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<mods:title id="A3F1E9023FAAEE46B26D3844F897E93C">On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata)</mods:title>
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<mods:namePart id="F6FF2E931D329BD7E859E5A8ED6E446F">Ng, Bee Yan Lee Ngan Kee</mods:namePart>
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<mods:affiliation id="5313B49CB3A6E5F73A03DDCE2A6BEDD4">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<emphasis id="7E957179FFF3FFD0FF30FF29FE66E9FC" bold="true" box="[151,439,151,178]" italics="true" pageId="4" pageNumber="221">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFF3FFD0FE19FF29FD4FE9FC" author="Edmondson, C. H." box="[446,670,151,177]" pageId="4" pageNumber="221" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
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(
<figureCitation id="D4DAB1EEFFF3FFD0FF38FF05FED2E998" box="[159,259,187,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FEA8FF05FE9EE998" box="[271,335,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">3A–F</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FEFCFF05FE71E998" box="[347,416,187,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A–C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FE0CFF05FE20E99B" box="[427,497,187,214]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5C, D</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FE5BFF05FD93E998" box="[508,578,187,213]" captionStart="FIGURE 6" captionStartId="11.[151,250,1114,1138]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-18@11.[151,1435,181,1089]" captionTargetPageId="11" captionText="FIGURE 6. P5 dorsal view. A–C, Clistocoeloma suvaense Edmondson, 1951; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, C. villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, C. nobile n. sp.; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea." figureDoi="http://doi.org/10.5281/zenodo.8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="4" pageNumber="221">6A–C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FDEAFF05FD43E99B" box="[589,658,187,214]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="4" pageNumber="221">8A, B</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD39FF05FD33E998" box="[670,738,187,213]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="4" pageNumber="221">9E–H</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD4AFF05FC90E998" box="[749,833,187,213]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[216,1372,183,1917]" captionTargetId="figure-18@15.[214,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 10. Left G1. A–D, Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, C. suvaense Edmondson, 1951, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="4" pageNumber="221">10A–H</figureCitation>
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<treatmentCitation id="CD408B7AFFF3FFD0FF30FEBEFD62E855" author="Edmondson, C. H." box="[151,691,255,280]" page="241" pageId="4" pageNumber="221" year="1951">
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<emphasis id="7E957179FFF3FFD0FF30FEBEFE5AE85A" box="[151,395,256,280]" italics="true" pageId="4" pageNumber="221">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFF3FFD0FE35FEBEFD62E855" author="Edmondson, C. H." box="[402,691,255,280]" pageId="4" pageNumber="221" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951: 238–241</bibRefCitation>
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, figs. 37a–i, 38 [
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locality: reef in Suva Habor, 
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<taxonomicName id="8BE1D6E8FFF3FFD0FF30FEA0FE5AE87B" ID-CoL="5YZJP" authorityName="Edmondson" authorityYear="1951" box="[151,395,286,310]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF3FFD0FF30FEA0FE5AE87B" box="[151,395,286,310]" italics="true" pageId="4" pageNumber="221">Clistocoeloma suvaense</emphasis>
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— 
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(list).— 
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<bibRefCitation id="2870D09AFFF3FFD0FCAAFEA0FC3FE87B" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S." box="[781,1006,286,310]" pageId="4" pageNumber="221" pagination="33 - 63" refId="ref10699" refString="Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. &amp; Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63." type="journal article" year="2004">
Komai 
<emphasis id="7E957179FFF3FFD0FCFBFEA1FC5EE87B" box="[860,911,286,310]" italics="true" pageId="4" pageNumber="221">et al.</emphasis>
2004: 41
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(list).— 
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<bibRefCitation id="2870D09AFFF3FFD0FBE5FEA0FAD9E87B" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1090,1288,286,310]" pageId="4" pageNumber="221" pagination="1 - 286" refId="ref11595" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="7E957179FFF3FFD0FBCEFEA1FB4DE87B" box="[1129,1180,286,310]" italics="true" pageId="4" pageNumber="221">et al.</emphasis>
2008: 220
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(list).
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<emphasis id="7E957179FFF3FFD0FF30FEDBFE94E833" bold="true" box="[151,325,356,383]" pageId="4" pageNumber="221">Type material.</emphasis>
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<typeStatus id="935A13C9FFF3FFD0FEEBFEDBFE6CE832" box="[332,445,357,383]" pageId="4" pageNumber="221">Holotype</typeStatus>
:
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male (21.0 × 
<quantity id="8B19008EFFF3FFD0FDF8FEDBFD15E832" box="[607,708,357,383]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="4" pageNumber="221" unit="mm" value="18.8">18.8 mm</quantity>
) (
<specimenCode id="1C470510FFF3FFD0FD7DFEDBFCBCE833" box="[730,877,356,383]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="4" pageNumber="221" type="Museum">BPBM 3976</specimenCode>
), 
<location id="493EFBB0FFF3FFD0FC27FEDBFC6AE832" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FC27FEDBFC6AE832" box="[896,955,357,383]" country="Fiji" name="Suva" pageId="4" pageNumber="221">Suva</location>
, 
<collectingCountry id="34F6EDFBFFF3FFD0FC61FEDAFC21E833" box="[966,1008,356,382]" name="Fiji" pageId="4" pageNumber="221">Fiji</collectingCountry>
, no collection data.
</materialsCitation>
</paragraph>
<paragraph id="4C5EAD6BFFF3FFD0FF60FE36FACCEA46" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
<emphasis id="7E957179FFF3FFD0FF60FE36FE2EE8EF" bold="true" box="[199,511,392,418]" pageId="4" pageNumber="221">Other material examined:</emphasis>
<materialsCitation id="FC89A736FFF3FFD0FDA1FE36FB24E88A" collectingDate="2019-07-22" collectingDateMax="2019-07-26" collectingDateMin="2019-07-22" collectionCode="ZRC" collectorName="B. Y. Lee &amp; S. Choy &amp; B. Rashni" country="Fiji" county="Fiji" latitude="-16.088612" location="Navaca" longLatPrecision="21" longitude="178.04501" municipality="Taveuni Island" pageId="4" pageNumber="221" specimenCode="ZRC 2022.0985, ZRC 2022.0986" specimenCount="2" specimenCount-male="2">
<emphasis id="7E957179FFF3FFD0FDA1FE36FDE8E8EF" bold="true" box="[518,569,392,418]" pageId="4" pageNumber="221">
<collectingCountry id="34F6EDFBFFF3FFD0FDA1FE36FDE5E8EF" box="[518,564,392,418]" name="Fiji" pageId="4" pageNumber="221">Fiji</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFF3FFD0FDE6FE37FD5DE8EF" box="[577,652,392,419]" pageId="4" pageNumber="221" type="male">1 male</specimenCount>
(18.8 × 
<quantity id="8B19008EFFF3FFD0FD4AFE36FC83E8EE" box="[749,850,392,419]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="4" pageNumber="221" unit="mm" value="16.9">16.9 mm</quantity>
) (
<specimenCode id="1C470510FFF3FFD0FCCEFE37FBF4E8EE" box="[873,1061,392,419]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC 2022.0985</specimenCode>
), river mouth, 
<location id="493EFBB0FFF3FFD0FB6DFE37FACFE8EE" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FB6DFE37FACFE8EE" box="[1226,1310,393,419]" country="Fiji" county="Fiji" latitude="-16.088612" longLatPrecision="21" longitude="178.04501" municipality="Taveuni Island" name="Navaca" pageId="4" pageNumber="221">Navaca</location>
mangrove, 
<collectingMunicipality id="AC3A3711FFF3FFD0FF30FE12FEEEE88B" box="[151,319,428,454]" pageId="4" pageNumber="221">Taveuni Island</collectingMunicipality>
, 
<collectingCounty id="A53FD5E7FFF3FFD0FEECFE12FEA4E88B" box="[331,373,428,454]" pageId="4" pageNumber="221">Fiji</collectingCounty>
, 
<geoCoordinate id="29D5CBACFFF3FFD0FE26FE12FDD7E88A" box="[385,518,428,455]" degrees="16" direction="south" minutes="05" orientation="latitude" pageId="4" pageNumber="221" precision="15" seconds="19" value="-16.088612">16°05’19”S</geoCoordinate>
<geoCoordinate id="29D5CBACFFF3FFD0FDAAFE12FD79E88A" box="[525,680,428,455]" degrees="179" direction="east" minutes="05" orientation="longitude" pageId="4" pageNumber="221" precision="15" seconds="48" value="179.09666">179°05’48”E</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FD4BFE13FC83E88A" box="[748,850,429,455]" pageId="4" pageNumber="221">B.Y. Lee</collectorName>
, 
<collectorName id="E114C8BDFFF3FFD0FCF9FE13FC68E88B" box="[862,953,428,455]" pageId="4" pageNumber="221">S. Choy</collectorName>
&amp; 
<collectorName id="E114C8BDFFF3FFD0FC7AFE13FB9EE88B" box="[989,1103,428,455]" pageId="4" pageNumber="221">B. Rashni</collectorName>
, 
<date id="385F8BABFFF3FFD0FBFDFE12FB20E88A" box="[1114,1265,428,455]" pageId="4" pageNumber="221" value="2019-07-26">
<collectingDate id="281B7243FFF3FFD0FBFDFE12FB20E88A" box="[1114,1265,428,455]" pageId="4" pageNumber="221" value="2019-07-26">26 July 2019</collectingDate>
</date>
.
</materialsCitation>
— 
<materialsCitation id="FC89A736FFF3FFD0FAB3FE13FBCAEB42" collectingDate="2019-07-22" collectingDateMax="2019-07-26" collectingDateMin="2019-07-22" collectionCode="ZRC" collectorName="B. Y. Lee &amp; B. Rashni" country="Fiji" county="Fiji" latitude="-16.088612" location="Navaca" longLatPrecision="21" longitude="178.04501" municipality="Taveuni Island" pageId="4" pageNumber="221" specimenCode="ZRC 2022.0985, ZRC 2022.0986" specimenCount="2" specimenCount-male="2">
<specimenCount id="5AE766E2FFF3FFD0FAB3FE13FA8BE88B" box="[1300,1370,428,455]" pageId="4" pageNumber="221" type="male">1 male</specimenCount>
(17.8 × 
<quantity id="8B19008EFFF3FFD0FF08FE6EFEC4E8A6" box="[175,277,464,491]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="4" pageNumber="221" unit="mm" value="16.3">16.3 mm</quantity>
) (
<specimenCode id="1C470510FFF3FFD0FE8AFE6FFE38E8A7" box="[301,489,464,491]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC 2022.0986</specimenCode>
), mangrove at Suva Point Beach, near My Suva Picnic Park and Suva City Council Park, Fiji, 
<geoCoordinate id="29D5CBACFFF3FFD0FEA8FE4BFE49EB42" box="[271,408,501,527]" degrees="18" direction="south" minutes="00" orientation="latitude" pageId="4" pageNumber="221" precision="15" seconds="28" value="-18.007778">18°00’28”S</geoCoordinate>
<geoCoordinate id="29D5CBACFFF3FFD0FE06FE4BFDE6EB42" box="[417,567,501,527]" degrees="178" direction="east" minutes="02" orientation="longitude" pageId="4" pageNumber="221" precision="15" seconds="42" value="178.04501">178°02’42”E</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FDDBFE4BFD31EB42" box="[636,736,501,527]" pageId="4" pageNumber="221">B.Y. Lee</collectorName>
&amp; 
<collectorName id="E114C8BDFFF3FFD0FCA2FE4BFCA9EB43" box="[773,888,500,527]" pageId="4" pageNumber="221">B. Rashni</collectorName>
, 
<date id="385F8BABFFF3FFD0FC24FE4BFBCAEB42" box="[899,1051,500,527]" pageId="4" pageNumber="221" value="2019-07-22">
<collectingDate id="281B7243FFF3FFD0FC24FE4BFBCAEB42" box="[899,1051,500,527]" pageId="4" pageNumber="221" value="2019-07-22">22 July 2019</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="FC89A736FFF3FFD0FB99FE4BFD1EEB1A" collectingDate="2019-07-25" collectionCode="ZRC" collectorName="B. Y. Lee &amp; S. Choy &amp; B. Rashni" country="Fiji" county="Fiji" latitude="-16.074446" location="Bure Bure" longLatPrecision="21" longitude="-179.08777" municipality="Taveuni Island" pageId="4" pageNumber="221" specimenCode="ZRC 2022.0987" specimenCount="1" specimenCount-male="1">
<specimenCount id="5AE766E2FFF3FFD0FB99FE4BFB54EB43" box="[1086,1157,500,527]" pageId="4" pageNumber="221" type="male">1 male</specimenCount>
(12.1 × 
<quantity id="8B19008EFFF3FFD0FB40FE4BFA9CEB42" box="[1255,1357,501,527]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="4" pageNumber="221" unit="mm" value="10.9">10.9 mm</quantity>
) (
<specimenCode id="1C470510FFF3FFD0FAC3FE4BFEC2EB7E" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC 2022.0987</specimenCode>
), 
<location id="493EFBB0FFF3FFD0FE83FDA7FE46EB7E" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FE83FDA7FE46EB7E" box="[292,407,537,563]" country="Fiji" county="Fiji" latitude="-16.074446" longLatPrecision="21" longitude="-179.08777" municipality="Taveuni Island" name="Bure Bure" pageId="4" pageNumber="221">Bure Bure</location>
mangrove, 
<location id="493EFBB0FFF3FFD0FDBDFDA6FD78EB7F" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FDBDFDA6FD78EB7F" box="[538,681,536,562]" country="Fiji" county="Fiji" latitude="-16.074446" longLatPrecision="21" longitude="-179.08777" municipality="Taveuni Island" name="Naqai Creek" pageId="4" pageNumber="221">Naqai Creek</location>
, near 
<location id="493EFBB0FFF3FFD0FD4EFDA6FC50EB7F" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FD4EFDA6FC50EB7F" box="[745,897,536,562]" country="Fiji" county="Fiji" latitude="-16.074446" longLatPrecision="21" longitude="-179.08777" municipality="Taveuni Island" name="Naqai Bridge" pageId="4" pageNumber="221">Naqai Bridge</location>
, 
<collectingMunicipality id="AC3A3711FFF3FFD0FC2CFDA6FBE3EB7F" box="[907,1074,536,562]" pageId="4" pageNumber="221">Taveuni Island</collectingMunicipality>
, 
<collectingCounty id="A53FD5E7FFF3FFD0FB9BFDA6FBB7EB7F" box="[1084,1126,536,562]" pageId="4" pageNumber="221">Fiji</collectingCounty>
, 
<geoCoordinate id="29D5CBACFFF3FFD0FBD7FDA6FB25EB7F" box="[1136,1268,536,562]" degrees="16" direction="south" minutes="04" orientation="latitude" pageId="4" pageNumber="221" precision="15" seconds="28" value="-16.074446">16°04’28”S</geoCoordinate>
<geoCoordinate id="29D5CBACFFF3FFD0FB5DFDA6FA46EB7E" box="[1274,1431,536,563]" degrees="179" direction="west" minutes="05" orientation="longitude" pageId="4" pageNumber="221" precision="15" seconds="16" value="-179.08777">179°05’16”W</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FF68FD83FEE4EB1A" box="[207,309,573,599]" pageId="4" pageNumber="221">B.Y. Lee</collectorName>
, 
<collectorName id="E114C8BDFFF3FFD0FEE7FD83FE4AEB1B" box="[320,411,572,599]" pageId="4" pageNumber="221">S. Choy</collectorName>
&amp; 
<collectorName id="E114C8BDFFF3FFD0FE18FD83FDE0EB1B" box="[447,561,572,599]" pageId="4" pageNumber="221">B. Rashni</collectorName>
, 
<date id="385F8BABFFF3FFD0FD9BFD82FD1EEB1A" box="[572,719,572,599]" pageId="4" pageNumber="221" value="2019-07-25">
<collectingDate id="281B7243FFF3FFD0FD9BFD82FD1EEB1A" box="[572,719,572,599]" pageId="4" pageNumber="221" value="2019-07-25">25 July 2019</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFF3FFD0FD56FD83FC97EB1B" box="[753,838,572,599]" pageId="4" pageNumber="221" type="male">2 males</specimenCount>
(13.7 × 
<quantity id="8B19008EFFF3FFD0FC01FD83FBDCEB1A" box="[934,1037,573,599]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="4" pageNumber="221" unit="mm" value="12.2">12.2 mm</quantity>
, 12.5 × 
<quantity id="8B19008EFFF3FFD0FBCFFD83FB1EEB1A" box="[1128,1231,573,599]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="4" pageNumber="221" unit="mm" value="11.4">11.4 mm</quantity>
), 
<materialsCitation id="FC89A736FFF3FFD0FB45FD83FD4CEBD2" collectingDate="2019-07-26" collectionCode="ZRC" collectorName="B. Y. Lee &amp; S. Choy &amp; B. Rashni" country="Fiji" county="Fiji" latitude="-16.088612" location="Navaca" longLatPrecision="21" longitude="179.09666" municipality="Taveuni Island" pageId="4" pageNumber="221" specimenCode="ZRC 2022.0988" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFF3FFD0FB45FD83FA92EB1B" box="[1250,1347,572,599]" pageId="4" pageNumber="221" type="female">1 female</specimenCount>
(13.3 × 
<quantity id="8B19008EFFF3FFD0FF30FDDFFF2DEB36" box="[151,252,609,635]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="4" pageNumber="221" unit="mm" value="11.9">11.9 mm</quantity>
) (
<specimenCode id="1C470510FFF3FFD0FEB4FDDFFE01EB36" box="[275,464,609,635]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC 2022.0988</specimenCode>
), river mouth, 
<location id="493EFBB0FFF3FFD0FDD1FDDFFD1BEB36" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FDD1FDDFFD1BEB36" box="[630,714,609,635]" country="Fiji" county="Fiji" latitude="-16.088612" longLatPrecision="21" longitude="179.09666" municipality="Taveuni Island" name="Navaca" pageId="4" pageNumber="221">Navaca</location>
mangrove, 
<collectingMunicipality id="AC3A3711FFF3FFD0FCF6FDDEFC28EB37" box="[849,1017,608,634]" pageId="4" pageNumber="221">Taveuni Island</collectingMunicipality>
, 
<collectingCounty id="A53FD5E7FFF3FFD0FBA1FDDEFBE1EB37" box="[1030,1072,608,634]" pageId="4" pageNumber="221">Fiji</collectingCounty>
, 
<geoCoordinate id="29D5CBACFFF3FFD0FB9BFDDEFB10EB36" box="[1084,1217,608,635]" degrees="16" direction="south" minutes="05" orientation="latitude" pageId="4" pageNumber="221" precision="15" seconds="19" value="-16.088612">16°05’19”S</geoCoordinate>
<geoCoordinate id="29D5CBACFFF3FFD0FB6FFDDEFA8FEB36" box="[1224,1374,608,635]" degrees="179" direction="east" minutes="05" orientation="longitude" pageId="4" pageNumber="221" precision="15" seconds="48" value="179.09666">179°05’48”E</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FF30FD3BFF2CEBD2" box="[151,253,645,671]" pageId="4" pageNumber="221">B.Y. Lee</collectorName>
, 
<collectorName id="E114C8BDFFF3FFD0FEAEFD3BFEB4EBD3" box="[265,357,644,671]" pageId="4" pageNumber="221">S. Choy</collectorName>
&amp; 
<collectorName id="E114C8BDFFF3FFD0FE2DFD3BFE2CEBD3" box="[394,509,644,671]" pageId="4" pageNumber="221">B. Rashni</collectorName>
, 
<date id="385F8BABFFF3FFD0FDAEFD3AFD4CEBD2" box="[521,669,644,671]" pageId="4" pageNumber="221" value="2019-07-26">
<collectingDate id="281B7243FFF3FFD0FDAEFD3AFD4CEBD2" box="[521,669,644,671]" pageId="4" pageNumber="221" value="2019-07-26">26 July 2019</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="5AE766E2FFF3FFD0FD18FD3BFCC4EBD3" box="[703,789,644,671]" pageId="4" pageNumber="221" type="male">2 males</specimenCount>
(17.1 × 
<quantity id="8B19008EFFF3FFD0FCDFFD3AFC0EEBD2" box="[888,991,644,671]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="221" unit="mm" value="15.5">15.5 mm</quantity>
, 15.2 × 
<quantity id="8B19008EFFF3FFD0FB9BFD3BFB74EBD2" box="[1084,1189,645,671]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="221" unit="mm" value="14.2">14.2 mm</quantity>
), 2 ovigerous females (16.6 × 15.0 mm, 16.2 × 
<quantity id="8B19008EFFF3FFD0FE15FD16FDCAEB8E" box="[434,539,680,707]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="221" unit="mm" value="14.5">14.5 mm</quantity>
), 
<materialsCitation id="FC89A736FFF3FFD0FD89FD17FEFDEA46" collectingDate="2019-07-22" collectionCode="ZRC" collectorName="B. Y. Lee &amp; B. Rashni" country="Fiji" county="Fiji" latitude="-18.007778" location="Suva Point Beach" longLatPrecision="21" longitude="178.04501" municipality="Suva City Council Park" pageId="4" pageNumber="221" specimenCode="ZRC 2022.0989" specimenCount="1" specimenCount-female="1">
<specimenCount id="5AE766E2FFF3FFD0FD89FD17FD5EEB8F" box="[558,655,680,707]" pageId="4" pageNumber="221" type="female">1 female</specimenCount>
(18.7 × 17.0 mm) (
<specimenCode id="1C470510FFF3FFD0FCCBFD17FBF9EB8E" box="[876,1064,681,707]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC 2022.0989</specimenCode>
), mangrove at 
<location id="493EFBB0FFF3FFD0FB6AFD17FA49EB8F" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FB6AFD17FA49EB8F" box="[1229,1432,680,707]" country="Fiji" county="Fiji" latitude="-18.007778" longLatPrecision="21" longitude="178.04501" municipality="Suva City Council Park" name="Suva Point Beach" pageId="4" pageNumber="221">Suva Point Beach</location>
, near 
<location id="493EFBB0FFF3FFD0FF69FD73FE6AEBAB" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FF69FD73FE6AEBAB" box="[206,443,716,743]" country="Fiji" county="Fiji" latitude="-18.007778" longLatPrecision="21" longitude="178.04501" municipality="Suva City Council Park" name="My Suva Picnic Park" pageId="4" pageNumber="221">My Suva Picnic Park</location>
and 
<collectingMunicipality id="AC3A3711FFF3FFD0FE56FD73FD2DEBAB" box="[497,764,716,743]" pageId="4" pageNumber="221">Suva City Council Park</collectingMunicipality>
, 
<collectingCounty id="A53FD5E7FFF3FFD0FCAFFD72FCE3EBAB" box="[776,818,716,742]" pageId="4" pageNumber="221">Fiji</collectingCounty>
, 
<geoCoordinate id="29D5CBACFFF3FFD0FC9BFD73FC10EBAA" box="[828,961,717,743]" degrees="18" direction="south" minutes="00" orientation="latitude" pageId="4" pageNumber="221" precision="15" seconds="28" value="-18.007778">18°00’28”S</geoCoordinate>
<geoCoordinate id="29D5CBACFFF3FFD0FC6FFD73FB8FEBAA" box="[968,1118,717,743]" degrees="178" direction="east" minutes="02" orientation="longitude" pageId="4" pageNumber="221" precision="15" seconds="42" value="178.04501">178°02’42”E</geoCoordinate>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FB06FD73FAD5EBAA" box="[1185,1284,717,743]" pageId="4" pageNumber="221">B.Y. Lee</collectorName>
&amp; 
<collectorName id="E114C8BDFFF3FFD0FA80FD73FA49EBAB" box="[1319,1432,716,743]" pageId="4" pageNumber="221">B. Rashni</collectorName>
, 
<date id="385F8BABFFF3FFD0FF30FD4FFEFDEA46" box="[151,300,752,779]" pageId="4" pageNumber="221" value="2019-07-22">
<collectingDate id="281B7243FFF3FFD0FF30FD4FFEFDEA46" box="[151,300,752,779]" pageId="4" pageNumber="221" value="2019-07-22">22 July 2019</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FC89A736FFF3FFD0FE91FD4EFACCEA46" box="[310,1309,752,779]" collectionCode="ZRC" collectorName="Godeffroy Museum." country="Samoa" location="Upolu" pageId="4" pageNumber="221" specimenCount="1" specimenCount-male="1">
<emphasis id="7E957179FFF3FFD0FE91FD4EFE41EA47" bold="true" box="[310,400,752,778]" pageId="4" pageNumber="221">
<collectingCountry id="34F6EDFBFFF3FFD0FE91FD4EFE5AEA47" box="[310,395,752,778]" name="Samoa" pageId="4" pageNumber="221">Samoa</collectingCountry>
:
</emphasis>
<specimenCount id="5AE766E2FFF3FFD0FE30FD4FFE33EA47" box="[407,482,752,779]" pageId="4" pageNumber="221" type="male">1 male</specimenCount>
(16.2 × 
<quantity id="8B19008EFFF3FFD0FDE5FD4FFD76EA46" box="[578,679,753,779]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="4" pageNumber="221" unit="mm" value="14.3">14.3 mm</quantity>
) (
<collectionCode id="2AF035AEFFF3FFD0FD1AFD4FFD24EA46" box="[701,757,753,779]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="221">ZRC</collectionCode>
1973.11.2.490) 
<location id="493EFBB0FFF3FFD0FC0AFD4EFC27EA47" LSID="urn:lsid:plazi:treatment:C4481C7DFFF3FFD3FF30FF29FDD1EE90:493EFBB0FFF3FFD0FC0AFD4EFC27EA47" box="[941,1014,752,778]" country="Samoa" name="Upolu" pageId="4" pageNumber="221">Upolu</location>
, coll. 
<collectorName id="E114C8BDFFF3FFD0FB9EFD4EFACCEA46" box="[1081,1309,752,779]" pageId="4" pageNumber="221">Godeffroy Museum.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFF3FFD0FF60FCAAFBFFEC4E" pageId="4" pageNumber="221" type="diagnosis">
<paragraph id="4C5EAD6BFFF3FFD0FF60FCAAFBFFEC4E" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
<emphasis id="7E957179FFF3FFD0FF60FCAAFE93EA63" bold="true" box="[199,322,788,814]" pageId="4" pageNumber="221">Diagnosis.</emphasis>
Carapace slightly wider than long, almost square; postfrontal lobes distinct, divided by longitudinal grooves; median pair broad, prominent, nearly equal in proportion; distant pair broad, prominent, nearly equal in proportion (
<figureCitation id="D4DAB1EEFFF3FFD0FEB9FCE2FE52EA3A" box="[286,387,860,887]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FE37FCE3FE65EA3A" box="[400,436,861,887]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FE65FCE3FE39EA3A" box="[450,488,861,887]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A</figureCitation>
). Anterolateral margin of carapace with distinct external orbital angle, blunt second tooth blunt separated by distinct shallow groove from external orbital tooth; posterolateral margin convex (
<figureCitation id="D4DAB1EEFFF3FFD0FAC3FC3EFF6DEAF2" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FF6FFC1BFF3DEAF2" box="[200,236,933,959]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FF5EFC1BFECEEAF2" box="[249,287,933,959]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A</figureCitation>
). Inner orbital angle present, truncate, slight gap between inner orbital angle and frontal margin (
<figureCitation id="D4DAB1EEFFF3FFD0FAC3FC1AFF6BEAAE" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FF6EFC77FF3FEAAE" box="[201,238,969,995]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4C</figureCitation>
). Chelipeds dorsal surface of dactylus of chela with 22–27 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (
<figureCitation id="D4DAB1EEFFF3FFD0FCC4FC52FC17ED4A" box="[867,966,1004,1031]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FC72FC52FC29ED4A" box="[981,1016,1004,1031]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5C</figureCitation>
); absent or weak granules in female and juveniles; presence of single longitudinal pectinated ridge with 33–36 chitinous comb-like tubercles on dorsal surface of chela; row of 9–11 granules behind pectinated ridge on chela (
<figureCitation id="D4DAB1EEFFF3FFD0FC54FB8AFB86ED02" box="[1011,1111,1076,1103]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FBC1FB8AFB5BED02" box="[1126,1162,1076,1103]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5C</figureCitation>
); inner surface of palm granulated, with single vertical row of 6–10 granules in adult males (
<figureCitation id="D4DAB1EEFFF3FFD0FC30FBE6FC22ED3E" box="[919,1011,1112,1139]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3F</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FC58FBE6FBF5ED3E" box="[1023,1060,1112,1139]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5D</figureCitation>
); absent in females and juveniles. P5 merus relatively short, wide, length approximately 0.39–0.46 times width (
<figureCitation id="D4DAB1EEFFF3FFD0FC5CFBC2FBA7EDDB" box="[1019,1142,1148,1174]" captionStart="FIGURE 6" captionStartId="11.[151,250,1114,1138]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-18@11.[151,1435,181,1089]" captionTargetPageId="11" captionText="FIGURE 6. P5 dorsal view. A–C, Clistocoeloma suvaense Edmondson, 1951; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, C. villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, C. nobile n. sp.; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea." figureDoi="http://doi.org/10.5281/zenodo.8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="4" pageNumber="221">Fig. 6A–C</figureCitation>
). G1 straight, distal region relatively wide; chitinous tip wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (
<figureCitation id="D4DAB1EEFFF3FFD0FD32FB7AFCCBED92" box="[661,794,1220,1247]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="4" pageNumber="221">Figs. 9E–H</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FC80FB7BFCAEED92" box="[807,895,1221,1247]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[216,1372,183,1917]" captionTargetId="figure-18@15.[214,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 10. Left G1. A–D, Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, C. suvaense Edmondson, 1951, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="4" pageNumber="221">10A–H</figureCitation>
). Female vulvae with central operculum, ovate, slightly protruded; sternal vulvar cover slightly raised above operculum (
<figureCitation id="D4DAB1EEFFF3FFD0FC6EFB56FBF0EC4E" box="[969,1057,1256,1283]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="4" pageNumber="221">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFF3FFD3FF60FAB2FE38E89C" lastPageId="7" lastPageNumber="224" pageId="4" pageNumber="221" type="description">
<paragraph id="4C5EAD6BFFF3FFD0FF60FAB2FD3BECB2" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
<emphasis id="7E957179FFF3FFD0FF60FAB2FE88EC6B" bold="true" box="[199,345,1292,1318]" pageId="4" pageNumber="221">Description.</emphasis>
Carapace slightly wider than long, almost square; covered with dense coat of setae, with small tufts of setae resembling tubercles; carapace margin fringed with dense, short setae; carapace smooth when denuded. Postfrontal lobes distinct, divided by longitudinal grooves; median pair broad, prominent, nearly equal in proportion; distant pair broad, prominent, nearly equal in proportion (
<figureCitation id="D4DAB1EEFFF3FFD0FCBBFAC6FC51ECDE" box="[796,896,1400,1427]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FC2CFAC7FC7EECDE" box="[907,943,1401,1427]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FC1CFAC7FC33ECDE" box="[955,994,1401,1427]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A</figureCitation>
). Anterolateral margin of carapace with distinct external orbital angle, blunt second tooth blunt separated by distinct shallow groove from external orbital tooth; posterolateral margin convex (
<figureCitation id="D4DAB1EEFFF3FFD0FD9AFA7EFD72EC96" box="[573,675,1472,1499]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD16FA7FFD04EC96" box="[689,725,1473,1499]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD44FA7FFCDBEC96" box="[739,778,1473,1499]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A</figureCitation>
). Inner orbital angle present, truncate, slight gap between inner orbital angle and frontal margin (
<figureCitation id="D4DAB1EEFFF3FFD0FDECFA5AFD7CECB2" box="[587,685,1508,1535]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD1EFA5BFD0CECB2" box="[697,733,1509,1535]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4C</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFF3FFD0FF60F9B7FD8DEF0A" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
Antenna short. Antennules folding transversely. Third maxilliped with merus relatively ovate. Epistomial margin relatively narrow (
<figureCitation id="D4DAB1EEFFF3FFD0FE1BF992FDCFEF0A" box="[444,542,1580,1607]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD8DF993FD9EEF0A" box="[554,591,1581,1607]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4C</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFF3FFD0FF60F9EEFD79EE0F" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
Chelipeds subequal in size; large, robust in male; outer surface of palm smooth when denuded; dorsal surface of carpus with small tufts of setae resembling tubercles, smooth when denuded; dorsal surface of dactylus of chela with 22–27 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (
<figureCitation id="D4DAB1EEFFF3FFD0FF38F902FED3EF9A" box="[159,258,1724,1751]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FEA8F902FEE2EF9A" box="[271,307,1724,1751]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5C</figureCitation>
); absent or weak granules in female and juveniles. Presence of single longitudinal pectinated ridge with 33–36 chitinous comb-like tubercles on dorsal surface of chela; row of 9–11 granules behind pectinated ridge on chela (
<figureCitation id="D4DAB1EEFFF3FFD0FEA2F8BAFEB7EE52" box="[261,358,1796,1823]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FED5F8BAFE46EE52" box="[370,407,1796,1823]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5C</figureCitation>
). Inner surface of palm granulated, with single vertical row of 6–10 granules in adult males (
<figureCitation id="D4DAB1EEFFF3FFD0FF38F896FF2DEE0E" box="[159,252,1832,1859]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="4" pageNumber="221">Figs. 3F</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FEAFF896FEFCEE0E" box="[264,301,1832,1859]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="4" pageNumber="221">5D</figureCitation>
); absent in females and juveniles.
</paragraph>
<paragraph id="4C5EAD6BFFF3FFD0FF60F8F2FDF8EEE3" blockId="4.[151,1437,356,1966]" pageId="4" pageNumber="221">
P2–P5 with dense coat of setae, with small tufts of setae resembling tubercles, longer setae on dorsal and ventral margins except distal portion of dactylus (
<figureCitation id="D4DAB1EEFFF3FFD0FDD8F8CEFD37EEC6" box="[639,742,1904,1931]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="4" pageNumber="221">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF3FFD0FD53F8CFFCCAEEC6" box="[756,795,1905,1931]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="4" pageNumber="221">4A</figureCitation>
); P5 merus relatively short, wide, length approximately 0.39–0.46 times width (
<figureCitation id="D4DAB1EEFFF3FFD0FE06F82AFDCDEEE3" box="[417,540,1940,1966]" captionStart="FIGURE 6" captionStartId="11.[151,250,1114,1138]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-18@11.[151,1435,181,1089]" captionTargetPageId="11" captionText="FIGURE 6. P5 dorsal view. A–C, Clistocoeloma suvaense Edmondson, 1951; A, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji; B, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; C, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa; D, C. villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–K, C. nobile n. sp.; E, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; F, paratype male (15.0 × 13.0 mm) (ZRC 2002.0155) Singapore; G, male (16.0 × 13.5 mm) (ZRC 2022.0977), Vietnam; H, male (17.4 × 14.9 mm) (ZRC 2022.0973), Philippines; I, male (17.3 × 15.0 mm) (ZRC 2022.0994), Hong Kong; J, male (18.2 × 15.0 mm), Japan; K, male (16.2 × 14.2 mm) (WS898), Korea." figureDoi="http://doi.org/10.5281/zenodo.8162504" httpUri="https://zenodo.org/record/8162504/files/figure.png" pageId="4" pageNumber="221">Fig. 6A–C</figureCitation>
).
</paragraph>
<caption id="189EFDE3FFF2FFD1FF30F833FEA4EEA0" ID-DOI="http://doi.org/10.5281/zenodo.8162492" ID-Zenodo-Dep="8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="5" pageNumber="222" startId="5.[151,250,1933,1957]" targetBox="[227,1360,181,1909]" targetPageId="5" targetType="figure">
<paragraph id="4C5EAD6BFFF2FFD1FF30F833FEA4EEA0" blockId="5.[151,1436,1933,2029]" pageId="5" pageNumber="222">
<emphasis id="7E957179FFF2FFD1FF30F833FEC2EEE8" bold="true" box="[151,275,1933,1957]" pageId="5" pageNumber="222">FIGURE 2.</emphasis>
Colour in life. A, 
<taxonomicName id="8BE1D6E8FFF2FFD1FE6DF833FCAEEEE8" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[458,895,1933,1957]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF2FFD1FE6DF833FD6FEEE8" box="[458,702,1933,1957]" italics="true" pageId="5" pageNumber="222">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFF2FFD1FD63F833FCAEEEE8" author="Edmondson, C. H." box="[708,895,1933,1957]" pageId="5" pageNumber="222" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, 
<taxonomicName id="8BE1D6E8FFF2FFD1FA22F833FF07EE84" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="nobile" status="n. sp.">
<emphasis id="7E957179FFF2FFD1FA22F833FA47EEE9" box="[1413,1430,1933,1956]" italics="true" pageId="5" pageNumber="222">C</emphasis>
. 
<emphasis id="7E957179FFF2FFD1FF30F80FFF07EE84" box="[151,214,1969,1993]" italics="true" pageId="5" pageNumber="222">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFF2FFD1FF7AF80FFEC7EE84" bold="true" box="[221,278,1969,1993]" pageId="5" pageNumber="222">
<taxonomicNameLabel id="65A6CC02FFF2FFD1FF7AF80FFEC7EE84" box="[221,278,1969,1993]" pageId="5" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, 
<taxonomicName id="8BE1D6E8FFF2FFD1FC6DF80FFBF7EE84" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[970,1062,1969,1993]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="nobile" status="n. sp.">
<emphasis id="7E957179FFF2FFD1FC6DF80FFC0AEE85" box="[970,987,1969,1992]" italics="true" pageId="5" pageNumber="222">C</emphasis>
. 
<emphasis id="7E957179FFF2FFD1FC40F80FFBF7EE84" box="[999,1062,1969,1993]" italics="true" pageId="5" pageNumber="222">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFF2FFD1FB8AF80FFBB7EE84" bold="true" box="[1069,1126,1969,1993]" pageId="5" pageNumber="222">
<taxonomicNameLabel id="65A6CC02FFF2FFD1FB8AF80FFBB7EE84" box="[1069,1126,1969,1993]" pageId="5" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam.
</paragraph>
</caption>
<caption id="189EFDE3FFF1FFD2FF30F8BCFDD5EE2F" ID-DOI="http://doi.org/10.5281/zenodo.8162494" ID-Zenodo-Dep="8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="6" pageNumber="223" startId="6.[151,250,1794,1818]" targetBox="[151,1435,181,1770]" targetPageId="6" targetType="figure">
<paragraph id="4C5EAD6BFFF1FFD2FF30F8BCFDD5EE2F" blockId="6.[151,1436,1794,1890]" pageId="6" pageNumber="223">
<emphasis id="7E957179FFF1FFD2FF30F8BCFEC2EE57" bold="true" box="[151,275,1794,1818]" pageId="6" pageNumber="223">FIGURE 3.</emphasis>
<taxonomicName id="8BE1D6E8FFF1FFD2FEBEF8BCFD1FEE57" authority="Edmondson, 1951" authorityName="Edmondson" authorityYear="1951" box="[281,718,1794,1818]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="223" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF1FFD2FEBEF8BCFDDCEE57" box="[281,525,1794,1818]" italics="true" pageId="6" pageNumber="223">Clistocoeloma suvaense</emphasis>
<bibRefCitation id="2870D09AFFF1FFD2FDB3F8BCFD1FEE57" author="Edmondson, C. H." box="[532,718,1794,1818]" pageId="6" pageNumber="223" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
</taxonomicName>
, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view.
</paragraph>
</caption>
<paragraph id="4C5EAD6BFFF1FFD3FF60F80AFC3FE998" blockId="6.[151,1436,1972,2035]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="224" pageId="6" pageNumber="223">
Male thoracic sternum covered with short setae, smooth when denuded; sternites 1 and 2 fused, with suture between fused sternites 3 and 4; sternopleonal cavity extends to edge sternites 1 and 2. Male pleon wide, stout, long setae fringed pleon margin; somite 3 widest; somite 6 with curved edge; telson wide, dome-shaped (
<figureCitation id="D4DAB1EEFFF0FFD3FAB7FF29FA49E9FC" box="[1296,1432,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="7" pageNumber="224">Figs. 3B, D</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FF30FF05FF6AE998" box="[151,187,187,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="7" pageNumber="224">4B</figureCitation>
); male pleonal locking mechanism absent, without tubercle on sternite 5.
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60FF61FE05E80C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
G1 straight, distal region relatively wide; chitinous tip wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (
<figureCitation id="D4DAB1EEFFF0FFD3FC19FEBDFB93E850" box="[958,1090,259,285]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="7" pageNumber="224">Figs. 9E–H</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FBE8FEBDFB77E850" box="[1103,1190,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[216,1372,183,1917]" captionTargetId="figure-18@15.[214,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 10. Left G1. A–D, Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, C. suvaense Edmondson, 1951, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="7" pageNumber="224">10A–H</figureCitation>
). G2 shorter than G1, slightly curved, tip rounded.
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60FEF5FBA8E8C4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
Female pleon rounded, relatively wider than long; telson dome-shaped (
<figureCitation id="D4DAB1EEFFF0FFD3FC7EFEF5FBE0E828" box="[985,1073,331,357]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="7" pageNumber="224">Fig. 8A</figureCitation>
). Vulvae with central operculum, ovate, slightly protruded; sternal vulvar cover slightly raised above operculum (
<figureCitation id="D4DAB1EEFFF0FFD3FBB3FED1FBBAE8C4" box="[1044,1131,367,393]" captionStart="FIGURE 8" captionStartId="13.[151,250,1067,1091]" captionTargetBox="[151,1435,181,1044]" captionTargetId="figure-18@13.[151,1435,181,1044]" captionTargetPageId="13" captionText="FIGURE 8. Female pleon, thoracic sternums and vulvae.A, B, Clistocoeloma suvaense Edmondson, 1951, female (18.7 × 17.0 mm) (ZRC 2022.0989), Fiji; C, D, Clistocoeloma nobile n. sp., paratype female, (18.7 × 15.2 mm) (ZRC 2022.0971), Singapore. A, C, overall ventral view; B, D, female thoracic sternums and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8162510" httpUri="https://zenodo.org/record/8162510/files/figure.png" pageId="7" pageNumber="224">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60FE2CFE38E89C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
<emphasis id="7E957179FFF0FFD3FF60FE2CFE8EE8E1" bold="true" box="[199,351,402,428]" pageId="7" pageNumber="224">Colouration.</emphasis>
Chelipeds pale yellow fingers with pale orange to reddish palm (
<figureCitation id="D4DAB1EEFFF0FFD3FBFFFE2DFB64E8E0" box="[1112,1205,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="7" pageNumber="224">Fig. 2A</figureCitation>
). In life, entire crab covered in thick layer of mud.
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFF0FFD3FF60FE65FE79EED8" pageId="7" pageNumber="224" type="discussion">
<paragraph id="4C5EAD6BFFF0FFD3FF60FE65FA8DEA74" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
<emphasis id="7E957179FFF0FFD3FF60FE65FEEBE8B8" bold="true" box="[199,314,475,501]" pageId="7" pageNumber="224">Remarks.</emphasis>
<taxonomicName id="8BE1D6E8FFF0FFD3FEE3FE65FD83E8B9" authorityName="Edmondson" authorityYear="1951" box="[324,594,475,501]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FEE3FE65FD83E8B9" box="[324,594,475,501]" italics="true" pageId="7" pageNumber="224">Clistocoeloma suvaense</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="2870D09AFFF0FFD3FC8DFE65FBD3E8B8" author="Edmondson, C. H." box="[810,1026,474,501]" pageId="7" pageNumber="224" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951)</bibRefCitation>
based on 
<specimenCount id="5AE766E2FFF0FFD3FBDCFE65FB32E8B8" box="[1147,1251,475,501]" pageId="7" pageNumber="224" type="male">one male</specimenCount>
and 
<specimenCount id="5AE766E2FFF0FFD3FAB8FE65FA4DE8B8" box="[1311,1436,475,501]" pageId="7" pageNumber="224" type="female">one female</specimenCount>
specimen collected from a reef in Suva Harbour. He compared it with other species that were recognised in 
<taxonomicName id="8BE1D6E8FFF0FFD3FF30FD9DFEE6EB70" authorityName="A. Milne-Edwards" authorityYear="1873" box="[151,311,547,573]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="7E957179FFF0FFD3FF30FD9DFEE6EB70" box="[151,311,547,573]" italics="true" pageId="7" pageNumber="224">Clistocoeloma</emphasis>
</taxonomicName>
at the time, i.e., 
<taxonomicName id="8BE1D6E8FFF0FFD3FE53FD9DFDA8EB70" authorityName="A. Milne-Edwards" authorityYear="1868" box="[500,633,547,573]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="balansae">
<emphasis id="7E957179FFF0FFD3FE53FD9DFDD6EB70" box="[500,519,547,573]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FDB2FD9DFDA8EB70" box="[533,633,547,573]" italics="true" pageId="7" pageNumber="224">balansae</emphasis>
</taxonomicName>
. 
<taxonomicName id="8BE1D6E8FFF0FFD3FD2FFD9DFCF9EB71" authorityName="De Man" authorityYear="1888" box="[648,808,547,573]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="merguiense">
<emphasis id="7E957179FFF0FFD3FD2FFD9DFD4AEB70" box="[648,667,547,573]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FD0EFD9DFCF9EB71" box="[681,808,547,572]" italics="true" pageId="7" pageNumber="224">merguiense</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BE1D6E8FFF0FFD3FCF8FD9DFB5AEB70" authority="(Rathbun, 1914)" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[863,1163,547,573]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="tectum">
<emphasis id="7E957179FFF0FFD3FCF8FD9DFCA3EB70" box="[863,882,547,573]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC27FD9DFC1BEB71" box="[896,970,547,572]" italics="true" pageId="7" pageNumber="224">tectum</emphasis>
(
<bibRefCitation id="2870D09AFFF0FFD3FC7EFD9DFB52EB70" author="Rathbun, M. J." box="[985,1155,547,573]" pageId="7" pageNumber="224" pagination="69 - 85" refId="ref11996" refString="Rathbun, M. J. (1914) New species of crabs of the families Grapsidae and Ocypodida. Proceedings of the United Stated National Museum, 47 (2044), 69 - 85. https: // doi. org / 10.5479 / si. 00963801.2044.69" type="journal article" year="1914">Rathbun, 1914</bibRefCitation>
)
</taxonomicName>
. 
<taxonomicName id="8BE1D6E8FFF0FFD3FB30FD9DFA4DEB70" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[1175,1436,547,573]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FB30FD9DFA4DEB70" box="[1175,1436,547,573]" italics="true" pageId="7" pageNumber="224">Clistoceoloma villosum</emphasis>
</taxonomicName>
was not treated or discussed as it was not in the genus at the time. 
<bibRefCitation id="2870D09AFFF0FFD3FCD9FDF9FB85EB2C" author="Edmondson, C. H." box="[894,1108,582,609]" pageId="7" pageNumber="224" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951)</bibRefCitation>
actually had 
<specimenCount id="5AE766E2FFF0FFD3FB4DFDF9FA4CEB2C" box="[1258,1437,583,609]" pageId="7" pageNumber="224" type="generic">three specimens</specimenCount>
from the Bishop Museum, two of which were collected from Suva Habour, 
<collectingCountry id="34F6EDFBFFF0FFD3FC66FDD5FC3BEBC8" box="[961,1002,619,645]" name="Fiji" pageId="7" pageNumber="224">Fiji</collectingCountry>
, while 
<specimenCount id="5AE766E2FFF0FFD3FB93FDD5FB46EBC8" box="[1076,1175,619,645]" pageId="7" pageNumber="224" type="male">one male</specimenCount>
specimen was collected from a Samoan reef. While 
<bibRefCitation id="2870D09AFFF0FFD3FE63FD31FD49EBE4" author="Edmondson, C. H." box="[452,664,654,681]" pageId="7" pageNumber="224" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951)</bibRefCitation>
was certain that the two Fijian specimens belonged to his new species, he was more doubtful about the Samoan one; noting that the latter had no distinct lobes on the anterolateral margins (lobes present on the anterolateral margins of 
<taxonomicName id="8BE1D6E8FFF0FFD3FD3DFD69FCCEEBBD" authorityName="Edmondson" authorityYear="1951" box="[666,799,727,753]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FD3DFD69FD7CEBBC" box="[666,685,727,753]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FD1CFD69FCCEEBBD" box="[699,799,727,752]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A6CC02FFF0FFD3FC80FD69FCB2EBBC" box="[807,867,727,753]" pageId="7" pageNumber="224" sensu="stricto">s. str.</taxonomicNameLabel>
) and the dorsal surface of the dactylus of the male chela with 16 granules (with 25 granules in 
<taxonomicName id="8BE1D6E8FFF0FFD3FDDDFD45FD2FEA59" authorityName="Edmondson" authorityYear="1951" box="[634,766,763,789]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FDDDFD45FD5CEA58" box="[634,653,763,789]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FD3DFD45FD2FEA59" box="[666,766,763,788]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A6CC02FFF0FFD3FCA3FD45FC90EA58" box="[772,833,763,789]" pageId="7" pageNumber="224" sensu="stricto">s. str.</taxonomicNameLabel>
). We have not examined 
<bibRefCitation id="2870D09AFFF0FFD3FBF2FD45FAEDEA58" author="Edmondson, C. H." box="[1109,1340,762,789]" pageId="7" pageNumber="224" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson’s (1951)</bibRefCitation>
Samoan specimens, but his observations match exactly what is known for 
<taxonomicName id="8BE1D6E8FFF0FFD3FCCAFCA1FC3AEA74" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[877,1003,799,825]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FCCAFCA1FC51EA74" box="[877,896,799,825]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC2AFCA1FC3AEA74" box="[909,1003,799,825]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
, we believe they are conspecific.
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60FCFDFAFBECD0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
<taxonomicName id="8BE1D6E8FFF0FFD3FF60FCFDFE05EA11" authorityName="Edmondson" authorityYear="1951" box="[199,468,835,861]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FF60FCFDFE05EA11" box="[199,468,835,861]" italics="true" pageId="7" pageNumber="224">Clistocoeloma suvaense</emphasis>
</taxonomicName>
is known with certainty only from its type locality, although there is 
<specimenCount id="5AE766E2FFF0FFD3FB77FCFDFAB8EA10" box="[1232,1385,835,861]" pageId="7" pageNumber="224" type="generic">one specimen</specimenCount>
also from 
<collectingCountry id="34F6EDFBFFF0FFD3FF72FCD9FEF0EACC" box="[213,289,871,897]" name="Samoa" pageId="7" pageNumber="224">Samoa</collectingCountry>
(see below) and published records from 
<collectingCountry id="34F6EDFBFFF0FFD3FD56FCD9FCB2EACC" box="[753,867,871,897]" name="Singapore" pageId="7" pageNumber="224">Singapore</collectingCountry>
(
<bibRefCitation id="2870D09AFFF0FFD3FCD6FCD9FBFEEACC" author="Wee, Y. C. &amp; Ng, P. K. L." box="[881,1071,871,897]" pageId="7" pageNumber="224" refId="ref12677" refString="Wee, Y. C. &amp; Ng, P. K. L. (Eds.) (1994) A first look at biodiversity in Singapore. National council on the Environment, Singapore, 163 pp." type="book" year="1994">Wee &amp; Ng 1994</bibRefCitation>
; 
<bibRefCitation id="2870D09AFFF0FFD3FB9CFCD9FB21EACC" author="Tan, C. G. S. &amp; Ng, P. K. L." box="[1083,1264,871,897]" pageId="7" pageNumber="224" pagination="75 - 84" refId="ref12558" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655" type="journal article" year="1994">Tan &amp; Ng 1994</bibRefCitation>
). In addition to the 
<typeStatus id="935A13C9FFF0FFD3FF66FC35FEF9EAE8" box="[193,296,907,933]" pageId="7" pageNumber="224">lectotype</typeStatus>
specimen of 
<taxonomicName id="8BE1D6E8FFF0FFD3FE18FC35FDEEEAE8" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[447,575,907,933]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FE18FC35FE03EAE8" box="[447,466,907,933]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FE47FC35FDEEEAE8" box="[480,575,907,933]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
, another Wroblewsky specimen originally from the Godeffroy Museum was found in the ZRC, although how it got there is not known, possibly from the time R. Serène was based in 
<collectingCountry id="34F6EDFBFFF0FFD3FA8CFC11FA4CEA84" box="[1323,1437,943,969]" name="Singapore" pageId="7" pageNumber="224">Singapore</collectingCountry>
in the 1960s. It is noteworthy that during that time, Serène was revising the IWP 
<taxonomicName id="8BE1D6E8FFF0FFD3FB8CFC6DFB7DEAA0" authorityName="Dana" authorityYear="1851" box="[1067,1196,979,1005]" class="Malacostraca" family="Sesarmidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="family">Sesarmidae</taxonomicName>
with Soh Cheng Lan (e.g., see Serène &amp; Soh 1970) and he had gathered material in 
<collectingCountry id="34F6EDFBFFF0FFD3FCCAFC49FC0EED5C" box="[877,991,1015,1041]" name="Singapore" pageId="7" pageNumber="224">Singapore</collectingCountry>
for this work. The morphology of this Wroblewsky specimen, however, clearly matches what is here defined as 
<taxonomicName id="8BE1D6E8FFF0FFD3FC7BFBA5FBB3ED79" authorityName="Edmondson" authorityYear="1951" box="[988,1122,1051,1077]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FC7BFBA5FC3EED78" box="[988,1007,1051,1077]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC5AFBA5FBB3ED79" box="[1021,1122,1051,1076]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
and not 
<taxonomicName id="8BE1D6E8FFF0FFD3FB6FFBA5FA98ED78" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[1224,1353,1051,1077]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FB6FFBA5FB0AED78" box="[1224,1243,1051,1077]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FB4DFBA5FA98ED78" box="[1258,1353,1051,1077]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
. Based on the specimen label, it has the same locality information as the 
<typeStatus id="935A13C9FFF0FFD3FC26FB81FC39ED14" box="[897,1000,1087,1113]" pageId="7" pageNumber="224">lectotype</typeStatus>
of 
<taxonomicName id="8BE1D6E8FFF0FFD3FBA8FB81FB41ED14" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[1039,1168,1087,1113]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FBA8FB81FBF3ED14" box="[1039,1058,1087,1113]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FB96FB81FB41ED14" box="[1073,1168,1087,1113]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
. In addition to the very different morphology, the measurements of this specimen also do not match those listed in A. 
<bibRefCitation id="2870D09AFFF0FFD3FB39FBDDFA4DED30" author="Milne-Edwards, A." box="[1182,1436,1123,1149]" pageId="7" pageNumber="224" pagination="25 - 31" refId="ref11446" refString="Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]" type="journal article" year="1869">Milne-Edwards (1869)</bibRefCitation>
in his original description. It is unclear whether A. Milne-Edwards actually examined this specimen or if there is a mistake in the labels. There is also no mention of additional specimens of “ 
<taxonomicName id="8BE1D6E8FFF0FFD3FC71FB15FB84ED88" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[982,1109,1195,1221]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FC71FB15FC38ED88" box="[982,1001,1195,1221]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC51FB15FB84ED88" box="[1014,1109,1195,1221]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
” in the eight sales catalogues of Godeffroy Museum that were published between 1864 and 1881 by curator J.D.E. Schmeltz other than that for the original specimen in A. Milne-Edwards (1896) or later papers by the author on the Godeffroy Museum collections (e.g. A. Mine-Edwards 1873a). For the moment, we accept the record of 
<taxonomicName id="8BE1D6E8FFF0FFD3FC1FFAA9FBECEC7D" authorityName="Edmondson" authorityYear="1951" box="[952,1085,1303,1329]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FC1FFAA9FC1AEC7C" box="[952,971,1303,1329]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC7FFAA9FBECEC7D" box="[984,1085,1303,1328]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
as being also present in 
<collectingCountry id="34F6EDFBFFF0FFD3FAEEFAA9FA48EC7C" box="[1353,1433,1303,1329]" name="Samoa" pageId="7" pageNumber="224">Samoa</collectingCountry>
. Even though both 
<taxonomicName id="8BE1D6E8FFF0FFD3FEC3FA85FE32EC19" authorityName="Edmondson" authorityYear="1951" box="[356,483,1339,1365]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FEC3FA85FE32EC19" box="[356,483,1339,1365]" italics="true" pageId="7" pageNumber="224">C suvaense</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BE1D6E8FFF0FFD3FDBEFA85FD49EC18" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[537,664,1339,1365]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FDBEFA85FDFDEC18" box="[537,556,1339,1365]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FD9EFA85FD49EC18" box="[569,664,1339,1365]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
are found together in 
<collectingCountry id="34F6EDFBFFF0FFD3FC29FA85FC0CEC18" box="[910,989,1339,1365]" name="Samoa" pageId="7" pageNumber="224">Samoa</collectingCountry>
, it is important to note that currently, no specimens of 
<taxonomicName id="8BE1D6E8FFF0FFD3FE94FAE1FE65EC34" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[307,436,1375,1401]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FE94FAE1FE97EC34" box="[307,326,1375,1401]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FEF2FAE1FE65EC34" box="[341,436,1375,1401]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
has been found in 
<collectingCountry id="34F6EDFBFFF0FFD3FD2AFAE1FD69EC34" box="[653,696,1375,1401]" name="Fiji" pageId="7" pageNumber="224">Fiji</collectingCountry>
. Recent collections there only found 
<taxonomicName id="8BE1D6E8FFF0FFD3FBFAFAE1FB35EC35" authorityName="Edmondson" authorityYear="1951" box="[1117,1252,1375,1401]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FBFAFAE1FBA1EC34" box="[1117,1136,1375,1401]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FBD8FAE1FB35EC35" box="[1151,1252,1375,1400]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
. Records of “ 
<taxonomicName id="8BE1D6E8FFF0FFD3FA24FAE1FF2DECD1" authorityName="Edmondson" authorityYear="1951" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FA24FAE1FA47EC34" box="[1411,1430,1375,1401]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FF30FA3DFF2DECD1" box="[151,252,1411,1436]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
” from 
<collectingCountry id="34F6EDFBFFF0FFD3FEEAFA3DFE6EECD0" box="[333,447,1411,1437]" name="Singapore" pageId="7" pageNumber="224">Singapore</collectingCountry>
by 
<bibRefCitation id="2870D09AFFF0FFD3FE4FFA3DFD60ECD0" author="Wee, Y. C. &amp; Ng, P. K. L." box="[488,689,1411,1437]" pageId="7" pageNumber="224" refId="ref12677" refString="Wee, Y. C. &amp; Ng, P. K. L. (Eds.) (1994) A first look at biodiversity in Singapore. National council on the Environment, Singapore, 163 pp." type="book" year="1994">Wee &amp; Ng (1994)</bibRefCitation>
and 
<bibRefCitation id="2870D09AFFF0FFD3FD40FA3DFC79ECD0" author="Tan, C. G. S. &amp; Ng, P. K. L." box="[743,936,1411,1437]" pageId="7" pageNumber="224" pagination="75 - 84" refId="ref12558" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655" type="journal article" year="1994">Tan &amp; Ng (1994)</bibRefCitation>
belong to 
<taxonomicName id="8BE1D6E8FFF0FFD3FBB8FA3DFB57ECD0" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[1055,1158,1411,1437]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="nobile" status="n. sp.">
<emphasis id="7E957179FFF0FFD3FBB8FA3DFBE3ECD0" box="[1055,1074,1411,1437]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FBE7FA3DFB57ECD0" box="[1088,1158,1411,1437]" italics="true" pageId="7" pageNumber="224">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFF0FFD3FB2AFA3DFB1AECD0" bold="true" box="[1165,1227,1411,1437]" pageId="7" pageNumber="224">
<taxonomicNameLabel id="65A6CC02FFF0FFD3FB2AFA3DFB1AECD0" box="[1165,1227,1411,1437]" pageId="7" pageNumber="224" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
instead.
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60FA19FE45EE48" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
Although 
<taxonomicName id="8BE1D6E8FFF0FFD3FE9DFA19FE13EC8D" authorityName="Edmondson" authorityYear="1951" box="[314,450,1447,1473]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FE9DFA19FE9CEC8C" box="[314,333,1447,1473]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FEFAFA19FE13EC8D" box="[349,450,1447,1472]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
is superficially similar to 
<taxonomicName id="8BE1D6E8FFF0FFD3FD54FA19FCA4EC8C" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[755,885,1447,1473]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FD54FA19FCD7EC8C" box="[755,774,1447,1473]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FCB1FA19FCA4EC8C" box="[790,885,1447,1473]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BE1D6E8FFF0FFD3FC17FA19FBC8EC8C" authority="Ng &amp; Ng, 2023" authorityName="Ng &amp; Ng" authorityYear="2023" box="[944,1049,1447,1473]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="nobile" status="n. sp.">
<emphasis id="7E957179FFF0FFD3FC17FA19FC12EC8C" box="[944,963,1447,1473]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC74FA19FBC8EC8C" box="[979,1049,1447,1473]" italics="true" pageId="7" pageNumber="224">nobile</emphasis>
</taxonomicName>
<emphasis id="7E957179FFF0FFD3FB85FA19FBB2EC8C" bold="true" box="[1058,1123,1447,1473]" pageId="7" pageNumber="224">
<taxonomicNameLabel id="65A6CC02FFF0FFD3FB85FA19FBB2EC8C" box="[1058,1123,1447,1473]" pageId="7" pageNumber="224" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, they are relatively easy to distinguish, especially adult males. 
<taxonomicName id="8BE1D6E8FFF0FFD3FD85FA75FCFEECA9" authorityName="Edmondson" authorityYear="1951" box="[546,815,1483,1509]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FD85FA75FCFEECA9" box="[546,815,1483,1509]" italics="true" pageId="7" pageNumber="224">Clistocoeloma suvaense</emphasis>
</taxonomicName>
clearly differs from 
<taxonomicName id="8BE1D6E8FFF0FFD3FBB4FA75FB43ECA8" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[1043,1170,1483,1509]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FBB4FA75FBF7ECA8" box="[1043,1062,1483,1509]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FB94FA75FB43ECA8" box="[1075,1170,1483,1509]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A6CC02FFF0FFD3FB3FFA75FB02ECA8" box="[1176,1235,1483,1509]" pageId="7" pageNumber="224" sensu="stricto">s. str.</taxonomicNameLabel>
in the presence of a distinct external orbital angle on the anterolateral margin of the carapace (
<figureCitation id="D4DAB1EEFFF0FFD3FC4CFA51FB9EEF44" box="[1003,1103,1519,1545]" captionStart="FIGURE 2" captionStartId="5.[151,250,1933,1957]" captionTargetBox="[227,1360,181,1909]" captionTargetId="figure-18@5.[227,1360,181,1909]" captionTargetPageId="5" captionText="FIGURE 2. Colour in life. A, Clistocoeloma suvaense Edmondson, 1951, male (17.8 × 16.3 mm) (ZRC 2022.0986), Fiji; B, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore; C, C. nobile n. sp., male (17.4 × 15.2 mm) (ZRC 2022.0975), Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8162492" httpUri="https://zenodo.org/record/8162492/files/figure.png" pageId="7" pageNumber="224">Figs. 2A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FBFCFA51FBAEEF44" box="[1115,1151,1519,1545]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="7" pageNumber="224">3A</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FB2BFA51FB7EEF44" box="[1164,1199,1519,1545]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="7" pageNumber="224">4A</figureCitation>
) (the external orbital angle on the anterolateral carapace margin is not distinct and low in 
<taxonomicName id="8BE1D6E8FFF0FFD3FC32F9ADFBC4EF60" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[917,1045,1555,1581]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FC32F9ADFC79EF60" box="[917,936,1555,1581]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC11F9ADFBC4EF60" box="[950,1045,1555,1581]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
; 
<figureCitation id="D4DAB1EEFFF0FFD3FB84F9ADFBAAEF60" box="[1059,1147,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[164,1423,181,1773]" captionTargetId="figure-32@2.[164,1423,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162488" httpUri="https://zenodo.org/record/8162488/files/figure.png" pageId="7" pageNumber="224">Fig. 1A</figureCitation>
); the presence of a broad, rounded inner orbital angle (
<figureCitation id="D4DAB1EEFFF0FFD3FE47F989FD95EF1C" box="[480,580,1591,1617]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="7" pageNumber="224">Figs. 3C</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FDF4F989FDA5EF1C" box="[595,628,1591,1617]" captionStart="FIGURE 4" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[151,1435,181,1837]" captionTargetId="figure-32@8.[151,1435,181,1837]" captionTargetPageId="8" captionText="FIGURE 4. Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162498" httpUri="https://zenodo.org/record/8162498/files/figure.png" pageId="7" pageNumber="224">4C</figureCitation>
) (the inner orbital angle is narrow and ovate in 
<taxonomicName id="8BE1D6E8FFF0FFD3FB05F989FAF5EF1C" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[1186,1316,1591,1617]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FB05F989FB64EF1C" box="[1186,1205,1591,1617]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FB62F989FAF5EF1C" box="[1221,1316,1591,1617]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
; 
<figureCitation id="D4DAB1EEFFF0FFD3FA92F989FA5DEF1C" box="[1333,1420,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1822]" captionTargetBox="[164,1423,181,1773]" captionTargetId="figure-32@2.[164,1423,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8162488" httpUri="https://zenodo.org/record/8162488/files/figure.png" pageId="7" pageNumber="224">Fig. 1C</figureCitation>
); possessing 25–26 granules on the dorsal surface of the dactylus of the adult male chela (
<figureCitation id="D4DAB1EEFFF0FFD3FBD5F9E5FB02EF38" box="[1138,1235,1627,1653]" captionStart="FIGURE 3" captionStartId="6.[151,250,1794,1818]" captionTargetBox="[151,1435,181,1770]" captionTargetId="figure-32@6.[151,1435,181,1770]" captionTargetPageId="6" captionText="FIGURE 3. Clistocoeloma suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, overall dorsal view; B, overall ventral view; C, frontal view of carapace; D, ventral view of pleon and thoracic sternum; E, right cheliped top view; F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162494" httpUri="https://zenodo.org/record/8162494/files/figure.png" pageId="7" pageNumber="224">Figs. 3E</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FB79F9E4FAD3EF38" box="[1246,1282,1626,1653]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="7" pageNumber="224">5D</figureCitation>
) (16 granules in 
<taxonomicName id="8BE1D6E8FFF0FFD3FF14F9C1FEE3EFD4" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[179,306,1663,1689]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FF14F9C1FF17EFD4" box="[179,198,1663,1689]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FF74F9C1FEE3EFD4" box="[211,306,1663,1689]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
; 
<figureCitation id="D4DAB1EEFFF0FFD3FEE6F9C1FE48EFD4" box="[321,409,1662,1689]" captionStart="FIGURE 5" captionStartId="10.[151,250,1726,1750]" captionTargetBox="[151,1435,417,1702]" captionTargetId="figure-164@10.[151,1435,417,1702]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; C, D, C. suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E, F, C. nobile n. sp., holotype male, (15.3 × 12.7 mm) (ZRC 2022.0970), Singapore.A, C, E, right cheliped top view; B, D, F, right cheliped ventral view." figureDoi="http://doi.org/10.5281/zenodo.8162500" httpUri="https://zenodo.org/record/8162500/files/figure.png" pageId="7" pageNumber="224">Fig. 5A</figureCitation>
); and the G1 is straighter with a proportionately broader distal portion (
<figureCitation id="D4DAB1EEFFF0FFD3FB14F9C1FAE7EFD4" box="[1203,1334,1663,1689]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="7" pageNumber="224">Figs. 9E–H</figureCitation>
, 
<figureCitation id="D4DAB1EEFFF0FFD3FAE7F9C1FA45EFD4" box="[1344,1428,1663,1689]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[216,1372,183,1917]" captionTargetId="figure-18@15.[214,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 10. Left G1. A–D, Clistocoeloma suvaense Edmondson, 1951, male (18.8 × 16.9 mm) (ZRC 2022.0985), Fiji; E–H, C. suvaense Edmondson, 1951, male (16.2 × 14.3 mm) (ZRC 1973.11.2.490), Samoa, A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162514" httpUri="https://zenodo.org/record/8162514/files/figure.png" pageId="7" pageNumber="224">10A–H</figureCitation>
) (G1 slightly curved with a narrower distal portion for 
<taxonomicName id="8BE1D6E8FFF0FFD3FCA2F91DFC56EFF0" baseAuthorityName="A.Milne-Edwards" baseAuthorityYear="1869" box="[773,903,1699,1725]" class="Malacostraca" family="Sesarmidae" genus="Clistoceoloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="7E957179FFF0FFD3FCA2F91DFCC9EFF0" box="[773,792,1699,1725]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FC8FF91DFC56EFF0" box="[808,903,1699,1725]" italics="true" pageId="7" pageNumber="224">villosum</emphasis>
</taxonomicName>
; 
<figureCitation id="D4DAB1EEFFF0FFD3FC3FF91DFBC7EFF0" box="[920,1046,1699,1725]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[212,1375,183,1917]" captionTargetId="figure-32@14.[210,1377,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Left G1. A–D, Clistocoeloma villosum (A. Milne-Edwards, 1869), lectotype male (18.6 × 17.0 mm) (ZMH K4188), Samoa; E–H, C. suvaense Edmondson, 1951, holotype male (21.0 × 18.8 mm) (BPBM 3976), Fiji. A, E, dorsal view; B, F, left lateral view; C, G, ventral view; D, H, right lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8162512" httpUri="https://zenodo.org/record/8162512/files/figure.png" pageId="7" pageNumber="224">Fig. 9A–D</figureCitation>
). Fresh collections of 
<taxonomicName id="8BE1D6E8FFF0FFD3FAB3F91DFA4DEFF1" authorityName="Edmondson" authorityYear="1951" box="[1300,1436,1699,1725]" class="Malacostraca" family="Sesarmidae" genus="Clistocoeloma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="suvaense">
<emphasis id="7E957179FFF0FFD3FAB3F91DFAF6EFF0" box="[1300,1319,1699,1725]" italics="true" pageId="7" pageNumber="224">C</emphasis>
. 
<emphasis id="7E957179FFF0FFD3FA90F91DFA4DEFF1" box="[1335,1436,1699,1724]" italics="true" pageId="7" pageNumber="224">suvaense</emphasis>
</taxonomicName>
from the 
<typeStatus id="935A13C9FFF0FFD3FF58F979FEFEEFAC" box="[255,303,1735,1761]" pageId="7" pageNumber="224">type</typeStatus>
locality show that the species is not uncommon in 
<collectingCountry id="34F6EDFBFFF0FFD3FCD4F979FC4CEFAC" box="[883,925,1735,1761]" name="Fiji" pageId="7" pageNumber="224">Fiji</collectingCountry>
, and all the recent specimens agree very well with the 
<typeStatus id="935A13C9FFF0FFD3FF5EF955FEF8EE48" box="[249,297,1771,1797]" pageId="7" pageNumber="224">type</typeStatus>
material.
</paragraph>
<paragraph id="4C5EAD6BFFF0FFD3FF60F8B0FE79EED8" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
<emphasis id="7E957179FFF0FFD3FF60F8B0FE41EE64" bold="true" box="[199,400,1806,1833]" pageId="7" pageNumber="224">Ecological notes.</emphasis>
It was mentioned that the specimens were collected from “a reef in Suva Harbor in 1933” (
<bibRefCitation id="2870D09AFFF0FFD3FF38F88DFE75EE00" author="Edmondson, C. H." box="[159,420,1842,1869]" pageId="7" pageNumber="224" pagination="183 - 243" refId="ref10601" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson 1951: 240</bibRefCitation>
). Fresh material was collected from the back mangrove at a higher elevation and found on muddy substrate under large loose rocks in Suva and Taveuni Island, 
<collectingCountry id="34F6EDFBFFF0FFD3FC3BF8E9FC16EE3C" box="[924,967,1879,1905]" name="Fiji" pageId="7" pageNumber="224">Fiji</collectingCountry>
. Specimens were found with layer of mud covering the entire crab.
</paragraph>
</subSubSection>
<subSubSection id="04FBFEE0FFF0FFD3FF60F820FDD1EE90" pageId="7" pageNumber="224" type="distribution">
<paragraph id="4C5EAD6BFFF0FFD3FF60F820FDD1EE90" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="224">
<emphasis id="7E957179FFF0FFD3FF60F820FEB0EEF5" bold="true" box="[199,353,1950,1976]" pageId="7" pageNumber="224">Distribution.</emphasis>
Currently only known for certain from the 
<typeStatus id="935A13C9FFF0FFD3FCE3F821FCA5EEF4" box="[836,884,1951,1977]" pageId="7" pageNumber="224">type</typeStatus>
locality, Suva, as well as Taveuni Island, 
<collectingCountry id="34F6EDFBFFF0FFD3FAE2F821FAA1EEF4" box="[1349,1392,1951,1977]" name="Fiji" pageId="7" pageNumber="224">Fiji</collectingCountry>
; the record from 
<collectingCountry id="34F6EDFBFFF0FFD3FE84F87DFEBEEE90" box="[291,367,1987,2013]" name="Samoa" pageId="7" pageNumber="224">Samoa</collectingCountry>
is uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>