1630 lines
352 KiB
XML
1630 lines
352 KiB
XML
<document id="326F6E10A1782E2005490D31EFD438E6" ID-DOI="10.5281/zenodo.178296" ID-GBIF-Dataset="0366c4ab-3941-4376-806f-6cf6e3d8c403" ID-ISSN="1175-5326" ID-Zenodo-Dep="178296" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459889844187" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Manuel-Santos, Marivene R." docDate="2007" docId="03898795FFC6FFACEEA2FD9B9FD8F813" docLanguage="en" docName="zt01558p068.pdf" docOrigin="Zootaxa 1558" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Vultocinidae" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="FFB0FFEDFFC7FFA4EE35FFA49A55FF94" masterDocTitle="Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae" masterLastPageNumber="68" masterPageNumber="39" pageNumber="40" updateTime="1698227947623" updateUser="plazi">
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<mods:titleInfo id="3A5CD786FE13710E25D30294C7BF1D8C">
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<mods:title id="8D292003CB773FE77C91AA42DA330EF1">Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae</mods:title>
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<mods:namePart id="51705471926C10293AAF447F01B82474">Ng, Peter K. L.</mods:namePart>
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<mods:name id="EBF5F738FEC2466D30D76EAD6D49B5E1" type="personal">
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<mods:namePart id="7F88422CE91BB567FED8CC0B310FD9E5">Manuel-Santos, Marivene R.</mods:namePart>
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<mods:typeOfResource id="71648D3B8161E46AAA052A7C317BBE87">text</mods:typeOfResource>
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<mods:title id="D781C7DA177C2BEA44C4438DB178AB74">Zootaxa</mods:title>
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<mods:part id="352FACA5B92DAE996D0C57D9B1777073">
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<mods:date id="7795015C9CAB944D8D7D656E1BB2B545">2007</mods:date>
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<mods:number id="99A8D49805CA56F6735FA00FDF8EE0AF">1558</mods:number>
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<mods:classification id="B667A3CC60F2BDE8F045AFF3EC50B9C4">journal article</mods:classification>
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<mods:identifier id="28082AFB70CC70E6A802A2F7B6A341B3" type="DOI">10.5281/zenodo.178296</mods:identifier>
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<mods:identifier id="B55968299F229F9844788DE70054C17E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F31C49F83BF5825A783E62DEEB2F65C6" type="Zenodo-Dep">178296</mods:identifier>
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<treatment id="03898795FFC6FFACEEA2FD9B9FD8F813" ID-DOI="http://doi.org/10.5281/zenodo.6250886" ID-GBIF-Taxon="119371809" ID-Zenodo-Dep="6250886" LSID="urn:lsid:plazi:treatment:03898795FFC6FFACEEA2FD9B9FD8F813" httpUri="http://treatment.plazi.org/id/03898795FFC6FFACEEA2FD9B9FD8F813" lastPageId="8" lastPageNumber="47" pageId="1" pageNumber="40">
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<subSubSection id="C33A6508FFC6FFA5EEA2FD9B9B9EFDCD" box="[151,459,575,601]" pageId="1" pageNumber="40" type="nomenclature">
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<paragraph id="8B9F3683FFC6FFA5EEA2FD9B9B9EFDCD" blockId="1.[151,459,575,601]" box="[151,459,575,601]" pageId="1" pageNumber="40">
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<heading id="D0D781EFFFC6FFA5EEA2FD9B9B9EFDCD" bold="true" box="[151,459,575,601]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
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<emphasis id="B954EA91FFC6FFA5EEA2FD9B9B9EFDCD" bold="true" box="[151,459,575,601]" pageId="1" pageNumber="40">
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<taxonomicName id="4C204D00FFC6FFA5EEA2FD9B9B63FDCD" ID-CoL="7NLK6" box="[151,310,575,601]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
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, new family
|
||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A6508FFC6FFA5EEA2FD2E9997FAB0" pageId="1" pageNumber="40" type="diagnosis">
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<paragraph id="8B9F3683FFC6FFA5EEA2FD2E9997FAB0" blockId="1.[151,1437,650,2036]" pageId="1" pageNumber="40">
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<emphasis id="B954EA91FFC6FFA5EEA2FD2E9B45FD30" bold="true" box="[151,272,650,676]" pageId="1" pageNumber="40">Diagnosis</emphasis>
|
||
. Dorsal surface of carapace subquadrate, with complex pattern of ridges, grooves, dense short setae (
|
||
<figureCitation id="131B2A06FFC6FFA5EEAAFD169AA3FD58" box="[159,246,690,716]" captionStart="FIGURE 2" captionStartId="18.[151,255,1936,1960]" captionTargetBox="[264,1322,182,1924]" captionTargetId="figure@18.[264,1323,182,1926]" captionTargetPageId="18" captionText="FIGURE 2. Vultocinus anfractus, new genus, new species. Overall dorsal views. A, holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines; B, paratype male (cl 19.9 mm, cw 22.0 mm) (ZRC), Philippines; C, male (cl 15.6 mm, cw 16.8 mm) (ZRC), Vanuatu." httpUri="https://zenodo.org/record/178298/files/figure.png" pageId="1" pageNumber="40">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A06FFC6FFA5EF30FD169B46FD58" box="[261,275,690,716]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="1" pageNumber="40">3</figureCitation>
|
||
A). Mandibles prominently dark coloured even after prolonged preservation (
|
||
<figureCitation id="131B2A06FFC6FFA5EA8EFD169F57FD58" box="[1211,1282,690,716]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="1" pageNumber="40">Fig. 8</figureCitation>
|
||
A). Thoracic sternum relatively narrow transversely (
|
||
<figureCitation id="131B2A06FFC6FFA5EC47FD7E98EFFD60" box="[626,698,730,756]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="1" pageNumber="40">Fig. 9</figureCitation>
|
||
A); episternites 4–6 not prominently expanded laterally (
|
||
<figureCitation id="131B2A06FFC6FFA5EB54FD7E9AF3FC88" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="1" pageNumber="40">Figs. 9</figureCitation>
|
||
A, 10A, 12A); sternite 7 very narrow longitudinally, tapering abruptly (
|
||
<figureCitation id="131B2A06FFC6FFA5EDDBFCA69E16FC88" box="[1006,1091,770,796]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="1" pageNumber="40">Fig. 10</figureCitation>
|
||
A), posterior part of episternite 7 prominently prolonged backwards to form long spur which covers penis and just touches coxa of P5 (
|
||
<figureCitation id="131B2A06FFC6FFA5EEAAFCF69AAAFCF8" box="[159,255,850,876]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="1" pageNumber="40">Figs. 13</figureCitation>
|
||
A, B); sutures 4/5 and 5/6 prominently interrupted medially (
|
||
<figureCitation id="131B2A06FFC6FFA5EDFAFCF69E70FCF8" box="[975,1061,850,876]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="1" pageNumber="40">Fig. 12</figureCitation>
|
||
A); suture 6/7 appears complete but with median portion very shallow but discernible (
|
||
<figureCitation id="131B2A06FFC6FFA5ED28FCDE9926FC00" box="[797,883,890,916]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="1" pageNumber="40">Fig. 12</figureCitation>
|
||
A); suture 7/8 distinctly complete with median part deep (
|
||
<figureCitation id="131B2A06FFC6FFA5EF26FC069B32FC28" box="[275,359,930,956]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="1" pageNumber="40">Fig. 12</figureCitation>
|
||
A); sternite 8 not exposed when abdomen closed (
|
||
<figureCitation id="131B2A06FFC6FFA5ED85FC069E51FC28" box="[944,1028,930,956]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="1" pageNumber="40">Fig. 10</figureCitation>
|
||
A). Ventral margins of meri of P2– P4 with 2 or 3 strong perpendicular spines (
|
||
<figureCitation id="131B2A06FFC6FFA5ECA6FC6E988FFC70" box="[659,730,970,996]" captionStart="FIGURE 4" captionStartId="20.[151,255,1144,1168]" captionTargetBox="[151,1435,184,1120]" captionTargetId="figure@20.[151,1436,184,1120]" captionTargetPageId="20" captionText="FIGURE 4. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, outer surface of right chela; B, outer surface of left chela; C, right side of thoracic sternum showing ventral surface of chelipeds and spines on ambulatory meri; D, right P 4." httpUri="https://zenodo.org/record/178300/files/figure.png" pageId="1" pageNumber="40">Fig. 4</figureCitation>
|
||
C, D). Sterno-abdominal cavity almost reaches anterior edge of sternite 4 (
|
||
<figureCitation id="131B2A06FFC6FFA5EF04FC569BD4FB98" box="[305,385,1010,1036]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="1" pageNumber="40">Figs. 9</figureCitation>
|
||
A, 12A); press button of male abdominal locking mechanism distinct, on posterior edge of thoracic sternite 5 (
|
||
<figureCitation id="131B2A06FFC6FFA5EF42FBBE9B9EFBA0" box="[375,459,1050,1076]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="1" pageNumber="40">Fig. 12</figureCitation>
|
||
A). Penis soft, effectively coxal, exits at base of coxa of P5 but covered by posterior prolongation of episternite 7 (
|
||
<figureCitation id="131B2A06FFC6FFA5EFC4FBE69805FBC8" box="[497,592,1090,1116]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="1" pageNumber="40">Figs. 13</figureCitation>
|
||
A, B). Male abdomen relatively narrow overall (
|
||
<figureCitation id="131B2A06FFC6FFA5EAB1FBE69E81FBC8" box="[1156,1236,1090,1116]" captionStart="FIGURE 3" captionStartId="19.[151,255,1088,1112]" captionTargetBox="[151,1435,194,1064]" captionTargetId="figure@19.[151,1436,194,1064]" captionTargetPageId="19" captionText="FIGURE 3. Vultocinus anfractus, new genus, new species. Holotype male (cl 19.5 mm, cw 20.3 mm) (NMCR 19114), Philippines. A, dorsal view of carapace; B, ventral view of carapace; C, buccal area showing third maxillipeds; D, frontal view of carapace and chelipeds." httpUri="https://zenodo.org/record/178299/files/figure.png" pageId="1" pageNumber="40">Figs. 3</figureCitation>
|
||
B, 5B, 9A, 10A), sutures for all somites and telson clearly visible; somite 1 transversely and longitudinally broad, trapezoidal in shape, exposed, not covered by posterior carapace margin (
|
||
<figureCitation id="131B2A06FFC6FFA5ED56FB3699EDFB38" box="[867,952,1170,1196]" captionStart="FIGURE 11" captionStartId="27.[151,255,1445,1469]" captionTargetBox="[151,1435,194,1420]" captionTargetId="figure@27.[151,1436,194,1421]" captionTargetPageId="27" captionText="FIGURE 11. Posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 1 – 3 = male abdominal somites 1 to 3 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178307/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
|
||
A), somites 2–6 relatively narrow transversely, somite 6 as long as broad (
|
||
<figureCitation id="131B2A06FFC6FFA5EC1AFB1E982AFB40" box="[559,639,1210,1236]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="1" pageNumber="40">Figs. 9</figureCitation>
|
||
A, 10A), somites 3, 4 immovable, fused, other somites freely articulating (
|
||
<figureCitation id="131B2A06FFC6FFA5EEDEFB469B66FB68" box="[235,307,1250,1276]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
|
||
B). G1 relatively slender, gently curved, surface with simple spines (
|
||
<figureCitation id="131B2A06FFC6FFA5EA5DFB469EFBFB68" box="[1128,1198,1250,1276]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
|
||
C-F); G2 with short distal segment, about 0.2 times length of basal segment (
|
||
<figureCitation id="131B2A06FFC6FFA5ED1BFAAE9927FAB0" box="[814,882,1290,1316]" captionStart="FIGURE 5" captionStartId="21.[151,255,1379,1403]" captionTargetBox="[158,1421,201,1348]" captionTargetId="figure@21.[151,1436,194,1355]" captionTargetPageId="21" captionText="FIGURE 5. Vultocinus anfractus, new genus, new species. Paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC), Philippines. A, left third maxilliped (setae denuded); B, male abdomen (partially denuded); C, left G 1, dorsal view; D, left G 1, ventral view; E, distal part of left G 1, dorsal view; F, distal part of left G 1, ventral view; G, left G 2; H, distal part of left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/178301/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
|
||
G, H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6508FFC6FFA5EEF3FA96983AFAD8" box="[198,623,1330,1356]" pageId="1" pageNumber="40" type="materials_examined">
|
||
<paragraph id="8B9F3683FFC6FFA5EEF3FA96983AFAD8" blockId="1.[151,1437,650,2036]" box="[198,623,1330,1356]" pageId="1" pageNumber="40">
|
||
<emphasis id="B954EA91FFC6FFA5EEF3FA969B06FAD8" bold="true" box="[198,339,1330,1356]" pageId="1" pageNumber="40">
|
||
<typeStatus id="549B8821FFC6FFA5EEF3FA969B56FAD8" box="[198,259,1330,1356]" pageId="1" pageNumber="40">Type</typeStatus>
|
||
genus
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D00FFC6FFA5EF54FA979B89FAD8" box="[353,476,1331,1356]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B954EA91FFC6FFA5EF54FA979B89FAD8" box="[353,476,1331,1356]" italics="true" pageId="1" pageNumber="40">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26757EAFFC6FFA5EFDEFA96983EFAD8" box="[491,619,1330,1356]" pageId="1" pageNumber="40" rank="genus">new genus</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6508FFC6FFA1EEF3FAFE9F46F8FE" lastPageId="5" lastPageNumber="44" pageId="1" pageNumber="40" type="discussion">
|
||
<paragraph id="8B9F3683FFC6FFA5EEF3FAFE9E74F918" blockId="1.[151,1437,650,2036]" pageId="1" pageNumber="40">
|
||
<emphasis id="B954EA91FFC6FFA5EEF3FAFE9B6DFAE0" bold="true" box="[198,312,1370,1396]" pageId="1" pageNumber="40">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D00FFC6FFA5EF7DFAFF9B96FAE0" box="[328,451,1371,1396]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B954EA91FFC6FFA5EF7DFAFF9B96FAE0" box="[328,451,1371,1396]" italics="true" pageId="1" pageNumber="40">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26757EAFFC6FFA5EFE1FAFE9800FAE0" box="[468,597,1370,1396]" pageId="1" pageNumber="40" rank="genus">new genus</taxonomicNameLabel>
|
||
, possesses a suite of unusual characters that make its precise affinities difficult to ascertain. Superficially, its carapace shape and setation pattern resembles species of
|
||
<taxonomicName id="4C204D00FFC6FFA5EB2BFA279B72FA50" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC6FFA5EB2BFA279FCEFA08" box="[1310,1435,1411,1436]" italics="true" pageId="1" pageNumber="40">Lophoplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EEA2FA0E9B72FA50" author="Tesch" box="[151,295,1450,1476]" pageId="1" pageNumber="55" refString="Tesch, J. (1918) The Decapoda Brachyura of the Siboga-Expedition. II. Goneplacidae and Pinnotheridae. In: Uitkomstem op Zoologisch, Botanisch, Oceanographisch en Geologisch gebied verzameld in Nederlansch Oost-Indie 1899 - 1900 aan boord H. M. Siboga onder commando van Luitenant ter zee Ie kl. G. F. Tydeman uitgeven door Dr. Max Webber. Siboga - Expeditie, 39 c 1 (84), 149 - 295, 12 pls." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
, but members of this genus have slender and sinuous G1s with a very short sigmoid G2, and are currently classified in the subfamily
|
||
<taxonomicName id="4C204D00FFC6FFA5EC73FA7699DBFA78" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[582,910,1490,1516]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">
|
||
Rhizopinae
|
||
<bibRefCitation id="EFB14B72FFC6FFA5ECE6FA7699DBFA78" author="Stimpson" box="[723,910,1490,1516]" pageId="1" pageNumber="55" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part V, Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 93 - 110." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
, family
|
||
<taxonomicName id="4C204D00FFC6FFA5EDDAFA769E21FA78" box="[1007,1140,1490,1516]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Pilumnidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EA86FA769F18FA78" author="Takeda" box="[1203,1357,1490,1516]" pageId="1" pageNumber="55" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, Tokyo, 86 (2), 120 - 123." type="journal article" year="1977">Takeda 1977</bibRefCitation>
|
||
; Ng &
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EEA2FA5E9B76F980" author="Davie" box="[151,291,1530,1556]" pageId="1" pageNumber="54" refString="Ng, P. K. L. & Davie, P. J. F. (1991) The Indo-Pacific Pilumnidae VII. Notes on the taxonomy of Heteropilumnus sasekumari (Serene, 1971) and Cryptolutea Ward, 1936 (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 30 (3), 517 - 524." type="journal article" year="1991">Davie 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EF1AFA5E985BF980" author="Ng" box="[303,526,1530,1556]" pageId="1" pageNumber="54" refString="Ng, P. K. L. & Huang, J. - F. (2002) The Indo-Pacific Pilumnidae XV. On two rare species of Rhizopinae (Decapoda, Brachyura) from Taiwan. Crustaceana, 74, 11, 1379 - 1385." type="journal article" year="2002">Ng & Huang 2002</bibRefCitation>
|
||
; review by Ng 1987). The general carapace shape and setose surfaces of
|
||
<taxonomicName id="4C204D00FFC6FFA5EB59FA5F9AB9F9A8" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC6FFA5EB59FA5F9AB9F9A8" italics="true" pageId="1" pageNumber="40">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
also resembles members of the genus
|
||
<taxonomicName id="4C204D00FFC6FFA5EC82F98799A8F9A8" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[695,1021,1570,1596]" class="Malacostraca" family="Planopilumnidae" genus="Planopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC6FFA5EC82F987993CF9A8" box="[695,873,1571,1596]" italics="true" pageId="1" pageNumber="40">Planopilumnus</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC6FFA5ED44F98699A8F9A8" author="Balss" box="[881,1021,1570,1596]" pageId="1" pageNumber="52" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattungen Pilumnus (Crustacea Dekapoda) und verwandter Gattungen. Capita Zoologica, 4, 1 - 47, pls. 1 - 7." type="journal article" year="1933">Balss, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
, of the subfamily Planopilumninae
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EEA2F9EE9B67F9F0" author="Serene" box="[151,306,1610,1636]" pageId="1" pageNumber="55" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge, Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par Crosnier (A.): Carpiliidae et Menippidae. Faune Tropicale, 24, 1 - 349, figs A-C + 1 - 243, pls 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
|
||
, family
|
||
<taxonomicName id="4C204D00FFC6FFA5EFA4F9EE9865F9F0" box="[401,560,1610,1636]" class="Malacostraca" family="Pseudoziidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Pseudoziidae</taxonomicName>
|
||
; but members of this family have stout G1s and a G2 which is only half the length of the G1 (see Ng &
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EFE8F9D69825F918" author="Clark" box="[477,624,1650,1676]" pageId="1" pageNumber="54" refString="Ng, P. K. L. & Clark, P. F. (2000 a) The Indo-Pacific Pilumnidae XII. On the familial placement of Chlorodiella bidentata (Nobili, 1901) and Tanaocheles stenochilus Kropp, 1984 using adult and larval characters with the establishment of a new subfamily, Tanaochelinae (Crustacea: Decapoda: Brachyura). Journal of Natural History, 34, 207 - 245." type="journal article" year="2000" yearSuffix="a">Clark 2000a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EC49F9D698D8F918" author="Clark" box="[636,653,1650,1676]" pageId="1" pageNumber="54" refString="Ng, P. K. L. & Clark, P. F. (2000 b) The eumedonid file: a case study of systematic compatibility using larval and adult characters (Crustacea: Decapoda: Brachyura). Invertebrate Reproduction and Development, 38 (3), 225 - 252." type="journal article" year="2000" yearSuffix="b">b</bibRefCitation>
|
||
; Ng et al. 2001;
|
||
<bibRefCitation id="EFB14B72FFC6FFA5ED7AF9D69E41F918" author="Ng" box="[847,1044,1650,1676]" pageId="1" pageNumber="54" refString="Ng, P. K. L. & Huang, J. - F. (2002) The Indo-Pacific Pilumnidae XV. On two rare species of Rhizopinae (Decapoda, Brachyura) from Taiwan. Crustaceana, 74, 11, 1379 - 1385." type="journal article" year="2002">Ng & Liao 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC6FFA6EEF3F93E9936FEEE" blockId="1.[151,1437,650,2036]" lastBlockId="2.[151,1437,152,2018]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
|
||
The possession of male abdominal somites 2 and 3 which are immovable, of a long and relatively stout G1 and of a long G2 which is subequal in length to the G1, appears to ally
|
||
<taxonomicName id="4C204D00FFC6FFA5EDD5F9679E0EF948" box="[992,1115,1731,1756]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC6FFA5EDD5F9679E0EF948" box="[992,1115,1731,1756]" italics="true" pageId="1" pageNumber="40">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
with some members of the
|
||
<taxonomicName id="4C204D00FFC6FFA5EEA2F94E9BAFF890" authority="MacLeay, 1838" authorityName="MacLeay" authorityYear="1838" box="[151,506,1770,1796]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">
|
||
Goneplacidae
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EF77F94E9BAFF890" author="MacLeay" box="[322,506,1770,1796]" lastPageId="1" lastPageNumber="54" pageId="1" pageNumber="53" refString="MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In A. Smith, Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa': together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe, Published under the Authority of the Lords Commissioners of Her Majesty's Treasury, Invertebratae: [1849] pp. 53 - 71, pls 2, 3." type="journal article" year="1838">MacLeay, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B954EA91FFC6FFA5EC31F94F9829F890" box="[516,636,1771,1796]" italics="true" pageId="1" pageNumber="40">
|
||
<taxonomicName id="4C204D00FFC6FFA5EC31F94F9810F890" box="[516,581,1771,1796]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
. While the taxonomy of this family has been a major problem for many years (see
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EF68F8B69839F8B8" author="Guinot" box="[349,620,1810,1836]" pageId="1" pageNumber="53" refString="Guinot, D. & Breton, G. (2006) Lithophylax trigeri A. Milne-Edwards & Brocchi, 1879 from the French Cretaceous (Cenomanian) and placement of the family Lithophylacidae Van Straelen, 1936 (Crustacea, Decapoda, Brachyura). Geodiversitas, 28 (4), 591 - 633." type="journal article" year="2006">Guinot & Breton 2006</bibRefCitation>
|
||
), recent studies which built on the key contributions made by
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EB7EF8B69ABCF8C0" author="Guinot" pageId="1" pageNumber="53" refString="Guinot, D. (1969 a) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae. Bulletin du Museum d'Histoire naturelle, Paris, (2) 41 (1), 241 - 265." type="journal article" year="1969" yearSuffix="a">Guinot (1969a</bibRefCitation>
|
||
–
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EECFF89E9B52F8C0" author="Guinot" box="[250,263,1850,1876]" pageId="1" pageNumber="53" refString="Guinot, D. (1969 c) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae (suite et fin). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 41 (3), 688 - 724." type="journal article" year="1969" yearSuffix="c">c</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EF23F89E9B07F8C0" author="Guinot" box="[278,338,1850,1876]" pageId="1" pageNumber="53" refString="Guinot, D. (1971) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VIII. Synthese et bibliographie. Bulletin du Museum national d'Histoire naturelle, Paris, (2) 42 (5), 1063 - 1090." type="journal article" year="1971">1971</bibRefCitation>
|
||
) have made the systematic position of the taxon much clearer. This is especially so when using the suite of many new characters pioneered by her over the years (see
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EA24F8C69EF2F8E8" author="Guinot" box="[1041,1191,1890,1916]" pageId="1" pageNumber="53" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum national d'Histoire naturelle, (A) Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EA86F8C69A83F830" author="Guinot" pageId="1" pageNumber="53" refString="Guinot, D. & Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot & Bouchard 1998</bibRefCitation>
|
||
). Over the years, many taxa and subfamilies have been transferred from the
|
||
<taxonomicName id="4C204D00FFC6FFA5EA49F82E9F48F830" box="[1148,1309,1930,1956]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
into other families (see Castro, in press, for review) and until 2003, the
|
||
<taxonomicName id="4C204D00FFC6FFA5ED41F8169E40F858" box="[884,1045,1970,1996]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
was believed to be composed of five subfamilies, viz. the Goneplacinae
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EC52F87E994AF860" author="MacLeay" box="[615,799,2010,2036]" lastPageId="1" lastPageNumber="54" pageId="1" pageNumber="53" refString="MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In A. Smith, Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa': together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe, Published under the Authority of the Lords Commissioners of Her Majesty's Treasury, Invertebratae: [1849] pp. 53 - 71, pls 2, 3." type="journal article" year="1838">MacLeay, 1838</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC6FFA5ED19F87E99BFF860" box="[812,1002,2010,2036]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
H.
