354 lines
45 KiB
XML
354 lines
45 KiB
XML
<document id="A1239EAA77A9164A2CE0846B3EC0DFC1" ID-CLB-Dataset="217329" ID-DOI="10.37520/aemnp.2022.017" ID-GBIF-Dataset="795cf556-a4a9-477b-b55d-cb80ed8a0f2d" ID-ISSN="1804-6487" ID-Zenodo-Dep="7399702" ID-ZooBank="28D55112-98B1-49A5-B382-58B1B068570B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670240514228" checkinUser="felipe" docAuthor="Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas & Solodovnikov, Alexey" docDate="2022" docId="038987A0FFC94B2CB3CEA3A917E9F8CA" docLanguage="en" docName="ActaEntMusNatPra.62.1.225-299.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Quedius lanei Hatch 1957" docType="treatment" docVersion="7" lastPageNumber="278" masterDocId="FFB0FFD8FFFA4B19B016A005151AFF9F" masterDocTitle="Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)" masterLastPageNumber="299" masterPageNumber="225" pageNumber="276" updateTime="1699412962807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="B1506DE3BF234AEBA6C5C01944DD9938">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="6F08C5A921A650FE158656B2935AB5A0">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="43B008CE7BF8016399395BA2FF167B82">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark & Department of Bioscience, University of Aarhus, Ny Munkegade 116, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:namePart id="B8D00BD936335BA28448DE4CEA7934D1">Brunke, Adam</mods:namePart>
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<mods:affiliation id="B325CD9BA13D40953A57F56F0028E9EC">Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:affiliation id="4F460CD438FA37961EBBEB8F44A5E886">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Zoological Institute, Russian Academy of Science, Universitetskaja nab.</mods:affiliation>
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<mods:title id="2664E17A834E35F7AA2CFCB7E407A6E7">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<taxonomicName id="4C204D35FFC94B2AB3CEA3A9101AFC5B" ID-CoL="4R2YM" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[984,1280,940,964]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="276" phylum="Arthropoda" rank="species" species="lanei">
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<emphasis id="B954EAA4FFC94B2AB3CEA3A9101AFC5B" bold="true" box="[984,1280,940,964]" pageId="51" pageNumber="276">
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<emphasis id="B954EAA4FFC94B2AB3CEA3A9116BFC5B" bold="true" box="[984,1137,940,964]" italics="true" pageId="51" pageNumber="276">Quedius lanei</emphasis>
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<bibRefCitation id="EFB14B47FFC94B2AB46EA3A8101AFC5B" author="HATCH M. H." box="[1144,1280,941,964]" pageId="51" pageNumber="276" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
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</emphasis>
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<paragraph id="8B9F36B6FFC94B2AB3FEA3CE11EAFC41" blockId="51.[828,1436,971,1024]" box="[1000,1264,971,990]" pageId="51" pageNumber="276">
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(
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<figureCitation id="131B2A33FFC94B2AB3F8A3CE113BFC41" box="[1006,1057,971,990]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="51" pageNumber="276">Fig. 1</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB43DA3CE112DFC41" box="[1067,1079,971,990]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="51" pageNumber="276">4</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB457A3CE1157FC41" box="[1089,1101,971,990]" captionStart="Fig" captionStartId="15.[151,182,1229,1248]" captionTargetBox="[166,1431,196,1208]" captionTargetId="figure-13@15.[151,1436,186,1218]" captionTargetPageId="15" captionText="Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal)." figureDoi="http://doi.org/10.5281/zenodo.7399714" httpUri="https://zenodo.org/record/7399714/files/figure.png" pageId="51" pageNumber="276">5</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB440A3CE1161FC41" box="[1110,1147,971,990]" captionStart="Fig" captionStartId="20.[151,182,2030,2049]" captionTargetBox="[202,1343,201,1961]" captionTargetId="figure-13@20.[182,1406,186,1971]" captionTargetPageId="20" captionText="Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965." figureDoi="http://doi.org/10.5281/zenodo.7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="51" pageNumber="276">10B</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB492A3CE11B1FC41" box="[1156,1195,971,990]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="51" pageNumber="276">16D</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB4A2A3CE11D1FC41" box="[1204,1227,971,990]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="51" pageNumber="276">17</figureCitation>
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,
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<figureCitation id="131B2A33FFC94B2AB4C2A3CE11F0FC41" box="[1236,1258,971,990]" captionStart="Fig" captionStartId="60.[151,182,2032,2051]" captionTargetBox="[193,1394,189,2003]" captionTargetId="figure-13@60.[193,1394,189,2003]" captionTargetPageId="60" captionText="Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +." figureDoi="http://doi.org/10.5281/zenodo.7399750" httpUri="https://zenodo.org/record/7399750/files/figure.png" pageId="51" pageNumber="276">21</figureCitation>
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)
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</paragraph>
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<treatmentCitationGroup id="AB301198FFC94B2AB32AA3E81086FB9F" box="[828,1436,1005,1024]" pageId="51" pageNumber="276">
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<treatmentCitation id="0A8110A7FFC94B2AB32AA3E81151FB9F" author="HATCH M. H." box="[828,1099,1005,1024]" page="216" pageId="51" pageNumber="276" year="1957">
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<taxonomicName id="4C204D35FFC94B2AB32AA3E81151FB9F" ID-CoL="4R2YM" authority="Hatch, 1957: 216" authorityName="Hatch" authorityPageNumber="216" authorityYear="1957" box="[828,1099,1005,1024]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="276" phylum="Arthropoda" rank="species" species="lanei">
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<emphasis id="B954EAA4FFC94B2AB32AA3E816A9FB9F" box="[828,947,1005,1024]" italics="true" pageId="51" pageNumber="276">Quedius lanei</emphasis>
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<bibRefCitation id="EFB14B47FFC94B2AB3AEA3E81151FB9F" author="HATCH M. H." box="[952,1099,1005,1024]" pageId="51" pageNumber="276" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957: 216</bibRefCitation>
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[
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<typeStatus id="549B8814FFC94B2AB442A3E8119BFB9F" box="[1108,1153,1005,1024]" pageId="51" pageNumber="276">Type</typeStatus>
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locality:
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<collectingRegion id="49E4F854FFC94B2AB4C7A3E81020FB9F" box="[1233,1338,1005,1024]" country="United States of America" name="Washington" pageId="51" pageNumber="276">Washington</collectingRegion>
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: Blue Mts.]
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9F36B6FFC94B2AB32AA412108FFBDA" blockId="51.[828,1436,1047,1093]" pageId="51" pageNumber="276">
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<emphasis id="B954EAA4FFC94B2AB32AA41216B2FBB5" bold="true" box="[828,936,1047,1066]" pageId="51" pageNumber="276">References.</emphasis>
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||
KඈඋGൾ (1962b): 333 (characters and notes); Sආൾඍൺඇൺ (1971a): 144 (characters and distribution); (1976a): 174 (distribution).
|
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</paragraph>
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<subSubSection id="C33A653DFFC94B2AB32AA4591102FA7B" pageId="51" pageNumber="276" type="materials_examined">
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<paragraph id="8B9F36B6FFC94B2AB32AA4591102FA7B" blockId="51.[828,1438,1116,1508]" pageId="51" pageNumber="276">
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<emphasis id="B954EAA4FFC94B2AB32AA45911BFFBF0" bold="true" box="[828,1189,1116,1135]" pageId="51" pageNumber="276">
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Material examined.
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<collectingCountry id="F3377626FFC94B2AB3EFA459113DFBF0" box="[1017,1063,1116,1135]" name="United States of America" pageId="51" pageNumber="276">USA</collectingCountry>
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||
: Cൺඅංൿඈඋඇංൺ:
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||
</emphasis>
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||
Fresno County, Huntington Lake, [37.24, -119.18],
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||
<date id="FF9E1076FFC94B2AB41FA4731170FB16" box="[1033,1130,1142,1161]" pageId="51" pageNumber="276" value="1960-04-25">25.IV.1960</date>
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, leg. L. G. Ingles (2
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||
<collectionCode id="ED31AE73FFC94B2AB537A473104BFB16" box="[1313,1361,1142,1161]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
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); Lassen National Park,Summit Lake, [40.49, -121.42],
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<quantity id="4CD89B53FFC94B2AB4A8A49411E7FB3B" box="[1214,1277,1169,1188]" metricMagnitude="3" metricUnit="m" metricValue="2.042" pageId="51" pageNumber="276" unit="m" value="2042.0">2042 m</quantity>
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,
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||
<date id="FF9E1076FFC94B2AB512A4941076FB3B" box="[1284,1388,1169,1188]" pageId="51" pageNumber="276" value="1979-07-15">15.VII.1979</date>
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, J. M. & B.A. Campbell (3
|
||
<collectionCode id="ED31AE73FFC94B2AB3F8A4A91104FB20" box="[1006,1054,1196,1215]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFC94B2AB43DA4A9116DFB20" bold="true" box="[1067,1143,1196,1215]" pageId="51" pageNumber="276">Nൾඏൺൽൺ:</emphasis>
|
||
Lake Tahoe, Zephyr Cove, [39.01, -119.94],
|
||
<quantity id="4CD89B53FFC94B2AB39BA4C316D4FB46" box="[909,974,1222,1241]" metricMagnitude="3" metricUnit="m" metricValue="1.889" pageId="51" pageNumber="276" unit="m" value="1889.0">1889 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB3C3A4C3112EFB46" box="[981,1076,1222,1241]" pageId="51" pageNumber="276" value="1986-12-09">9.XII.1986</date>
|
||
, leg.A.Smetana (1
|
||
<collectionCode id="ED31AE73FFC94B2AB4CFA4C31012FB46" box="[1241,1288,1222,1241]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFC94B2AB503A4C31072FB46" bold="true" box="[1301,1384,1222,1241]" pageId="51" pageNumber="276">OඋൾGඈඇ:</emphasis>
|
||
<quantity id="4CD89B53FFC94B2AB57AA4C31087FB46" box="[1388,1437,1222,1241]" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" pageId="51" pageNumber="276" unit="mi" value="16.0">16 mi</quantity>
|
||
NE Bly Deming Creek Road, [42.55, -120.76],
|
||
<quantity id="4CD89B53FFC94B2AB4D2A4E4101FFB6B" box="[1220,1285,1249,1268]" metricMagnitude="3" metricUnit="m" metricValue="1.828" pageId="51" pageNumber="276" unit="m" value="1828.0">1828 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB51AA4E4106CFB6B" box="[1292,1398,1249,1268]" pageId="51" pageNumber="276" value="1979-07-21">21.VII.1979</date>
|
||
, leg. J. M. Campbell & J. Schuh (3
|
||
<collectionCode id="ED31AE73FFC94B2AB456A4F9116AFA90" box="[1088,1136,1276,1295]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
); Grant County, Strawberry Range, Strawberry Lake, [44.3, -118.68],
|
||
<quantity id="4CD89B53FFC94B2AB475A51311BDFAB6" box="[1123,1191,1302,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="51" pageNumber="276" unit="m" value="1920.0">1920 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB4A7A5131013FAB6" box="[1201,1289,1302,1321]" pageId="51" pageNumber="276" value="1989-06-04">4.VI.1989</date>
|
||
, leg. A. Smetana (20
|
||
<collectionCode id="ED31AE73FFC94B2AB34BA5341697FADB" box="[861,909,1329,1348]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
); Klamath County, Gearhart Mountain, [42.51, -120.86],
|
||
<quantity id="4CD89B53FFC94B2AB564A5341655FAC0" metricMagnitude="3" metricUnit="m" metricValue="2.194" pageId="51" pageNumber="276" unit="m" value="2194.0">2194 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB340A54916A3FAC0" box="[854,953,1356,1375]" pageId="51" pageNumber="276" value="1974-06-24">24.VI.1974</date>
|
||
, leg.A. & D. Smetana (1
|
||
<collectionCode id="ED31AE73FFC94B2AB499A54911A5FAC0" box="[1167,1215,1356,1375]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
); Klamath County, Mare’s Egg Springs,
|
||
<quantity id="4CD89B53FFC94B2AB3A2A56316F7FAE6" box="[948,1005,1382,1401]" metricMagnitude="4" metricUnit="m" metricValue="1.17482112" pageId="51" pageNumber="276" unit="mi" value="7.3">7.3 mi</quantity>
|
||
SW Fort Klamath, [42.66, -122.08],
|
||
<date id="FF9E1076FFC94B2AB520A5631083FAE6" box="[1334,1433,1382,1401]" pageId="51" pageNumber="276" value="1974-06-25">25.VI.1974</date>
|
||
, leg. A. & D. Smetana (2
|
||
<collectionCode id="ED31AE73FFC94B2AB41AA5841126FA0B" box="[1036,1084,1409,1428]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
); Union County, Blue Mountain Road 62, Jarboe Creek, [45.80, -117.85],
|
||
<quantity id="4CD89B53FFC94B2AB45BA599118BFA30" box="[1101,1169,1436,1455]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="51" pageNumber="276" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB48CA59911E9FA30" box="[1178,1267,1436,1455]" pageId="51" pageNumber="276" value="1989-05-29">29.V.1989</date>
|
||
, leg. A. Smetana (1
|
||
<collectionCode id="ED31AE73FFC94B2AB32AA5B31671FA56" box="[828,875,1462,1481]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFC94B2AB36EA5B316EEFA56" bold="true" box="[888,1012,1462,1481]" pageId="51" pageNumber="276">WൺඌHංඇGඍඈඇ:</emphasis>
|
||
Blue Mountain, [45.5, -118],
|
||
<quantity id="4CD89B53FFC94B2AB4FDA5B3103BFA56" box="[1259,1313,1462,1481]" metricMagnitude="2" metricUnit="m" metricValue="9.14" pageId="51" pageNumber="276" unit="m" value="914.0">914 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFC94B2AB53EA5B31083FA56" box="[1320,1433,1462,1481]" pageId="51" pageNumber="276" value="1923-08-29">29.VIII.1923</date>
|
||
, leg. M. C. Lane (1
|
||
<collectionCode id="ED31AE73FFC94B2AB3C8A5D41114FA7B" box="[990,1038,1489,1508]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="276">CNC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFC94B2CB32AA5F8171BFE4A" lastPageId="53" lastPageNumber="278" pageId="51" pageNumber="276" type="description">
|
||
<paragraph id="8B9F36B6FFC94B2AB32AA5F8108DF8EB" blockId="51.[828,1437,1527,2068]" pageId="51" pageNumber="276">
|
||
<emphasis id="B954EAA4FFC94B2AB32AA5F816FBF98B" bold="true" box="[828,993,1533,1556]" pageId="51" pageNumber="276">Redescription.</emphasis>
|
||
Measurements JJ (n = 3): HW = 1.56– 1.78 (1.63); HL = 1.33–1.49 (1.39); HL/HW 0.84–0.87 (0.86); PW = 2.04–2.22 (2.11); PL = 1.80–2.00 (1.87); PL/PW 0.88–0.90 (0.89); EW = 2.16–2.29 (2.20); EL = 2.02–2.11 (2.06); EL/EW 0.92–0.95 (0.94); EL/PL 1.06– 1.14 (1.10); PW/HW 1.49–1.53 (1.52); forebody length 5.18–5.60 (5.33).
|
||
<specimenCount id="9D26FD3FFFC94B2AB417A6B0113DF94B" box="[1025,1063,1717,1748]" pageId="51" pageNumber="276" type="female">♀♀</specimenCount>
|
||
(n = 6): HW = 1.53–1.73 (1.64); HL = 1.27–1.40 (1.35); HL/HW 0.77–0.86 (0.82); PW = 1.96–2.29 (2.09); PL = 1.73–2.00 (1.85); PL/PW 0.87–0.91 (0.89); EW = 2.00–2.44 (2.18); EL = 1.93–2.29 (2.07); EL/EW 0.92–1.02 (0.95); EL/PL 1.09–1.18 (1.12); PW/ HW 1.46–1.63 (1.55); forebody length 4.98–5.69 (5.27).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFC94B2AB34CA7781662F82B" blockId="51.[828,1437,1527,2068]" pageId="51" pageNumber="276">
|
||
Medium sized species; body dark brown to black (
|
||
<figureCitation id="131B2A33FFC94B2AB567A7781671F82B" captionStart="Fig" captionStartId="20.[151,182,2030,2049]" captionTargetBox="[202,1343,201,1961]" captionTargetId="figure-13@20.[182,1406,186,1971]" captionTargetPageId="20" captionText="Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965." figureDoi="http://doi.org/10.5281/zenodo.7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="51" pageNumber="276">Fig. 10B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFC94B2DB34CA7B81411F8E4" blockId="51.[828,1437,1527,2068]" lastBlockId="52.[151,760,1796,2075]" lastPageId="52" lastPageNumber="277" pageId="51" pageNumber="276">
|
||
Head black, distinctly transverse with eyes medium sized (EyL/TL = 1.81–2.14 (1.99)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (
|
||
<figureCitation id="131B2A33FFCE4B2DB231A701179BF884" box="[551,641,1796,1819]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="52" pageNumber="277">Fig. 6G</figureCitation>
|
||
); antennae and palpi dark with joints pale reddish, all antennomeres elongate, except last and penultimate antennomere almost transverse.
|
||
</paragraph>
|
||
<caption id="DF5F663EFFCE4B2DB081A6871192F930" ID-DOI="http://doi.org/10.5281/zenodo.7399742" ID-Zenodo-Dep="7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="52" pageNumber="277" startId="52.[151,182,1666,1685]" targetBox="[170,1418,189,1655]" targetPageId="52">
|
||
<paragraph id="8B9F36B6FFCE4B2DB081A6871192F930" blockId="52.[151,1437,1666,1711]" pageId="52" pageNumber="277">
|
||
Fig. 17. Variation in genitalic morphology of species from
|
||
<taxonomicName id="4C204D35FFCE4B2DB269A687165CF90A" authorityName="Smetana" authorityYear="1965" box="[639,838,1666,1685]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="277" phylum="Arthropoda" rank="species" species="molochinoides">
|
||
<emphasis id="B954EAA4FFCE4B2DB269A687165CF90A" box="[639,838,1666,1685]" italics="true" pageId="52" pageNumber="277">Quedius molochinoides</emphasis>
|
||
</taxonomicName>
|
||
-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36B6FFCE4B2DB0A3A781115BF844" blockId="52.[151,760,1796,2075]" lastBlockId="52.[828,1437,1796,2075]" pageId="52" pageNumber="277">Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra variable in coloration, from all black or reddish brown to occasionally dark with reddish sutural margins, uniformly pubescent, punctures clearly separated, quadrate or slightly wider than long, slightly longer than pronotum; legs reddish brown with inner face of tibia and inner surface of femur darkened and tarsi paler.</paragraph>
|
||
<paragraph id="8B9F36B6FFCE4B2DB34CA7E1116AF784" blockId="52.[828,1437,1796,2075]" pageId="52" pageNumber="277">Abdomen dark brown to black, tergites uniformly punctured, with clear iridescence.</paragraph>
|
||
<paragraph id="8B9F36B6FFCF4B2CB0A3A0B8171BFE4A" blockId="53.[151,760,189,1877]" pageId="53" pageNumber="278">
|
||
<emphasis id="B954EAA4FFCF4B2CB0A3A0B815EEFF4A" bold="true" box="[181,244,189,213]" italics="true" pageId="53" pageNumber="278">Male.</emphasis>
|
||
Aedeagus (
|
||
<figureCitation id="131B2A33FFCF4B2CB162A0BB14C1FF4A" box="[372,475,190,213]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="53" pageNumber="278">Figs 16D</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A33FFCF4B2CB1FEA0BB171CFF4A" box="[488,518,190,213]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="53" pageNumber="278">17</figureCitation>
|
||
): paramere lanceolate, rather broad, without clear basal attenuation, extending to apex of median lobe, with small sensory peg setae forming two irregular rows meeting in an irregular cluster below apex; median lobe lanceolate, broad, with sides slightly constricting until abrupt constriction near pointed apex, on parameral side with two blunt teeth simply protruding inward and slightly basad; internal sac with two larger oval and two smaller rounder sclerites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFCF4B2CB081A1DB171DFC0A" pageId="53" pageNumber="278" type="diagnosis">
|
||
<paragraph id="8B9F36B6FFCF4B2CB081A1DB171DFC0A" blockId="53.[151,760,189,1877]" pageId="53" pageNumber="278">
|
||
<emphasis id="B954EAA4FFCF4B2CB081A1DB1490FE6A" bold="true" box="[151,394,478,501]" pageId="53" pageNumber="278">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C204D35FFCF4B2CB198A1DB173AFE6A" authorityName="Hatch" authorityYear="1957" box="[398,544,478,501]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB198A1DB173AFE6A" box="[398,544,478,501]" italics="true" pageId="53" pageNumber="278">Quedius lanei</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C204D35FFCF4B2CB0C4A1FB144AFD8A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[210,336,510,533]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFCF4B2CB0C4A1FB144AFD8A" box="[210,336,510,533]" italics="true" pageId="53" pageNumber="278">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D35FFCF4B2CB192A1FB14E4FD8A" authorityName="Casey" authorityYear="1915" box="[388,510,510,533]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFCF4B2CB192A1FB14E4FD8A" box="[388,510,510,533]" italics="true" pageId="53" pageNumber="278">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from
|
||
<taxonomicName id="4C204D35FFCF4B2CB255A23B17E2FDCA" authorityName="Smetana" authorityYear="1965" box="[579,760,574,597]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="labradorensis">
|
||
<emphasis id="B954EAA4FFCF4B2CB255A23B17E2FDCA" box="[579,760,574,597]" italics="true" pageId="53" pageNumber="278">Q. labradorensis</emphasis>
|
||
</taxonomicName>
|
||
by the darker body and the clearly darkened basal three antennomeres. It can be distinguished from other members of the
|
||
<taxonomicName id="4C204D35FFCF4B2CB0CAA29B1462FD2A" authorityName="Smetana" authorityYear="1965" box="[220,376,670,693]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="molochinoides">
|
||
<emphasis id="B954EAA4FFCF4B2CB0CAA29B1462FD2A" box="[220,376,670,693]" italics="true" pageId="53" pageNumber="278">molochinoides</emphasis>
|
||
</taxonomicName>
|
||
-group (
|
||
<emphasis id="B954EAA4FFCF4B2CB1D8A29B17B3FD2A" box="[462,681,670,693]" italics="true" pageId="53" pageNumber="278">
|
||
<taxonomicName id="4C204D35FFCF4B2CB1D8A29B1727FD2A" authorityName="Hansen & Brunke & Simonsen & Solodovnikov" authorityYear="2022" box="[462,573,670,693]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="altanai">Q. altanai</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D35FFCF4B2CB25EA29B17BFFD2A" authorityName="Hatch" authorityYear="1957" box="[584,677,670,693]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="horni">Q. horni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C204D35FFCF4B2CB2C8A29B142EFD4A" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="molochinoides">
|
||
<emphasis id="B954EAA4FFCF4B2CB2C8A29B142EFD4A" italics="true" pageId="53" pageNumber="278">Q. molochinoides</emphasis>
|
||
</taxonomicName>
|
||
) only by the male genitalia, specifically by the paramere, which is parallel-sided to constricted at base, and the broad median lobe with a constricted apex and two blunt teeth pointing in and downwards. However,
|
||
<taxonomicName id="4C204D35FFCF4B2CB081A33B15F6FCCA" authorityName="Hatch" authorityYear="1957" box="[151,236,830,853]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB081A33B15F6FCCA" box="[151,236,830,853]" italics="true" pageId="53" pageNumber="278">Q. lanei</emphasis>
|
||
</taxonomicName>
|
||
does not seem to co-occur with any other species of the
|
||
<emphasis id="B954EAA4FFCF4B2CB0CFA35B1467FCEA" box="[217,381,862,885]" italics="true" pageId="53" pageNumber="278">molochinoides-</emphasis>
|
||
group, this fact reducing the need of genitalia investigation in all cases.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFCF4B2CB081A39B17FEFBCA" pageId="53" pageNumber="278" type="discussion">
|
||
<paragraph id="8B9F36B6FFCF4B2CB081A39B17FEFBCA" blockId="53.[151,760,189,1877]" pageId="53" pageNumber="278">
|
||
<emphasis id="B954EAA4FFCF4B2CB081A39B1403FC2A" bold="true" box="[151,281,926,949]" pageId="53" pageNumber="278">Comments.</emphasis>
|
||
A series of specimens in Sආൾඍൺඇൺ (1971) including one Hatch
|
||
<typeStatus id="549B8814FFCF4B2CB158A3BB14B3FC4A" box="[334,425,958,981]" pageId="53" pageNumber="278" type="paratype">paratype</typeStatus>
|
||
from Fish Lake in Summerland,
|
||
<collectingRegion id="49E4F854FFCF4B2CB081A3DB1454FC6A" box="[151,334,990,1013]" country="Canada" name="British Columbia" pageId="53" pageNumber="278">British Columbia</collectingRegion>
|
||
ascribed to
|
||
<taxonomicName id="4C204D35FFCF4B2CB1DCA3DB1704FC6A" authorityName="Hatch" authorityYear="1957" box="[458,542,990,1013]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB1DCA3DB1704FC6A" box="[458,542,990,1013]" italics="true" pageId="53" pageNumber="278">Q. lanei</emphasis>
|
||
</taxonomicName>
|
||
was found to belong to
|
||
<taxonomicName id="4C204D35FFCF4B2CB0A6A3FB147CFB8A" authorityName="Smetana" authorityYear="1965" box="[176,358,1022,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="molochinoides">
|
||
<emphasis id="B954EAA4FFCF4B2CB0A6A3FB147CFB8A" box="[176,358,1022,1045]" italics="true" pageId="53" pageNumber="278">Q. molochinoides</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C204D35FFCF4B2CB164A3FB1718FB8A" authorityName="Hatch" authorityYear="1957" box="[370,514,1022,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB164A3FB1718FB8A" box="[370,514,1022,1045]" italics="true" pageId="53" pageNumber="278">Quedius lanei</emphasis>
|
||
</taxonomicName>
|
||
does not occur in
|
||
<collectingCountry id="F3377626FFCF4B2CB2AFA3FB15A9FBAA" name="Canada" pageId="53" pageNumber="278">Canada</collectingCountry>
|
||
.
|
||
<typeStatus id="549B8814FFCF4B2CB0AAA41B143EFBAA" box="[188,292,1054,1077]" pageId="53" pageNumber="278" type="paratype">Paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C204D35FFCF4B2CB152A41B1482FBAA" authorityName="Hatch" authorityYear="1957" box="[324,408,1054,1077]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB152A41B1482FBAA" box="[324,408,1054,1077]" italics="true" pageId="53" pageNumber="278">Q. lanei</emphasis>
|
||
</taxonomicName>
|
||
from Idaho were found to belong to
|
||
<taxonomicName id="4C204D35FFCF4B2CB0A5A43B1405FBCA" authorityName="Hansen & Brunke & Simonsen & Solodovnikov" authorityYear="2022" box="[179,287,1086,1109]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="altanai">
|
||
<emphasis id="B954EAA4FFCF4B2CB0A5A43B1405FBCA" box="[179,287,1086,1109]" italics="true" pageId="53" pageNumber="278">Q. altanai</emphasis>
|
||
</taxonomicName>
|
||
and have thus been moved to this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFCF4B2CB081A45B1701FA2A" pageId="53" pageNumber="278" type="description">
|
||
<paragraph id="8B9F36B6FFCF4B2CB081A45B1701FA2A" blockId="53.[151,760,189,1877]" pageId="53" pageNumber="278">
|
||
<emphasis id="B954EAA4FFCF4B2CB081A45B140FFBEA" bold="true" box="[151,277,1118,1141]" pageId="53" pageNumber="278">Bionomics.</emphasis>
|
||
<taxonomicName id="4C204D35FFCF4B2CB10CA45B14B4FBEA" authorityName="Hatch" authorityYear="1957" box="[282,430,1118,1141]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB10CA45B14B4FBEA" box="[282,430,1118,1141]" italics="true" pageId="53" pageNumber="278">Quedius lanei</emphasis>
|
||
</taxonomicName>
|
||
prefers mountainous areas and occurs only at high elevations with records from roughly
|
||
<quantity id="4CD89B53FFCF4B2CB081A49B15FDFB2A" box="[151,231,1182,1205]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="278" unit="m" value="1000.0">1000 m</quantity>
|
||
in the northern part of its range to up to
|
||
<quantity id="4CD89B53FFCF4B2CB298A49B17C4FB2A" box="[654,734,1182,1205]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="53" pageNumber="278" unit="m" value="2200.0">2200 m</quantity>
|
||
in the southern part. Little is known about the prefered habitat of the species, but based on the examined records it seems to be found mainly in wet debris near creeks or mountain lakes. For example, the
|
||
<collectingRegion id="49E4F854FFCF4B2CB183A51B14FFFAAA" box="[405,485,1310,1333]" country="United States of America" name="Oregon" pageId="53" pageNumber="278">Oregon</collectingRegion>
|
||
specimens from Klamath County were collected by sifting wet moss and deciduous leaf litter (willow) at the edges of fast-running creeks or by sifting wet debris on the bottom of a dried-out pond in a coniferous forest (Sආൾඍൺඇൺ 1976).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFCF4B2CB081A5BB17E9F8CA" pageId="53" pageNumber="278" type="distribution">
|
||
<paragraph id="8B9F36B6FFCF4B2CB081A5BB17E9F8CA" blockId="53.[151,760,189,1877]" pageId="53" pageNumber="278">
|
||
<emphasis id="B954EAA4FFCF4B2CB081A5BB1435FA4A" bold="true" box="[151,303,1470,1493]" pageId="53" pageNumber="278">Distribution.</emphasis>
|
||
<taxonomicName id="4C204D35FFCF4B2CB12FA5BB14CDFA4A" authorityName="Hatch" authorityYear="1957" box="[313,471,1470,1493]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB12FA5BB14CDFA4A" box="[313,471,1470,1493]" italics="true" pageId="53" pageNumber="278">Quedius lanei</emphasis>
|
||
</taxonomicName>
|
||
is currently known from high elevation sites on the eastern slopes and foothills of Cascades, the Sierra
|
||
<collectingRegion id="49E4F854FFCF4B2CB18FA5FB14F0F98A" box="[409,490,1534,1557]" country="United States of America" name="Nevada" pageId="53" pageNumber="278">Nevada</collectingRegion>
|
||
and the Blue Mountains (ecoregions 6.2.8, 6.2.9 and 6.2.12,
|
||
<figureCitation id="131B2A33FFCF4B2CB236A61B176BF9AA" box="[544,625,1566,1589]" captionStart="Fig" captionStartId="60.[151,182,2032,2051]" captionTargetBox="[193,1394,189,2003]" captionTargetId="figure-13@60.[193,1394,189,2003]" captionTargetPageId="60" captionText="Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +." figureDoi="http://doi.org/10.5281/zenodo.7399750" httpUri="https://zenodo.org/record/7399750/files/figure.png" pageId="53" pageNumber="278">Fig. 21</figureCitation>
|
||
). It is found from sutheastern
|
||
<collectingRegion id="49E4F854FFCF4B2CB159A63B14D4F9CA" box="[335,462,1598,1621]" country="United States of America" name="Washington" pageId="53" pageNumber="278">Washington</collectingRegion>
|
||
through
|
||
<collectingRegion id="49E4F854FFCF4B2CB238A63B1764F9CA" box="[558,638,1598,1621]" country="United States of America" name="Oregon" pageId="53" pageNumber="278">Oregon</collectingRegion>
|
||
to much of eastern
|
||
<collectingRegion id="49E4F854FFCF4B2CB0FEA65B144EF9EA" box="[232,340,1630,1653]" country="United States of America" name="California" pageId="53" pageNumber="278">California</collectingRegion>
|
||
and limited areas of western
|
||
<collectingRegion id="49E4F854FFCF4B2CB292A65B17C3F9EA" box="[644,729,1630,1653]" country="United States of America" name="Nevada" pageId="53" pageNumber="278">Nevada</collectingRegion>
|
||
. In the north it does not seem to extend into the northern part of the Cascades, although based on the ecoregions it may be be present quite far north on the eastern slopes. To the northwest it is replaced by
|
||
<taxonomicName id="4C204D35FFCF4B2CB1D4A6DB1705F96A" authorityName="Hatch" authorityYear="1957" box="[450,543,1758,1781]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="horni">
|
||
<emphasis id="B954EAA4FFCF4B2CB1D4A6DB1705F96A" box="[450,543,1758,1781]" italics="true" pageId="53" pageNumber="278">Q. horni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C204D35FFCF4B2CB238A6DB17DFF96A" authorityName="Hatch" authorityYear="1957" box="[558,709,1758,1781]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="lanei">
|
||
<emphasis id="B954EAA4FFCF4B2CB238A6DB17DFF96A" box="[558,709,1758,1781]" italics="true" pageId="53" pageNumber="278">Quedius lanei</emphasis>
|
||
</taxonomicName>
|
||
was formerly considered to also include specimens from central and southern Rockies, but we here show that these belong to a different species
|
||
<taxonomicName id="4C204D35FFCF4B2CB16CA73B14FCF8CA" authorityName="Hansen & Brunke & Simonsen & Solodovnikov" authorityYear="2022" box="[378,486,1854,1877]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="278" phylum="Arthropoda" rank="species" species="altanai">
|
||
<emphasis id="B954EAA4FFCF4B2CB16CA73B14FCF8CA" box="[378,486,1854,1877]" italics="true" pageId="53" pageNumber="278">Q. altanai</emphasis>
|
||
</taxonomicName>
|
||
described as new herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |