 27.65], 
<date id="FF9E1076FFE14B02B482A7E81001F79F" box="[1172,1307,2029,2048]" pageId="27" pageNumber="252" value="1971-05-28" valueMax="1971-06-09" valueMin="1971-05-28">28.V-9.VI.1971</date>
, leg. Wallis (1 J 
<collectionCode id="ED31AE73FFE14B02B344A802169EF785" box="[850,900,2055,2074]" pageId="27" pageNumber="252">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFE14B02B382A80216E7F785" bold="true" box="[916,1021,2055,2074]" pageId="27" pageNumber="252">
<collectingCountry id="F3377626FFE14B02B382A80216E3F785" box="[916,1017,2055,2074]" name="Croatia" pageId="27" pageNumber="252">CROATIA</collectingCountry>
:
</emphasis>
Capela [Kapela], [44.89, 15.52], leg. Heyden (1 
<collectionCode id="ED31AE73FFE64B05B081A0BB15D3FF4E" box="[151,201,190,209]" pageId="28" pageNumber="253">SDEI</collectionCode>
); Velebit Mountains, [44.52, 15.22], wine traps, 
<date id="FF9E1076FFE64B05B24DA0BB178CFF4E" box="[603,662,190,209]" pageId="28" pageNumber="253" value="2005-05">V.2005</date>
(1 J cJen). 
<emphasis id="B954EAA4FFE64B05B081A0DD15EEFF74" bold="true" box="[151,244,216,235]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B081A0DD15EAFF74" box="[151,240,216,235]" name="Cyprus" pageId="28" pageNumber="253">CYPRUS</collectingCountry>
:
</emphasis>
Agios Nikolaios, [34.86, 32.75], 
<date id="FF9E1076FFE64B05B206A0DD1774FF74" box="[528,622,216,235]" pageId="28" pageNumber="253" value="1995-04-16">16.IV.1995</date>
, leg. H. Schmid (1 J 
<collectionCode id="ED31AE73FFE64B05B0D4A0F615E6FE99" box="[194,252,243,262]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
); Troodos Mountain, Stavros tis Psokas, [35.02, 32.63], 
<quantity id="4CD89B53FFE64B05B2C7A0F615D3FEBE" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="28" pageNumber="253" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">800- 950m</quantity>
, 
<date id="FF9E1076FFE64B05B0C5A10B1448FEBE" box="[211,338,270,289]" pageId="28" pageNumber="253" value="2010-04-12" valueMax="2010-04-18" valueMin="2010-04-12">12.-18.IV.2010</date>
, leg. I. Wolf (1 J cSch). 
<emphasis id="B954EAA4FFE64B05B238A10B17BFFEBE" bold="true" box="[558,677,270,289]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B238A10B17BAFEBE" box="[558,672,270,289]" name="Denmark" pageId="28" pageNumber="253">DENMARK</collectingCountry>
:
</emphasis>
Ovestrup Hede, 
<geoCoordinate id="EE145071FFE64B05B0D8A12D140EFEA4" box="[206,276,296,315]" degrees="56.2412" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="56.2412">56.2412</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B10BA12D1443FEA4" box="[285,345,296,315]" degrees="8.9297" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="8.9297">8.9297</geoCoordinate>
, pitfall trap, heathland grazed part, 
<date id="FF9E1076FFE64B05B294A12D17EEFEA4" box="[642,756,296,315]" pageId="28" pageNumber="253" value="2017-06-07" valueMax="2017-06-22" valueMin="2017-06-07">7-22.VI.2017</date>
, leg. E. Kristensen, D. Byriel, &amp; M. Justesen (1 J 
<collectionCode id="ED31AE73FFE64B05B250A1461790FEC9" box="[582,650,323,342]" pageId="28" pageNumber="253">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B28FA14617E2FEC9" bold="true" box="[665,760,323,342]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B28FA14617EEFEC9" box="[665,756,323,342]" name="France" pageId="28" pageNumber="253">FRANCE</collectingCountry>
:
</emphasis>
Avignon, [43.94, 4.81], 
<date id="FF9E1076FFE64B05B17AA15B14DCFEEE" box="[364,454,350,369]" pageId="28" pageNumber="253" value="1949-05-22">22.V.1949</date>
, leg. R. Skovgaard (1 J 
<collectionCode id="ED31AE73FFE64B05B2BFA15B17F4FEEE" box="[681,750,350,369]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Chigny n. Oise, [49.91, 3.76], 
<date id="FF9E1076FFE64B05B1B2A17D14E5FE14" box="[420,511,376,395]" pageId="28" pageNumber="253" value="1918-10-19">19.X.1918</date>
, leg. Salchert (
<specimenCount id="9D26FD3FFFE64B05B290A17D17BDFE14" box="[646,679,370,395]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B2B8A17D17F5FE14" box="[686,751,376,395]" pageId="28" pageNumber="253">ZMHB</collectionCode>
); Paris, [48.83, 2.41], 
<date id="FF9E1076FFE64B05B156A1961482FE39" box="[320,408,403,422]" pageId="28" pageNumber="253" value="1949-05-19">19.V.1949</date>
, leg. R. Skovgaard (1 J 
<collectionCode id="ED31AE73FFE64B05B272A19617B2FE39" box="[612,680,403,422]" pageId="28" pageNumber="253">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B2A3A19617E2FE39" bold="true" box="[693,760,403,422]" pageId="28" pageNumber="253">Cඈඋඌൾ:</emphasis>
Bocognano,[42.08,9.06],1905, leg.O.Leonhard (1 
<collectionCode id="ED31AE73FFE64B05B228A1AB1775FE5E" box="[574,623,430,449]" pageId="28" pageNumber="253">SDEI</collectionCode>
);Col de Bavella, [41.79, 9.23], 
<quantity id="4CD89B53FFE64B05B11AA1CD1452FE44" box="[268,328,456,475]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="28" pageNumber="253" unit="m" value="1240.0">1240m</quantity>
, East slope, 
<date id="FF9E1076FFE64B05B1B9A1CD170FFE44" box="[431,533,456,475]" pageId="28" pageNumber="253" value="1994-07-25">25.VII.1994</date>
leg. L. Zerche (1 J 
<collectionCode id="ED31AE73FFE64B05B2AAA1CD17F4FE44" box="[700,750,456,475]" pageId="28" pageNumber="253">SDEI</collectionCode>
); Foret de Valdo-Niello, Evisa, [42.28, 8.91], 
<quantity id="4CD89B53FFE64B05B219A1E61798FE69" box="[527,642,483,502]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="28" pageNumber="253" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000-1200 m</quantity>
, 
<date id="FF9E1076FFE64B05B29AA1E617EEFE69" box="[652,756,483,502]" pageId="28" pageNumber="253" value="1967-07-27">27.VII.1967</date>
, leg. Langemark-Lomholdt (1 J 
<collectionCode id="ED31AE73FFE64B05B1B5A1FB14FDFD8E" box="[419,487,510,529]" pageId="28" pageNumber="253">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B1E2A1FB1773FD8E" bold="true" box="[500,617,510,529]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B1E2A1FB177FFD8E" box="[500,613,510,529]" name="Germany" pageId="28" pageNumber="253">GERMANY</collectingCountry>
:
</emphasis>
Berlin-Marzahn, Hellersdorfer Berg, [52.53, 13.58], 
<date id="FF9E1076FFE64B05B1ABA21D173BFDB4" box="[445,545,536,555]" pageId="28" pageNumber="253" value="1997-08-08">8.VIII.1997</date>
, leg. Hieke (
<specimenCount id="9D26FD3FFFE64B05B29AA21D17B3FDB4" box="[652,681,530,555]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B2B8A21D17F5FDB4" box="[686,751,536,555]" pageId="28" pageNumber="253">ZMHB</collectionCode>
); 
<collectingRegion id="49E4F854FFE64B05B081A23615D7FDD9" box="[151,205,563,582]" country="Germany" name="Berlin" pageId="28" pageNumber="253">Berlin</collectingRegion>
, Prenzlauerberg Hof, [52.53, 13.46], 
<date id="FF9E1076FFE64B05B210A2361744FDD9" box="[518,606,563,582]" pageId="28" pageNumber="253" value="1985-05-29">29.V.1985</date>
, leg. I. Uhlig (1 J 
<collectionCode id="ED31AE73FFE64B05B081A24B15CDFDFE" box="[151,215,590,609]" pageId="28" pageNumber="253">ZMHB</collectionCode>
); Grunewald n. Zehlendorf, [52.48, 13.22], leg.Wagner (1 
<collectionCode id="ED31AE73FFE64B05B2ABA24B17F5FDFE" box="[701,751,590,609]" pageId="28" pageNumber="253">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B081A26D15E0FDE4" bold="true" box="[151,250,616,635]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B081A26D15EFFDE4" box="[151,245,616,635]" name="Greece" pageId="28" pageNumber="253">GREECE</collectingCountry>
:
</emphasis>
<collectingRegion id="49E4F854FFE64B05B117A26D1422FDE4" box="[257,312,616,635]" country="Greece" name="Attiki" pageId="28" pageNumber="253">Attica</collectingRegion>
, [38.05, 23.84], 
<date id="FF9E1076FFE64B05B1C6A26D1728FDE4" box="[464,562,616,635]" pageId="28" pageNumber="253" value="1922-04-17">17.IV.1922</date>
, leg. W. Liebmann (1 
<collectionCode id="ED31AE73FFE64B05B081A28615D3FD09" box="[151,201,643,662]" pageId="28" pageNumber="253">SDEI</collectionCode>
); Paleokastro, Halkidiki, [40.43, 23.42], 
<quantity id="4CD89B53FFE64B05B20FA2861751FD09" box="[537,587,643,662]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="253" unit="m" value="800.0">800m</quantity>
, 
<date id="FF9E1076FFE64B05B244A28617A3FD09" box="[594,697,643,662]" pageId="28" pageNumber="253" value="1983-07-11">11.VII.1983</date>
, leg.A. Casale (
<specimenCount id="9D26FD3FFFE64B05B0CAA29B15E0FD2F" box="[220,250,663,689]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cBor); 
<quantity id="4CD89B53FFE64B05B12FA29B147EFD2E" box="[313,356,670,689]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="253" unit="km" value="2.0">2 km</quantity>
W Panagia, Trikala, [39.79, 21.33], 
<date id="FF9E1076FFE64B05B285A29B17EEFD2E" box="[659,756,670,689]" pageId="28" pageNumber="253" value="1982-06-15">15.VI.1982</date>
, leg. R Danielsson (1 J 
<collectionCode id="ED31AE73FFE64B05B14BA2BD1486FD54" box="[349,412,696,715]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
); Voria Pindos, Morfi, [40.21, 21.21], dry slope, 
<date id="FF9E1076FFE64B05B0D8A2D61435FD79" box="[206,303,723,742]" pageId="28" pageNumber="253" value="1997-06-26">26.VI.1997</date>
, leg. Winkelmann (
<specimenCount id="9D26FD3FFFE64B05B1C2A2D614E8FD79" box="[468,498,717,742]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch); 
<emphasis id="B954EAA4FFE64B05B224A2D6176CFD79" bold="true" box="[562,630,723,742]" pageId="28" pageNumber="253">Cඈඋൿඎ:</emphasis>
Pandokratoras, 
<geoCoordinate id="EE145071FFE64B05B081A2EB15C7FC9E" box="[151,221,750,769]" degrees="39.7492" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="39.7492">39.7492</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B0F5A2EB1432FC9E" box="[227,296,750,769]" degrees="19.8701" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="19.8701">19.8701</geoCoordinate>
,N-slope with scattered pine,decidious trees,and bushes, litter and roots between rocks sifted, 
<quantity id="4CD89B53FFE64B05B1DFA30D14E5FC84" box="[457,511,776,795]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="253" unit="m" value="800.0">800 m</quantity>
, 
<date id="FF9E1076FFE64B05B21EA30D1744FC84" box="[520,606,776,795]" pageId="28" pageNumber="253" value="2017-06-04">4.VI.2017</date>
, leg. Schülke (1 J cSch);Pantokrator,[39.74,19.87], 
<date id="FF9E1076FFE64B05B1A5A32614EAFCA9" box="[435,496,803,822]" pageId="28" pageNumber="253" value="1964-05">V.1964</date>
, leg.Palm (
<specimenCount id="9D26FD3FFFE64B05B258A3261773FCA9" box="[590,617,797,822]" pageId="28" pageNumber="253" type="female">1♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B278A32617B7FCA9" box="[622,685,803,822]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B2AEA32617E2FCA9" bold="true" box="[696,760,803,822]" pageId="28" pageNumber="253">Cඋൾඍൾ:</emphasis>
Georgioupouli, [35.36, 24.26], 
<date id="FF9E1076FFE64B05B18CA33B1703FCCE" box="[410,537,830,849]" pageId="28" pageNumber="253" value="1995-04-24" valueMax="1995-04-25" valueMin="1995-04-24">24.-25.IV.1995</date>
, leg. B. Jaeger (
<specimenCount id="9D26FD3FFFE64B05B2B6A33B17A7FCCF" box="[672,701,823,849]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch); Ida mts., ca. 
<quantity id="4CD89B53FFE64B05B117A35D1431FCF4" box="[257,299,856,875]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="253" unit="km" value="6.0">6 km</quantity>
S Analipsis, 
<geoCoordinate id="EE145071FFE64B05B18EA35D14C4FCF4" box="[408,478,856,875]" degrees="35.1752" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="35.1752">35.1752</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B1F0A35D1736FCF4" box="[486,556,856,875]" degrees="24.8391" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="24.8391">24.8391</geoCoordinate>
, 
<taxonomicName id="4C204D35FFE64B05B222A35D1760FCF4" box="[564,634,856,875]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="251" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
forest, 
<quantity id="4CD89B53FFE64B05B2AEA35D17EFFCF4" box="[696,757,856,875]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="28" pageNumber="253" unit="m" value="1260.0">1260m</quantity>
, 
<date id="FF9E1076FFE64B05B081A37615EBFC19" box="[151,241,883,902]" pageId="28" pageNumber="253" value="2006-10-21">21.X.2006</date>
, leg. M. Schülke (1 J cSch); Mt.Ida, 
<quantity id="4CD89B53FFE64B05B233A376174FFC19" box="[549,597,883,902]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="28" pageNumber="253" unit="km" value="28.0">28km</quantity>
SW Iraklio, [35.23, 24.77], 
<quantity id="4CD89B53FFE64B05B0C1A38B1408FC3E" box="[215,274,910,929]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="253" unit="m" value="1600.0">1600m</quantity>
, 
<date id="FF9E1076FFE64B05B10CA38B146BFC3E" box="[282,369,910,929]" pageId="28" pageNumber="253" value="1991-09-07">7.IX.1991</date>
, leg. N. Esser (
<specimenCount id="9D26FD3FFFE64B05B1F8A38B1710FC3F" box="[494,522,903,929]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B219A38B1749FC3E" box="[527,595,910,929]" pageId="28" pageNumber="253">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B276A38B17D7FC3E" bold="true" box="[608,717,910,929]" pageId="28" pageNumber="253">KൺඋඉൺඍHඈඌ:</emphasis>
Karpathos, 
<geoCoordinate id="EE145071FFE64B05B0CEA3AD1404FC24" box="[216,286,936,955]" degrees="35.5097" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="35.5097">35.5097</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B130A3AD1476FC24" box="[294,364,936,955]" degrees="27.2189" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="27.2189">27.2189</geoCoordinate>
, 
<quantity id="4CD89B53FFE64B05B165A3AD1485FC24" box="[371,415,936,955]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="253" unit="m" value="40.0">40 m</quantity>
, terracod grassy fallow, 
<date id="FF9E1076FFE64B05B27FA3AD17A1FC24" box="[617,699,936,955]" pageId="28" pageNumber="253" value="2016-01-01">01.I.2016</date>
, leg. V. Assing (
<collectionCode id="ED31AE73FFE64B05B0CBA3C61404FC49" box="[221,286,963,982]" pageId="28" pageNumber="253">ZMHB</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B13BA3C6146CFC49" bold="true" box="[301,374,963,982]" pageId="28" pageNumber="253">Lൾඌൻඈඌ:</emphasis>
SW Agiassos, road to Olympos peak, [39.06, 26.36], 
<quantity id="4CD89B53FFE64B05B0CCA3DB1409FC6E" box="[218,275,990,1009]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="28" pageNumber="253" unit="m" value="700.0">700 m</quantity>
, 
<date id="FF9E1076FFE64B05B108A3DB149AFC6E" box="[286,384,990,1009]" pageId="28" pageNumber="253" value="1996-06-10">10.VI.1996</date>
, leg. Jäch (1 J 
<specimenCount id="9D26FD3FFFE64B05B218A3DB172AFC6F" box="[526,560,983,1009]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B221A3DB176BFC6E" box="[567,625,990,1009]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B297A3DB17E2FC6E" bold="true" box="[641,760,990,1009]" pageId="28" pageNumber="253">Pൾඅඈඉඈඇඇൾඌ:</emphasis>
Korinthia, Kyllini, Zoria, [37.94, 22.43], 
<quantity id="4CD89B53FFE64B05B1FBA3FD1734FB94" box="[493,558,1016,1035]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="253" unit="m" value="2000.0">2000 m</quantity>
, 
<date id="FF9E1076FFE64B05B221A3FD1797FB94" box="[567,653,1016,1035]" pageId="28" pageNumber="253" value="1995-06-03">3.VI.1995</date>
, leg. Giachino &amp; Vailati (1 J 
<specimenCount id="9D26FD3FFFE64B05B13BA4161446FBB9" box="[301,348,1037,1062]" pageId="28" pageNumber="253" type="female">3 ♀♀</specimenCount>
cBor); Kyllini Mountain, [37.97, 22.39], 
<quantity id="4CD89B53FFE64B05B2AEA41617EFFBB9" box="[696,757,1043,1062]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="28" pageNumber="253" unit="m" value="2100.0">2100m</quantity>
, North side, below summit over valley, 
<date id="FF9E1076FFE64B05B1F7A42B172DFBDE" box="[481,567,1070,1089]" pageId="28" pageNumber="253" value="1996-06-09">9.VI.1996</date>
, leg. P. Wunderle (
<specimenCount id="9D26FD3FFFE64B05B2CFA42B17E3FBDF" box="[729,761,1063,1089]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch); Sparta, [37.06, 22.43], 
<date id="FF9E1076FFE64B05B18DA44D14F2FBC4" box="[411,488,1096,1115]" pageId="28" pageNumber="253" value="1962-05-03">3.V.1962</date>
, leg. Skovgaard (1 J 
<collectionCode id="ED31AE73FFE64B05B2BDA44D17F5FBC4" box="[683,751,1096,1115]" pageId="28" pageNumber="253">NHMD</collectionCode>
); N Taygetos, 
<quantity id="4CD89B53FFE64B05B110A4661429FBE9" box="[262,307,1123,1142]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="253" unit="km" value="6.0">6 km</quantity>
NE Thouria, 
<geoCoordinate id="EE145071FFE64B05B1BAA46614E8FBE9" box="[428,498,1123,1142]" degrees="37.1401" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="37.1401">37.1401</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B1EBA4661759FBE9" box="[509,579,1123,1142]" degrees="22.1284" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="22.1284">22.1284</geoCoordinate>
, litter and soil under bushes, sifted, 
<quantity id="4CD89B53FFE64B05B105A47B1450FB0E" box="[275,330,1150,1169]" metricMagnitude="2" metricUnit="m" metricValue="5.84" pageId="28" pageNumber="253" unit="m" value="584.0">584 m</quantity>
, 
<date id="FF9E1076FFE64B05B145A47B14ABFB0E" box="[339,433,1150,1169]" pageId="28" pageNumber="253" value="2015-04-20">20.IV.2015</date>
, leg. M. Schülke (
<specimenCount id="9D26FD3FFFE64B05B25DA47B1770FB0F" box="[587,618,1143,1169]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch). 
<emphasis id="B954EAA4FFE64B05B2BEA47B17E2FB0E" bold="true" box="[680,760,1150,1169]" pageId="28" pageNumber="253">RHඈൽൾඌ:</emphasis>
Prophitis Ilias, 
<geoCoordinate id="EE145071FFE64B05B100A49D1446FB34" box="[278,348,1176,1195]" degrees="36.2734" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="36.2734">36.2734</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B173A49D14B1FB34" box="[357,427,1176,1195]" degrees="27.9287" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="27.9287">27.9287</geoCoordinate>
, pine forest, under stones &amp; sifted from litter &amp; moss, 
<quantity id="4CD89B53FFE64B05B106A4B61459FB59" box="[272,323,1203,1222]" metricMagnitude="2" metricUnit="m" metricValue="6.85" pageId="28" pageNumber="253" unit="m" value="685.0">685m</quantity>
, 
<date id="FF9E1076FFE64B05B15DA4B614B9FB59" box="[331,419,1203,1222]" pageId="28" pageNumber="253" value="2015-05-17">17.V.2015</date>
, leg. Wrase &amp; Laser (1 J cSch). 
<emphasis id="B954EAA4FFE64B05B2ADA4B617E2FB59" bold="true" box="[699,760,1203,1222]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B2ADA4B617EEFB59" box="[699,756,1203,1222]" name="Iran" pageId="28" pageNumber="253">IRAN</collectingCountry>
:
</emphasis>
Kohgiluyech-Va-Boyern-Ahmad province, 
<quantity id="4CD89B53FFE64B05B212A4CB1720FB7E" box="[516,570,1230,1249]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="253" unit="km" value="20.0">20 km</quantity>
SW Yasuk, 
<quantity id="4CD89B53FFE64B05B2B5A4CB17C3FB7E" box="[675,729,1230,1249]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="253" unit="km" value="10.0">10 km</quantity>
SE Sepidor, [30.56, 51.51], 
<date id="FF9E1076FFE64B05B177A4ED14DEFB64" box="[353,452,1256,1275]" pageId="28" pageNumber="253" value="2007-05-05" valueMax="2007-05-06" valueMin="2007-05-05">5.-6.V.2007</date>
, leg.A. Anistchschenko (
<specimenCount id="9D26FD3FFFE64B05B28CA4ED17A2FB64" box="[666,696,1250,1275]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B2ABA4ED17F5FB64" box="[701,751,1256,1275]" pageId="28" pageNumber="253">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B081A50615E5FA89" bold="true" box="[151,255,1283,1302]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B081A50615E1FA89" box="[151,251,1283,1302]" name="Ireland" pageId="28" pageNumber="253">IRELAND</collectingCountry>
:
</emphasis>
Inch Dingle Eire, [52.14, -9.97], 
<date id="FF9E1076FFE64B05B206A506174DFA89" box="[528,599,1283,1302]" pageId="28" pageNumber="253" value="2012-09">IX.2012</date>
, leg. J.J. Shaw (1 J cJen); Pembury, Kent 
<geoCoordinate id="EE145071FFE64B05B158A51B148EFAAE" box="[334,404,1310,1329]" degrees="51.1625" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="51.1625">51.1625</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B18AA51B14C2FAAE" box="[412,472,1310,1329]" degrees="0.3431" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="0.3431">0.3431</geoCoordinate>
, 
<date id="FF9E1076FFE64B05B1C9A51B175CFAAE" box="[479,582,1310,1329]" pageId="28" pageNumber="253" value="2011-07-24">24.VII.2011</date>
, leg. J.J. Shaw (2 JJ cJen). 
<emphasis id="B954EAA4FFE64B05B0DDA53D143AFAD4" bold="true" box="[203,288,1336,1355]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B0DDA53D1406FAD4" box="[203,284,1336,1355]" name="Israel" pageId="28" pageNumber="253">ISRAEL</collectingCountry>
:
</emphasis>
Upper Galilee,Har Avivim,W Avivim, 
<geoCoordinate id="EE145071FFE64B05B275A53D17B2FAD4" box="[611,680,1336,1355]" degrees="33.0833" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="33.0833">33.0833</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B2B9A53D17EFFAD4" box="[687,757,1336,1355]" degrees="35.4923" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="35.4923">35.4923</geoCoordinate>
, open stony grazing land limestone, 
<quantity id="4CD89B53FFE64B05B1AAA55614F7FAF9" box="[444,493,1363,1382]" metricMagnitude="2" metricUnit="m" metricValue="7.68" pageId="28" pageNumber="253" unit="m" value="768.0">768m</quantity>
, 
<date id="FF9E1076FFE64B05B1E3A5561749FAF9" box="[501,595,1363,1382]" pageId="28" pageNumber="253" value="2006-04-21">21.IV.2006</date>
, leg. D.W.Wrase (1 J cSch); Upper Galilee, Meron Mts. Har Meron, Kamin Rom, 
<geoCoordinate id="EE145071FFE64B05B2B9A56B17EFFA1E" box="[687,757,1390,1409]" degrees="32.9908" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="32.9908">32.9908</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B081A58D15C1FA04" box="[151,219,1416,1435]" degrees="35.4111" direction="east" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="35.4111">35.4111</geoCoordinate>
, open stony grazing land, limestone, 
<quantity id="4CD89B53FFE64B05B213A58D175AFA04" box="[517,576,1416,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="28" pageNumber="253" unit="m" value="1100.0">1100m</quantity>
, 
<date id="FF9E1076FFE64B05B250A58D17B9FA04" box="[582,675,1416,1435]" pageId="28" pageNumber="253" value="2006-04-28">28.IV.2006</date>
,leg.D.W. Wrase (1J cSch). 
<emphasis id="B954EAA4FFE64B05B138A5A61468FA29" bold="true" box="[302,370,1443,1462]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B138A5A61474FA29" box="[302,366,1443,1462]" name="Italy" pageId="28" pageNumber="253">ITALY</collectingCountry>
:
</emphasis>
CalabriaAspromonte,[38.21, 15.99],1905,leg. Pagametti (1 
<collectionCode id="ED31AE73FFE64B05B112A5BB142CFA4E" box="[260,310,1470,1489]" pageId="28" pageNumber="253">SDEI</collectionCode>
); 
<collectingRegion id="49E4F854FFE64B05B152A5BB1497FA4E" box="[324,397,1470,1489]" country="Italy" name="Calabria" pageId="28" pageNumber="253">Calabria</collectingRegion>
, Roseto Capo Spulico, [39.98, 16.61], 
<quantity id="4CD89B53FFE64B05B2DFA5BB17EFFA4E" box="[713,757,1470,1489]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="253" unit="m" value="20.0">20 m</quantity>
, 
<date id="FF9E1076FFE64B05B081A5DD15FEFA74" box="[151,228,1496,1515]" pageId="28" pageNumber="253" value="1998-10-06">6.X.1998</date>
leg.F.Angelini(
<specimenCount id="9D26FD3FFFE64B05B17AA5DD149FFA74" box="[364,389,1490,1515]" pageId="28" pageNumber="253" type="female">1♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B19DA5DD14D4FA74" box="[395,462,1496,1515]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Dicomano, 
<collectingRegion id="49E4F854FFE64B05B221A5DD1766FA74" box="[567,636,1496,1515]" country="Italy" name="Toscana" pageId="28" pageNumber="253">Toscana</collectingRegion>
,[43.89,11.52], 
<date id="FF9E1076FFE64B05B081A5F615D4F999" box="[151,206,1523,1542]" pageId="28" pageNumber="253" value="1971-01">I.1971</date>
, leg Bordoni (1J cBor); Macerata, [43.29, 13.45], 
<date id="FF9E1076FFE64B05B27AA5F617A1F999" box="[620,699,1523,1542]" pageId="28" pageNumber="253" value="1955-10-01">1.X.1955</date>
, leg.D. Caccamicelo (
<specimenCount id="9D26FD3FFFE64B05B118A60B1433F9BF" box="[270,297,1543,1569]" pageId="28" pageNumber="253" type="female">1♀</specimenCount>
cBor); 
<collectingRegion id="49E4F854FFE64B05B170A60B14B2F9BE" box="[358,424,1550,1569]" country="Italy" name="Marche" pageId="28" pageNumber="253">Marche</collectingRegion>
,Montemonaco, Isola San Biagio,[42.91, 13.31], fields, 
<quantity id="4CD89B53FFE64B05B11AA62D145BF9A4" box="[268,321,1576,1595]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="253" unit="m" value="900.0">900 m</quantity>
, 
<date id="FF9E1076FFE64B05B15EA62D14F8F9A4" box="[328,482,1576,1595]" pageId="28" pageNumber="253" value="2004-07-27" valueMax="2004-09-01" valueMin="2004-07-27">27.VII.-1.IX.2004</date>
, leg.M.Rismondo &amp; R. Fabbri (2 JJ 
<specimenCount id="9D26FD3FFFE64B05B0ACA64615FCF9C9" box="[186,230,1597,1622]" pageId="28" pageNumber="253" type="female">2♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B0FDA6461435F9C9" box="[235,303,1603,1622]" pageId="28" pageNumber="253">NHMD</collectionCode>
); same locality, black hornbeam forest, 
<date id="FF9E1076FFE64B05B265A64617EFF9C9" box="[627,757,1603,1622]" pageId="28" pageNumber="253" value="2004-09-01" valueMax="2004-10-07" valueMin="2004-09-01">1.IX.-7.X.2004</date>
, leg. M. Rismondo &amp; R. Fabbri (1 J 
<collectionCode id="ED31AE73FFE64B05B1CEA65B1706F9EE" box="[472,540,1630,1649]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Modena, Prignano sulla Secchia, La Quercia, [44.45, 10.73], pitfall traps, 
<quantity id="4CD89B53FFE64B05B220A67D1777F914" box="[566,621,1656,1675]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="253" unit="m" value="400.0">400 m</quantity>
, 
<date id="FF9E1076FFE64B05B260A67D17EEF914" box="[630,756,1656,1675]" pageId="28" pageNumber="253" value="2006-07-05" valueMax="2006-07-18" valueMin="2006-07-05">5.-18.VII.2006</date>
, leg. R. Fabbri (1 J 
<collectionCode id="ED31AE73FFE64B05B120A6961460F939" box="[310,378,1683,1702]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Reggio di 
<collectingRegion id="49E4F854FFE64B05B1CBA696173CF939" box="[477,550,1683,1702]" country="Italy" name="Calabria" pageId="28" pageNumber="253">Calabria</collectingRegion>
, river Sant Agata, [38.09, 15.66], 
<date id="FF9E1076FFE64B05B0C1A6AB142EF95E" box="[215,308,1710,1729]" pageId="28" pageNumber="253" value="1959-03-11">11.III.1959</date>
(1 J cBor); Settecamini, Rome, 
<collectingRegion id="49E4F854FFE64B05B252A6AB176CF95E" box="[580,630,1710,1729]" country="Italy" name="Lazio" pageId="28" pageNumber="253">Lazio</collectingRegion>
, [41.93, 12.62], 
<date id="FF9E1076FFE64B05B081A6CD15EFF944" box="[151,245,1736,1755]" pageId="28" pageNumber="253" value="1969-11-23">23.Xi.1969</date>
(1 J 
<specimenCount id="9D26FD3FFFE64B05B130A6CD145FF944" box="[294,325,1730,1755]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cBor); Sovara, Anghiari, [43.53, 12.03], 
<date id="FF9E1076FFE64B05B28BA6CD17EEF944" box="[669,756,1736,1755]" pageId="28" pageNumber="253" value="1930-05-19">19.V.1930</date>
, leg. A. Andreini (
<specimenCount id="9D26FD3FFFE64B05B138A6E61455F969" box="[302,335,1757,1782]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cBor). 
<emphasis id="B954EAA4FFE64B05B186A6E614F2F969" bold="true" box="[400,488,1763,1782]" pageId="28" pageNumber="253">Sൺඋൽංඇංൺ:</emphasis>
Aritzo Mountain, Gennargentu, [40.01, 9.29], leg. C.Kruger (2 
<collectionCode id="ED31AE73FFE64B05B180A6FB14D2F88E" box="[406,456,1790,1809]" pageId="28" pageNumber="253">SDEI</collectionCode>
); Ottana, [40.23, 9.04], 
<date id="FF9E1076FFE64B05B29CA6FB17F9F88E" box="[650,739,1790,1809]" pageId="28" pageNumber="253" value="1974-10-15">15.X.1974</date>
(3 JJ 
<specimenCount id="9D26FD3FFFE64B05B0ACA71D15CFF8B4" box="[186,213,1810,1835]" pageId="28" pageNumber="253" type="female">1♀</specimenCount>
cBor);Oristano,[39.89,8.59], 
<date id="FF9E1076FFE64B05B1C7A71D1707F8B4" box="[465,541,1816,1835]" pageId="28" pageNumber="253" value="1975-05-03">3.V.1975</date>
,leg.S. Ripze (
<specimenCount id="9D26FD3FFFE64B05B285A71D17A4F8B4" box="[659,702,1810,1835]" pageId="28" pageNumber="253" type="female">2♀♀</specimenCount>
cBor); 
<emphasis id="B954EAA4FFE64B05B081A73615C9F8D9" bold="true" box="[151,211,1843,1862]" pageId="28" pageNumber="253">Sංർංඅඒ:</emphasis>
Ficuzza, [37.85, 13.38], 1906, leg. O.Leonhard (1 
<collectionCode id="ED31AE73FFE64B05B279A73617BBF8D9" box="[623,673,1843,1862]" pageId="28" pageNumber="253">SDEI</collectionCode>
);Messina, [38.18, 15.54], leg. Schiödte (
<specimenCount id="9D26FD3FFFE64B05B181A74B14D0F8FF" box="[407,458,1863,1889]" pageId="28" pageNumber="253" type="female">4 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B1D8A74B1708F8FE" box="[462,530,1870,1889]" pageId="28" pageNumber="253">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B234A74B1786F8FE" bold="true" box="[546,668,1870,1889]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B234A74B1782F8FE" box="[546,664,1870,1889]" name="Morocco" pageId="28" pageNumber="253">MOROCCO</collectingCountry>
:
</emphasis>
Ain Leuh, [33.28, -5.33], 
<date id="FF9E1076FFE64B05B100A76D1474F8E4" box="[278,366,1896,1915]" pageId="28" pageNumber="253" value="1925-05-17">17.V.1925</date>
, leg. M. Escalera (1 J 
<specimenCount id="9D26FD3FFFE64B05B222A76D177CF8E4" box="[564,614,1890,1915]" pageId="28" pageNumber="253" type="female">4 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B27CA76D17B7F8E4" box="[618,685,1896,1915]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="28" pageNumber="253" type="Museum">MNCN</collectionCode>
); Ar Rif, Chefchaounen reg., env.Bad Bessen, 
<geoCoordinate id="EE145071FFE64B05B1D1A7861716F809" box="[455,524,1923,1942]" degrees="34.9913" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="34.9913">34.9913</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B204A786174EF809" box="[530,596,1923,1942]" degrees="4.8555" direction="west" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="-4.8555">-4.8555</geoCoordinate>
, 
<quantity id="4CD89B53FFE64B05B24CA7861782F809" box="[602,664,1923,1942]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="28" pageNumber="253" unit="m" value="1124.0">1124 m</quantity>
, 
<date id="FF9E1076FFE64B05B289A78617EFF809" box="[671,757,1923,1942]" pageId="28" pageNumber="253" value="2007-06-03">3.VI.2007</date>
, leg. P. Hlaváč (1 J cSch); Ar Rif, Chefchaounen region, rd. Bad Berret - Bad Bessen, 
<quantity id="4CD89B53FFE64B05B115A7BD145EF854" box="[259,324,1976,1995]" metricMagnitude="3" metricUnit="m" metricValue="1.392" pageId="28" pageNumber="253" unit="m" value="1392.0">1392 m</quantity>
, 
<geoCoordinate id="EE145071FFE64B05B15BA7BD1489F854" box="[333,403,1976,1995]" degrees="35.0169" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="35.0169">35.0169</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B18DA7BD14C7F854" box="[411,477,1976,1995]" degrees="4.0035" direction="west" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="-4.0035">-4.0035</geoCoordinate>
, 
<date id="FF9E1076FFE64B05B1F3A7BD1748F854" box="[485,594,1976,1995]" pageId="28" pageNumber="253" value="2007-06-03" valueMax="2007-06-06" valueMin="2007-06-03">3.-6.VI.2007</date>
, leg. P. Hlaváč (
<specimenCount id="9D26FD3FFFE64B05B2CCA7BD17E2F854" box="[730,760,1970,1995]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch); Taza, [34.22, -4.00], leg. F. Schubert (
<specimenCount id="9D26FD3FFFE64B05B206A7D61734F879" box="[528,558,1997,2022]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B225A7D61777F879" box="[563,621,2003,2022]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B26CA7D617E2F879" bold="true" box="[634,760,2003,2022]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B26CA7D617EEF879" box="[634,756,2003,2022]" name="Portugal" pageId="28" pageNumber="253">PORTUGAL</collectingCountry>
:
</emphasis>
Penedo da Meditação, 
<collectingRegion id="49E4F854FFE64B05B144A7EB1487F79E" box="[338,413,2030,2049]" country="Portugal" name="Coimbra" pageId="28" pageNumber="253">Coimbra</collectingRegion>
, [40.224371, -8.40], 
<date id="FF9E1076FFE64B05B25EA7EB17BDF79E" box="[584,679,2030,2049]" pageId="28" pageNumber="253" value="1961-04-24">24.IV.1961</date>
, leg.R.N. F. (1 J 
<collectionCode id="ED31AE73FFE64B05B0CDA80D140FF784" box="[219,277,2056,2075]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B135A80D1466F784" bold="true" box="[291,380,2056,2075]" pageId="28" pageNumber="253">Mൺൽൾංඋൺ:</emphasis>
Faja da Nogueira, [32.74, -16.90], laurisilva with big 
<emphasis id="B954EAA4FFE64B05B392A0BB16D2FF4E" box="[900,968,190,209]" italics="true" pageId="28" pageNumber="253">Ocoteas</emphasis>
, 
<quantity id="4CD89B53FFE64B05B3C4A0BB1112FF4E" box="[978,1032,190,209]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="28" pageNumber="253" unit="m" value="750.0">750 m</quantity>
, 
<date id="FF9E1076FFE64B05B406A0BB1168FF4E" box="[1040,1138,190,209]" pageId="28" pageNumber="253" value="1980-09-20">20.IX.1980</date>
, leg. H. Enghoff &amp; O. Martin (2 JJ 
<specimenCount id="9D26FD3FFFE64B05B32AA0DD1673FF74" box="[828,873,210,235]" pageId="28" pageNumber="253" type="female">3 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B37BA0DD16ABFF74" box="[877,945,216,235]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Paul da Serra, [32.73, -17.05], 
<date id="FF9E1076FFE64B05B4AFA0DD103BFF74" box="[1209,1313,216,235]" pageId="28" pageNumber="253" value="1987-12-27">27.XII.1987</date>
, leg.Gillerfors (1 J 
<collectionCode id="ED31AE73FFE64B05B37EA0F616BDFE99" box="[872,935,243,262]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B3A0A0F616E0FE99" bold="true" box="[950,1018,243,262]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B3A0A0F616EDFE99" box="[950,1015,243,262]" name="Spain" pageId="28" pageNumber="253">SPAIN</collectingCountry>
:
</emphasis>
Castanos de Calabazas, 
<quantity id="4CD89B53FFE64B05B4DCA0F611EFFE99" box="[1226,1269,243,262]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="253" unit="km" value="1.0">1 km</quantity>
S Castanar de Ibor, Extemadura, 
<geoCoordinate id="EE145071FFE64B05B3A6A10B16E2FEBE" box="[944,1016,270,289]" degrees="39.6235" direction="north" orientation="latitude" pageId="28" pageNumber="253" precision="5" value="39.6235">39.6235</geoCoordinate>
, 
<geoCoordinate id="EE145071FFE64B05B415A10B115DFEBE" box="[1027,1095,270,289]" degrees="5.4128" direction="west" orientation="longitude" pageId="28" pageNumber="253" precision="5" value="-5.4128">-5.4128</geoCoordinate>
, 
<quantity id="4CD89B53FFE64B05B444A10B1197FEBE" box="[1106,1165,270,289]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="28" pageNumber="253" unit="m" value="860.0">860 m</quantity>
, sifted mossy rock scree litter, chestnut and oak forest /w rock screes and small stream, 
<date id="FF9E1076FFE64B05B51EA12D107AFEA4" box="[1288,1376,296,315]" pageId="28" pageNumber="253" value="2019-05-10">10.V.2019</date>
, leg.A. K. Hansen, J. J. Shaw, &amp; J. Kypke (
<collectionCode id="ED31AE73FFE64B05B470A14611B1FEC9" box="[1126,1195,323,342]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Cavarrubias [Covarrubias], Burgos,[42.05, -3.51], 
<date id="FF9E1076FFE64B05B3E1A15B114CFEEE" box="[1015,1110,350,369]" pageId="28" pageNumber="253" value="1995-11-10">10.XI.1995</date>
, leg.A.Anichtchenko(1J cSha); Lloret de Mar, [41.69, 2.83], 
<date id="FF9E1076FFE64B05B3E3A17D119BFE14" box="[1013,1153,376,395]" pageId="28" pageNumber="253" value="1961-09-17" valueMax="1961-10-06" valueMin="1961-09-17">17.IX.-6.X.1961</date>
, leg. Palm (1 J 
<collectionCode id="ED31AE73FFE64B05B513A17D105EFE14" box="[1285,1348,376,395]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
); Sierra de Ancares,A Degrada vill., [42.81, -6.87], 
<quantity id="4CD89B53FFE64B05B49CA19611E0FE39" box="[1162,1274,403,422]" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="28" pageNumber="253" unit="m" value="1175.0" valueMax="1250.0" valueMin="1100.0">1100-1250 m</quantity>
, 
<date id="FF9E1076FFE64B05B514A196106CFE39" box="[1282,1398,403,422]" pageId="28" pageNumber="253" value="1996-07-08" valueMax="1996-07-09" valueMin="1996-07-08">8.-9.VII.1996</date>
, leg. Wrase (2 JJ cSch); Sierra de Guadarrama, Navacerrada,Arroyo de Peca [Peña] Cabrita, [40.77, -3.99], 
<quantity id="4CD89B53FFE64B05B420A1CD116FFE44" box="[1078,1141,456,475]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="253" unit="m" value="1800.0">1800 m</quantity>
, 
<date id="FF9E1076FFE64B05B46BA1CD11C9FE44" box="[1149,1235,456,475]" pageId="28" pageNumber="253" value="2001-09-05">5.IX.2001</date>
, leg.A.Anichtchenko (1 J 
<collectionCode id="ED31AE73FFE64B05B359A1E61689FE69" box="[847,915,483,502]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Sierra Nevada,rio Lanjaron,Tello,[36.92, -3.47], 
<quantity id="4CD89B53FFE64B05B523A1E6106EFE69" box="[1333,1396,483,502]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="253" unit="m" value="1500.0">1500 m</quantity>
, 
<date id="FF9E1076FFE64B05B56DA1E61680FD8E" pageId="28" pageNumber="253" value="2010-03-29" valueMax="2010-03-31" valueMin="2010-03-29">29.- 31.III.2010</date>
, leg.Shavrin &amp;Anichtchenko (1J 
<collectionCode id="ED31AE73FFE64B05B4A1A1FB11E1FD8E" box="[1207,1275,510,529]" pageId="28" pageNumber="253">NHMD</collectionCode>
);Toledo,El Pielago Navamorcruende, [40.15, -4.79], 
<date id="FF9E1076FFE64B05B443A21D11ADFDB4" box="[1109,1207,536,555]" pageId="28" pageNumber="253" value="2005-08-04">4.VIII.2005</date>
leg. Lencina &amp; Andujar (1 
<collectionCode id="ED31AE73FFE64B05B32AA2361666FDD9" box="[828,892,563,582]" pageId="28" pageNumber="253">ZMHB</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B39EA23616CBFDD9" bold="true" box="[904,977,563,582]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B39EA23616D6FDD9" box="[904,972,563,582]" name="Syria" pageId="28" pageNumber="253">SYRIA</collectingCountry>
:
</emphasis>
Krak des Chevaliers, [34.75, 36.29], 
<quantity id="4CD89B53FFE64B05B516A236102FFDD9" box="[1280,1333,563,582]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="253" unit="m" value="600.0">600 m</quantity>
, 
<date id="FF9E1076FFE64B05B52AA236109DFDD9" box="[1340,1415,563,582]" pageId="28" pageNumber="253" value="1982-05-07">7.V.1982</date>
(1 J 
<specimenCount id="9D26FD3FFFE64B05B346A24B166AFDFF" box="[848,880,583,609]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B363A24B16BCFDFE" box="[885,934,590,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="253" type="Museum">MCZ</collectionCode>
); MashtalHelu [Mashta Al Hilu], E of Safita, [34.87, 36.26], 
<date id="FF9E1076FFE64B05B32AA26D1680FDE4" box="[828,922,616,635]" pageId="28" pageNumber="253" value="2008-04-30">30.IV.2008</date>
, leg.Skoupý (1 J cSch); Zainie, 
<quantity id="4CD89B53FFE64B05B4BDA26D11C5FDE4" box="[1195,1247,616,635]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="253" unit="km" value="45.0">45 km</quantity>
NE of 
<collectingRegion id="49E4F854FFE64B05B50FA26D1040FDE4" box="[1305,1370,616,635]" country="Syria" name="Al Ladhiqiyah" pageId="28" pageNumber="253">Latakia</collectingRegion>
, [35.78, 36.19], 
<date id="FF9E1076FFE64B05B36DA28616DDFD09" box="[891,967,643,662]" pageId="28" pageNumber="253" value="1982-05-04">4.V.1982</date>
, leg. M. Dvorák (
<specimenCount id="9D26FD3FFFE64B05B441A2861199FD09" box="[1111,1155,637,662]" pageId="28" pageNumber="253" type="female">3♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B49FA28611A0FD09" box="[1161,1210,643,662]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="253" type="Museum">MCZ</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B4D0A2861078FD09" bold="true" box="[1222,1378,643,662]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B4D0A286103FFD09" box="[1222,1317,643,662]" name="Sweden" pageId="28" pageNumber="253">SWEDEN</collectingCountry>
: Fൺඋද:
</emphasis>
Farön, [57.94, 19.14] (1 
<collectionCode id="ED31AE73FFE64B05B3C7A29B110AFD2E" box="[977,1040,670,689]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B409A29B11A6FD2E" bold="true" box="[1055,1212,670,689]" pageId="28" pageNumber="253">GඈඍඌKൺ Sൺඇൽදඇ:</emphasis>
Sandön, [58.36, 19.24] (1 
<collectionCode id="ED31AE73FFE64B05B32AA2BD1661FD54" box="[828,891,696,715]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B39EA2BD16D6FD54" bold="true" box="[904,972,696,715]" pageId="28" pageNumber="253">Öඅൺඇൽ:</emphasis>
Stenåsa, [56.51, 16.59] (1 
<collectionCode id="ED31AE73FFE64B05B4BEA2BD11FDFD54" box="[1192,1255,696,715]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B4E2A2BD104EFD54" bold="true" box="[1268,1364,696,715]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B4E2A2BD1055FD54" box="[1268,1359,696,715]" name="Turkey" pageId="28" pageNumber="253">TURKEY</collectingCountry>
:
</emphasis>
Amanos [Nur Mountains], [36.74, 36.33], 
<date id="FF9E1076FFE64B05B447A2D611B3FD79" box="[1105,1193,723,742]" pageId="28" pageNumber="253" value="1987-05-24">24.V.1987</date>
, leg. Jäch (
<specimenCount id="9D26FD3FFFE64B05B513A2D61038FD79" box="[1285,1314,717,742]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B531A2D6107BFD79" box="[1319,1377,723,742]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
); 
<collectingRegion id="49E4F854FFE64B05B578A2D61083FD79" box="[1390,1433,723,742]" country="Turkey" name="Bolu" pageId="28" pageNumber="253">Bolu</collectingRegion>
, [40.72, 31.61], leg. F. Schubert (
<specimenCount id="9D26FD3FFFE64B05B447A2EB116BFC9F" box="[1105,1137,743,769]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B460A2EB11AAFC9E" box="[1142,1200,750,769]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
); Çıkrıkçı, Turgutlu, [38.47, 27.81], 
<date id="FF9E1076FFE64B05B36DA30D16CEFC84" box="[891,980,776,795]" pageId="28" pageNumber="253" value="2006-02-15">15.II.2006</date>
, leg. S.Anlaş (1 J 
<collectionCode id="ED31AE73FFE64B05B467A30D11AFFC84" box="[1137,1205,776,795]" pageId="28" pageNumber="253">NHMD</collectionCode>
); Çıkrıkçı, Turgutlu, [38.47, 27.81], 
<date id="FF9E1076FFE64B05B36FA32616CBFCA9" box="[889,977,803,822]" pageId="28" pageNumber="253" value="2006-02-17">17.II.2006</date>
, leg.S.Anlaş (1J 
<specimenCount id="9D26FD3FFFE64B05B470A326118BFCA9" box="[1126,1169,797,822]" pageId="28" pageNumber="253" type="female">3♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B480A32611C3FCA9" box="[1174,1241,803,822]" pageId="28" pageNumber="253">NHMD</collectionCode>
);Danin-Danin, 
<collectingRegion id="49E4F854FFE64B05B54CA3261087FCA9" box="[1370,1437,803,822]" country="Turkey" name="Hakkari" pageId="28" pageNumber="253">Hakkari</collectingRegion>
Prov, [37.44,42.86], 
<quantity id="4CD89B53FFE64B05B3F2A33B1139FCCE" box="[996,1059,830,849]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="253" unit="m" value="2200.0">2200 m</quantity>
, 
<date id="FF9E1076FFE64B05B43CA33B1165FCCE" box="[1066,1151,830,849]" pageId="28" pageNumber="253" value="1987-06-01">1.VI.1987</date>
, leg.Schönemann &amp; Schillhammer (2 JJ 
<specimenCount id="9D26FD3FFFE64B05B362A35D16BAFCF4" box="[884,928,850,875]" pageId="28" pageNumber="253" type="female">7♀♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B3B0A35D16FAFCF4" box="[934,992,856,875]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
); 
<collectingRegion id="49E4F854FFE64B05B3FAA35D1101FCF4" box="[1004,1051,856,875]" country="Turkey" name="Izmir" pageId="28" pageNumber="253">Izmir</collectingRegion>
, Boz daglari, Boz dag köy env., [38.34, 27.75], 
<quantity id="4CD89B53FFE64B05B32AA37616B5FC19" box="[828,943,883,902]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="28" pageNumber="253" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500-1700 m</quantity>
, 
<date id="FF9E1076FFE64B05B3AFA3761124FC19" box="[953,1086,883,902]" pageId="28" pageNumber="253" value="2003-05-30" valueMax="2003-06-03" valueMin="2003-05-30">30.V-3.VI.2003</date>
, leg. R. Lohaj (1 J 
<specimenCount id="9D26FD3FFFE64B05B4FAA3761015FC19" box="[1260,1295,877,902]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
cSch); 
<quantity id="4CD89B53FFE64B05B546A376109CFC19" box="[1360,1414,883,902]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="253" unit="km" value="50.0">50 km</quantity>
N 
<collectingRegion id="49E4F854FFE64B05B32AA38B169AFC3E" box="[828,896,910,929]" country="Turkey" name="Antalya" pageId="28" pageNumber="253">Antalya</collectingRegion>
, [37.33, 30.77], 
<date id="FF9E1076FFE64B05B413A38B117BFC3E" box="[1029,1121,910,929]" pageId="28" pageNumber="253" value="2015-04-18">18.IV.2015</date>
leg. Snizek (
<specimenCount id="9D26FD3FFFE64B05B4DBA38B11F3FC3F" box="[1229,1257,903,929]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B4F8A38B102AFC3E" box="[1262,1328,910,929]" country="Germany" httpUri="http://grbio.org/cool/3082-3vhh" name="Naturkundesmuseum" pageId="28" pageNumber="253">NMEG</collectionCode>
); Sogukoluk, Belen, 
<collectingRegion id="49E4F854FFE64B05B362A3AD16BCFC24" box="[884,934,936,955]" country="Turkey" name="Hatay" pageId="28" pageNumber="253">Hatay</collectingRegion>
, [36.49,36.14], 
<date id="FF9E1076FFE64B05B43EA3AD1164FC24" box="[1064,1150,936,955]" pageId="28" pageNumber="253" value="1987-05-24">24.V.1987</date>
, leg.Schönemann &amp;Schillhammer (
<specimenCount id="9D26FD3FFFE64B05B352A3C6167AFC49" box="[836,864,957,982]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B372A3C61687FC49" box="[868,925,963,982]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
). 
<emphasis id="B954EAA4FFE64B05B3BEA3C611C9FC49" bold="true" box="[936,1235,963,982]" pageId="28" pageNumber="253">
<collectingCountry id="F3377626FFE64B05B3BEA3C61171FC49" box="[936,1131,963,982]" name="United Kingdom" pageId="28" pageNumber="253">UNITED KINGDOM</collectingCountry>
: Sർඈඍඅൺඇൽ:
</emphasis>
Bridge of Orchy, [56.51, -4.76], 
<date id="FF9E1076FFE64B05B36DA3DB16F8FC6E" box="[891,994,990,1009]" pageId="28" pageNumber="253" value="1987-07-23">23.VII.1987</date>
, leg. Gillerfors (1 J 
<specimenCount id="9D26FD3FFFE64B05B48DA3DB11A7FC6F" box="[1179,1213,983,1009]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B4D2A3DB1019FC6E" box="[1220,1283,990,1009]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="28" pageNumber="253">MZLU</collectionCode>
); Hawock [55.42, -2.78], 
<date id="FF9E1076FFE64B05B36EA3FD16A4FB94" box="[888,958,1016,1035]" pageId="28" pageNumber="253" value="1964-06">VI.1964</date>
, leg. M. Paoletti (
<specimenCount id="9D26FD3FFFE64B05B443A3FD1169FB94" box="[1109,1139,1010,1035]" pageId="28" pageNumber="253" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFE64B05B461A3FD11ABFB94" box="[1143,1201,1016,1035]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="253">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE64B04B32AA421145FFE8A" lastPageId="29" lastPageNumber="254" pageId="28" pageNumber="253" type="description">
<paragraph id="8B9F36B6FFE64B05B32AA421108DFA04" blockId="28.[828,1437,1054,2075]" pageId="28" pageNumber="253">
<emphasis id="B954EAA4FFE64B05B32AA42116FBFBA4" bold="true" box="[828,993,1060,1083]" pageId="28" pageNumber="253">Redescription.</emphasis>
Measurements JJ (n = 6): HW = 1.78– 1.87 (1.81); HL = 1.47–1.56 (1.51); HL/HW 0.81–0.85 (0.83); PW = 2.27–2.36 (2.31); PL = 2.22–2.24 (2.23); PL/PW 0.94–0.98 (0.96); EW = 2.60–2.69 (2.63); EL = 2.49–2.60 (2.54); EL/EW 0.95–0.98 (0.96); EL/PL 1.12– 1.16 (1.14); PW/HW 1.49–1.58 (1.53); forebody length 6.20–6.33 (6.28). 
<specimenCount id="9D26FD3FFFE64B05B417A4D8113DFB63" box="[1025,1063,1245,1276]" pageId="28" pageNumber="253" type="female">♀♀</specimenCount>
(n = 4): HW = 1.80–1.87 (1.84); HL = 1.49–1.56 (1.52); HL/HW 0.81–0.86 (0.83); PW = 2.38–2.51 (2.44); PL = 2.27–2.36 (2.31); PL/PW 0.94–0.95 (0.95); EW = 2.73–2.89 (2.82); EL = 2.58–2.69 (2.64); EL/EW 0.92–0.97 (0.94); EL/PL 1.09–1.17 (1.14); PW/ HW 1.53–1.69 (1.61); forebody length 6.42–6.56 (6.47).
</paragraph>
<paragraph id="8B9F36B6FFE64B05B34CA5A1100AFA24" blockId="28.[828,1437,1054,2075]" box="[858,1296,1444,1467]" pageId="28" pageNumber="253">
Very large species; body black (
<figureCitation id="131B2A33FFE64B05B4B8A5A1101EFA24" box="[1198,1284,1444,1467]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="28" pageNumber="253">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFE64B05B34CA5C11087F9C4" blockId="28.[828,1437,1054,2075]" pageId="28" pageNumber="253">
Head black, slightly transverse, with eyes large (EyL/TL = 3.64–4.78 (4.22)) following contour of head; microsculpture of transverse waves, 4 to 6 punctures between anterior frontal punctures on frons (
<figureCitation id="131B2A33FFE64B05B444A62111BFF9A4" box="[1106,1189,1572,1595]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="28" pageNumber="253">Fig. 6D</figureCitation>
); antennae pale reddish, long, all antennomeres clearly elongate; palpi pale reddish.
</paragraph>
<paragraph id="8B9F36B6FFE64B05B34CA661115DF8E4" blockId="28.[828,1437,1054,2075]" pageId="28" pageNumber="253">Thorax: pronotum black, almost quadrate, as wide as long, clearly wider than head, microsculpture of transverse waves, three punctures in dorsal row and three in sublateral row with the most posterior one reaching the level of third puncture of dorsal row; scutellum punctured and setiferous, elytra black, densely punctured and pubescent, slightly wider than long, slightly longer than pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</paragraph>
<paragraph id="8B9F36B6FFE64B05B34CA78116B1F824" blockId="28.[828,1437,1054,2075]" pageId="28" pageNumber="253">Abdomen black, tergites uniformly punctured, slightly iridescent.</paragraph>
<paragraph id="8B9F36B6FFE64B04B34CA7C6145FFE8A" blockId="28.[828,1437,1054,2075]" lastBlockId="29.[151,761,190,1333]" lastPageId="29" lastPageNumber="254" pageId="28" pageNumber="253">
<emphasis id="B954EAA4FFE64B05B34CA7C61683F844" bold="true" box="[858,921,1987,2011]" italics="true" pageId="28" pageNumber="253">Male.</emphasis>
Aedeagus (
<figureCitation id="131B2A33FFE64B05B40DA7C11164F844" box="[1051,1150,1988,2011]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="28" pageNumber="253">Fig. 11B</figureCitation>
): Paramere slender, lanceolate, reaching apex of median lobe, sensory peg setae forming two clear rows; median lobe with protrusion at apices and two small but distinct teeth near lower level of parameres peg setae; internal sac with two small distal medial sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE74B04B081A11B17E3FDCA" pageId="29" pageNumber="254" type="diagnosis">
<paragraph id="8B9F36B6FFE74B04B081A11B17E3FDCA" blockId="29.[151,761,190,1333]" pageId="29" pageNumber="254">
<emphasis id="B954EAA4FFE74B04B081A11B1490FEAA" bold="true" box="[151,394,286,309]" pageId="29" pageNumber="254">Differential diagnosis.</emphasis>
<taxonomicName id="4C204D35FFE74B04B185A11B174AFEAA" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[403,592,286,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B185A11B174AFEAA" box="[403,592,286,309]" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
</taxonomicName>
is easily distinguished from other members of the 
<taxonomicName id="4C204D35FFE74B04B20BA13B1795FECA" authorityName="Gravenhorst" authorityYear="1802" box="[541,655,318,341]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFE74B04B20BA13B1795FECA" box="[541,655,318,341]" italics="true" pageId="29" pageNumber="254">fuliginosus</emphasis>
</taxonomicName>
-group by the punctured and setiferous scutellum and three punctures in the sublateral row of pronotum with the posteriormost one reaching the level of the third (basalmost) puncture of the dorsal row. It can be additionally distinguished from 
<taxonomicName id="4C204D35FFE74B04B2C8A1BB1412FE6A" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFE74B04B2C8A1BB1412FE6A" italics="true" pageId="29" pageNumber="254">Q. fuliginosus</emphasis>
</taxonomicName>
by the pale antennae, and from 
<taxonomicName id="4C204D35FFE74B04B251A1DB17C3FE6A" authorityName="Bernhauer" authorityYear="1908" box="[583,729,478,501]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFE74B04B251A1DB17C3FE6A" box="[583,729,478,501]" italics="true" pageId="29" pageNumber="254">Q. curtipennis</emphasis>
</taxonomicName>
by the slender paramere and median lobe with apical protrusion. 
<taxonomicName id="4C204D35FFE74B04B081A21B1457FDAA" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[151,333,542,565]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B081A21B1457FDAA" box="[151,333,542,565]" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
</taxonomicName>
is mainly found in drier habitats and rarely co-occurs with any of the other species in the species group.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE74B04B081A25B11D5FE2A" pageId="29" pageNumber="254" type="synonymic_list">
<paragraph id="8B9F36B6FFE74B04B081A25B14C7FC2A" blockId="29.[151,761,190,1333]" pageId="29" pageNumber="254">
<emphasis id="B954EAA4FFE74B04B081A25B1440FDEA" bold="true" box="[151,346,606,629]" pageId="29" pageNumber="254">Synonymic notes.</emphasis>
The name 
<taxonomicName id="4C204D35FFE74B04B1DCA25B15CCFD0A" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B954EAA4FFE74B04B1DCA25B1778FDEA" box="[458,610,606,629]" italics="true" pageId="29" pageNumber="254">Quedius tristis</emphasis>
(
<bibRefCitation id="EFB14B47FFE74B04B278A25B15D4FD0A" author="GRAVENHORST J. L. C." pageId="29" pageNumber="254" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
has long been applied to this species. GඋൺඏൾඇHඈඋඌඍ (1802) did not describe 
<taxonomicName id="4C204D35FFE74B04B19BA29B1753FD2A" authorityName="Gravenhorst" authorityYear="1802" box="[397,585,670,693]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B954EAA4FFE74B04B19BA29B1753FD2A" box="[397,585,670,693]" italics="true" pageId="29" pageNumber="254">Staphylinus tristis</emphasis>
</taxonomicName>
as a new species but attributed it to Fabricius. The Fabrician species is now in 
<taxonomicName id="4C204D35FFE74B04B0A6A2DB14BBFD6A" box="[176,417,734,757]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="subGenus" subGenus="Pseudocypus">
<emphasis id="B954EAA4FFE74B04B0A6A2DB15E5FD6A" box="[176,255,734,757]" italics="true" pageId="29" pageNumber="254">Ocypus</emphasis>
(
<emphasis id="B954EAA4FFE74B04B11BA2DB148CFD6A" box="[269,406,734,757]" italics="true" pageId="29" pageNumber="254">Pseudocypus</emphasis>
)
</taxonomicName>
. Nൾඐඍඈඇ (1995) made an appeal to the International Commission of Zoological Nomenclature requesting that the species takes the name 
<taxonomicName id="4C204D35FFE74B04B289A31B15EDFCCA" baseAuthorityName="Brulle" baseAuthorityYear="1832" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B289A31B15EDFCCA" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
</taxonomicName>
, as this was the next available name associated with this species. As a result, 
<taxonomicName id="4C204D35FFE74B04B1CAA35B15CCFC0A" authority="(Brulle, 1832)" baseAuthorityName="Brulle" baseAuthorityYear="1832" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B1CAA35B1787FCEA" box="[476,669,862,885]" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
(
<bibRefCitation id="EFB14B47FFE74B04B2BBA35B15D4FC0A" author="BRULLE G. A." pageId="29" pageNumber="254" pagination="289 - 400" refId="ref80797" refString="BRULLE G. A. 1832 - 1833: Expedition scientifique de Moree. Section des Sciences Physiques. Tome III. 1. re Partie. Zoologie. Deuxieme Section. Des animales articules. F. G. Levrault, Paris, 400 pp. [Pp. 1 - 288 published in 1832, pp. 289 - 400 in 1833.]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
)
</taxonomicName>
was accepted on the Official List of Specific Names in Zoology (
<bibRefCitation id="EFB14B47FFE74B04B10DA39B14CBFC2A" author="ICZN" box="[283,465,926,949]" pageId="29" pageNumber="254" pagination="215 - 217" refId="ref84977" refString="ICZN 1996: Opinion 1851. Xantholinini Erichson, 1839 and Quediini Kraatz, [1857] (Insecta, Coleoptera): given precedence over some senior synonyms; Quedius Stephens, 1829: Staphylinus levicollis Brulle, 1832 designated as the type species. Bulletin of Zoological Nomenclature 53: 215 - 217." type="journal article" year="1996">ICZN 1996: 215</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFE74B04B0A3A3BB1488FB4A" blockId="29.[151,761,190,1333]" pageId="29" pageNumber="254">
BൾඋඇHൺඎൾඋ &amp; SർHඎൻൾඋඍ (1916) placed 
<taxonomicName id="4C204D35FFE74B04B26FA3BB14D8FC6A" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B954EAA4FFE74B04B26FA3BB141DFC6A" italics="true" pageId="29" pageNumber="254">Staphylinus nigritarsis</emphasis>
<bibRefCitation id="EFB14B47FFE74B04B107A3DB14D8FC6A" author="MENETRIES E." box="[273,450,990,1013]" pageId="29" pageNumber="254" refId="ref87999" refString="MENETRIES E. 1832: Catalogue raisonne des objets de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse entrepris par ordre de S. M. L'Empereur. L'Academie Imperiale des Sciences, St. Petersbourg, 271 + xxxii + iv pp." type="book" year="1832">
Ménétriés, 
<quantity id="4CD89B53FFE74B04B19BA3DB14D8FC6A" box="[397,450,990,1013]" metricMagnitude="1" metricUnit="m" metricValue="4.65328" pageId="29" pageNumber="254" unit="in" value="1832.0">1832</quantity>
</bibRefCitation>
</taxonomicName>
in synonymy to 
<taxonomicName id="4C204D35FFE74B04B292A3DB15E4FB8A" baseAuthorityName="Fabricius" baseAuthorityYear="1792" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="splendens">
<emphasis id="B954EAA4FFE74B04B292A3DB15E4FB8A" italics="true" pageId="29" pageNumber="254">Philonthus splendens</emphasis>
</taxonomicName>
. Gඎඌൺඋඈඏ (1993) revised the type series of 
<taxonomicName id="4C204D35FFE74B04B2D4A3FB15F5FBAA" authorityName="Menetries" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B954EAA4FFE74B04B2D4A3FB15F5FBAA" italics="true" pageId="29" pageNumber="254">S. nigritarsis</emphasis>
</taxonomicName>
and designated a 
<typeStatus id="549B8814FFE74B04B1B5A41B171CFBAA" box="[419,518,1054,1077]" pageId="29" pageNumber="254" type="lectotype">lectotype</typeStatus>
. He found 
<taxonomicName id="4C204D35FFE74B04B265A41B17EDFBAA" authorityName="Menetries" authorityYear="1832" box="[627,759,1054,1077]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B954EAA4FFE74B04B265A41B17EDFBAA" box="[627,759,1054,1077]" italics="true" pageId="29" pageNumber="254">S. nigritarsis</emphasis>
</taxonomicName>
to be conspecific with 
<taxonomicName id="4C204D35FFE74B04B196A43B178CFBCA" authority="(Brulle, 1832)" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[384,662,1086,1109]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B196A43B14E7FBCA" box="[384,509,1086,1109]" italics="true" pageId="29" pageNumber="254">Q. levicollis</emphasis>
(
<bibRefCitation id="EFB14B47FFE74B04B21CA43B1795FBCA" author="BRULLE G. A." box="[522,655,1086,1109]" pageId="29" pageNumber="254" pagination="289 - 400" refId="ref80797" refString="BRULLE G. A. 1832 - 1833: Expedition scientifique de Moree. Section des Sciences Physiques. Tome III. 1. re Partie. Zoologie. Deuxieme Section. Des animales articules. F. G. Levrault, Paris, 400 pp. [Pp. 1 - 288 published in 1832, pp. 289 - 400 in 1833.]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
)
</taxonomicName>
based on both external characters and the shape of the aedeagus. We confirm Gusarov’s identification through examination of photos of the type from ZIN provided by M. Salnitska (see supplementary 
<figureCitation id="131B2A33FFE74B04B121A4BB1499FB4A" box="[311,387,1214,1237]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="29" pageNumber="254">Fig. S1</figureCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFE74B04B0A3A4DB11D5FE2A" blockId="29.[151,761,190,1333]" lastBlockId="29.[828,1437,190,1333]" pageId="29" pageNumber="254">
We were not able to examine the 
<typeStatus id="549B8814FFE74B04B231A4DB1744FB6A" box="[551,606,1246,1269]" pageId="29" pageNumber="254">types</typeStatus>
of 
<taxonomicName id="4C204D35FFE74B04B291A4DB14B2FA8A" authority="Nordmann, 1837" authorityName="Nordmann" authorityYear="1837" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B954EAA4FFE74B04B291A4DB15F4FA8A" italics="true" pageId="29" pageNumber="254">Philonthus frontalis</emphasis>
<bibRefCitation id="EFB14B47FFE74B04B0E0A4FB14B2FA8A" author="NORDMANN A." box="[246,424,1278,1301]" pageId="29" pageNumber="254" refId="ref88588" refString="NORDMANN A. 1837: Symbolae ad monographiam staphylinorum. Academiae Caesareae Scientiarun, Petropoli, 167 pp + 2 pls." type="book" year="1837">Nordmann, 1837</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFE74B04B1CAA4FB15D4FAAA" authority="Maran, 1935" authorityName="Maran" authorityYear="1935" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="balthasari">
<emphasis id="B954EAA4FFE74B04B1CAA4FB17BFFA8A" box="[476,677,1278,1301]" italics="true" pageId="29" pageNumber="254">Quedius balthasari</emphasis>
<bibRefCitation id="EFB14B47FFE74B04B2BBA4FB15D4FAAA" author="MARAN J." pageId="29" pageNumber="254" pagination="8 - 9" refId="ref87754" refString="MARAN J. 1935: Nove a zajimave formy rodu Quedius Steph. z ostrova Krety. De novis et raris formis generis Quedius Steph. ex insula Creta. Casopis Ceskoslovenske Spolecnosti Entomologicke 32: 8 - 9 (in Czech)." type="journal article" year="1935">Mařan, 1935</bibRefCitation>
</taxonomicName>
, both currently accepted synonyms of 
<emphasis id="B954EAA4FFE74B04B267A51B17E3FAAA" box="[625,761,1310,1333]" italics="true" pageId="29" pageNumber="254">
<taxonomicName id="4C204D35FFE74B04B267A51B17EEFAAA" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[625,756,1310,1333]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">Q. levicollis</taxonomicName>
.
</emphasis>
Based on the original descriptions, both synonymies appear to be correct. 
<taxonomicName id="4C204D35FFE74B04B3DEA0DB118CFF6A" authorityName="Nordmann" authorityYear="1837" box="[968,1174,222,245]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B954EAA4FFE74B04B3DEA0DB118CFF6A" box="[968,1174,222,245]" italics="true" pageId="29" pageNumber="254">Philonthus frontalis</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3377626FFE74B04B57AA0DB1086FF6A" box="[1388,1436,222,245]" name="Italy" pageId="29" pageNumber="254">Italy</collectingCountry>
and 
<collectingCountry id="F3377626FFE74B04B37DA0FB16AFFE8A" box="[875,949,254,277]" name="Greece" pageId="29" pageNumber="254">Greece</collectingCountry>
and fits the description of 
<taxonomicName id="4C204D35FFE74B04B4CBA0FB1044FE8A" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[1245,1374,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B4CBA0FB1044FE8A" box="[1245,1374,254,277]" italics="true" pageId="29" pageNumber="254">Q. levicollis</emphasis>
</taxonomicName>
, with interorbital punctures and punctate scutellum. 
<taxonomicName id="4C204D35FFE74B04B554A11B112DFECA" authority="Maran, 1935" authorityName="Maran" authorityYear="1935" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="balthasari">
<emphasis id="B954EAA4FFE74B04B554A11B16BDFECA" italics="true" pageId="29" pageNumber="254">Quedius balthasari</emphasis>
<bibRefCitation id="EFB14B47FFE74B04B3B8A13B112DFECA" author="MARAN J." box="[942,1079,318,341]" pageId="29" pageNumber="254" pagination="8 - 9" refId="ref87754" refString="MARAN J. 1935: Nove a zajimave formy rodu Quedius Steph. z ostrova Krety. De novis et raris formis generis Quedius Steph. ex insula Creta. Casopis Ceskoslovenske Spolecnosti Entomologicke 32: 8 - 9 (in Czech)." type="journal article" year="1935">Mařan, 1935</bibRefCitation>
</taxonomicName>
, described from central 
<collectingRegion id="49E4F854FFE74B04B522A13B1077FECA" box="[1332,1389,318,341]" country="Greece" name="Kriti" pageId="29" pageNumber="254">Crete</collectingRegion>
as a species different from 
<taxonomicName id="4C204D35FFE74B04B408A15B1183FEEA" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[1054,1177,350,373]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B408A15B1183FEEA" box="[1054,1177,350,373]" italics="true" pageId="29" pageNumber="254">Q. levicollis</emphasis>
</taxonomicName>
in sparser punctation and reddish margins of elytra is likely a color aberration because 
<taxonomicName id="4C204D35FFE74B04B32AA19B16A0FE2A" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[828,954,414,437]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B32AA19B16A0FE2A" box="[828,954,414,437]" italics="true" pageId="29" pageNumber="254">Q. levicollis</emphasis>
</taxonomicName>
is also known from 
<collectingRegion id="49E4F854FFE74B04B498A19B11D0FE2A" box="[1166,1226,414,437]" country="Greece" name="Kriti" pageId="29" pageNumber="254">Crete</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE74B04B32AA1BB11A9FC2A" pageId="29" pageNumber="254" type="description">
<paragraph id="8B9F36B6FFE74B04B32AA1BB16A2FCEA" blockId="29.[828,1437,190,1333]" pageId="29" pageNumber="254">
<emphasis id="B954EAA4FFE74B04B32AA1BB16A3FE4A" bold="true" box="[828,953,446,469]" pageId="29" pageNumber="254">Bionomics.</emphasis>
<taxonomicName id="4C204D35FFE74B04B3D6A1BB119AFE4A" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[960,1152,446,469]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B3D6A1BB119AFE4A" box="[960,1152,446,469]" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
</taxonomicName>
is a clearly thermophilous eurytopic species, which prefers sandy soils and is found in both open habitats and dry forests. It occurs in various ground debris and under rocks, e.g., it has been found in forest litter at different elevations (
<quantity id="4CD89B53FFE74B04B4A4A23B1021FDCA" box="[1202,1339,574,597]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.9" metricValueMin="0.2" pageId="29" pageNumber="254" unit="m" value="1050.0" valueMax="1900.0" valueMin="200.0">200–1900 m</quantity>
) in 
<collectingCountry id="F3377626FFE74B04B571A23B1647FDEA" name="Cyprus" pageId="29" pageNumber="254">Cyprus</collectingCountry>
(AඌඌංඇG &amp; Wඎඇൽൾඋඅൾ 2001), under rocks in 
<collectingCountry id="F3377626FFE74B04B548A25B1086FDEA" box="[1374,1436,606,629]" name="France" pageId="29" pageNumber="254">Corse</collectingCountry>
(MൺඋർHൺඅ &amp; 
<collectingCountry id="F3377626FFE74B04B3C2A27B16EBFD0A" box="[980,1009,638,661]" name="Switzerland" pageId="29" pageNumber="254">CH</collectingCountry>
ൺඋൽඈඇඇൾඍ 2000), and from sifted wet leaf-litter under oleander trees, at 
<quantity id="4CD89B53FFE74B04B4A4A29B1020FD2A" box="[1202,1338,670,693]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="29" pageNumber="254" unit="m" value="800.0" valueMax="1100.0" valueMin="500.0">500–1100 m</quantity>
in 
<collectingCountry id="F3377626FFE74B04B577A29B1086FD2A" box="[1377,1436,670,693]" name="Israel" pageId="29" pageNumber="254">Israel</collectingCountry>
(Sආൾඍൺඇൺ 1978b). We have encountered this species under rocks in open oak and chestnut forest in 
<collectingCountry id="F3377626FFE74B04B4F9A2DB1036FD6A" box="[1263,1324,734,757]" name="Spain" pageId="29" pageNumber="254">Spain</collectingCountry>
(
<figureCitation id="131B2A33FFE74B04B52DA2DB1095FD6A" box="[1339,1423,734,757]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="29" pageNumber="254">Fig. 2B</figureCitation>
), and between the roots of 
<taxonomicName id="4C204D35FFE74B04B457A2FB1166FC8A" box="[1089,1148,766,789]" class="Magnoliopsida" family="Ericaceae" genus="Erica" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="254" phylum="Tracheophyta" rank="genus">
<emphasis id="B954EAA4FFE74B04B457A2FB1166FC8A" box="[1089,1148,766,789]" italics="true" pageId="29" pageNumber="254">Erica</emphasis>
</taxonomicName>
in a heathland in 
<collectingCountry id="F3377626FFE74B04B524A2FB1083FC8A" box="[1330,1433,766,789]" name="Denmark" pageId="29" pageNumber="254">Denmark</collectingCountry>
. It has been recorded from caves in Sardinia (Bඈඋൽඈඇං 1982), although this most likely represents an accidental occurrence.
</paragraph>
<paragraph id="8B9F36B6FFE74B04B34CA37B11A9FC2A" blockId="29.[828,1437,190,1333]" pageId="29" pageNumber="254">
WൾඅඌH (1993) found this species to contain six ovaries with a fasciculata 
<typeStatus id="549B8814FFE74B04B3EBA39B1131FC2A" box="[1021,1067,926,949]" pageId="29" pageNumber="254">type</typeStatus>
of structure.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFE74B07B32AA3BB17ADFF6A" lastPageId="30" lastPageNumber="255" pageId="29" pageNumber="254" type="distribution">
<paragraph id="8B9F36B6FFE74B07B32AA3BB17ADFF6A" blockId="29.[828,1437,190,1333]" lastBlockId="30.[151,760,190,245]" lastPageId="30" lastPageNumber="255" pageId="29" pageNumber="254">
<emphasis id="B954EAA4FFE74B04B32AA3BB16C9FC4A" bold="true" box="[828,979,958,981]" pageId="29" pageNumber="254">Distribution.</emphasis>
<taxonomicName id="4C204D35FFE74B04B3C8A3BB11B2FC4A" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[990,1192,958,981]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B3C8A3BB11B2FC4A" box="[990,1192,958,981]" italics="true" pageId="29" pageNumber="254">Quedius levicollis</emphasis>
</taxonomicName>
is widely distributed across the Western Palearctic from the Atlantic coast, including British Islands through the Mediterranean, including North Africa, in the west to the Russian Black Sea coast and the Middle East (
<figureCitation id="131B2A33FFE74B04B440A43B11B9FBCA" box="[1110,1187,1086,1109]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="29" pageNumber="254">Fig. 19</figureCitation>
). In the Middle East the species extends into the Levant region, but not into the Nile delta. The south-easternmost record of 
<taxonomicName id="4C204D35FFE74B04B4C6A47B1057FB0A" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[1232,1357,1150,1173]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="254" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFE74B04B4C6A47B1057FB0A" box="[1232,1357,1150,1173]" italics="true" pageId="29" pageNumber="254">Q. levicollis</emphasis>
</taxonomicName>
is from South 
<collectingCountry id="F3377626FFE74B04B36BA49B16B0FB2A" box="[893,938,1182,1205]" name="Iran" pageId="29" pageNumber="254">Iran</collectingCountry>
. The species is found on all major Mediterranean islands, including 
<collectingCountry id="F3377626FFE74B04B414A4BB1148FB4A" box="[1026,1106,1214,1237]" name="Cyprus" pageId="29" pageNumber="254">Cyprus</collectingCountry>
, Crete, 
<collectingCountry id="F3377626FFE74B04B4BEA4BB11FCFB4A" box="[1192,1254,1214,1237]" name="France" pageId="29" pageNumber="254">Corse</collectingCountry>
and Sardinia, as well on the island of Madeira in the Atlantic Ocean. It is most common in the southern part of its distribution range gradually becoming rarer towards the north-east where it occurs at xerothermic sites reaching its northernmost limit on the Swedish island of Gotland in the Baltic Sea.
</paragraph>
</subSubSection>
</treatment>
</document>