|
||
<bibRefCitation id="EFB14B72FFC6FFA5EA20F87E9F41F860" author="Milne" box="[1045,1300,2010,2036]" pageId="1" pageNumber="54" refString="Milne Edwards, H. (1852) De la famille des Ocypodides (Ocypodidae). Second Memoire, in, Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences Naturelles (Zoologie), 18, 128 - 166 pls. 3, 4." type="journal article" year="1852">Milne Edwards, 1852</bibRefCitation>
|
||
, Euryplacinae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEFDFF3C9BD1FF26" author="Stimpson" box="[200,388,152,178]" pageId="2" pageNumber="55" refString="Stimpson W. (1871) Brachyura. Preliminary report on the Crustacea dredged in the Gulf Stream in the Straits of Florida, by L. F. Pourtales, Assist. U. S. Coast Surveys, Part 1. Bulletin of the Museum of Comparative Zoology at Harvard College, 2, 109 - 160." type="journal article" year="1871">Stimpson, 1871</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6EFA7FF3C9944FF26" authority="Serene, 1964" authorityName="Serene" authorityYear="1964" box="[402,785,152,178]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Chasmocarcininae">
|
||
Chasmocarcininae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC41FF3C9944FF26" author="Serene" box="[628,785,152,178]" pageId="2" pageNumber="54" refString="Serene R. (1964) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sousfamille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187, figs. 1 - 6." type="journal article" year="1964">Serène, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC5FFA6ED67FF3C9EFDFF26" authority="Guinot, 1986" authorityName="Guinot" authorityYear="1986" box="[850,1192,152,178]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Trogloplacinae">
|
||
Trogloplacinae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA3EFF3C9EFDFF26" author="Guinot" box="[1035,1192,152,178]" pageId="2" pageNumber="53" refString="Guinot, D. (1986) Description d'un crabe cavernicole aveugle de Nouvelle-Bretagne (Papouasie Nouvelle-Guinee), Trogloplax joliveti n. gen. n. sp., et etablissement d'une sous-famille nouvelle, Trogloplacinae subfam. nov. Comptes Rendus des Seances de Academie des Sciences, Paris, (3) 308, 307 - 312." type="journal article" year="1986">Guinot, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
(see Ng et al. 2001;
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2FF649B71FF4E" author="Davie" box="[151,292,192,218]" pageId="2" pageNumber="53" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia, 19.3 B, pp. 1 - 641." type="book chapter" year="2002">Davie 2002</bibRefCitation>
|
||
). Recent studies have substantially changed the landscape of goneplacid taxonomy –
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EB3FFF649B78FE96" author="Karasawa" pageId="2" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 a) The family Goneplacidae MacLeay, 1838 (Crustacea: Decapoda: Brachyura): systematics, phylogeny and fossil records. Paleontological Research, 7 (2), 129 - 151." type="journal article" year="2003" yearSuffix="a">Karasawa & Kato (2003a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF0EFF4C9B19FE96" author="Karasawa" box="[315,332,232,258]" pageId="2" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 b) The phylogeny, systematics and fossil record of the Goneplacidae MacLeay (Crustacea, Decapoda, Brachyura) revisited. Contributions in Zoology to the Mesozoic and Cenozoic Decapod Crustaceans, 72 (2 - 3), 147 - 152." type="journal article" year="2003" yearSuffix="b">b</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF51FF439840FE96" author="Stevcic" box="[356,533,231,258]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC16FF4C99CCFE96" author="Karasawa" box="[547,921,232,258]" pageId="2" pageNumber="53" refString="Karasawa, H. & Schweitzer, M. (2006) A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology, 75 (1 / 2), 23 - 73." type="journal article" year="2006">Karasawa & Schweitzer (2006)</bibRefCitation>
|
||
and Castro (in press); which are discussed individually below. These discussions are not always restricted to the extant fauna, a major shortcoming since there is often a need to use many important characters like male abdominal structure, as well as detailed sternal and penial features which are rarely observable in fossils.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC5FFA6EEF3FE2C9B69FC3E" blockId="2.[151,1437,152,2018]" pageId="2" pageNumber="41">
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEF3FE2C9BA7FE36" author="Karasawa" box="[198,498,392,418]" pageId="2" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 a) The family Goneplacidae MacLeay, 1838 (Crustacea: Decapoda: Brachyura): systematics, phylogeny and fossil records. Paleontological Research, 7 (2), 129 - 151." type="journal article" year="2003" yearSuffix="a">Karasawa & Kato (2003a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EFCBFE2C985BFE36" author="Karasawa" box="[510,526,392,418]" pageId="2" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 b) The phylogeny, systematics and fossil record of the Goneplacidae MacLeay (Crustacea, Decapoda, Brachyura) revisited. Contributions in Zoology to the Mesozoic and Cenozoic Decapod Crustaceans, 72 (2 - 3), 147 - 152." type="journal article" year="2003" yearSuffix="b">b</bibRefCitation>
|
||
) reappraised the phylogeny of the
|
||
<taxonomicName id="4C204D00FFC5FFA6ED96FE2C9E11FE36" box="[931,1092,392,418]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
and rearranged the subfamilial system based on selected extant and fossil taxa. They synonymised the
|
||
<taxonomicName id="4C204D00FFC5FFA6EA26FE149E84FE5E" box="[1043,1233,432,458]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
under the Goneplacinae, and also established a new subfamily, Mathildellinae, for
|
||
<taxonomicName id="4C204D00FFC5FFA6EDF9FE7C9F41FE66" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[972,1300,472,498]" class="Malacostraca" family="Mathildellidae" genus="Beuroisia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EDF9FE7C9E68FE65" box="[972,1085,472,497]" italics="true" pageId="2" pageNumber="41">Beuroisia</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EB70FE7C9A81FD8E" author="Forges" pageId="2" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6EED4FDA49859FD8E" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[225,524,512,538]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EED4FDA49B69FD8D" box="[225,316,512,537]" italics="true" pageId="2" pageNumber="41">Intesius</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC0CFDA49880FD8E" author="Forges" box="[569,725,512,538]" pageId="2" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6ECD6FDA49E6EFD8E" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[739,1083,512,538]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6ECD6FDA49939FD8D" box="[739,876,512,537]" italics="true" pageId="2" pageNumber="41">Mathildella</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA5DFDA49F51FD8E" author="Forges" box="[1128,1284,512,538]" pageId="2" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6EB27FDA49B3CFDD6" authority="Alcock, 1894" authorityName="Alcock" authorityYear="1894" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EB27FDA49A95FDD5" italics="true" pageId="2" pageNumber="41">Platypilumnus</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEFDFD8C9B3CFDD6" author="Alcock" box="[200,361,552,578]" pageId="2" pageNumber="52" refString="Alcock, A. (1894). Natural history notes from Indian Marine Survey Steamer " Investigator ". Ser. II, no. 1. On the result of deep-sea dredging during the season of 1890 - 1891. Annals and Magazine of Natural History, (6) 13, 400 - 411" type="journal article" year="1894">Alcock, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC5FFA6EF9FFD8C9944FDD6" authority="Serene, 1969" authorityName="Serene" authorityYear="1969" box="[426,785,552,578]" class="Malacostraca" family="Mathildellidae" genus="Neopilumnoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EF9FFD8C9839FDD5" box="[426,620,552,577]" italics="true" pageId="2" pageNumber="41">Neopilumnoplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC41FD8C9944FDD6" author="Serene" box="[628,785,552,578]" pageId="2" pageNumber="54" refString="Serene R. (1969) Description de deux especes nouvelles et redecouverte d'une espece de Brachyoure (Decapoda Crustacea) dans les eaux d'Indonesie. Cahiers du Pacifique, 13, 281 - 293, figs. 1 - 24, pls. 1 - 4." type="journal article" year="1969">Serène, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, which had originally been placed in the
|
||
<taxonomicName id="4C204D00FFC5FFA6EAC9FD8C9A96FDFE" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
. Although Karasawa & Kato (2003) define the Mathildellinae with a suite of carapace characters, one of the most diagnostic characters (but unfortunately only usable for fresh specimens) is whether somites 3 and 4 are mobile. All known extant adult male goneplacines (and carcinoplacines) have somites 3 and 4 freely mobile (with one exception,
|
||
<taxonomicName id="4C204D00FFC5FFA6EFDBFD6C99AAFD76" authority="Takeda & Miyake, 1969" authorityName="Takeda & Miyake" authorityYear="1969" box="[494,1023,712,738]" class="Malacostraca" family="Goneplacidae" genus="Neommatocarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EFDBFD6C9884FD75" box="[494,721,712,737]" italics="true" pageId="2" pageNumber="41">Neommatocarcinus</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ECEEFD6C99AAFD76" author="Takeda" box="[731,1023,712,738]" pageId="2" pageNumber="55" refString="Takeda, M. & Miyake, S. (1969) A small collection of crabs from New Zealand. OHMU, Occasional Papers of Zoological Laboratory, Faculty of Agriculture, Kyushu University 2, 157 - 193, pls. 1 - 3." type="journal article" year="1969">Takeda & Miyake, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has somites 3–5 fused, see also Castro, in press). On the other hand, all extant adult male mathildellines have abdominal somites 3 and 4 immobile although the sutures separating them are still visible (see also Ng &
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA7FFCBC9E85FCA6" author="Chan" box="[1098,1232,792,818]" pageId="2" pageNumber="54" refString="Ng, P. K. L. & Chan, T. - Y. (2000) Note on Mathildella serrata (Sakai, 1974) (Crustacea: Decapoda: Brachyura: Goneplacidae) from deep waters in Taiwan. National Taiwan Museum Special Publication Series, 10, 149 - 153." type="journal article" year="2000">Chan 2000</bibRefCitation>
|
||
; Ng &
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EB19FCBC9FC2FCA6" author="Ho" box="[1324,1431,792,818]" pageId="2" pageNumber="54" refString="Ng, P. K. L. & Ho, P. - H. (2003) Mathildella rubra, a new species of deep-water carcinoplacine crab (Decapoda, Brachyura) from the Philippines. Crustaceana, 76 (3), 333 - 342." type="journal article" year="2003">Ho 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2FCE49BC4FCCE" author="Crosnier" box="[151,401,832,858]" pageId="2" pageNumber="53" refString="Crosnier, A. & Ng, P. K. L. (2004) Remarques sur le genre Intesius (Crustacea, Decapoda, Brachyura, Goneplacidae) et description de deux especes nouvelles. Zoosystema, 26 (2), 263 - 277." type="journal article" year="2004">Crosnier & Ng 2004</bibRefCitation>
|
||
). The articulation between somites 4 and 5 is usually more difficult to ascertain. In the mathildellines we have examined, there is much more limited movement between these somites compared to other somites.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC5FFA7EEF3FC1C9F68FDD6" blockId="2.[151,1437,152,2018]" lastBlockId="3.[151,1437,151,2018]" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
|
||
In a major reappraisal,
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EFE0FC1398D5FC46" author="Stevcic" box="[469,640,951,978]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
substantially reorganized the family but with minimal or no discussion. He restricted what he defined as the superfamily Goneplacoidea to just three families:
|
||
<taxonomicName id="4C204D00FFC5FFA6EACEFC449FC9FC6E" box="[1275,1436,992,1018]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
(with two subfamilies, Goneplacinae and
|
||
<taxonomicName id="4C204D00FFC5FFA6ECA8FBAC990AFBB6" box="[669,863,1032,1058]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
, with two tribes—Carcinoplacini, Psopheticini
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2FB8B9B6AFBDE" author="Stevcic" box="[151,319,1071,1098]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C204D00FFC5FFA6EF6DFB9498D2FBDE" authority="Colosi, 1923" authorityName="Colosi" authorityYear="1923" box="[344,647,1072,1098]" class="Malacostraca" family="Geryonidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">
|
||
Geryonidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EFD9FB9498D2FBDE" author="Colosi" box="[492,647,1072,1098]" pageId="2" pageNumber="53" refString="Colosi, G. (1923) Una specie fossile di Gerionide (Decapodi brachiuri). Bollettino dei Societe Naturale di Napoli, 2 (15 I), 248 - 255." type="journal article" year="1923">Colosi, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
(with three subfamilies, Geryoninae with six tribes—Geryonini, Progeryonini
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF02FBF39B8CFBE6" author="Stevcic" box="[311,473,1111,1138]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, Paragalenini
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC4AFBF39974FBE6" author="Stevcic" box="[639,801,1111,1138]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, Mathildellini, Intesiini
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA09FBF39E8BFBE6" author="Stevcic" box="[1084,1246,1111,1138]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, Platycheloniini
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2FBDB9B68FB0E" author="Stevcic" box="[151,317,1151,1178]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
; Platypilumninae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EC22FBDB98EBFB0E" author="Stevcic" box="[535,702,1151,1178]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
; and
|
||
<taxonomicName id="4C204D00FFC5FFA6ED37FB249E09FB0E" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[770,1116,1151,1178]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Bathyplacinae">
|
||
Bathyplacinae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ED81FBDB9E09FB0E" author="Stevcic" box="[948,1116,1151,1178]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
), and
|
||
<taxonomicName id="4C204D00FFC5FFA6EA9DFB249F3AFB0E" box="[1192,1391,1152,1178]" class="Malacostraca" family="Planopilumnidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Planopilumnidae</taxonomicName>
|
||
(as Planopilumninae [
|
||
<emphasis id="B954EA91FFC5FFA6EF5BFB0C9BDAFB55" box="[366,399,1192,1217]" italics="true" pageId="2" pageNumber="41">sic</emphasis>
|
||
])
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF9FFB0C9813FB56" author="Serene" box="[426,582,1192,1218]" pageId="2" pageNumber="55" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge, Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par Crosnier (A.): Carpiliidae et Menippidae. Faune Tropicale, 24, 1 - 349, figs A-C + 1 - 243, pls 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
|
||
. Many other taxa which had been placed in the
|
||
<taxonomicName id="4C204D00FFC5FFA6EA43FB0C9F42FB56" box="[1142,1303,1192,1218]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
up to 2005 were transferred to other groups or whole new families and subfamilies. His recognition of some of them as families, e.g. the
|
||
<taxonomicName id="4C204D00FFC5FFA6EF59FB5C9852FA86" box="[364,519,1272,1298]" class="Malacostraca" family="Euryplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Euryplacidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC5FFA6EC7DFB5C9974FA86" box="[584,801,1272,1298]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Chasmocarcininae">Chasmocarcininae</taxonomicName>
|
||
as distinct families was expected as there had been much prior discussion on their status, although his unexplained elevation of both groups to superfamily status seems unwarranted. The same applies for placing
|
||
<taxonomicName id="4C204D00FFC5FFA6ECE8FAEC9E4EFAF6" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[733,1051,1352,1378]" class="Malacostraca" family="Scalopidiidae" genus="Scalopidia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6ECE8FAEC990FFAF5" box="[733,858,1352,1377]" italics="true" pageId="2" pageNumber="41">Scalopidia</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ED54FAEC9E4EFAF6" author="Stimpson" box="[865,1051,1352,1378]" pageId="2" pageNumber="55" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part V, Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 93 - 110." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
, into its own family, the
|
||
<taxonomicName id="4C204D00FFC5FFA6EB0FFAEC9BC7FA1E" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" class="Malacostraca" family="Scalopidiidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">
|
||
Scalopidiidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEDFFACB9BC7FA1E" author="Stevcic" box="[234,402,1391,1418]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, in the superfamily Chasmocarcinoidea. The subfamily
|
||
<taxonomicName id="4C204D00FFC5FFA6EA75FAD49FC9FA1E" authority="Guinot, 1986" authorityName="Guinot" authorityYear="1986" box="[1088,1436,1392,1418]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Trogloplacinae">
|
||
Trogloplacinae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EAC8FAD49FC9FA1E" author="Guinot" box="[1277,1436,1392,1418]" pageId="2" pageNumber="53" refString="Guinot, D. (1986) Description d'un crabe cavernicole aveugle de Nouvelle-Bretagne (Papouasie Nouvelle-Guinee), Trogloplax joliveti n. gen. n. sp., et etablissement d'une sous-famille nouvelle, Trogloplacinae subfam. nov. Comptes Rendus des Seances de Academie des Sciences, Paris, (3) 308, 307 - 312." type="journal article" year="1986">Guinot, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
(regarded as a family by
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF8FFA3C9812FA26" author="Davie" box="[442,583,1432,1458]" pageId="2" pageNumber="53" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia, 19.3 B, pp. 1 - 641." type="book chapter" year="2002">Davie 2002</bibRefCitation>
|
||
), has clear chasmocarcinid affinities (see
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA18FA3C9F65FA26" author="Guinot" box="[1069,1328,1432,1458]" pageId="2" pageNumber="53" refString="Davie, P. J. F. & Guinot, D. (1996) Two new freshwater crabs in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae MacLeay (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39 (2), 277 - 287." type="journal article" year="1996">Guinot & Davie 1996</bibRefCitation>
|
||
), but this was ignored and
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF6AFA1B985FFA4E" author="Stevcic" box="[351,522,1471,1498]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
recognised it as a distinct superfamily, separate from his Chasmocarcinoidea. Two genera which had been allied to the
|
||
<taxonomicName id="4C204D00FFC5FFA6ECA6FA4C993AF996" box="[659,879,1512,1538]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6ED48FA4C9EACF996" authority="Kossmann, 1877" authorityName="Kossmann" authorityYear="1877" box="[893,1273,1512,1538]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6ED48FA4C9E76F995" box="[893,1059,1512,1537]" italics="true" pageId="2" pageNumber="41">Parapilumnus</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA1BFA4C9EACF996" author="Kossmann" box="[1070,1273,1512,1538]" pageId="2" pageNumber="53" refString="Kossmann, R. (1877) Malacostraca (1. Theil: Brachyura). Chapter Erste Halfte, III. In: Kossmann, R. (editor), Zoologische Ergebnisse einer im Auftrage der Koniglichen Academie der Wissenschaften zu Berlin ausgefuhrten Reise in die Kustengebiete des Rothen Meeres. Pp. 1 - 66, pls. 1 - 3." type="book chapter" year="1877">Kossmann, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC5FFA6EB08FA4C9B04F9BE" authority="Stimpson, 1871" authorityName="Stimpson" authorityYear="1871" class="Malacostraca" family="Acidopsidae" genus="Acidops" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EB08FA4C9FC9F995" box="[1341,1436,1512,1537]" italics="true" pageId="2" pageNumber="41">Acidops</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2F9B49B04F9BE" author="Stimpson" box="[151,337,1552,1578]" pageId="2" pageNumber="55" refString="Stimpson W. (1871) Brachyura. Preliminary report on the Crustacea dredged in the Gulf Stream in the Straits of Florida, by L. F. Pourtales, Assist. U. S. Coast Surveys, Part 1. Bulletin of the Museum of Comparative Zoology at Harvard College, 2, 109 - 160." type="journal article" year="1871">Stimpson, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
, were inexplicably referred to their own families (Parapilumnidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EA69F9AB9EABF9BE" author="Stevcic" box="[1116,1278,1551,1578]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC5FFA6EB09F9B49B2BF9C6" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" class="Malacostraca" family="Acidopsidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">
|
||
Acidopsidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEEEF9939B2BF9C6" author="Stevcic" box="[219,382,1591,1618]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, respectively) and superfamilies in two different parts of his paper, although the affinities between them are very clear and should be classified together (
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EDBEF9C499ACF9EE" author="Ng" box="[907,1017,1632,1658]" pageId="2" pageNumber="54" refString="Ng, P. K. L. & Huang, J. - F. (2002) The Indo-Pacific Pilumnidae XV. On two rare species of Rhizopinae (Decapoda, Brachyura) from Taiwan. Crustaceana, 74, 11, 1379 - 1385." type="journal article" year="2002">Ng 2002</bibRefCitation>
|
||
). Two other genera linked with the
|
||
<taxonomicName id="4C204D00FFC5FFA6EEA2F92C9B26F936" box="[151,371,1672,1698]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6EFB5F92C980CF936" authority="Ng, 1987" authorityName="Ng" authorityYear="1987" box="[384,601,1672,1698]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EFB5F92C9B88F935" box="[384,477,1672,1697]" italics="true" pageId="2" pageNumber="41">Raoulia</emphasis>
|
||
Ng, 1987
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC5FFA6ECA9F92C9E4DF936" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[668,1048,1672,1698]" class="Malacostraca" family="Chasmocarcinidae" genus="Typhlocarcinodes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6ECA9F92C993EF935" box="[668,875,1672,1697]" italics="true" pageId="2" pageNumber="41">Typhlocarcinodes</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ED40F92C9E4DF936" author="Alcock" box="[885,1048,1672,1698]" pageId="2" pageNumber="52" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3) 3, 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, were also referred to their own families (Raoulidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EFB3F90B987CF95E" author="Stevcic" box="[390,553,1711,1738]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, and Typhlocarcinodidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ED6CF90B99A8F95E" author="Stevcic" box="[857,1021,1711,1738]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, respectively), and own superfamilies. One genus clearly affiliated with the
|
||
<taxonomicName id="4C204D00FFC5FFA6ECA9F97C9968F966" box="[668,829,1752,1778]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
and Goneplacinae
|
||
<emphasis id="B954EA91FFC5FFA6EA1EF97C9EF1F965" box="[1067,1188,1752,1777]" italics="true" pageId="2" pageNumber="41">
|
||
<taxonomicName id="4C204D00FFC5FFA6EA1EF97C9E39F965" box="[1067,1132,1752,1777]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C204D00FFC5FFA6EA82F97C9F4CF965" box="[1207,1305,1752,1777]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EA82F97C9F4CF965" box="[1207,1305,1752,1777]" italics="true" pageId="2" pageNumber="41">Notonyx</emphasis>
|
||
</taxonomicName>
|
||
A. Milne-
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2F8A49B1FF88E" author="Edwards" box="[151,330,1792,1818]" pageId="2" pageNumber="54" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie, Deuxieme partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18." type="journal article" year="1873">Edwards, 1873</bibRefCitation>
|
||
, was removed to its own family (Notonycidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6ED49F95B9E4BF88E" author="Stevcic" box="[892,1054,1791,1818]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
) and superfamily Notonycoidea
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EEA2F8839B6CF8D6" author="Stevcic" box="[151,313,1831,1858]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
(see also
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EF86F88C98DDF8D6" author="Clark" box="[435,648,1832,1858]" lastPageId="2" lastPageNumber="53" pageId="2" pageNumber="52" refString="Clark, P. F. & Ng, P. K. L. (2005) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea, Brachyura, Goneplacidae) from the intertidal zone of Phuket, Thailand. Zoosystema, 28 (2), 539 - 551." type="journal article" year="2005">Clark & Ng 2005</bibRefCitation>
|
||
; Castro in press). The genus
|
||
<taxonomicName id="4C204D00FFC5FFA6EDD2F88C9F49F8D6" authority="Ng & Ng, 2003" authorityName="Ng & Ng" authorityYear="2003" box="[999,1308,1832,1858]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EDD2F88C9E01F8D5" box="[999,1108,1832,1857]" italics="true" pageId="2" pageNumber="41">Conleyus</emphasis>
|
||
Ng & Ng, 2003
|
||
</taxonomicName>
|
||
, originally placed in the
|
||
<taxonomicName id="4C204D00FFC5FFA6EF00F8F49BA2F8FE" box="[309,503,1872,1898]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
, was refered to its own family (
|
||
<taxonomicName id="4C204D00FFC5FFA6ED59F8F49EF5F8FE" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[876,1184,1871,1898]" class="Malacostraca" family="Conleyidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">
|
||
Conleyidae
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EDCEF8EB9EF5F8FE" author="Stevcic" box="[1019,1184,1871,1898]" pageId="2" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
). Some of his actions are very difficult to rationalize, e.g. Parapilumoidea and Acidoposidea, even if both are not true chasmocarcinids as defined by
|
||
<bibRefCitation id="EFB14B72FFC5FFA6EFB8F80498FDF82E" author="Guinot" box="[397,680,1952,1978]" pageId="2" pageNumber="53" refString="Davie, P. J. F. & Guinot, D. (1996) Two new freshwater crabs in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae MacLeay (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39 (2), 277 - 287." type="journal article" year="1996">Guinot & Davie (1996)</bibRefCitation>
|
||
. The same situation prevails for
|
||
<taxonomicName id="4C204D00FFC5FFA6EA18F8049EDFF82D" box="[1069,1162,1952,1977]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EA18F8049EDFF82D" box="[1069,1162,1952,1977]" italics="true" pageId="2" pageNumber="41">Raoulia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC5FFA6EAF3F8049FC0F82D" box="[1222,1429,1952,1977]" class="Malacostraca" family="Chasmocarcinidae" genus="Typhlocarcinodes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC5FFA6EAF3F8049FC0F82D" box="[1222,1429,1952,1977]" italics="true" pageId="2" pageNumber="41">Typhlocarcinodes</emphasis>
|
||
</taxonomicName>
|
||
. Both genera may not be chasmocarcinids, but to classify them as belonging to separate superfamilies is difficult to understand. The case with
|
||
<taxonomicName id="4C204D00FFC4FFA7EC03FF3C98CDFF25" box="[566,664,152,177]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EC03FF3C98CDFF25" box="[566,664,152,177]" italics="true" pageId="3" pageNumber="42">Notonyx</emphasis>
|
||
</taxonomicName>
|
||
and Notonycoidea also suggests that
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EA50FF339F41FF26" author="Stevcic" box="[1125,1300,151,178]" pageId="3" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Stevčić (2005)</bibRefCitation>
|
||
placed too much emphasis on general external morphology. While
|
||
<taxonomicName id="4C204D00FFC4FFA7ED07FF6499CAFF4D" box="[818,927,192,217]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7ED07FF6499CAFF4D" box="[818,927,192,217]" italics="true" pageId="3" pageNumber="42">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
has many peculiar features, the need for a separate superfamily is also unwarranted because the genus has many affinities with goneplacids. Stevčić’s (2005) inclusion of
|
||
<taxonomicName id="4C204D00FFC4FFA7EFB0FEB4985BFEBE" box="[389,526,272,298]" class="Malacostraca" family="Geryonidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Geryonidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EC7EFEB49947FEBE" box="[587,786,272,298]" class="Malacostraca" family="Planopilumnidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Planopilumnidae</taxonomicName>
|
||
into his Goneplacoidea is also difficult to understand. This is also true for his transfer of the Mathildellinae (as a tribe) into his Geryoninae and the removal of genera previously included in it like
|
||
<taxonomicName id="4C204D00FFC4FFA7EC22FEC49827FEED" box="[535,626,352,377]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EC22FEC49827FEED" box="[535,626,352,377]" italics="true" pageId="3" pageNumber="42">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EC9FFEC49901FEED" box="[682,852,352,377]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EC9FFEC49901FEED" box="[682,852,352,377]" italics="true" pageId="3" pageNumber="42">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
into its own tribe and subfamily respectively. The affinities of the geryonids with portunoids has been well argued (see
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EDD2FE2C9EBFFE36" author="Ng" box="[999,1258,392,418]" pageId="3" pageNumber="54" refString="Ng, P. K. L. & Manning, R. B. (1998) A new deep-water crab from Belau, Micronesia, with a key to the Pacific species of Chaceon (Crustacea: Decapoda: Brachyura: Geryonidae). Proceedings of the Biological Society of Washington, 111 (2), 389 - 397." type="journal article" year="1998">Ng & Manning 1998</bibRefCitation>
|
||
; Ng &
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EB7EFE2C9A80FE5E" author="Guinot" pageId="3" pageNumber="54" refString="Ng, P. K. L. & Guinot, D. (1999). On a new species of deep-water crab of the genus Progeryon (Decapoda, Brachyura, Geryonidae) from Hawaii. Crustaceana, 72 (7), 685 - 692." type="journal article" year="1999">Guinot 1999</bibRefCitation>
|
||
), while planopilumnids have been included in the
|
||
<taxonomicName id="4C204D00FFC4FFA7ED2BFE149E08FE5E" authority="Alcock, 1898" authorityName="Alcock" authorityYear="1898" box="[798,1117,432,458]" class="Malacostraca" family="Pseudoziidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">
|
||
Pseudoziidae
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EDF5FE149E08FE5E" author="Alcock" box="[960,1117,432,458]" pageId="3" pageNumber="52" refString="Alcock, A. (1898) Materials for a Carcinological Fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The Family Xanthidae. Journal of the Asiatic Society of Bengal, Calcutta, 67 (2) (1), 67 - 233." type="journal article" year="1898">Alcock, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
by Ng et al. (2001) and
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EB4DFE149B16FE66" author="Ng" pageId="3" pageNumber="54" refString="Ng, P. K. L. & Huang, J. - F. (2002) The Indo-Pacific Pilumnidae XV. On two rare species of Rhizopinae (Decapoda, Brachyura) from Taiwan. Crustaceana, 74, 11, 1379 - 1385." type="journal article" year="2002">Ng & Liao (2002)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C204D00FFC4FFA7EF7BFE7C9BFCFE65" box="[334,425,472,497]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF7BFE7C9BFCFE65" box="[334,425,472,497]" italics="true" pageId="3" pageNumber="42">Intesius</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C204D00FFC4FFA7EFDEFE7C98C1FE65" box="[491,660,472,497]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EFDEFE7C98C1FE65" box="[491,660,472,497]" italics="true" pageId="3" pageNumber="42">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
) is closely allied to
|
||
<emphasis id="B954EA91FFC4FFA7EDB5FE7C9E54FE65" box="[896,1025,472,497]" italics="true" pageId="3" pageNumber="42">Mathidella</emphasis>
|
||
(see also
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EA41FE7C9FC2FE66" author="Karasawa" box="[1140,1431,472,498]" pageId="3" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 a) The family Goneplacidae MacLeay, 1838 (Crustacea: Decapoda: Brachyura): systematics, phylogeny and fossil records. Paleontological Research, 7 (2), 129 - 151." type="journal article" year="2003" yearSuffix="a">Karasawa & Kato 2003a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EEA2FDA49BD9FD8E" author="Crosnier" box="[151,396,512,538]" pageId="3" pageNumber="53" refString="Crosnier, A. & Ng, P. K. L. (2004) Remarques sur le genre Intesius (Crustacea, Decapoda, Brachyura, Goneplacidae) et description de deux especes nouvelles. Zoosystema, 26 (2), 263 - 277." type="journal article" year="2004">Crosnier & Ng 2004</bibRefCitation>
|
||
) and separating them makes little sense. The lack of any explanation or justification by
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EEA2FD839B17FDD6" author="Stevcic" box="[151,322,551,578]" pageId="3" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Stevčić (2005)</bibRefCitation>
|
||
makes these taxonomic actions and transfers difficult to accept, refute or even discuss.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC4FFA7EEF3FDF49B3DFBB6" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EEF3FDF4986AFDFE" author="Karasawa" box="[198,575,592,618]" pageId="3" pageNumber="53" refString="Karasawa, H. & Schweitzer, M. (2006) A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology, 75 (1 / 2), 23 - 73." type="journal article" year="2006">Karasawa & Schweitzer (2006)</bibRefCitation>
|
||
, in another reappraisal of the
|
||
<taxonomicName id="4C204D00FFC4FFA7ED93FDF49E1FFDFE" box="[934,1098,592,618]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
, this time comparing it with the Xanthoidea, followed some aspects of Stevčić’s (2005) system, but argued that the Mathildellinae was actually a portunoid and recognized it as a separate family there. They regarded the Goneplacinae as synonymous with the
|
||
<taxonomicName id="4C204D00FFC4FFA7EF7EFD6C9858FD76" box="[331,525,712,738]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
; and placed both the
|
||
<taxonomicName id="4C204D00FFC4FFA7ED3AFD6C99EFFD76" box="[783,954,712,738]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
(as a family) and the
|
||
<taxonomicName id="4C204D00FFC4FFA7EAF3FD6C9F1AFD76" box="[1222,1359,712,738]" class="Malacostraca" family="Geryonidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Geryonidae</taxonomicName>
|
||
in the Portunoidea. The transfer of the Mathildellinae is difficult to contend with as with regards to the general form of the carapace, sternal, abdominal and gonopodal characters, it has many affinities with the goneplacids. Other parts of their Goneplacoidea appear to be more reasonable, with the
|
||
<taxonomicName id="4C204D00FFC4FFA7EA27FCE49EE5FCCE" box="[1042,1200,832,858]" class="Malacostraca" family="Euryplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Euryplacidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EA89FCE49FCDFCCE" box="[1212,1432,832,858]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2FCCC9B12FC16" box="[151,327,872,898]" class="Malacostraca" family="Trogloplacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Trogloplacidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EFB0FCCC98E8FC16" authority="Miers, 1886" authorityName="Miers" authorityYear="1886" box="[389,701,872,898]" class="Malacostraca" family="Hexapodidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">
|
||
Hexapodidae
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EC1CFCCC98E8FC16" author="Miers" box="[553,701,872,898]" pageId="3" pageNumber="54" refString="Miers, E. J. (1886) Part II. Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 76. In: Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, N. R., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. & c. Regius Professor of Natural history in the University of Edinburgh of the civilian scientific staff on board and now of John Murray one of the naturalists of the Expedition. Zoology. Published by Order of Her Majesty's Government. London, Edinburgh and Dublin, HMSO, 17, i - l + 1 - 362, pls. 1 - 29." type="journal article" year="1886">Miers, 1886</bibRefCitation>
|
||
</taxonomicName>
|
||
. More difficult to accommodate is their transfer of the genus
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2FC349B9FFC3E" authority="Bouvier, 1922" authorityName="Bouvier" authorityYear="1922" box="[151,458,912,938]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EEA2FC349B46FC3D" box="[151,275,912,937]" italics="true" pageId="3" pageNumber="42">Progeryon</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EF2BFC349B9FFC3E" author="Bouvier" box="[286,458,912,938]" pageId="3" pageNumber="52" refString="Bouvier, E. - L. (1922) Observations complementaires sur les crustaces decapodes (abstraction faite des Carides) provenant des Campagnes de S. A. S le Prince de Monaco. Resultats Campagne Scientifiques, Monaco, 62, 1 - 106, pl. 1 - 6." type="journal article" year="1922">Bouvier, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
, to its own family (i.e. the
|
||
<taxonomicName id="4C204D00FFC4FFA7ED29FC349E23FC3E" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[796,1142,911,938]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">
|
||
Progeryonidae
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EDE7FC2B9E23FC3E" author="Stevcic" box="[978,1142,911,938]" pageId="3" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
) and superfamily which they justified stating that the form of the male thoracic sternum is different from those in other portunoids, including mathildellids. They left the matter of
|
||
<taxonomicName id="4C204D00FFC4FFA7ECE4FC449E79FC6E" authority="Kossmann, 1878" authorityName="Kossmann" authorityYear="1878" box="[721,1068,992,1018]" class="Malacostraca" family="Progeryonidae" genus="Paragalene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7ECE4FC44990FFC6D" box="[721,858,992,1017]" italics="true" pageId="3" pageNumber="42">Paragalene</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC4FFA7ED56FC449E79FC6E" author="Kossmann" box="[867,1068,992,1018]" pageId="3" pageNumber="53" refString="Kossmann, R. (1878). Kurze Notizen ber einige neue Crustaceen sowie uber neue Fundorte einiger bereits beschriebenen. Archiv fur Naturgeschichte, 44 (1), 251 - 258" type="journal article" year="1878">Kossmann, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
, and the Paragaleninii
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EB0BFC7B9A81FBB6" author="Stevcic" pageId="3" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, unresolved,
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC4FFA7EEF3FB949B39FAAE" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
|
||
Most recently, Castro (in press) reviewed and revised the members of the species which have been placed in Goneplacinae
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EF54FBFC984EFBE6" author="MacLeay" box="[353,539,1112,1138]" lastPageId="3" lastPageNumber="54" pageId="3" pageNumber="53" refString="MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In A. Smith, Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa': together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe, Published under the Authority of the Lords Commissioners of Her Majesty's Treasury, Invertebratae: [1849] pp. 53 - 71, pls 2, 3." type="journal article" year="1838">MacLeay, 1838</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC4FFA7EC69FBFC994FFBE6" box="[604,794,1112,1138]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="subFamily" subFamily="Carcinoplacinae">Carcinoplacinae</taxonomicName>
|
||
H.
|
||
<bibRefCitation id="EFB14B72FFC4FFA7ED7CFBFC9E1EFBE6" author="Milne" box="[841,1099,1112,1138]" pageId="3" pageNumber="54" refString="Milne Edwards, H. (1852) De la famille des Ocypodides (Ocypodidae). Second Memoire, in, Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences Naturelles (Zoologie), 18, 128 - 166 pls. 3, 4." type="journal article" year="1852">Milne Edwards, 1852</bibRefCitation>
|
||
, and supported Karasawa & Kato’s (2003) synonymisation of these two subfamilies. He also recognised the Mathildellinae as a separate subfamily within the
|
||
<taxonomicName id="4C204D00FFC4FFA7EFA0FB0C986CFB56" box="[405,569,1192,1218]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
, and synonymised Notonycidae under the Goneplacinae. As to the status of genera like
|
||
<taxonomicName id="4C204D00FFC4FFA7EF08FB74988AFB7E" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[317,735,1232,1258]" class="Malacostraca" family="Goneplacidae" genus="Bathyplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF08FB749BE1FB7D" box="[317,436,1232,1257]" italics="true" pageId="3" pageNumber="42">Bathyplax</emphasis>
|
||
A. Milne-Edwards, 1880
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7ECDEFB749932FB7D" box="[747,871,1232,1257]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7ECDEFB749932FB7D" box="[747,871,1232,1257]" italics="true" pageId="3" pageNumber="42">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7ED40FB7499ABFB7D" box="[885,1022,1232,1257]" class="Malacostraca" family="Progeryonidae" genus="Paragalene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7ED40FB7499ABFB7D" box="[885,1022,1232,1257]" italics="true" pageId="3" pageNumber="42">Paragalene</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EA39FB749E2CFB7D" box="[1036,1145,1232,1257]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EA39FB749E2CFB7D" box="[1036,1145,1232,1257]" italics="true" pageId="3" pageNumber="42">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
, etc., he argued that they were not certainly not goneplacines in the strict sense, but may perhaps be retained in the
|
||
<taxonomicName id="4C204D00FFC4FFA7EA87FB5C9F06FA86" box="[1202,1363,1272,1298]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
<emphasis id="B954EA91FFC4FFA7EB6EFB5C9A90FAAD" italics="true" pageId="3" pageNumber="42">
|
||
<taxonomicName id="4C204D00FFC4FFA7EB6EFB5C9FC9FA85" box="[1371,1436,1272,1297]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
provisionally.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC4FFA7EEF3FAEC9926F936" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
|
||
How does
|
||
<taxonomicName id="4C204D00FFC4FFA7EF70FAEC9B94FAF5" box="[325,449,1352,1377]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF70FAEC9B94FAF5" box="[325,449,1352,1377]" italics="true" pageId="3" pageNumber="42">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
fit into all these recent developments? The problem is that many of the subfamilies or families defined by the earlier workers have omitted key characters associated with the thoracic sternum, abdominal locking mechanism, position of the penis and associated structures. The most comprehensive study (Castro, in press) has many of the characters that are deemed useful in a modern classification but it is mostly restricted to the Goneplacinae
|
||
<emphasis id="B954EA91FFC4FFA7EC31FA4C98CCF995" box="[516,665,1512,1537]" italics="true" pageId="3" pageNumber="42">
|
||
<taxonomicName id="4C204D00FFC4FFA7EC31FA4C9810F995" box="[516,581,1512,1537]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, although he does briefly discuss many of the other “goneplacid allies”. To resolve the systematic position of
|
||
<taxonomicName id="4C204D00FFC4FFA7EC97F9B49948F9BD" box="[674,797,1552,1577]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EC97F9B49948F9BD" box="[674,797,1552,1577]" italics="true" pageId="3" pageNumber="42">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
, it was important that representative genera of the various supra-generic groupings discussed above be re-examined. To this effect, we examined in detail representative genera from the Goneplacinae s. str., Mathildellinae s. str., as well as two of the problematic genera,
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2F92C9B46F935" box="[151,275,1672,1697]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EEA2F92C9B46F935" box="[151,275,1672,1697]" italics="true" pageId="3" pageNumber="42">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EF79F92C9BECF935" box="[332,441,1672,1697]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF79F92C9BECF935" box="[332,441,1672,1697]" italics="true" pageId="3" pageNumber="42">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
, and compared them with
|
||
<taxonomicName id="4C204D00FFC4FFA7ECC4F92C9939F935" box="[753,876,1672,1697]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7ECC4F92C9939F935" box="[753,876,1672,1697]" italics="true" pageId="3" pageNumber="42">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC4FFA0EEF3F9149B88FE66" blockId="3.[151,1437,151,2018]" lastBlockId="4.[151,1437,152,2018]" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42">
|
||
The position of the press button of the abdominal locking mechanism (see
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EA0AF9149F3BF95E" author="Guinot" box="[1087,1390,1712,1738]" pageId="3" pageNumber="53" refString="Guinot, D. & Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot & Bouchard 1998</bibRefCitation>
|
||
) on or along thoracic sternite 5 is important. While all of the genera considered here have the button on thoracic sternite 5 at the lateral edges of the sterno-abdominal cavity,there are two distinct patterns. In one group with
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2F88C9BF3F8D6" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[151,422,1832,1858]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EEA2F88C9B5CF8D5" box="[151,265,1832,1857]" italics="true" pageId="3" pageNumber="42">Goneplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EF24F88C9BF3F8D6" author="Leach" box="[273,422,1832,1858]" pageId="3" pageNumber="53" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (editor), The Edinburgh Encyclopaedia, 7, 383 - 437, pl. 221." type="journal article" year="1814">Leach, 1814</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC4FFA7EFD3F88C99FDF8D6" authority="H. Milne Edwards, 1852" authorityName="H. Milne Edwards" authorityYear="1852" box="[486,936,1832,1858]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EFD3F88C9822F8D5" box="[486,631,1832,1857]" italics="true" pageId="3" pageNumber="42">Carcinoplax</emphasis>
|
||
H.
|
||
<bibRefCitation id="EFB14B72FFC4FFA7EC93F88C99FDF8D6" author="Milne" box="[678,936,1832,1858]" pageId="3" pageNumber="54" refString="Milne Edwards, H. (1852) De la famille des Ocypodides (Ocypodidae). Second Memoire, in, Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences Naturelles (Zoologie), 18, 128 - 166 pls. 3, 4." type="journal article" year="1852">Milne Edwards, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
, the press button is on the anterior part of sternite 5, close to sternite 4 (
|
||
<figureCitation id="131B2A06FFC4FFA7EC32F8F49835F8FE" box="[519,608,1872,1898]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="3" pageNumber="42">Fig. 12</figureCitation>
|
||
B, C). In all the other genera—
|
||
<taxonomicName id="4C204D00FFC4FFA7EDD7F8F49E3EF8FD" box="[994,1131,1872,1897]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EDD7F8F49E3EF8FD" box="[994,1131,1872,1897]" italics="true" pageId="3" pageNumber="42">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EA48F8F49E8DF8FD" box="[1149,1240,1872,1897]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EA48F8F49E8DF8FD" box="[1149,1240,1872,1897]" italics="true" pageId="3" pageNumber="42">Intesius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EADEF8F49FC1F8FD" box="[1259,1428,1872,1897]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EADEF8F49FC1F8FD" box="[1259,1428,1872,1897]" italics="true" pageId="3" pageNumber="42">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2F8DC9B46F805" box="[151,275,1912,1937]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EEA2F8DC9B46F805" box="[151,275,1912,1937]" italics="true" pageId="3" pageNumber="42">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC4FFA7EF16F8DC9BC5F805" box="[291,400,1912,1937]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF16F8DC9BC5F805" box="[291,400,1912,1937]" italics="true" pageId="3" pageNumber="42">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EFFEF8DC9813F805" box="[459,582,1912,1937]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EFFEF8DC9813F805" box="[459,582,1912,1937]" italics="true" pageId="3" pageNumber="42">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
, the press button is on the posterior part of sternite 5, close to sternite 6 (
|
||
<figureCitation id="131B2A06FFC4FFA7EEAAF8049AA0F82E" box="[159,245,1952,1978]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="3" pageNumber="42">Fig. 12</figureCitation>
|
||
A, D–H). This correlates somewhat with the extent of the sterno-abdominal cavity in some cases. In
|
||
<taxonomicName id="4C204D00FFC4FFA7EEA2F86C9B5CF875" box="[151,265,1992,2017]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EEA2F86C9B5CF875" box="[151,265,1992,2017]" italics="true" pageId="3" pageNumber="42">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC4FFA7EF71F86C9B80F875" box="[324,469,1992,2017]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC4FFA7EF71F86C9B80F875" box="[324,469,1992,2017]" italics="true" pageId="3" pageNumber="42">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, the sterno-abdominal cavity is usually relatively deep, and extends almost to the anterior edge of thoracic sternite 4 (
|
||
<figureCitation id="131B2A06FFC3FFA0EC7AFF3C98F0FF26" box="[591,677,152,178]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Figs. 9</figureCitation>
|
||
B, C, 12B, C). In
|
||
<taxonomicName id="4C204D00FFC3FFA0EDB3FF3C9E5BFF25" box="[902,1038,152,177]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EDB3FF3C9E5BFF25" box="[902,1038,152,177]" italics="true" pageId="4" pageNumber="43">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0EA14FF3C9E29FF25" box="[1057,1148,152,177]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA14FF3C9E29FF25" box="[1057,1148,152,177]" italics="true" pageId="4" pageNumber="43">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EA89FF3C9F30FF25" box="[1212,1381,152,177]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA89FF3C9F30FF25" box="[1212,1381,152,177]" italics="true" pageId="4" pageNumber="43">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
, the sterno-abdominal cavity is usually relatively shallow, and extends only to the middle part of thoracic sternite 4 (
|
||
<figureCitation id="131B2A06FFC3FFA0EEAAFF4C9AA1FE96" box="[159,244,232,258]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Figs. 9</figureCitation>
|
||
D–F, 12D–F). The position of the sterno-abdominal cavity in
|
||
<taxonomicName id="4C204D00FFC3FFA0EDDBFF4C9E3FFE95" box="[1006,1130,232,257]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EDDBFF4C9E3FFE95" box="[1006,1130,232,257]" italics="true" pageId="4" pageNumber="43">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
, however, is like that of
|
||
<taxonomicName id="4C204D00FFC3FFA0EEA2FEB49B5CFEBD" box="[151,265,272,297]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EEA2FEB49B5CFEBD" box="[151,265,272,297]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EF75FEB49B84FEBD" box="[320,465,272,297]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EF75FEB49B84FEBD" box="[320,465,272,297]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, almost reaching the anterior edge of sternite 4 (
|
||
<figureCitation id="131B2A06FFC3FFA0EA31FEB49E1DFEBE" box="[1028,1096,272,298]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
G) but the cavity is relatively shallower (
|
||
<figureCitation id="131B2A06FFC3FFA0EF2DFE9C9B3BFEC6" box="[280,366,312,338]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
G). The position of the sterno-abdominal cavity in
|
||
<taxonomicName id="4C204D00FFC3FFA0ED8AFE9C9E79FEC5" box="[959,1068,312,337]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED8AFE9C9E79FEC5" box="[959,1068,312,337]" italics="true" pageId="4" pageNumber="43">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
is intermediate between that of
|
||
<taxonomicName id="4C204D00FFC3FFA0EEA2FEC49B5CFEED" box="[151,265,352,377]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EEA2FEC49B5CFEED" box="[151,265,352,377]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0EF22FEC49BFDFEED" box="[279,424,352,377]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EF22FEC49BFDFEED" box="[279,424,352,377]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EFD7FEC4980BFEED" box="[482,606,352,377]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EFD7FEC4980BFEED" box="[482,606,352,377]" italics="true" pageId="4" pageNumber="43">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
on one hand, and
|
||
<taxonomicName id="4C204D00FFC3FFA0ED01FEC499E8FEED" box="[820,957,352,377]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED01FEC499E8FEED" box="[820,957,352,377]" italics="true" pageId="4" pageNumber="43">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0EDF9FEC49E72FEED" box="[972,1063,352,377]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EDF9FEC49E72FEED" box="[972,1063,352,377]" italics="true" pageId="4" pageNumber="43">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EA6AFEC49F5CFEED" box="[1119,1289,352,377]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA6AFEC49F5CFEED" box="[1119,1289,352,377]" italics="true" pageId="4" pageNumber="43">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
on the other (
|
||
<figureCitation id="131B2A06FFC3FFA0EEAAFE2C9AB2FE36" box="[159,231,392,418]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
H), although the cavity itself is relatively deep (
|
||
<figureCitation id="131B2A06FFC3FFA0ED10FE2C9928FE36" box="[805,893,392,418]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
H).
|
||
<taxonomicName id="4C204D00FFC3FFA0ED9FFE2C9E73FE35" box="[938,1062,392,417]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED9FFE2C9E73FE35" box="[938,1062,392,417]" italics="true" pageId="4" pageNumber="43">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
stands out in that it has a deep sterno-abdominal cavity which is relatively narrow transversely (
|
||
<figureCitation id="131B2A06FFC3FFA0ED90FE1499AEFE5E" box="[933,1019,432,458]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
A) and almost reaches the anterior edge of sternite 4 (
|
||
<figureCitation id="131B2A06FFC3FFA0EF44FE7C9BE2FE66" box="[369,439,472,498]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC3FFA0EEF3FDA49815FCA6" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="43">
|
||
A note on the use of the extent of the sterno-abdominal cavity as a character is useful. In some taxa, the anterior-most point of the cavity extends beyond the tip of the telson, even when the male abdomen is fully closed and locked into position by the press button mechanism. It is most distinct in genera like
|
||
<taxonomicName id="4C204D00FFC3FFA0EB16FDF49FC0FDFD" box="[1315,1429,592,617]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EB16FDF49FC0FDFD" box="[1315,1429,592,617]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0EEA2FDDC9B7CFD05" box="[151,297,632,657]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EEA2FDDC9B7CFD05" box="[151,297,632,657]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EF56FDDC9B8BFD05" box="[355,478,632,657]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EF56FDDC9B8BFD05" box="[355,478,632,657]" italics="true" pageId="4" pageNumber="43">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B2A06FFC3FFA0EFDAFDDC9860FD06" box="[495,565,632,658]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
A–C), and to a lesser extent by genera like
|
||
<taxonomicName id="4C204D00FFC3FFA0EA02FDDC9EE6FD05" box="[1079,1203,632,657]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA02FDDC9EE6FD05" box="[1079,1203,632,657]" italics="true" pageId="4" pageNumber="43">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EADBFDDC9F0EFD05" box="[1262,1371,632,657]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EADBFDDC9F0EFD05" box="[1262,1371,632,657]" italics="true" pageId="4" pageNumber="43">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B2A06FFC3FFA0EB59FDDC9AF0FD2E" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
G, H). It is not so obvious in
|
||
<taxonomicName id="4C204D00FFC3FFA0EFC0FD04982BFD2D" box="[501,638,672,697]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EFC0FD04982BFD2D" box="[501,638,672,697]" italics="true" pageId="4" pageNumber="43">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0ECB9FD0498B2FD2D" box="[652,743,672,697]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ECB9FD0498B2FD2D" box="[652,743,672,697]" italics="true" pageId="4" pageNumber="43">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0ED2AFD04999DFD2D" box="[799,968,672,697]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED2AFD04999DFD2D" box="[799,968,672,697]" italics="true" pageId="4" pageNumber="43">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131B2A06FFC3FFA0EDECFD049E48FD2E" box="[985,1053,672,698]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="4" pageNumber="43">Fig. 9</figureCitation>
|
||
D–F) because the sterno-abdominal cavity is relatively shallower and the male abdomen always locks onto the sternum very tightly. The gap is presumably to enable the animals to defecate without moving the rest of the abdomen or even the telson (see also
|
||
<bibRefCitation id="EFB14B72FFC3FFA0EF31FCBC9866FCA6" author="Guinot" box="[260,563,792,818]" pageId="4" pageNumber="53" refString="Guinot, D. & Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot & Bouchard 1998</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC3FFA0EEF3FCE49AB4FAAE" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="43">
|
||
The structure of the sutures 6/7 and 7/8 are useful in understanding the relationships between the groups. In the two species of
|
||
<taxonomicName id="4C204D00FFC3FFA0EFA7FCCC9851FC15" box="[402,516,872,897]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EFA7FCCC9851FC15" box="[402,516,872,897]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
examined in this study, suture 6/7 can be regarded as complete even though it becomes rather shallow medially and appears interrupted (
|
||
<figureCitation id="131B2A06FFC3FFA0ED6CFC3499E6FC3E" box="[857,947,912,938]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
B). In
|
||
<taxonomicName id="4C204D00FFC3FFA0EA36FC349EC1FC3D" box="[1027,1172,912,937]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA36FC349EC1FC3D" box="[1027,1172,912,937]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, the same suture also appears to be complete but there is a narrow median gap (
|
||
<figureCitation id="131B2A06FFC3FFA0ED08FC1C99C4FC46" box="[829,913,952,978]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
C) and so may be regarded as interrupted. In the case of
|
||
<taxonomicName id="4C204D00FFC3FFA0EF16FC449BCBFC6D" box="[291,414,992,1017]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EF16FC449BCBFC6D" box="[291,414,992,1017]" italics="true" pageId="4" pageNumber="43">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
, this suture curves anteriorly, becoming very shallow medially so it is not as easy to ascertain if this is truly complete or a case of incipient interruption (
|
||
<figureCitation id="131B2A06FFC3FFA0ED99FBAC9E55FBB6" box="[940,1024,1032,1058]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
A). In the Goneplacinae
|
||
<taxonomicName id="4C204D00FFC3FFA0EB2BFBAC9AEDFBDE" authority="Castro" authorityName="Castro" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="sensu">
|
||
<emphasis id="B954EA91FFC3FFA0EB2BFBAC9F0AFBB5" box="[1310,1375,1032,1057]" italics="true" pageId="4" pageNumber="43">sensu</emphasis>
|
||
Castro
|
||
</taxonomicName>
|
||
(in press), suture 6/7 varies between complete and interrupted. In all the other genera examined here, suture 6/7 is clearly complete (
|
||
<figureCitation id="131B2A06FFC3FFA0EC31FBFC980CFBE6" box="[516,601,1112,1138]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
D–H). With regards to suture 7/8, in
|
||
<taxonomicName id="4C204D00FFC3FFA0EA3AFBFC9ED4FBE5" box="[1039,1153,1112,1137]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EA3AFBFC9ED4FBE5" box="[1039,1153,1112,1137]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
, it is prominently interrupted medially (
|
||
<figureCitation id="131B2A06FFC3FFA0EF56FB249BE9FB0E" box="[355,444,1152,1178]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
B). In
|
||
<taxonomicName id="4C204D00FFC3FFA0EC3CFB2498CFFB0D" box="[521,666,1152,1177]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EC3CFB2498CFFB0D" box="[521,666,1152,1177]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, the suture appears to be complete but there is a narrow median interruption (
|
||
<figureCitation id="131B2A06FFC3FFA0EF01FB0C9BD2FB56" box="[308,391,1192,1218]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="4" pageNumber="43">Figs. 7</figureCitation>
|
||
E, 12C) and so may be regarded as interrupted as well. According to Castro (in press), while most Goneplacinae are like
|
||
<taxonomicName id="4C204D00FFC3FFA0EC06FB7498F0FB7D" box="[563,677,1232,1257]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EC06FB7498F0FB7D" box="[563,677,1232,1257]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0ECD6FB749921FB7D" box="[739,884,1232,1257]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ECD6FB749921FB7D" box="[739,884,1232,1257]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, there some taxa which have suture 7/8 complete. In all the other genera examined here (including
|
||
<taxonomicName id="4C204D00FFC3FFA0ED10FB5C99CBFA85" box="[805,926,1272,1297]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED10FB5C99CBFA85" box="[805,926,1272,1297]" italics="true" pageId="4" pageNumber="43">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
), suture 7/8 is clearly complete (
|
||
<figureCitation id="131B2A06FFC3FFA0EB1FFB5C9F2AFA86" box="[1322,1407,1272,1298]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="4" pageNumber="43">Fig. 12</figureCitation>
|
||
A, D–H).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC3FFA0EEF3FAEC98D1F966" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="43">
|
||
Also of interest is the arrangement of thoracic sternite
|
||
<quantity id="4CD89B66FFC3FFA0ED61FAEC99D6FAF6" box="[852,899,1352,1378]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="4" pageNumber="43" unit="in" value="8.0">8 in</quantity>
|
||
relation to the adjacent coxa of P5 and male abdomen.
|
||
<bibRefCitation id="EFB14B72FFC3FFA0EF27FAD49851FA1E" author="Guinot" box="[274,516,1392,1418]" pageId="4" pageNumber="53" refString="Guinot, D. (1969 b) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae (suite). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 41 (3), 507 - 528." type="journal article" year="1969" yearSuffix="b">Guinot (1969b: 520)</bibRefCitation>
|
||
first highlighted this interesting character in her attempts to better understand goneplacoid relationships. In genera like
|
||
<taxonomicName id="4C204D00FFC3FFA0ECB6FA3C98A0FA25" box="[643,757,1432,1457]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ECB6FA3C98A0FA25" box="[643,757,1432,1457]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0ED07FA3C9991FA25" box="[818,964,1432,1457]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ED07FA3C9991FA25" box="[818,964,1432,1457]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, part of sternite 8 is still clearly visible even after the abdomen is completely closed. In species of
|
||
<emphasis id="B954EA91FFC3FFA0ED73FA649E07FA4D" box="[838,1106,1472,1497]" italics="true" pageId="4" pageNumber="43">
|
||
<taxonomicName id="4C204D00FFC3FFA0ED73FA649E55FA4D" box="[838,1024,1472,1497]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="sensu">
|
||
Goneplax
|
||
<taxonomicNameLabel id="A26757EAFFC3FFA0ED8AFA649E55FA4D" box="[959,1024,1472,1497]" pageId="4" pageNumber="43" sensu="stricto">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, the exposed part of sternite 8 is quite substantial (
|
||
<figureCitation id="131B2A06FFC3FFA0EFA3FA4C9B89F996" box="[406,476,1512,1538]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="4" pageNumber="43">Fig. 7</figureCitation>
|
||
C) (see also
|
||
<bibRefCitation id="EFB14B72FFC3FFA0EC58FA4C9941F996" author="Guinot" box="[621,788,1512,1538]" pageId="4" pageNumber="53" refString="Guinot, D. (1969 b) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae (suite). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 41 (3), 507 - 528." type="journal article" year="1969" yearSuffix="b">Guinot 1969b</bibRefCitation>
|
||
: Figs. 63, 64). In at least some of the Indo-West Pacific species of “
|
||
<taxonomicName id="4C204D00FFC3FFA0EF15F9B49BC7F9BD" box="[288,402,1552,1577]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EF15F9B49BC7F9BD" box="[288,402,1552,1577]" italics="true" pageId="4" pageNumber="43">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
”, the exposed part of sternite 8 is relatively small (
|
||
<figureCitation id="131B2A06FFC3FFA0EDD8F9B49E1BF9BE" box="[1005,1102,1552,1578]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="4" pageNumber="43">Figs. 10</figureCitation>
|
||
B, 11b), while in genera like
|
||
<taxonomicName id="4C204D00FFC3FFA0EEA2F99C9B7DF9C5" box="[151,296,1592,1617]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EEA2F99C9B7DF9C5" box="[151,296,1592,1617]" italics="true" pageId="4" pageNumber="43">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, the exposed part of sternite 8 varies from small (
|
||
<figureCitation id="131B2A06FFC3FFA0ED43F99C9982F9C6" box="[886,983,1592,1618]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="4" pageNumber="43">Figs. 10</figureCitation>
|
||
C, 11C) to relatively large (see
|
||
<bibRefCitation id="EFB14B72FFC3FFA0EB7FF99C9AB0F9EE" author="Guinot" pageId="4" pageNumber="53" refString="Guinot, D. (1969 b) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae (suite). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 41 (3), 507 - 528." type="journal article" year="1969" yearSuffix="b">Guinot 1969b</bibRefCitation>
|
||
: Figs. 60–62, 65). In genera like
|
||
<taxonomicName id="4C204D00FFC3FFA0EC53F9C498BBF9ED" box="[614,750,1632,1657]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EC53F9C498BBF9ED" box="[614,750,1632,1657]" italics="true" pageId="4" pageNumber="43">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC3FFA0ECC8F9C4990DF9ED" box="[765,856,1632,1657]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ECC8F9C4990DF9ED" box="[765,856,1632,1657]" italics="true" pageId="4" pageNumber="43">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0EDA4F9C49E6FF9ED" box="[913,1082,1632,1657]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EDA4F9C49E6FF9ED" box="[913,1082,1632,1657]" italics="true" pageId="4" pageNumber="43">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
, the exposed part of sternite 8 is either completely covered by the abdomen (
|
||
<figureCitation id="131B2A06FFC3FFA0EC84F92C9944F936" box="[689,785,1672,1698]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="4" pageNumber="43">Figs. 10</figureCitation>
|
||
D, F, 11D, F) or just visible with only a small part of the edge visible (
|
||
<figureCitation id="131B2A06FFC3FFA0EF01F9149BC1F95E" box="[308,404,1712,1738]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="4" pageNumber="43">Figs. 10</figureCitation>
|
||
E, 11E). In
|
||
<taxonomicName id="4C204D00FFC3FFA0EC2EF91498C2F95D" box="[539,663,1712,1737]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0EC2EF91498C2F95D" box="[539,663,1712,1737]" italics="true" pageId="4" pageNumber="43">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC3FFA0ECE5F9149968F95D" box="[720,829,1712,1737]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC3FFA0ECE5F9149968F95D" box="[720,829,1712,1737]" italics="true" pageId="4" pageNumber="43">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
, the lateral part of sternite 8 completely covered by the male abdomen (
|
||
<figureCitation id="131B2A06FFC3FFA0EF48F97C9B8EF966" box="[381,475,1752,1778]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="4" pageNumber="43">Figs. 10</figureCitation>
|
||
G, H, 11G, H).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC3FFA0EEF3F8A49898F806" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="43">The position of the penis is of fundamental importance (Guinot, D., pers. comm.). This character is closely associated with the shape of episternite 7 and the lateral margin of abdominal somite 3. In all cases, the exposed part of the penis is protected by abdominal somite 3 when the abdomen is closed. How this is accomplished is what distinguishes each of the groups.</paragraph>
|
||
<paragraph id="8B9F3683FFC3FFA1EEF3F8049927FE5E" blockId="4.[151,1437,152,2018]" lastBlockId="5.[151,1437,152,2018]" lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
|
||
In the
|
||
<taxonomicName id="4C204D00FFC3FFA0EF21F8049BE0F82E" box="[276,437,1952,1978]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
<emphasis id="B954EA91FFC3FFA0EF8BF8049806F82D" box="[446,595,1952,1977]" italics="true" pageId="4" pageNumber="43">
|
||
<taxonomicName id="4C204D00FFC3FFA0EF8BF8049BAAF82D" box="[446,511,1952,1977]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, the penis leaves the coxa at its base, but continues under the sternum for a short distance, only exiting and becoming completely exposed inside the sterno-abdominal cavity after the coxal condyle (
|
||
<figureCitation id="131B2A06FFC2FFA1EF40FF3C9B9DFF26" box="[373,456,152,178]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
C, D). Episternite 7 is expanded laterally to partially overlap the entire inner margin of the coxa of P5 (
|
||
<figureCitation id="131B2A06FFC2FFA1EF44FF649B97FF4E" box="[369,450,192,218]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="5" pageNumber="44">Figs. 7</figureCitation>
|
||
D, 10B, 13C) or at least most of the anterior part (
|
||
<figureCitation id="131B2A06FFC2FFA1EA25FF649E24FF4E" box="[1040,1137,192,218]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Figs. 10</figureCitation>
|
||
C, 12D). The lateral edge of abdominal somite 3 never reaches the coxa of P5, at most touching the condyle (
|
||
<figureCitation id="131B2A06FFC2FFA1EAA3FF4C9EBFFE96" box="[1174,1258,232,258]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="5" pageNumber="44">Figs. 7</figureCitation>
|
||
D, 10B, C). In
|
||
<taxonomicName id="4C204D00FFC2FFA1EEA2FEB49B5CFEBD" box="[151,265,272,297]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EEA2FEB49B5CFEBD" box="[151,265,272,297]" italics="true" pageId="5" pageNumber="44">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
, the penis is clearly of the coxosternal
|
||
<typeStatus id="549B8821FFC2FFA1ECE0FEB4995EFEBE" box="[725,779,272,298]" pageId="5" pageNumber="44">type</typeStatus>
|
||
, exiting prominently between sternites 7 and 8 at a submedially position (
|
||
<figureCitation id="131B2A06FFC2FFA1EF47FE9C9B97FEC6" box="[370,450,312,338]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="5" pageNumber="44">Figs. 7</figureCitation>
|
||
F, 13C). In
|
||
<taxonomicName id="4C204D00FFC2FFA1EC73FE9C9882FEC5" box="[582,727,312,337]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EC73FE9C9882FEC5" box="[582,727,312,337]" italics="true" pageId="5" pageNumber="44">Carcinoplax</emphasis>
|
||
</taxonomicName>
|
||
, the penis exits just after the condyle, at the beginning of the junction between sternites 7 and 8 (
|
||
<figureCitation id="131B2A06FFC2FFA1EC07FEC498D3FEEE" box="[562,646,352,378]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
D). As a result, the distal part of the penis, where it comes out of the coxa, is protected by the expanded part of episternite 7, and the exposed part of the penis, where it exits between sternites 7 and 8, is protected by the closed abdomen.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC2FFA1EEF3FE7C9B09FD06" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="44">
|
||
The same situation is true for
|
||
<taxonomicName id="4C204D00FFC2FFA1EC05FE7C98C8FE65" box="[560,669,472,497]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EC05FE7C98C8FE65" box="[560,669,472,497]" italics="true" pageId="5" pageNumber="44">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
, i.e. the penis is clearly coxosternal, exiting between sternites 7 and 8 at a submedian position (
|
||
<figureCitation id="131B2A06FFC2FFA1EC3BFDA49831FD8E" box="[526,612,512,538]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
F), episternite 7 is expanded laterally to partially overlap most of the anterior part (
|
||
<figureCitation id="131B2A06FFC2FFA1EF0CFD8C9BDBFDD6" box="[313,398,552,578]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
H) and the lateral edge of male abdominal somite 3 does not reach the coxa of P5 or its condyle (
|
||
<figureCitation id="131B2A06FFC2FFA1EF32FDF49B3EFDFE" box="[263,363,592,618]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Figs. 10</figureCitation>
|
||
H, 11F). The penis is therefore partially protected by the expanded episternite 7 and the closed abdomen.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC2FFA1EEF3FD049E4AFCCE" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="44">
|
||
In
|
||
<taxonomicName id="4C204D00FFC2FFA1EED3FD049B3AFD2D" box="[230,367,672,697]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EED3FD049B3AFD2D" box="[230,367,672,697]" italics="true" pageId="5" pageNumber="44">Mathildella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC2FFA1EF48FD049B8DFD2D" box="[381,472,672,697]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EF48FD049B8DFD2D" box="[381,472,672,697]" italics="true" pageId="5" pageNumber="44">Intesius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC2FFA1EC24FD0498D8FD2D" box="[529,653,672,697]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EC24FD0498D8FD2D" box="[529,653,672,697]" italics="true" pageId="5" pageNumber="44">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
, the penis is clearly coxal, opening near the condyle of the coxa of P5 (
|
||
<figureCitation id="131B2A06FFC2FFA1EEF3FD6C9B4CFD76" box="[198,281,712,738]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
E), episternite 7 is not expanded laterally to any major degree and either does not reach the coxa of P5 (
|
||
<figureCitation id="131B2A06FFC2FFA1EEFCFD549B75FC9E" box="[201,288,752,778]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
D) or only just touches the anterior part (
|
||
<figureCitation id="131B2A06FFC2FFA1ED2CFD549925FC9E" box="[793,880,752,778]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
E–G), and the lateral edge of male abdominal somite 3 covers the inner part of the coxa of P5, including the condyle (
|
||
<figureCitation id="131B2A06FFC2FFA1EDE8FCBC9E64FCA6" box="[989,1073,792,818]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
D–G). In such an arrangement, the exposed part of the penis is protected almost completely by the abdomen.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC2FFA1EEF3FCCC9BE8FB0E" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="44">
|
||
The configuration in
|
||
<taxonomicName id="4C204D00FFC2FFA1EF89FCCC9860FC15" box="[444,565,872,897]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EF89FCCC9860FC15" box="[444,565,872,897]" italics="true" pageId="5" pageNumber="44">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
is unique. The penis is clearly coxal in origin, opening near the condyle of the coxa of P5 (
|
||
<figureCitation id="131B2A06FFC2FFA1EF7BFC349BF7FC3E" box="[334,418,912,938]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
A, B) and the lateral edge of male abdominal somite 3 covers the inner part of the coxa of P5 but not the condyle (
|
||
<figureCitation id="131B2A06FFC2FFA1EFDCFC1C9816FC46" box="[489,579,952,978]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
A). Instead, the condyle of the coxa of P5 is partially covered by the expanded lateral margin of abdominal somite 2 (
|
||
<figureCitation id="131B2A06FFC2FFA1ECFAFC449971FC6E" box="[719,804,992,1018]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="5" pageNumber="44">Fig. 10</figureCitation>
|
||
A). The posterior edge of episternite 7 is very unusual in that is strongly produced posteriorly to form a spur-like projection which partially covers the anterior edge of the coxa of P5 and brackets the exposed penis (
|
||
<figureCitation id="131B2A06FFC2FFA1ECDBFB949916FBDE" box="[750,835,1072,1098]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="5" pageNumber="44">Fig. 13</figureCitation>
|
||
A, B). The penis is therefore protected laterally by the strongly produced episternite 7 and the closed abdomen. Because of this, the penis of
|
||
<taxonomicName id="4C204D00FFC2FFA1EAFFFBFC9F16FBE5" box="[1226,1347,1112,1137]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EAFFFBFC9F16FBE5" box="[1226,1347,1112,1137]" italics="true" pageId="5" pageNumber="44">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
can be described as coxosternal.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC2FFA1EEF3FB0C9F46F8FE" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="44">
|
||
Looking at these datasets altogether (Table 1), there seems little doubt that
|
||
<taxonomicName id="4C204D00FFC2FFA1EA06FB0C9EF9FB55" box="[1075,1196,1192,1217]" class="Malacostraca" family="Vultocinidae" genus="Vultocinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EA06FB0C9EF9FB55" box="[1075,1196,1192,1217]" italics="true" pageId="5" pageNumber="44">Vultocinus</emphasis>
|
||
</taxonomicName>
|
||
needs to be referred to its own family,
|
||
<taxonomicName id="4C204D00FFC2FFA1EF5EFB749850FB7E" box="[363,517,1232,1258]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
. Its combination of male sternal, abdominal and penial structures in particular, is unique. The data sets also strongly indicate that the subfamily Mathildellinae is quite distinct from the Goneplacinae (as defined by Castro, in press), and should be regarded as a distinct family as has been suggested by
|
||
<bibRefCitation id="EFB14B72FFC2FFA1EF27FAEC98DEFAF6" author="Karasawa" box="[274,651,1352,1378]" pageId="5" pageNumber="53" refString="Karasawa, H. & Schweitzer, M. (2006) A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology, 75 (1 / 2), 23 - 73." type="journal article" year="2006">Karasawa & Schweitzer (2006)</bibRefCitation>
|
||
, although we do not agree with its placement in the Portunoidea. The
|
||
<taxonomicName id="4C204D00FFC2FFA1EEF8FAD49B3BFA1E" box="[205,366,1392,1418]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
<emphasis id="B954EA91FFC2FFA1EF40FAD4985CFA1D" box="[373,521,1392,1417]" italics="true" pageId="5" pageNumber="44">
|
||
<taxonomicName id="4C204D00FFC2FFA1EF40FAD49BE3FA1D" box="[373,438,1392,1417]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
therefore now has just two subfamilies, Goneplacinae
|
||
<emphasis id="B954EA91FFC2FFA1EAA1FAD49F7DFA1D" box="[1172,1320,1392,1417]" italics="true" pageId="5" pageNumber="44">
|
||
<taxonomicName id="4C204D00FFC2FFA1EAA1FAD49E80FA1D" box="[1172,1237,1392,1417]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC2FFA1EB5CFAD49BE9FA26" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="subFamily" subFamily="Bathyplacinae">
|
||
Bathyplacinae
|
||
<bibRefCitation id="EFB14B72FFC2FFA1EF29FA339BE9FA26" author="Stevcic" box="[284,444,1431,1458]" pageId="5" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
(see discussion below). We also agree with
|
||
<bibRefCitation id="EFB14B72FFC2FFA1ED88FA3C9F79FA26" author="Karasawa" box="[957,1324,1432,1458]" pageId="5" pageNumber="53" refString="Karasawa, H. & Schweitzer, M. (2006) A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology, 75 (1 / 2), 23 - 73." type="journal article" year="2006">Karasawa & Schweitzer (2006)</bibRefCitation>
|
||
in recognizing the
|
||
<taxonomicName id="4C204D00FFC2FFA1EF21FA649BEAFA4E" box="[276,447,1472,1498]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Progeryonidae</taxonomicName>
|
||
as a distinct family, but certainly not at the superfamily level. The differences cited by them are not valid, but other characters discussed here suggest that it should not be placed in the
|
||
<taxonomicName id="4C204D00FFC2FFA1EB66FA4C9AACF9BE" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
as presently defined. For convenience, in lieu of examining specimens of
|
||
<taxonomicName id="4C204D00FFC2FFA1EA5FF9B49EA6F9BD" box="[1130,1267,1552,1577]" class="Malacostraca" family="Progeryonidae" genus="Paragalene" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EA5FF9B49EA6F9BD" box="[1130,1267,1552,1577]" italics="true" pageId="5" pageNumber="44">Paragalene</emphasis>
|
||
</taxonomicName>
|
||
, we also keep this genus with
|
||
<taxonomicName id="4C204D00FFC2FFA1EF7AF99C9B9EF9C5" box="[335,459,1592,1617]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EF7AF99C9B9EF9C5" box="[335,459,1592,1617]" italics="true" pageId="5" pageNumber="44">Progeryon</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C204D00FFC2FFA1EC2EF99C989FF9C6" box="[539,714,1592,1618]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Progeryonidae</taxonomicName>
|
||
. The data sets also argue for
|
||
<taxonomicName id="4C204D00FFC2FFA1EA2EF99C9EDDF9C5" box="[1051,1160,1592,1617]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC2FFA1EA2EF99C9EDDF9C5" box="[1051,1160,1592,1617]" italics="true" pageId="5" pageNumber="44">Conleyus</emphasis>
|
||
</taxonomicName>
|
||
to be in its own family, the
|
||
<taxonomicName id="4C204D00FFC2FFA1EEF4F9C49B1FF9EE" box="[193,330,1632,1658]" class="Malacostraca" family="Conleyidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Conleyidae</taxonomicName>
|
||
, as has been done by
|
||
<bibRefCitation id="EFB14B72FFC2FFA1EC76F9FB98BBF9EE" author="Stevcic" box="[579,750,1631,1658]" pageId="5" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
although the reasons cited by him (including for his superfamily) are mostly invalid.
|
||
<bibRefCitation id="EFB14B72FFC2FFA1EFD7F92398C5F936" author="Stevcic" box="[482,656,1671,1698]" pageId="5" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
did not examine material and had relied entirely on the original description by Ng & Ng (2003) who did not mention or detail any of the key characters presented in this paper. As with the case of many of Števčić’s (2005) suprageneric taxa, they have been effectively vindicated here by default, even though the characters originally cited by him as justification for these new taxa are incorrect. As a summary, the families may be briefly diagnosed as follows, the ones for
|
||
<taxonomicName id="4C204D00FFC2FFA1EA84F88C9F07F8D6" box="[1201,1362,1832,1858]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
<emphasis id="B954EA91FFC2FFA1EB6EF88C9AB7F8FD" italics="true" pageId="5" pageNumber="44">
|
||
<taxonomicName id="4C204D00FFC2FFA1EB6EF88C9FC9F8D5" box="[1371,1436,1832,1857]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC2FFA1EF17F8F49B98F8FE" box="[290,461,1872,1898]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
<emphasis id="B954EA91FFC2FFA1EFE1F8F4983DF8FD" box="[468,616,1872,1897]" italics="true" pageId="5" pageNumber="44">
|
||
<taxonomicName id="4C204D00FFC2FFA1EFE1F8F49840F8FD" box="[468,533,1872,1897]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, augmenting those by Castro (in press) for these two taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6508FFC2FFACEEF3F8D39BEEFF68" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44" type="diagnosis">
|
||
<paragraph id="8B9F3683FFC2FFA2EEF3F8D39FC8FCCE" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,2018]" lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44">
|
||
<emphasis id="B954EA91FFC2FFA1EEF3F8D39889F805" bold="true" box="[198,732,1911,1937]" pageId="5" pageNumber="44">
|
||
Diagnosis of
|
||
<taxonomicName id="4C204D00FFC2FFA1EF5CF8D39889F805" authority="MacLeay, 1838" authorityName="MacLeay" authorityYear="1838" box="[361,732,1911,1937]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="family">
|
||
Goneplacidae
|
||
<bibRefCitation id="EFB14B72FFC2FFA1EC28F8D39889F805" author="MacLeay" box="[541,732,1911,1937]" lastPageId="5" lastPageNumber="54" pageId="5" pageNumber="53" refString="MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In A. Smith, Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa': together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe, Published under the Authority of the Lords Commissioners of Her Majesty's Treasury, Invertebratae: [1849] pp. 53 - 71, pls 2, 3." type="journal article" year="1838">MacLeay, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
: Dorsal surface of carapace smooth to setose, regions usually discernible but never very prominent or with deep grooves (
|
||
<figureCitation id="131B2A06FFC2FFA1EDAEF80499B6F82E" box="[923,995,1952,1978]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="5" pageNumber="44">Fig. 6</figureCitation>
|
||
A–C). Mandible white to grey in life and preserved (
|
||
<figureCitation id="131B2A06FFC2FFA1EF7EF86C9BC9F876" box="[331,412,1992,2018]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="5" pageNumber="44">Figs. 7</figureCitation>
|
||
A, 8B). Thoracic sternum relatively broad to very broad; episternites 4–7 expanded laterally to form broad plate which covers anterior edge of coxa of P5 (
|
||
<figureCitation id="131B2A06FFC1FFA2ED82FF3C9E5CFF26" box="[951,1033,152,178]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
B, D, 9B, C, 10B, C, 12B, C, 13C, D); surface of sternite 3 without longitudinal median line (
|
||
<figureCitation id="131B2A06FFC1FFA2ED6FFF6499FAFF4E" box="[858,943,192,218]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
B, 9B, C); sternite 7 broad to very broad (
|
||
<figureCitation id="131B2A06FFC1FFA2EEAAFF4C9AA4FE96" box="[159,241,232,258]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
D, 10B, C); sutures 4/5 and 5/6 interrupted medially (
|
||
<figureCitation id="131B2A06FFC1FFA2ED5CFF4C99EEFE96" box="[873,955,232,258]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
E, 12B, C); sutures 6/7 complete and 7/8 distinctly interrupted medially (
|
||
<figureCitation id="131B2A06FFC1FFA2EC39FEB49806FEBE" box="[524,595,272,298]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Fig. 7</figureCitation>
|
||
E), sutures 6/7 and 7/8 with small gap medially suggesting interruption (
|
||
<figureCitation id="131B2A06FFC1FFA2EEAAFE9C9AAFFEC6" box="[159,250,312,338]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="6" pageNumber="45">Fig. 12</figureCitation>
|
||
B, C), or occasionally complete; lateral part of sternite 8 distinctly exposed when male abdomen closed, visible as a large plate between coxa and abdomen (
|
||
<figureCitation id="131B2A06FFC1FFA2ED55FEC499F2FEEE" box="[864,935,352,378]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Fig. 7</figureCitation>
|
||
D) or a small rounded or ovate plate (
|
||
<figureCitation id="131B2A06FFC1FFA2EB59FEC49AE3FE36" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="6" pageNumber="45">Fig. 10</figureCitation>
|
||
B, C). Ventral margins of meri of P2–4 unarmed or only with spinules or granules (
|
||
<figureCitation id="131B2A06FFC1FFA2EA84FE2C9EADFE36" box="[1201,1272,392,418]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
A-C). Sternoabdominal cavity almost reaches to anterior edge of sternite 4 (
|
||
<figureCitation id="131B2A06FFC1FFA2ED4CFE149999FE5E" box="[889,972,432,458]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
B, 9B, C, 12B, C); press button of male abdominal locking mechanism on anterior edge of sternite 5 (
|
||
<figureCitation id="131B2A06FFC1FFA2ED51FE7C99E0FE66" box="[868,949,472,498]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
E, 12B, C). Penis coxosternal, originating from coxa of P5 but exiting between sternites 7 and 8, beyond condyle of coxa (
|
||
<figureCitation id="131B2A06FFC1FFA2EA09FDA49ED9FD8E" box="[1084,1164,512,538]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
F, 13C, D). Male abdomen broadly triangular (
|
||
<figureCitation id="131B2A06FFC1FFA2EF80FD8C9852FDD6" box="[437,519,552,578]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
B, D, 9B, C, 10B, C), sutures for all somites and telson clearly visible, somite 1 transversely broad but very narrow longitudinally, sometimes partially hidden under posterior carapace margin (
|
||
<figureCitation id="131B2A06FFC1FFA2EEE5FDDC9B71FD06" box="[208,292,632,658]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
C, 11B, C), somites 2–6 prominently transversely broad, somite 6 distinctly broader than long (
|
||
<figureCitation id="131B2A06FFC1FFA2EEAAFD049AA5FD2E" box="[159,240,672,698]" captionStart="FIGURE 7" captionStartId="23.[151,255,1351,1375]" captionTargetBox="[151,1435,194,1327]" captionTargetId="figure@23.[151,1436,194,1327]" captionTargetPageId="23" captionText="FIGURE 7. Goneplax, new species, male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700). A, buccal field, showing mouthparts and mandibles; B, anterior part of thoracic sternum and male abdominal somites 5, 6 and telson; C, posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs; D, posterior part of thoracic sternum and male abdominal somites 3 – 5; E, sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrow) and sternal sutures; F, structure and position of penis, coxa of P 5 and episternite 7. Abbreviations: p = penis, pb = press button of abdominal locking mechanism, t = telson, a 1 - 6 = male abdominal somites 1 to 6 respectively, st 4 - 8 = sternites 4 to 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178303/files/figure.png" pageId="6" pageNumber="45">Figs. 7</figureCitation>
|
||
B, D, 9B, C, 10B, C), all somites freely mobile. G1 relatively straight or slightly curved, sinuous, slen- der to stout, broad basal part gradually tapering distally, tip usually dilated/truncated. G2 usually as long as G2 (occasionally much shorter in a few genera), distal segment usually long, as long as, subequal to or longer than basal segment (sometimes short). Constituent genera:
|
||
<taxonomicName id="4C204D00FFC1FFA2ED5FFCBC9ED5FCA6" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[874,1152,792,818]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2ED5FFCBC9989FCA5" box="[874,988,792,817]" italics="true" pageId="6" pageNumber="45">Goneplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EDDDFCBC9ED5FCA6" author="Leach" box="[1000,1152,792,818]" pageId="6" pageNumber="53" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (editor), The Edinburgh Encyclopaedia, 7, 383 - 437, pl. 221." type="journal article" year="1814">Leach, 1814</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2EAA5FCBC9B1DFCCE" authority="H. Milne Edwards, 1852" authorityName="H. Milne Edwards" authorityYear="1852" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EAA5FCBC9F77FCA5" box="[1168,1314,792,817]" italics="true" pageId="6" pageNumber="45">Carcinoplax</emphasis>
|
||
H.
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EB63FCBC9B1DFCCE" author="Milne" pageId="6" pageNumber="54" refString="Milne Edwards, H. (1852) De la famille des Ocypodides (Ocypodidae). Second Memoire, in, Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences Naturelles (Zoologie), 18, 128 - 166 pls. 3, 4." type="journal article" year="1852">Milne Edwards, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2EF61FCE498B7FCCE" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[340,738,832,858]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EF61FCE49BE3FCCD" box="[340,438,832,857]" italics="true" pageId="6" pageNumber="45">Notonyx</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EF88FCE498B7FCCE" author="Milne-Edwards" box="[445,738,832,858]" pageId="6" pageNumber="54" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie, Deuxieme partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18." type="journal article" year="1873">A. Milne-Edwards, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
, and genera listed in Castro (in press) for his Goneplacinae.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC1FFA2EEF3FCC39944F966" blockId="6.[151,1437,152,2018]" pageId="6" pageNumber="45">
|
||
<emphasis id="B954EA91FFC1FFA2EEF3FCC39925FC15" bold="true" box="[198,880,871,897]" pageId="6" pageNumber="45">
|
||
Diagnosis of
|
||
<taxonomicName id="4C204D00FFC1FFA2EF5EFCC39934FC15" authority="Karasawa & Kato, 2003" authorityName="Karasawa & Kato" authorityYear="2003" box="[363,865,871,897]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="family">
|
||
Mathildellidae
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EC1EFCC39934FC15" author="Karasawa" box="[555,865,871,897]" pageId="6" pageNumber="53" refString="Karasawa, H. & Kato, H. (2003 a) The family Goneplacidae MacLeay, 1838 (Crustacea: Decapoda: Brachyura): systematics, phylogeny and fossil records. Paleontological Research, 7 (2), 129 - 151." type="journal article" year="2003" yearSuffix="a">Karasawa & Kato, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
a
|
||
</emphasis>
|
||
: Dorsal surface of carapace smooth to setose, regions sometimes distinct but never prominent, grooves shallow (
|
||
<figureCitation id="131B2A06FFC1FFA2ED94FC3499B0FC3E" box="[929,997,912,938]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
D–F). Mandibles white to grey in life and preserved (
|
||
<figureCitation id="131B2A06FFC1FFA2EF66FC1C9BC9FC46" box="[339,412,952,978]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="6" pageNumber="45">Fig. 8</figureCitation>
|
||
B). Thoracic sternum usually broad, sometimes relatively narrower; episternites 4–6 expanded laterally to some degree or other, never prominently (
|
||
<figureCitation id="131B2A06FFC1FFA2EDB6FC449986FC6E" box="[899,979,992,1018]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
D–F, 10D–F, 12D–F); episternite 7 not expanded laterally, neither touching coxa of fourth ambulatory leg nor forming small plate which partially covers or just touches anterior edge of coxa of P5 (
|
||
<figureCitation id="131B2A06FFC1FFA2ECD0FB94996FFBDE" box="[741,826,1072,1098]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="6" pageNumber="45">Fig. 13</figureCitation>
|
||
E); surface of sternite 3 without longitudinal median line (
|
||
<figureCitation id="131B2A06FFC1FFA2EEE6FBFC9B42FBE6" box="[211,279,1112,1138]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Fig. 9</figureCitation>
|
||
D–F), sternite 7 relatively broad to somewhat narrow transversely (
|
||
<figureCitation id="131B2A06FFC1FFA2EA1EFBFC9E2AFBE6" box="[1067,1151,1112,1138]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="6" pageNumber="45">Fig. 10</figureCitation>
|
||
D–F); sutures 4/5 and 5/ 6 interrupted medially (
|
||
<figureCitation id="131B2A06FFC1FFA2EF9FFB249BA8FB0E" box="[426,509,1152,1178]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="6" pageNumber="45">Fig. 12</figureCitation>
|
||
D–F); sutures 6/7 and 7/8 complete, median part of suture especially deep, well demarcated (
|
||
<figureCitation id="131B2A06FFC1FFA2EF1BFB0C9BD7FB56" box="[302,386,1192,1218]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="6" pageNumber="45">Fig. 12</figureCitation>
|
||
D–F); lateral part of sternite 8 completely covered by closed male abdomen (
|
||
<figureCitation id="131B2A06FFC1FFA2EB3CFB0C9F0BFB56" box="[1289,1374,1192,1218]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="6" pageNumber="45">Fig. 10</figureCitation>
|
||
D, F) or barely visible with only the edge visible (
|
||
<figureCitation id="131B2A06FFC1FFA2EC85FB74995DFB7E" box="[688,776,1232,1258]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="6" pageNumber="45">Fig. 10</figureCitation>
|
||
E). Ventral margins of meri of P2–4 usually smooth or granulated (
|
||
<figureCitation id="131B2A06FFC1FFA2EF18FB5C9B21FA86" box="[301,372,1272,1298]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
D, E), rarely spiniform (except for
|
||
<taxonomicName id="4C204D00FFC1FFA2ED23FB5C99EAFA85" box="[790,959,1272,1297]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2ED23FB5C99EAFA85" box="[790,959,1272,1297]" italics="true" pageId="6" pageNumber="45">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="131B2A06FFC1FFA2EDFAFB5C9E40FA86" box="[975,1045,1272,1298]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
F) but if so, not like condition in
|
||
<taxonomicName id="4C204D00FFC1FFA2EEA2FA849B64FAAE" box="[151,305,1312,1338]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
. Sterno-abdominal cavity reaches to middle part of sternite 4, never approaching median area (
|
||
<figureCitation id="131B2A06FFC1FFA2EEAAFAEC9AA5FAF6" box="[159,240,1352,1378]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
D–F, 12D–F); press button of male abdominal locking mechanism on posterior edge of sternite 5 (
|
||
<figureCitation id="131B2A06FFC1FFA2EB59FAEC9AE1FA1E" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="6" pageNumber="45">Fig. 12</figureCitation>
|
||
D–F). Penis coxal, originating directly from base of coxa of P5 (
|
||
<figureCitation id="131B2A06FFC1FFA2ED9CFAD499ABFA1E" box="[937,1022,1392,1418]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="6" pageNumber="45">Fig. 13</figureCitation>
|
||
E). Male abdomen triangular (
|
||
<figureCitation id="131B2A06FFC1FFA2EB54FAD49AF3FA26" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
D, E, 10D, E) but may be somewhat quadrate (e.g.
|
||
<taxonomicName id="4C204D00FFC1FFA2ECC8FA3C99F3FA25" box="[765,934,1432,1457]" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2ECC8FA3C99F3FA25" box="[765,934,1432,1457]" italics="true" pageId="6" pageNumber="45">Platypilumnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="131B2A06FFC1FFA2ED80FA3C9E50FA26" box="[949,1029,1432,1458]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
F, 10F), sutures for all somites and telson clearly visible, somite 1 transversely broad but relatively narrow longitudinally, subrectangular in shape to somewhat sinuous, exposed, not covered by posterior carapace margin (
|
||
<figureCitation id="131B2A06FFC1FFA2EDC9FA4C9E18F996" box="[1020,1101,1512,1538]" captionStart="FIGURE 11" captionStartId="27.[151,255,1445,1469]" captionTargetBox="[151,1435,194,1420]" captionTargetId="figure@27.[151,1436,194,1421]" captionTargetPageId="27" captionText="FIGURE 11. Posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 1 – 3 = male abdominal somites 1 to 3 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178307/files/figure.png" pageId="6" pageNumber="45">Fig. 11</figureCitation>
|
||
D–F), somites 2–6 distinctly transversely broad, somite 6 distinctly broader than long (
|
||
<figureCitation id="131B2A06FFC1FFA2ED7BF9B499F5F9BE" box="[846,928,1552,1578]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
D–F, 10D–F), somites 3 and 4 immovable although suture evident, somite 4 and 5 sometimes immobile as well. G1 C-shaped, curving outwards, basal part very broad, abruptly tapering to slender median and distal parts. G2 as long as G1, distal segment long, two-thirds to three-quarters length of basal segment. Constituent genera:
|
||
<taxonomicName id="4C204D00FFC1FFA2EA21F92C9F20F936" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[1044,1397,1672,1698]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EA21F92C9EC8F935" box="[1044,1181,1672,1697]" italics="true" pageId="6" pageNumber="45">Mathildella</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EEA2F9149B67F95E" author="Forges" box="[151,306,1712,1738]" pageId="6" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2EF0BF914982EF95E" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[318,635,1712,1738]" class="Malacostraca" family="Mathildellidae" genus="Beuroisia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EF0BF9149BFAF95D" box="[318,431,1712,1737]" italics="true" pageId="6" pageNumber="45">Beuroisia</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EC90F9149915F95E" author="Forges" box="[677,832,1712,1738]" pageId="6" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2ED79F9149E21F95E" authority="Guinot & Richer" authorityName="Guinot & Richer" box="[844,1140,1712,1738]" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2ED79F91499F2F95D" box="[844,935,1712,1737]" italics="true" pageId="6" pageNumber="45">Intesius</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EAABF9149F6CF95E" author="Forges" box="[1182,1337,1712,1738]" pageId="6" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2EB70F9149BCEF966" authority="Alcock, 1894" authorityName="Alcock" authorityYear="1894" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EB70F9149AA1F965" italics="true" pageId="6" pageNumber="45">Platypilumnus</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EECEF97C9BCEF966" author="Alcock" box="[251,411,1752,1778]" pageId="6" pageNumber="52" refString="Alcock, A. (1894). Natural history notes from Indian Marine Survey Steamer " Investigator ". Ser. II, no. 1. On the result of deep-sea dredging during the season of 1890 - 1891. Annals and Magazine of Natural History, (6) 13, 400 - 411" type="journal article" year="1894">Alcock, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC1FFA2EF92F97C9958F966" authority="Serene, 1969" authorityName="Serene" authorityYear="1969" box="[423,781,1752,1778]" class="Malacostraca" family="Mathildellidae" genus="Neopilumnoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC1FFA2EF92F97C983CF965" box="[423,617,1752,1777]" italics="true" pageId="6" pageNumber="45">Neopilumnoplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC1FFA2EC45F97C9958F966" author="Serene" box="[624,781,1752,1778]" pageId="6" pageNumber="54" refString="Serene R. (1969) Description de deux especes nouvelles et redecouverte d'une espece de Brachyoure (Decapoda Crustacea) dans les eaux d'Indonesie. Cahiers du Pacifique, 13, 281 - 293, figs. 1 - 24, pls. 1 - 4." type="journal article" year="1969">Serène, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC1FFA2EEF3F95B9FC9F876" blockId="6.[151,1437,152,2018]" pageId="6" pageNumber="45">
|
||
<emphasis id="B954EA91FFC1FFA2EEF3F95B9831F88D" bold="true" box="[198,612,1791,1817]" pageId="6" pageNumber="45">
|
||
Diagnosis of
|
||
<taxonomicName id="4C204D00FFC1FFA2EF59F95B9831F88D" authority="Stev" authorityName="Stev" box="[364,612,1791,1817]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="family">Progeryonidae Štev</taxonomicName>
|
||
</emphasis>
|
||
ě
|
||
<emphasis id="B954EA91FFC1FFA2EC44F95B982CF88D" bold="true" box="[625,633,1791,1817]" pageId="6" pageNumber="45">i</emphasis>
|
||
ć,
|
||
<emphasis id="B954EA91FFC1FFA2ECADF95B9887F88D" bold="true" box="[664,722,1791,1817]" pageId="6" pageNumber="45">2005</emphasis>
|
||
: Dorsal surface of carapace with very short felt-like setae, regions discernible but low, grooves shallow (
|
||
<figureCitation id="131B2A06FFC1FFA2EC8FF88C9957F8D6" box="[698,770,1832,1858]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
G). Mandibles white in preservative (
|
||
<figureCitation id="131B2A06FFC1FFA2EA8AF88C9F52F8D6" box="[1215,1287,1832,1858]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="6" pageNumber="45">Fig. 8</figureCitation>
|
||
C). Thoracic sternum relatively broad; episternites 4–6 not prominently expanded laterally (
|
||
<figureCitation id="131B2A06FFC1FFA2EA1FF8F49E2FF8FE" box="[1066,1146,1872,1898]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Figs. 9</figureCitation>
|
||
G, 10G, 12G); episternite 7 not prominently expanded but with small plate which just touches anterior edge of coxa of P5 (as in
|
||
<figureCitation id="131B2A06FFC1FFA2EB58F8DC9AE0F82E" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="6" pageNumber="45">Fig. 13</figureCitation>
|
||
E); sternite 3 without longitudinal median line (
|
||
<figureCitation id="131B2A06FFC1FFA2ECD4F8049972F82E" box="[737,807,1952,1978]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="6" pageNumber="45">Fig. 9</figureCitation>
|
||
G); sternite 7 very narrow transversely, tapering gradually posteriorly (
|
||
<figureCitation id="131B2A06FFC1FFA2EF5FF86C9B94F876" box="[362,449,1992,2018]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="6" pageNumber="45">Fig. 10</figureCitation>
|
||
G); sutures 4/5 and 5/6 interrupted medially but median grooves very shallow (
|
||
<figureCitation id="131B2A06FFC1FFA2EB58F86C9FC9F876" box="[1389,1436,1992,2018]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="6" pageNumber="45">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC0FFA3EEA2FF3C9EDDFDD6" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="46">
|
||
12G); sutures 6/7 and 7/8 complete, median part of suture especially deep, well demarcated; lateral part of sternite 8 completely covered by closed male abdomen (
|
||
<figureCitation id="131B2A06FFC0FFA3ED08FF6499C0FF4E" box="[829,917,192,218]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="7" pageNumber="46">Fig. 10</figureCitation>
|
||
G). Ventral margins of meri of P2–4 gently granulated ﹝Fiɡ․ ₆G). Sterno-abdominal cavity almost reaches anterior edge of sternite 4 (
|
||
<figureCitation id="131B2A06FFC0FFA3EA91FF4C9EA1FE96" box="[1188,1268,232,258]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
G, 12G); press button of male abdominal locking mechanism on posterior edge of sternite 5 (
|
||
<figureCitation id="131B2A06FFC0FFA3EA17FEB49E22FEBE" box="[1058,1143,272,298]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="7" pageNumber="46">Fig. 12</figureCitation>
|
||
G). Penis coxal, originating directly from base of coxa of P5 (as in
|
||
<figureCitation id="131B2A06FFC0FFA3ECACFE9C98BBFEC6" box="[665,750,312,338]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="7" pageNumber="46">Fig. 13</figureCitation>
|
||
E). Male abdomen appears subovate due to broad somites and distinctly convex margins (
|
||
<figureCitation id="131B2A06FFC0FFA3EC33FEC49803FEEE" box="[518,598,352,378]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
G, 10G), sutures for all somites and telson clearly visible, somite 1 relatively narrow longitudinally but also relatively narrow transversely, sublunate in shape, completely exposed, not covered by posterior carapace margin (
|
||
<figureCitation id="131B2A06FFC0FFA3EC93FE1498A8FE5E" box="[678,765,432,458]" captionStart="FIGURE 11" captionStartId="27.[151,255,1445,1469]" captionTargetBox="[151,1435,194,1420]" captionTargetId="figure@27.[151,1436,194,1421]" captionTargetPageId="27" captionText="FIGURE 11. Posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 1 – 3 = male abdominal somites 1 to 3 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178307/files/figure.png" pageId="7" pageNumber="46">Fig. 11</figureCitation>
|
||
G), somites 2–6 distinctly transversely broad, somite 6 much broader than long (
|
||
<figureCitation id="131B2A06FFC0FFA3EFF6FE7C9841FE66" box="[451,532,472,498]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
G, 10G), all somites 3 mobile. G1 relatively stout, almost straight, gradually tapering to relatively sharp tip. G2 prominently longer than G1, distal segment long, almost as long as basal segment. Constituent genera:
|
||
<taxonomicName id="4C204D00FFC0FFA3EFC6FD8C9975FDD6" authority="Bouvier, 1922" authorityName="Bouvier" authorityYear="1922" box="[499,800,552,578]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EFC6FD8C983AFDD5" box="[499,623,552,577]" italics="true" pageId="7" pageNumber="46">Progeryon</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EC42FD8C9975FDD6" author="Bouvier" box="[631,800,552,578]" pageId="7" pageNumber="52" refString="Bouvier, E. - L. (1922) Observations complementaires sur les crustaces decapodes (abstraction faite des Carides) provenant des Campagnes de S. A. S le Prince de Monaco. Resultats Campagne Scientifiques, Monaco, 62, 1 - 106, pl. 1 - 6." type="journal article" year="1922">Bouvier, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3ED19FD8C9ED1FDD6" authority="Kossmann, 1878" authorityName="Kossmann" authorityYear="1878" box="[812,1156,552,578]" class="Malacostraca" family="Progeryonidae" genus="Paragalene" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3ED19FD8C99E0FDD5" box="[812,949,552,577]" italics="true" pageId="7" pageNumber="46">Paragalene</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC0FFA3ED89FD8C9ED1FDD6" author="Kossmann" box="[956,1156,552,578]" pageId="7" pageNumber="53" refString="Kossmann, R. (1878). Kurze Notizen ber einige neue Crustaceen sowie uber neue Fundorte einiger bereits beschriebenen. Archiv fur Naturgeschichte, 44 (1), 251 - 258" type="journal article" year="1878">Kossmann, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC0FFA3EEF3FDEB9B9CFB0E" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="46">
|
||
<emphasis id="B954EA91FFC0FFA3EEF3FDEB987BFDFD" bold="true" box="[198,558,591,617]" pageId="7" pageNumber="46">
|
||
Diagnosis of
|
||
<taxonomicName id="4C204D00FFC0FFA3EF50FDEB987BFDFD" authority="Stev" authorityName="Stev" box="[357,558,591,617]" class="Malacostraca" family="Conleyidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Conleyidae Štev</taxonomicName>
|
||
</emphasis>
|
||
ě
|
||
<emphasis id="B954EA91FFC0FFA3EC0EFDEB9816FDFD" bold="true" box="[571,579,591,617]" pageId="7" pageNumber="46">i</emphasis>
|
||
ć,
|
||
<emphasis id="B954EA91FFC0FFA3EC6AFDEB98CCFDFD" bold="true" box="[607,665,591,617]" pageId="7" pageNumber="46">2005</emphasis>
|
||
: Dorsal surface of carapace with poorly defined regions and shallow grooves, sparsely setose (
|
||
<figureCitation id="131B2A06FFC0FFA3EFCAFDD1981DFD1B" box="[511,584,629,655]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="7" pageNumber="46">Fig. 6</figureCitation>
|
||
H). Mandibles white in life and preserved (
|
||
<figureCitation id="131B2A06FFC0FFA3EA66FDD19ECFFD1B" box="[1107,1178,629,655]" captionStart="FIGURE 8" captionStartId="24.[151,255,1099,1123]" captionTargetBox="[151,1435,194,1073]" captionTargetId="figure@24.[151,1436,194,1075]" captionTargetPageId="24" captionText="FIGURE 8. Buccal field, showing mouthparts and mandibles. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); C, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC); D, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178304/files/figure.png" pageId="7" pageNumber="46">Fig. 8</figureCitation>
|
||
D). Thoracic sternum broad; episternites 4–6 expanded laterally slightly (
|
||
<figureCitation id="131B2A06FFC0FFA3ECC5FD3E9914FD20" box="[752,833,666,692]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
H, 10H, 12H); episternite 7 distinctly expanded laterally to form plate which covers anterior edge of coxa of P5 (
|
||
<figureCitation id="131B2A06FFC0FFA3ED93FD6499AAFD4E" box="[934,1023,704,730]" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="7" pageNumber="46">Fig. 13</figureCitation>
|
||
F); sternite 3 without longitudinal median line (
|
||
<figureCitation id="131B2A06FFC0FFA3EF00FD419B28FD6B" box="[309,381,741,767]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Fig. 9</figureCitation>
|
||
H); sternite 7 relatively broad (
|
||
<figureCitation id="131B2A06FFC0FFA3ECC6FD41991FFD6B" box="[755,842,741,767]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="7" pageNumber="46">Fig. 10</figureCitation>
|
||
H); sutures 4/5 and 5/6 interrupted medially (
|
||
<figureCitation id="131B2A06FFC0FFA3EB59FD419AE0FCB0" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="7" pageNumber="46">Fig. 12</figureCitation>
|
||
H); sutures 6/7 and 7/8 complete (as in
|
||
<taxonomicName id="4C204D00FFC0FFA3ECB2FCAE9962FCB0" box="[647,823,778,804]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
), suture more or less evenly prominent along length (
|
||
<figureCitation id="131B2A06FFC0FFA3EEAAFC949AA0FCDE" box="[159,245,816,842]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="7" pageNumber="46">Fig. 12</figureCitation>
|
||
H); lateral part of sternite 8 completely covered by closed male abdomen (
|
||
<figureCitation id="131B2A06FFC0FFA3EA6AFC949EE0FCDE" box="[1119,1205,816,842]" captionStart="FIGURE 10" captionStartId="26.[151,255,1725,1749]" captionTargetBox="[151,1436,194,1701]" captionTargetId="figure@26.[151,1436,194,1701]" captionTargetPageId="26" captionText="FIGURE 10. Posterior part of thoracic sternum and male abdominal somites 3 – 5. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003: male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 2 – 5 = male abdominal somites 2 to 5 respectively, st 7 = sternites 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178306/files/figure.png" pageId="7" pageNumber="46">Fig. 10</figureCitation>
|
||
H). Ventral margins of meri of P2–4 granuliform (
|
||
<figureCitation id="131B2A06FFC0FFA3EFC3FCF1986EFCFB" box="[502,571,853,879]" captionStart="FIGURE 6" captionStartId="22.[151,255,1696,1720]" captionTargetBox="[151,1436,194,1671]" captionTargetId="figure@22.[151,1436,194,1672]" captionTargetPageId="22" captionText="FIGURE 6. Overall views. A, Goneplax, new species: male (cl 21.3 mm cw 35.3 mm) (ZRC 2004.700); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH)." httpUri="https://zenodo.org/record/178302/files/figure.png" pageId="7" pageNumber="46">Fig. 6</figureCitation>
|
||
H). Sterno-abdominal cavity reaches to middle part of sternite 4 (
|
||
<figureCitation id="131B2A06FFC0FFA3EB1AFCF19FD5FCFB" box="[1327,1408,853,879]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
H, 12H); press button of male abdominal locking mechanism on posterior edge of sternite 5 (
|
||
<figureCitation id="131B2A06FFC0FFA3EAE3FCDE9F7BFC00" box="[1238,1326,890,916]" captionStart="FIGURE 12" captionStartId="28.[151,255,1724,1748]" captionTargetBox="[227,1361,180,1710]" captionTargetId="figure@28.[227,1361,180,1710]" captionTargetPageId="28" captionText="FIGURE 12. Sterno-abdominal cavity, showing position of press button of abdominal locking mechanism (indicated by arrows) and sternal sutures. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: pb = press button of abdominal locking mechanism, st 4 – 7 = sternites 4 to 7 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178308/files/figure.png" pageId="7" pageNumber="46">Fig. 12</figureCitation>
|
||
H). Penis coxosternal, originating from coxa of P5 but exiting between sternites 7 and 8, beyond condyle of coxa (
|
||
<figureCitation id="131B2A06FFC0FFA3EB58FC049AE0FC4B" captionStart="FIGURE 13" captionStartId="29.[151,255,1377,1401]" captionTargetBox="[264,1323,194,1353]" captionTargetId="figure@29.[264,1323,194,1353]" captionTargetPageId="29" captionText="FIGURE 13. Structure and position of penis, coxa of P 5 and episternite 7 (position of penis indicated by arrow). A, B, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); C, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); D, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Conleyus defodio Ng & Ng, 2003, holotype male (cl, 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: p = penis, st 7 and 8 = sternites 7 and 8 respectively, es 7 = episternite 7, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178309/files/figure.png" pageId="7" pageNumber="46">Fig. 13</figureCitation>
|
||
F). Male abdomen triangular (
|
||
<figureCitation id="131B2A06FFC0FFA3EC2AFC619827FC4B" box="[543,626,965,991]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Figs. 9</figureCitation>
|
||
H, 10H), sutures for all somites and telson clearly visible, somite 1 transversely broad but narrow longitudinally, sinuous, exposed, not covered by posterior carapace margin (
|
||
<figureCitation id="131B2A06FFC0FFA3EEAAFBB49AA0FBBE" box="[159,245,1040,1066]" captionStart="FIGURE 11" captionStartId="27.[151,255,1445,1469]" captionTargetBox="[151,1435,194,1420]" captionTargetId="figure@27.[151,1436,194,1421]" captionTargetPageId="27" captionText="FIGURE 11. Posterior part of carapace showing posterior carapace margin, male abdominal somites 1 and 2, and coxa of last ambulatory legs. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981, male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 1 – 3 = male abdominal somites 1 to 3 respectively, c 5 = coxa of P 5." httpUri="https://zenodo.org/record/178307/files/figure.png" pageId="7" pageNumber="46">Fig. 11</figureCitation>
|
||
H), somites 2–6 transversely broad, somite 6 broader than long (
|
||
<figureCitation id="131B2A06FFC0FFA3EA36FBB49E1EFBBE" box="[1027,1099,1040,1066]" captionStart="FIGURE 9" captionStartId="25.[151,255,1726,1750]" captionTargetBox="[224,1358,194,1697]" captionTargetId="figure@25.[224,1358,194,1697]" captionTargetPageId="25" captionText="FIGURE 9. Anterior part of thoracic sternum and male abdominal somites 5, 6 and telson. A, Vultocinus anfractus, new genus, new species, paratype male (cl 27.3 mm, cw 25.3 mm) (ZRC); B, “ Goneplax ” marivenae Komatsu & Takeda, 2004, male (cl 11.0 mm, cw 17.6 mm) (ZRC 2004.724); C, Carcinoplax spinosissima Rathbun, 1914, male (cl 24.0 mm, cw 29.1 mm) (ZRC 2001.535); D, Mathildella rubra Ng & Ho, 2003, male (cl 16.2 mm, cw 19.5 mm) (ZRC 2004.535); E, Intesius pilosus Guinot & Richer de Forges, 1981: male (cl 32.3 mm, cw 38.3 mm) (ZRC 2004.731); F, Platypilumnus gracilipes Alcock, 1894, male (cl 25.3 mm, cw 29.6 mm) (ZRC 2004.717); G, Progeryon mus Ng & Guinot, 1999, holotype male (cl 38.6 mm, cw 46.5 mm) (ZRC 1997.442); H, Conleyus defodio Ng & Ng, 2003, holotype male (cl 12.2 mm, cw 16.4 mm) (FLMNH). Abbreviations: t = telson, a 4 – 6 = male abdominal somites 4 to 6 respectively, st 4 – 7 = sternites 4 to 7 respectively." httpUri="https://zenodo.org/record/178305/files/figure.png" pageId="7" pageNumber="46">Fig. 9</figureCitation>
|
||
H, 10H), all somites mobile although somites 3 and 4 somewhat stiff but still movable. G1 relatively slender, sinuous, with tip directed upwards. G2 as long as G1, distal segment short, about 0.3 times length of basal segment. Constituent genus:
|
||
<taxonomicName id="4C204D00FFC0FFA3EEA2FB249B90FB0E" authority="Ng & Ng, 2003" authorityName="Ng & Ng" authorityYear="2003" box="[151,453,1152,1178]" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EEA2FB249B51FB0D" box="[151,260,1152,1177]" italics="true" pageId="7" pageNumber="46">Conleyus</emphasis>
|
||
Ng & Ng, 2003
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC0FFA3EEF3FB0198E0F9CE" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="46">
|
||
As to their superfamily status, we retain the
|
||
<taxonomicName id="4C204D00FFC0FFA3ECE7FB0199D5FB2B" box="[722,896,1189,1215]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3EDB9FB019E6FFB2B" box="[908,1082,1189,1215]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Progeryonidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3EA73FB019E99FB2B" box="[1094,1228,1189,1215]" class="Malacostraca" family="Conleyidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Conleyidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC0FFA3EB33FB019FCEFB2B" box="[1286,1435,1189,1215]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
in the Goneplacoidea for the time being. As for some of the other unusual suprageneric taxa established or recognized by
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EF7BFB4B9BAAFA9E" author="Stevcic" box="[334,511,1263,1290]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
discussed above, viz. Psopheticini
|
||
<bibRefCitation id="EFB14B72FFC0FFA3ED8CFB4B9E35FA9E" author="Stevcic" box="[953,1120,1263,1290]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, Platypilumninae
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EB0BFB4B9A81FABB" author="Stevcic" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3EED4FAB19863FABB" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[225,566,1301,1327]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Bathyplacinae">
|
||
Bathyplacinae
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EFA7FAB19863FABB" author="Stevcic" box="[402,566,1301,1327]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3EC77FAB199D8FABB" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[578,909,1301,1327]" class="Malacostraca" family="Scalopidiidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">
|
||
Scalopidiidae
|
||
<bibRefCitation id="EFB14B72FFC0FFA3ECDFFAB199D8FABB" author="Stevcic" box="[746,909,1301,1327]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, Platycheloniini
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EA60FAB19EACFABB" author="Stevcic" box="[1109,1273,1301,1327]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Stevcic, 2005</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C204D00FFC0FFA3EB0CFAB19BF2FAC0" authority="Serene, 1984" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Planopilumnidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">
|
||
Planopilumnidae
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EF3EFA9E9BF2FAC0" author="Serene" box="[267,423,1338,1364]" pageId="7" pageNumber="55" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge, Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par Crosnier (A.): Carpiliidae et Menippidae. Faune Tropicale, 24, 1 - 349, figs A-C + 1 - 243, pls 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
; the following decisions are taken. Psopheticini
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EDE8FA9E9E2AFAC0" author="Stevcic" box="[989,1151,1338,1364]" pageId="7" pageNumber="55" refString="Stevcic, Z. (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Brachyura). Fauna Croatica, 14, 1, 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, is clearly a synonym of Goneplacinae
|
||
<emphasis id="B954EA91FFC0FFA3EF77FAC49B8DFAED" box="[322,472,1376,1401]" italics="true" pageId="7" pageNumber="46">
|
||
<taxonomicName id="4C204D00FFC0FFA3EF77FAC49BD6FAED" box="[322,387,1376,1401]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
(see Castro, in press). The Platypilumninae is a synonym of the
|
||
<taxonomicName id="4C204D00FFC0FFA3EAC4FAC49FC9FAEE" box="[1265,1436,1376,1402]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
(see above). The
|
||
<taxonomicName id="4C204D00FFC0FFA3EF5DFA219845FA0B" box="[360,528,1413,1439]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Bathyplacinae">Bathyplacinae</taxonomicName>
|
||
has several unusual features and should be tentatively recognized as a subfamily of the
|
||
<taxonomicName id="4C204D00FFC0FFA3EF06FA0E9B81FA50" box="[307,468,1450,1476]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
(see above). The
|
||
<taxonomicName id="4C204D00FFC0FFA3EC96FA0E9917FA50" box="[675,834,1450,1476]" class="Malacostraca" family="Scalopidiidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Scalopidiidae</taxonomicName>
|
||
is probably also a valid family, but may be allied to the
|
||
<taxonomicName id="4C204D00FFC0FFA3EEF7FA749BCEFA7E" box="[194,411,1488,1514]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
(D. Guinot, pers. comm.). The Platycheloniini is clearly a synonym of the
|
||
<taxonomicName id="4C204D00FFC0FFA3EB3EFA749A8CF99B" class="Malacostraca" family="Planopilumnidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Planopilumnidae</taxonomicName>
|
||
, and is allied to the
|
||
<taxonomicName id="4C204D00FFC0FFA3EFFAFA51983FF99B" box="[463,618,1525,1551]" class="Malacostraca" family="Pseudoziidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Pseudoziidae</taxonomicName>
|
||
(see Ng 2003). The systematics of the
|
||
<taxonomicName id="4C204D00FFC0FFA3EA71FA519EBBF99B" box="[1092,1262,1525,1551]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Bathyplacinae">Bathyplacinae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D00FFC0FFA3EAC8FA519FC9F99B" box="[1277,1436,1525,1551]" class="Malacostraca" family="Scalopidiidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Scalopidiidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D00FFC0FFA3EEFCF9BE9B31F9A0" box="[201,356,1562,1588]" class="Malacostraca" family="Pseudoziidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="family">Pseudoziidae</taxonomicName>
|
||
is currently being reappraised in detail by the first author with D. Guinot and P. Davie). All are here regarded in any case as goneplacoids.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC0FFA3EEF3F9C19976F8AE" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="46">
|
||
A note on
|
||
<taxonomicName id="4C204D00FFC0FFA3EF7DF9C198F6F9EB" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[328,675,1637,1663]" class="Malacostraca" family="Euryplacidae" genus="Pilumnoplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EF7DF9C19B8BF9EA" box="[328,478,1637,1662]" italics="true" pageId="7" pageNumber="46">Pilumnoplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EFD2F9C198F6F9EB" author="Stimpson" box="[487,675,1637,1663]" pageId="7" pageNumber="55" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part V, Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 93 - 110." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
, is necessary. The genus was affiliated with the Mathildellinae, but is now a junior subjective synonym of
|
||
<taxonomicName id="4C204D00FFC0FFA3ECACF92F99E6F930" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[665,947,1674,1700]" class="Malacostraca" family="Euryplacidae" genus="Eucrate" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3ECACF92F98A3F930" box="[665,758,1675,1700]" italics="true" pageId="7" pageNumber="46">Eucrate</emphasis>
|
||
De Haan, 1835
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="549B8821FFC0FFA3EDF1F92E99ADF930" box="[964,1016,1674,1700]" pageId="7" pageNumber="46">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C204D00FFC0FFA3EA6AF92F9B04F95E" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="crenatus" subGenus="Eucrate">
|
||
<emphasis id="B954EA91FFC0FFA3EA6AF92F9EE0F930" box="[1119,1205,1675,1700]" italics="true" pageId="7" pageNumber="46">Cancer</emphasis>
|
||
(
|
||
<emphasis id="B954EA91FFC0FFA3EAFDF92F9F70F930" box="[1224,1317,1675,1700]" italics="true" pageId="7" pageNumber="46">Eucrate</emphasis>
|
||
)
|
||
<emphasis id="B954EA91FFC0FFA3EB0DF92F9FC9F930" box="[1336,1436,1675,1700]" italics="true" pageId="7" pageNumber="46">crenatus</emphasis>
|
||
De Haan, 1835
|
||
</taxonomicName>
|
||
), in the Euryplacinae
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EC69F914994FF95E" author="Stimpson" box="[604,794,1712,1738]" pageId="7" pageNumber="55" refString="Stimpson W. (1871) Brachyura. Preliminary report on the Crustacea dredged in the Gulf Stream in the Straits of Florida, by L. F. Pourtales, Assist. U. S. Coast Surveys, Part 1. Bulletin of the Museum of Comparative Zoology at Harvard College, 2, 109 - 160." type="journal article" year="1871">Stimpson, 1871</bibRefCitation>
|
||
. Its
|
||
<typeStatus id="549B8821FFC0FFA3ED65F91499D7F95E" box="[848,898,1712,1738]" pageId="7" pageNumber="46">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C204D00FFC0FFA3EDC6F9149A81F97B" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" class="Malacostraca" family="Euryplacidae" genus="Pilumnoplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="sulcatifrons">
|
||
<emphasis id="B954EA91FFC0FFA3EDC6F9149F49F95D" box="[1011,1308,1712,1737]" italics="true" pageId="7" pageNumber="46">Pilumnoplax sulcatifrons</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EB13F9149A81F97B" author="Stimpson" pageId="7" pageNumber="55" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part V, Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 93 - 110." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
, is now generally accepted as a species of
|
||
<taxonomicName id="4C204D00FFC0FFA3ECFAF971997EF97A" box="[719,811,1749,1774]" class="Malacostraca" family="Euryplacidae" genus="Eucrate" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3ECFAF971997EF97A" box="[719,811,1749,1774]" italics="true" pageId="7" pageNumber="46">Eucrate</emphasis>
|
||
</taxonomicName>
|
||
, and all the other species which have been placed in
|
||
<taxonomicName id="4C204D00FFC0FFA3EEA2F95F9B78F880" box="[151,301,1787,1812]" class="Malacostraca" family="Euryplacidae" genus="Pilumnoplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EEA2F95F9B78F880" box="[151,301,1787,1812]" italics="true" pageId="7" pageNumber="46">Pilumnoplax</emphasis>
|
||
</taxonomicName>
|
||
over the years have to be transferred to other genera, e.g.
|
||
<taxonomicName id="4C204D00FFC0FFA3EDFAF95F9F65F880" authority="Serene, 1969" authorityName="Serene" authorityYear="1969" box="[975,1328,1786,1812]" class="Malacostraca" family="Mathildellidae" genus="Neopilumnoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EDFAF95F9EC4F880" box="[975,1169,1787,1812]" italics="true" pageId="7" pageNumber="46">Neopilumnoplax</emphasis>
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EAA2F95E9F65F880" author="Serene" box="[1175,1328,1786,1812]" pageId="7" pageNumber="54" refString="Serene R. (1969) Description de deux especes nouvelles et redecouverte d'une espece de Brachyoure (Decapoda Crustacea) dans les eaux d'Indonesie. Cahiers du Pacifique, 13, 281 - 293, figs. 1 - 24, pls. 1 - 4." type="journal article" year="1969">Serène, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EEA2F8849B75F8AE" author="Tesch" box="[151,288,1824,1850]" pageId="7" pageNumber="55" refString="Tesch, J. (1918) The Decapoda Brachyura of the Siboga-Expedition. II. Goneplacidae and Pinnotheridae. In: Uitkomstem op Zoologisch, Botanisch, Oceanographisch en Geologisch gebied verzameld in Nederlansch Oost-Indie 1899 - 1900 aan boord H. M. Siboga onder commando van Luitenant ter zee Ie kl. G. F. Tydeman uitgeven door Dr. Max Webber. Siboga - Expeditie, 39 c 1 (84), 149 - 295, 12 pls." type="journal article" year="1918">Tesch 1918</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EF19F8849B85F8AE" author="Guinot" box="[300,464,1824,1850]" pageId="7" pageNumber="53" refString="Guinot, D. (1969 c) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. VII. Les Goneplacidae (suite et fin). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 41 (3), 688 - 724." type="journal article" year="1969" yearSuffix="c">Guinot 1969c</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB14B72FFC0FFA3EFE9F8849943F8AE" author="Manning" box="[476,790,1824,1850]" pageId="7" pageNumber="54" refString="Manning, R. B. & Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, i - xii, 1 - 379." type="journal article" year="1981">Manning & Holthuis 1981</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFC0FFACEEF3F8E19BEEFF68" blockId="7.[151,1437,152,2036]" lastBlockId="8.[151,1436,152,740]" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
|
||
The use of the genus
|
||
<taxonomicName id="4C204D00FFC0FFA3EF8AF8E19864F8CA" box="[447,561,1861,1886]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFC0FFA3EF8AF8E19864F8CA" box="[447,561,1861,1886]" italics="true" pageId="7" pageNumber="46">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
here is in the broad sense. Castro (in press) revises the genus and all of the strictly Indo-West Pacific species were transferred to new genera.
|
||
<emphasis id="B954EA91FFC0FFA3EDF6F8CF9E82F810" box="[963,1239,1899,1924]" italics="true" pageId="7" pageNumber="46">
|
||
<taxonomicName id="4C204D00FFC0FFA3EDF6F8CF9ED4F810" box="[963,1153,1899,1924]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="sensu">
|
||
Goneplax
|
||
<taxonomicNameLabel id="A26757EAFFC0FFA3EA75F8CF9ED4F810" box="[1088,1153,1899,1924]" pageId="7" pageNumber="46" sensu="stricto">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
is restricted for Atlantic species. An exception is a new taxon here labeled as
|
||
<emphasis id="B954EA91FFC0FFA3ED42F83499A5F83D" box="[887,1008,1936,1961]" italics="true" pageId="7" pageNumber="46">
|
||
<taxonomicName id="4C204D00FFC0FFA3ED42F83499B9F83D" box="[887,1004,1936,1961]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="genus" status="sp. nov.">Goneplax</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A26757EAFFC0FFA3EDC2F8349ED2F83E" box="[1015,1159,1936,1962]" pageId="7" pageNumber="46" rank="species">new species</taxonomicNameLabel>
|
||
, which is closely affiliated with
|
||
<taxonomicName id="4C204D00FFC0FFA3EF25F81198C1F85B" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[272,660,1973,1999]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="rhomboides">
|
||
<emphasis id="B954EA91FFC0FFA3EF25F8119BE9F85A" box="[272,444,1973,1998]" italics="true" pageId="7" pageNumber="46">G. rhomboides</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="549B8821FFC0FFA3ECE5F8119957F85B" box="[720,770,1973,1999]" pageId="7" pageNumber="46">type</typeStatus>
|
||
species of the genus, and currently being described by Guinot & Castro (in press). It is mostly Atlantic in distribution but it also found on the Indian Ocean coast of
|
||
<collectingCountry id="F3377613FFCFFFACEEA2FF3C9B61FF26" box="[151,308,152,178]" name="South Africa" pageId="8" pageNumber="47">South Africa</collectingCountry>
|
||
and hence in the Indo-West Pacific region. The Indo-West Pacific “
|
||
<taxonomicName id="4C204D00FFCFFFACEAA1FF3C9F53FF25" box="[1172,1286,152,177]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFCFFFACEAA1FF3C9F53FF25" box="[1172,1286,152,177]" italics="true" pageId="8" pageNumber="47">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<emphasis id="B954EA91FFCFFFACEB15FF3C9FCEFF25" box="[1312,1435,152,177]" italics="true" pageId="8" pageNumber="47">marivenae</emphasis>
|
||
Komatsu & Takeda, 2004, on the other hand, belongs to its own genus (Castro, in press). Both are considered and figured in this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6508FFCFFFACEEF3FEA398BBFCBE" pageId="8" pageNumber="47" type="materials_examined">
|
||
<paragraph id="8B9F3683FFCFFFACEEF3FEA398CCFD70" blockId="8.[151,1436,152,740]" pageId="8" pageNumber="47">
|
||
<emphasis id="B954EA91FFCFFFACEEF3FEA39B89FEB5" bold="true" box="[198,476,263,289]" pageId="8" pageNumber="47">Comparative material</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D00FFCFFFACEFDEFEAC9808FEB5" box="[491,605,264,289]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
<emphasis id="B954EA91FFCFFFACEFDEFEAC9808FEB5" box="[491,605,264,289]" italics="true" pageId="8" pageNumber="47">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26757EAFFCFFFACEC59FEAC98A8FEB6" box="[620,765,264,290]" pageId="8" pageNumber="47" rank="species">new species</taxonomicNameLabel>
|
||
: male (cl
|
||
<quantity id="4CD89B66FFCFFFACED5AFEAC998DFEB6" box="[879,984,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="8" pageNumber="47" unit="mm" value="21.3">21.3 mm</quantity>
|
||
cw
|
||
<quantity id="4CD89B66FFCFFFACEA3FFEAC9E21FEB6" box="[1034,1140,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="8" pageNumber="47" unit="mm" value="35.3">35.3 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEABEFEAC9E93FEB6" box="[1163,1222,264,290]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2004.700), Algoa Bay, Port Elisabeth,
|
||
<collectingCountry id="F3377613FFCFFFACEFA4FE8E987BFED0" box="[401,558,298,324]" name="South Africa" pageId="8" pageNumber="47">South Africa</collectingCountry>
|
||
, coll. trawlers,
|
||
<date id="FF9E1043FFCFFFACECD9FE8E9928FED0" box="[748,893,298,324]" pageId="8" pageNumber="47" value="2003-12-01">1 Dec 2003</date>
|
||
; “
|
||
<taxonomicName id="4C204D00FFCFFFACEDAFFE8E9E59FED7" box="[922,1036,298,323]" class="Malacostraca" family="Goneplacidae" genus="Goneplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EA91FFCFFFACEDAFFE8E9E59FED7" box="[922,1036,298,323]" italics="true" pageId="8" pageNumber="47">Goneplax</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<emphasis id="B954EA91FFCFFFACEA16FE8E9ECAFED7" box="[1059,1183,298,323]" italics="true" pageId="8" pageNumber="47">marivenae</emphasis>
|
||
Komatsu & Takeda, 2004: male (cl 11.0 mm, cw
|
||
<quantity id="4CD89B66FFCFFFACEFDFFEE99803FEF3" box="[490,598,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="8" pageNumber="47" unit="mm" value="17.6">17.6 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEC58FEE998FDFEF3" box="[621,680,333,359]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2004.724), Balicasag Island, Panglao, Bohol, Visayas, in tangle nets, coll. local shell fishermen,
|
||
<date id="FF9E1043FFCFFFACEC24FED4989CFE1E" box="[529,713,368,394]" pageId="8" pageNumber="47" value="2004-07-25" valueMax="2004-07-30" valueMin="2004-07-25">25–30 Jul 2004</date>
|
||
;
|
||
<taxonomicName id="4C204D00FFCFFFACECE0FED49EEEFE1E" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[725,1211,368,394]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="spinosissima">
|
||
<emphasis id="B954EA91FFCFFFACECE0FED49E56FE1D" box="[725,1027,368,393]" italics="true" pageId="8" pageNumber="47">Carcinoplax spinosissima</emphasis>
|
||
Rathbun, 1914
|
||
</taxonomicName>
|
||
: male (cl 24.0 mm, cw
|
||
<quantity id="4CD89B66FFCFFFACEEF5FE369B7FFE38" box="[192,298,402,428]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="8" pageNumber="47" unit="mm" value="29.1">29.1 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEF74FE369B29FE38" box="[321,380,402,428]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2001.535), Balicasag Island, Panglao, Bohol, Visayas, in tangle nets, coll. local shell fishermen,
|
||
<date id="FF9E1043FFCFFFACEEC5FE119BC5FE5B" box="[240,400,437,463]" pageId="8" pageNumber="47" value="2001-11-28">28 Nov 2001</date>
|
||
;
|
||
<taxonomicName id="4C204D00FFCFFFACEFA8FE11996FFE5B" authority="Ng & Ho, 2003" authorityName="Ng & Ho" authorityYear="2003" box="[413,826,437,463]" class="Malacostraca" family="Mathildellidae" genus="Mathildella" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="rubra">
|
||
<emphasis id="B954EA91FFCFFFACEFA8FE119827FE5A" box="[413,626,437,462]" italics="true" pageId="8" pageNumber="47">Mathildella rubra</emphasis>
|
||
Ng &
|
||
<bibRefCitation id="EFB14B72FFCFFFACECF2FE11996FFE5B" author="Ho" box="[711,826,437,463]" pageId="8" pageNumber="54" refString="Ng, P. K. L. & Ho, P. - H. (2003) Mathildella rubra, a new species of deep-water carcinoplacine crab (Decapoda, Brachyura) from the Philippines. Crustaceana, 76 (3), 333 - 342." type="journal article" year="2003">Ho, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
: male (cl
|
||
<quantity id="4CD89B66FFCFFFACED85FE119E48FE5B" box="[944,1053,437,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="8" pageNumber="47" unit="mm" value="16.2">16.2 mm</quantity>
|
||
, cw
|
||
<quantity id="4CD89B66FFCFFFACEA60FE119E95FE5B" box="[1109,1216,437,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="8" pageNumber="47" unit="mm" value="19.5">19.5 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEAECFE119F41FE5B" box="[1241,1300,437,463]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2004.535), Balicasag Island, Panglao, Bohol, Visayas, in tangle nets, coll. local shell fishermen,
|
||
<date id="FF9E1043FFCFFFACEAA6FE7C9F66FE66" box="[1171,1331,472,498]" pageId="8" pageNumber="47" value="2003-11-28">28 Nov 2003</date>
|
||
;
|
||
<taxonomicName id="4C204D00FFCFFFACEB74FE7C9BE0FD80" authority="Guinot & Richer" authorityName="Guinot & Richer" class="Malacostraca" family="Mathildellidae" genus="Intesius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="pilosus">
|
||
<emphasis id="B954EA91FFCFFFACEB74FE7C9ABFFD87" italics="true" pageId="8" pageNumber="47">Intesius pilosus</emphasis>
|
||
Guinot & Richer
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="EFB14B72FFCFFFACEFEAFE5E9829FD80" author="Forges" box="[479,636,506,532]" pageId="8" pageNumber="53" refString="Guinot, D. & Richer de Forges, B. (1981) Crabes de profondeur, nouveaux ou rares, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura) (Deuxieme partie). Bulletin du Museum National d'Histoire Naturelle, Paris, 1981, (4) 3 (A 1), 227 - 260." type="journal article" year="1981">Forges, 1981</bibRefCitation>
|
||
: male (cl
|
||
<quantity id="4CD89B66FFCFFFACECD9FE5E9902FD80" box="[748,855,506,532]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="8" pageNumber="47" unit="mm" value="32.3">32.3 mm</quantity>
|
||
, cw
|
||
<quantity id="4CD89B66FFCFFFACEDBEFE5E99A0FD80" box="[907,1013,506,532]" metricMagnitude="-2" metricUnit="m" metricValue="3.8299999999999996" pageId="8" pageNumber="47" unit="mm" value="38.3">38.3 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEA39FE5E9E12FD80" box="[1036,1095,506,532]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2004.731), Balicasag Island, Panglao, Bohol, Visayas, in tangle nets, coll. local shell fishermen,
|
||
<date id="FF9E1043FFCFFFACEDE5FDB99E36FDA3" box="[976,1123,541,567]" pageId="8" pageNumber="47" value="2004-03-02">2 Mar 2004</date>
|
||
;
|
||
<taxonomicName id="4C204D00FFCFFFACEA46FDB99B6FFDCE" authority="Alcock, 1894" authorityName="Alcock" authorityYear="1894" class="Malacostraca" family="Mathildellidae" genus="Platypilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="B954EA91FFCFFFACEA46FDB99FC9FDA2" box="[1139,1436,541,566]" italics="true" pageId="8" pageNumber="47">Platypilumnus gracilipes</emphasis>
|
||
<bibRefCitation id="EFB14B72FFCFFFACEEA2FDE49B6FFDCE" author="Alcock" box="[151,314,576,602]" pageId="8" pageNumber="52" refString="Alcock, A. (1894). Natural history notes from Indian Marine Survey Steamer " Investigator ". Ser. II, no. 1. On the result of deep-sea dredging during the season of 1890 - 1891. Annals and Magazine of Natural History, (6) 13, 400 - 411" type="journal article" year="1894">Alcock, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
: male (cl
|
||
<quantity id="4CD89B66FFCFFFACEF9AFDE49849FDCE" box="[431,540,576,602]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="8" pageNumber="47" unit="mm" value="25.3">25.3 mm</quantity>
|
||
, cw
|
||
<quantity id="4CD89B66FFCFFFACEC66FDE498EAFDCE" box="[595,703,576,602]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="8" pageNumber="47" unit="mm" value="29.6">29.6 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACECE2FDE49947FDCE" box="[727,786,576,602]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
2004.717), Balicasag Island, Panglao, Bohol, Visayas, in tangle nets, coll. local shell fishermen,
|
||
<date id="FF9E1043FFCFFFACECB8FDC69956FDE8" box="[653,771,610,636]" pageId="8" pageNumber="47" value="2003-12">Dec 2003</date>
|
||
;
|
||
<taxonomicName id="4C204D00FFCFFFACED24FDC69EEBFDE8" authority="Ng & Guinot, 1999" authorityName="Ng & Guinot" authorityYear="1999" box="[785,1214,610,636]" class="Malacostraca" family="Progeryonidae" genus="Progeryon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="mus">
|
||
<emphasis id="B954EA91FFCFFFACED24FDC69993FDEF" box="[785,966,610,635]" italics="true" pageId="8" pageNumber="47">Progeryon mus</emphasis>
|
||
Ng &
|
||
<bibRefCitation id="EFB14B72FFCFFFACEA28FDC69EEBFDE8" author="Guinot" box="[1053,1214,610,636]" pageId="8" pageNumber="54" refString="Ng, P. K. L. & Guinot, D. (1999). On a new species of deep-water crab of the genus Progeryon (Decapoda, Brachyura, Geryonidae) from Hawaii. Crustaceana, 72 (7), 685 - 692." type="journal article" year="1999">Guinot, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="549B8821FFCFFFACEAFEFDC69F64FDE8" box="[1227,1329,610,636]" pageId="8" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
male (cl
|
||
<quantity id="4CD89B66FFCFFFACEEA2FD219B57FD0B" box="[151,258,645,671]" metricMagnitude="-2" metricUnit="m" metricValue="3.86" pageId="8" pageNumber="47" unit="mm" value="38.6">38.6 mm</quantity>
|
||
, cw
|
||
<quantity id="4CD89B66FFCFFFACEF00FD219BCAFD0B" box="[309,415,645,671]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="8" pageNumber="47" unit="mm" value="46.5">46.5 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACEF83FD219BA4FD0B" box="[438,497,645,671]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
|
||
1997.442), Nihoa Island, Hawaii,
|
||
<quantity id="4CD89B66FFCFFFACEDB6FD219993FD0B" box="[899,966,645,671]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="8" pageNumber="47" unit="m" value="441.0">441m</quantity>
|
||
depth, coll. 1979;
|
||
<taxonomicName id="4C204D00FFCFFFACEA93FD219B73FD56" authority="Ng & Ng, 2003" authorityName="Ng & Ng" authorityYear="2003" class="Malacostraca" family="Conleyidae" genus="Conleyus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="defodio">
|
||
<emphasis id="B954EA91FFCFFFACEA93FD219F27FD0A" box="[1190,1394,645,670]" italics="true" pageId="8" pageNumber="47">Conleyus defodio</emphasis>
|
||
Ng & Ng, 2003
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="549B8821FFCFFFACEF05FD0C9BC2FD56" box="[304,407,680,706]" pageId="8" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
male (cl,
|
||
<quantity id="4CD89B66FFCFFFACEC3EFD0C9823FD56" box="[523,630,680,706]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="47" unit="mm" value="12.2">12.2 mm</quantity>
|
||
, cw
|
||
<quantity id="4CD89B66FFCFFFACEC9EFD0C9940FD56" box="[683,789,680,706]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="8" pageNumber="47" unit="mm" value="16.4">16.4 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED31AE46FFCFFFACED06FD0C99F5FD56" box="[819,928,680,706]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="47">FLMNH</collectionCode>
|
||
), Luminao fore-reef,
|
||
<quantity id="4CD89B66FFCFFFACEAADFD0C9EA2FD56" box="[1176,1271,680,706]" metricMagnitude="1" metricUnit="m" metricValue="6.7056" pageId="8" pageNumber="47" unit="ft" value="220.0">220 feet</quantity>
|
||
deep, in coral rubble,
|
||
<collectingCountry id="F3377613FFCFFFACEEC5FD6E9B6EFD70" box="[240,315,714,740]" name="Guam" pageId="8" pageNumber="47">Guam</collectingCountry>
|
||
, coll. H. T. Conley,
|
||
<date id="FF9E1043FFCFFFACEC10FD6E98C0FD70" box="[549,661,714,740]" pageId="8" pageNumber="47" value="1999-10">Oct 1999</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F3683FFCFFFACEEA2FCB598BBFCBE" blockId="8.[151,750,784,810]" box="[151,750,784,810]" pageId="8" pageNumber="47">
|
||
<emphasis id="B954EA91FFCFFFACEEA2FCB59B5DFCBD" bold="true" box="[151,264,785,809]" pageId="8" pageNumber="47">
|
||
<collectionCode id="ED31AE46FFCFFFACEEA2FCB59ABAFCBD" box="[151,239,785,809]" pageId="8" pageNumber="47">TABLE</collectionCode>
|
||
1.
|
||
</emphasis>
|
||
Differences between
|
||
<taxonomicName id="4C204D00FFCFFFACEFC7FCB5982FFCBC" box="[498,634,785,808]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
and allies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="8B9F3683FFCFFFACEF79FCEB9F2EF979" pageId="8" pageNumber="47">
|
||
<table id="F920C423FFCF005BEEA9FCEB9FD8F8CC" box="[156,1421,847,1880]" gridcols="6" gridrows="8" pageId="8" pageNumber="47">
|
||
<tr id="351034C1FFCF005BEEA9FCEB9FD8FCF0" box="[156,1421,847,868]" gridrow="0" pageId="8" pageNumber="47" rowspan-0="1">
|
||
<th id="76C15DBDFFCF005BEF79FCEB9844FCF0" box="[332,529,847,868]" gridcol="1" gridrow="0" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C204D00FFCFFFACEF79FCEB9B93FCF0" box="[332,454,847,868]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
</th>
|
||
<th id="76C15DBDFFCF005BEC1FFCEB98A5FCF0" box="[554,752,847,868]" gridcol="2" gridrow="0" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C204D00FFCFFFACEC1FFCEB98FBFCF0" box="[554,686,847,868]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
|
||
</th>
|
||
<th id="76C15DBDFFCF005BED3DFCEB999EFCF0" box="[776,971,847,868]" gridcol="3" gridrow="0" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C204D00FFCFFFACED3DFCEB99C1FCF0" box="[776,916,847,868]" class="Malacostraca" family="Mathildellidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Mathildellidae</taxonomicName>
|
||
</th>
|
||
<th id="76C15DBDFFCF005BEDD3FCEB9EFAFCF0" box="[998,1199,847,868]" gridcol="4" gridrow="0" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C204D00FFCFFFACEDD3FCEB9E27FCF0" box="[998,1138,847,868]" class="Malacostraca" family="Progeryonidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Progeryonidae</taxonomicName>
|
||
</th>
|
||
<th id="76C15DBDFFCF005BEAF1FCEB9FD8FCF0" box="[1220,1421,847,868]" gridcol="5" gridrow="0" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C204D00FFCFFFACEAF1FCEB9F64FCF0" box="[1220,1329,847,868]" class="Malacostraca" family="Conleyidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Conleyidae</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FC269FD8FC71" box="[156,1421,898,997]" gridrow="1" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FC269B7BFC71" box="[156,302,898,997]" gridcol="0" gridrow="1" pageId="8" pageNumber="47">Dorsal surface of carapace</th>
|
||
<td id="76C15DBDFFCF005BEF79FC269844FC71" box="[332,529,898,997]" gridcol="1" gridrow="1" pageId="8" pageNumber="47">With complex pattern of ridges, grooves and dense short setae (Figs. 2, 3A)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFC2698A5FC71" box="[554,752,898,997]" gridcol="2" gridrow="1" pageId="8" pageNumber="47">Smooth to setose, never with any distinct patterns or deep grooves (Figs. 6A–C)</td>
|
||
<td id="76C15DBDFFCF005BED3DFC26999EFC71" box="[776,971,898,997]" gridcol="3" gridrow="1" pageId="8" pageNumber="47">Smooth to setose, never with any distinct patterns or deep grooves (Fig. 6D–F)</td>
|
||
<td id="76C15DBDFFCF005BEDD3FC269EFAFC71" box="[998,1199,898,997]" gridcol="4" gridrow="1" pageId="8" pageNumber="47">With very short felt- like setae, without any distinct patterns or deep grooves (Fig. 6G)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FC269FD8FC71" box="[1220,1421,898,997]" gridcol="5" gridrow="1" pageId="8" pageNumber="47">Sparsely setose, with- out any distinct patterns or deep grooves (Fig. 6H)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FBA69FD8FBDE" box="[156,1421,1026,1098]" gridrow="2" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FBA69B7BFBDE" box="[156,302,1026,1098]" gridcol="0" gridrow="2" pageId="8" pageNumber="47">Mandibles</th>
|
||
<td id="76C15DBDFFCF005BEF79FBA69844FBDE" box="[332,529,1026,1098]" gridcol="1" gridrow="2" pageId="8" pageNumber="47">Dark brown in colour in life and in preserva- tive (Fig. 8A)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFBA698A5FBDE" box="[554,752,1026,1098]" gridcol="2" gridrow="2" pageId="8" pageNumber="47">White to grey in life and in preservative (Fig. 7A)</td>
|
||
<td id="76C15DBDFFCF005BED3DFBA6999EFBDE" box="[776,971,1026,1098]" gridcol="3" gridrow="2" pageId="8" pageNumber="47">White to grey in life and in preservative (Fig. 8B)</td>
|
||
<td id="76C15DBDFFCF005BEDD3FBA69EFAFBDE" box="[998,1199,1026,1098]" gridcol="4" gridrow="2" pageId="8" pageNumber="47">White in preservative (Fig. 8C)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FBA69FD8FBDE" box="[1220,1421,1026,1098]" gridcol="5" gridrow="2" pageId="8" pageNumber="47">White in life and in preservative (Fig.. 8D)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FBC39FD8FB23" box="[156,1421,1127,1207]" gridrow="3" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FBC39B7BFB23" box="[156,302,1127,1207]" gridcol="0" gridrow="3" pageId="8" pageNumber="47">Ventral mar- gins of meri of P2–4</th>
|
||
<td id="76C15DBDFFCF005BEF79FBC39844FB23" box="[332,529,1127,1207]" gridcol="1" gridrow="3" pageId="8" pageNumber="47">With 2 or 3 strong spines (Fig. 4C, D)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFBC398A5FB23" box="[554,752,1127,1207]" gridcol="2" gridrow="3" pageId="8" pageNumber="47">Smooth or granular (Fig. 6A-C)</td>
|
||
<td id="76C15DBDFFCF005BED3DFBC3999EFB23" box="[776,971,1127,1207]" gridcol="3" gridrow="3" pageId="8" pageNumber="47">Usually granular or smooth, occasionally spinous (Fig. 6D-F)</td>
|
||
<td id="76C15DBDFFCF005BEDD3FBC39EFAFB23" box="[998,1199,1127,1207]" gridcol="4" gridrow="3" pageId="8" pageNumber="47">Weakly granular (Fig. 6G)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FBC39FD8FB23" box="[1220,1421,1127,1207]" gridcol="5" gridrow="3" pageId="8" pageNumber="47">Granular (Fig. 6H)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FB719FD8FA89" box="[156,1421,1237,1309]" gridrow="4" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FB719B7BFA89" box="[156,302,1237,1309]" gridcol="0" gridrow="4" pageId="8" pageNumber="47">Thoracic stern- ite 3</th>
|
||
<td id="76C15DBDFFCF005BEF79FB719844FA89" box="[332,529,1237,1309]" gridcol="1" gridrow="4" pageId="8" pageNumber="47">With shallow but dis- tinct longitudinal median line (Fig. 9A)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFB7198A5FA89" box="[554,752,1237,1309]" gridcol="2" gridrow="4" pageId="8" pageNumber="47">Surface even, without longitudinal median line (Fig. 7B, 9B, C)</td>
|
||
<td id="76C15DBDFFCF005BED3DFB71999EFA89" box="[776,971,1237,1309]" gridcol="3" gridrow="4" pageId="8" pageNumber="47">Surface even, without longitudinal median line (Fig. 9D–F)</td>
|
||
<td id="76C15DBDFFCF005BEDD3FB719EFAFA89" box="[998,1199,1237,1309]" gridcol="4" gridrow="4" pageId="8" pageNumber="47">Surface even, without longitudinal median line (Fig. 9G)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FB719FD8FA89" box="[1220,1421,1237,1309]" gridcol="5" gridrow="4" pageId="8" pageNumber="47">Surface even, without longitudinal median line (Fig. 9H)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FA9E9FD8FA47" box="[156,1421,1338,1491]" gridrow="5" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FA9E9B7BFA47" box="[156,302,1338,1491]" gridcol="0" gridrow="5" pageId="8" pageNumber="47">Thoracic stern- ite 7</th>
|
||
<td id="76C15DBDFFCF005BEF79FA9E9844FA47" box="[332,529,1338,1491]" gridcol="1" gridrow="5" pageId="8" pageNumber="47">Very narrow trans- versely, especially towards posterior part which tapers off sud- denly (Fig. 10A)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFA9E98A5FA47" box="[554,752,1338,1491]" gridcol="2" gridrow="5" pageId="8" pageNumber="47">Broad to very broad transversely (Fig. 7D, 10B, C)</td>
|
||
<td id="76C15DBDFFCF005BED3DFA9E999EFA47" box="[776,971,1338,1491]" gridcol="3" gridrow="5" pageId="8" pageNumber="47">
|
||
Relatively broad to somewhat narrow transversely, but not reaching condition of
|
||
<taxonomicName id="4C204D00FFCFFFACED3DFA019920FA2C" box="[776,885,1445,1464]" class="Malacostraca" family="Vultocinidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="family">Vultocinidae</taxonomicName>
|
||
(Fig. 10D–F)
|
||
</td>
|
||
<td id="76C15DBDFFCF005BEDD3FA9E9EFAFA47" box="[998,1199,1338,1491]" gridcol="4" gridrow="5" pageId="8" pageNumber="47">Very narrow trans- versely, tapering gradu- ally posteriorly (Fig. 10G)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FA9E9FD8FA47" box="[1220,1421,1338,1491]" gridcol="5" gridrow="5" pageId="8" pageNumber="47">Relatively broad trans- versely (Fig. 10H)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9FA4B9FD8F9F9" box="[156,1421,1519,1645]" gridrow="6" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9FA4B9B7BF9F9" box="[156,302,1519,1645]" gridcol="0" gridrow="6" pageId="8" pageNumber="47">Episternites 4– 6</th>
|
||
<td id="76C15DBDFFCF005BEF79FA4B9844F9F9" box="[332,529,1519,1645]" gridcol="1" gridrow="6" pageId="8" pageNumber="47">Not prominently expanded laterally (Figs. 9A, 10A, 12A)</td>
|
||
<td id="76C15DBDFFCF005BEC1FFA4B98A5F9F9" box="[554,752,1519,1645]" gridcol="2" gridrow="6" pageId="8" pageNumber="47">Prominently expanded laterally (Figs. 7B, D, 9B, C, 10B, C, 12B, C)</td>
|
||
<td id="76C15DBDFFCF005BED3DFA4B999EF9F9" box="[776,971,1519,1645]" gridcol="3" gridrow="6" pageId="8" pageNumber="47">Expanded laterally to some degree or other, never prominently (Figs. 9D–F, 10D–F, 12D–F)</td>
|
||
<td id="76C15DBDFFCF005BEDD3FA4B9EFAF9F9" box="[998,1199,1519,1645]" gridcol="4" gridrow="6" pageId="8" pageNumber="47">Not prominently expanded laterally (Figs. 9G, 10G, 12G)</td>
|
||
<td id="76C15DBDFFCF005BEAF1FA4B9FD8F9F9" box="[1220,1421,1519,1645]" gridcol="5" gridrow="6" pageId="8" pageNumber="47">Expanded laterally slightly (Figs. 9H, 10H, 12H)</td>
|
||
</tr>
|
||
<tr id="351034C1FFCF005BEEA9F92E9FD8F8CC" box="[156,1421,1674,1880]" gridrow="7" pageId="8" pageNumber="47">
|
||
<th id="76C15DBDFFCF005BEEA9F92E9B7BF8CC" box="[156,302,1674,1880]" gridcol="0" gridrow="7" pageId="8" pageNumber="47">Episternite 7</th>
|
||
<td id="76C15DBDFFCF005BEF79F92E9844F8CC" box="[332,529,1674,1880]" gridcol="1" gridrow="7" pageId="8" pageNumber="47">Posterior angle promi- nently prolonged back- wards to form long spur which just touches coxa of P5 (Fig. 13A, B)</td>
|
||
<td id="76C15DBDFFCF005BEC1FF92E98A5F8CC" box="[554,752,1674,1880]" gridcol="2" gridrow="7" pageId="8" pageNumber="47">Expanded laterally to form broad plate which covers anterior edge of coxa of P5 (Fig. 7F, 13C, D)</td>
|
||
<td id="76C15DBDFFCF005BED3DF92E999EF8CC" box="[776,971,1674,1880]" gridcol="3" gridrow="7" pageId="8" pageNumber="47">Not expanded later- ally, not touching coxa of fourth ambulatory leg, or forming small plate which partially covers or just touches anterior edge of coxa of P5 (Fig. 13E)</td>
|
||
<td id="76C15DBDFFCF005BEDD3F92E9EFAF8CC" box="[998,1199,1674,1880]" gridcol="4" gridrow="7" pageId="8" pageNumber="47">Not prominently expanded but with small plate which just touches anterior edge of coxa of P5 (as in Fig. 13E)</td>
|
||
<td id="76C15DBDFFCF005BEAF1F92E9FD8F8CC" box="[1220,1421,1674,1880]" gridcol="5" gridrow="7" pageId="8" pageNumber="47">Expanded laterally to form plate which covers anterior edge of coxa of P5 (Fig. 13F)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C33A6508FFCFFFACEB21F8D19FD8F813" box="[1300,1421,1909,1927]" pageId="8" pageNumber="47" type="description">
|
||
<paragraph id="8B9F3683FFCFFFACEB21F8D19FD8F813" blockId="8.[1300,1421,1909,1927]" box="[1300,1421,1909,1927]" pageId="8" pageNumber="47">
|
||
<emphasis id="B954EA91FFCFFFACEB21F8D19FD8F813" box="[1300,1421,1909,1927]" italics="true" pageId="8" pageNumber="47">...... continued</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |