171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="C2EDF75F1E6CCA04ECA6FA3BB5909808" ID-DOI="10.11646/zootaxa.3946.2.1" ID-GBIF-Dataset="f56bb3a6-9852-4218-9c33-b1b3c79670b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="233167" ID-ZooBank="9EE92E94-8743-49E9-B96E-A057C77D9BC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032553967" checkinUser="plazi" docAuthor="Andrew Hamilton, K. G." docDate="2015" docId="B4209064536BFFE4FF5FFDE7FAC16AD7" docLanguage="en" docName="zt03946p189.pdf" docOrigin="Zootaxa 3946 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Taphrotylus Hamilton, 2015, gen.nov." docType="treatment" docVersion="7" lastPageNumber="173" masterDocId="4819E81C537DFFF2FFC8FFE7FFC26D40" masterDocTitle="A new tribe and species of Clastopterinae (Hemiptera: Cercopoidea: Clastopteridae) from Africa, Asia and North America" masterLastPageNumber="189" masterPageNumber="151" pageNumber="173" updateTime="1698573836090" updateUser="plazi">
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<mods:title id="8FE069FEE35510E46B3131A6340FCAEB">A new tribe and species of Clastopterinae (Hemiptera: Cercopoidea: Clastopteridae) from Africa, Asia and North America</mods:title>
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<mods:name id="7AA808F60FB758FABCA8C3354B9B17A5" type="personal">
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<mods:roleTerm id="898EB0F366EEB41F6A91A6E1C3A38B3E">Author</mods:roleTerm>
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<mods:namePart id="BB0710F95B006E9670EB45250CD0DBAE">Andrew Hamilton, K. G.</mods:namePart>
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<mods:title id="5D6B401F73DEA2EF200827ECA157291E">Zootaxa</mods:title>
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<mods:date id="2E3261DE344787325B71FDEFD02DF890">2015</mods:date>
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<mods:number id="9B1BAEB3D08932CC7EB7C38C88FEEC23">3946</mods:number>
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<treatment id="B4209064536BFFE4FF5FFDE7FAC16AD7" ID-DOI="http://doi.org/10.5281/zenodo.6106178" ID-GBIF-Taxon="119608429" ID-Zenodo-Dep="6106178" LSID="urn:lsid:plazi:treatment:B4209064536BFFE4FF5FFDE7FAC16AD7" httpUri="http://treatment.plazi.org/id/B4209064536BFFE4FF5FFDE7FAC16AD7" lastPageNumber="173" pageId="22" pageNumber="173">
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<subSubSection id="749372F9536BFFE4FF5FFDE7FE566F5A" box="[151,404,512,538]" pageId="22" pageNumber="173" type="nomenclature">
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<paragraph id="3C362172536BFFE4FF5FFDE7FE566F5A" blockId="22.[151,404,512,538]" box="[151,404,512,538]" pageId="22" pageNumber="173">
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<heading id="677E961E536BFFE4FF5FFDE7FE566F5A" bold="true" box="[151,404,512,538]" fontSize="11" level="1" pageId="22" pageNumber="173" reason="1">
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<emphasis id="0EFDFD60536BFFE4FF5FFDE7FE566F5A" bold="true" box="[151,404,512,538]" pageId="22" pageNumber="173">
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<taxonomicName id="FB895AF1536BFFE4FF5FFDE7FEE56F5A" box="[151,295,512,538]" class="Insecta" family="Clastopteridae" genus="Taphrotylus" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="0EFDFD60536BFFE4FF5FFDE7FEE56F5A" bold="true" box="[151,295,512,538]" italics="true" pageId="22" pageNumber="173">Taphrotylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="15CE401B536BFFE4FEE6FDE7FE566F5A" box="[302,404,512,538]" pageId="22" pageNumber="173" rank="genus">gen.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF5FFDA1FBDA6FC3" pageId="22" pageNumber="173" type="etymology">
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<paragraph id="3C362172536BFFE4FF5FFDA1FCD96F1F" blockId="22.[151,1437,582,1363]" box="[151,795,582,607]" pageId="22" pageNumber="173">
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<emphasis id="0EFDFD60536BFFE4FF5FFDA1FEDC6F1F" bold="true" box="[151,286,582,607]" pageId="22" pageNumber="173">Etymology.</emphasis>
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<emphasis id="0EFDFD60536BFFE4FEECFDA1FE406F1F" box="[292,386,582,607]" italics="true" pageId="22" pageNumber="173">taphros-</emphasis>
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(adjective) ditched;
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<emphasis id="0EFDFD60536BFFE4FDA1FDA1FD5C6F1F" box="[617,670,582,607]" italics="true" pageId="22" pageNumber="173">tylos</emphasis>
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(m), knob.
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</paragraph>
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<paragraph id="3C362172536BFFE4FF0FFD8DFBDA6FC3" blockId="22.[151,1437,582,1363]" box="[199,1048,618,643]" pageId="22" pageNumber="173">
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<emphasis id="0EFDFD60536BFFE4FF0FFD8DFEA46FC3" bold="true" box="[199,358,618,643]" pageId="22" pageNumber="173">Type-species:</emphasis>
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<taxonomicName id="FB895AF1536BFFE4FEA5FD8DFD456FC3" box="[365,647,618,643]" class="Insecta" family="Clastopteridae" genus="Taphrotylus" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="insignificans" status="sp. nov.">
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<emphasis id="0EFDFD60536BFFE4FEA5FD8DFD456FC3" box="[365,647,618,643]" italics="true" pageId="22" pageNumber="173">Taphrotylus insignificans</emphasis>
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</taxonomicName>
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<emphasis id="0EFDFD60536BFFE4FD46FD8DFD256FC3" bold="true" box="[654,743,618,643]" pageId="22" pageNumber="173">
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<taxonomicNameLabel id="15CE401B536BFFE4FD46FD8DFD256FC3" box="[654,743,618,643]" pageId="22" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, the only included species.
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF0FFD69FE376FE7" box="[199,501,654,679]" pageId="22" pageNumber="173" type="distribution">
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<paragraph id="3C362172536BFFE4FF0FFD69FE376FE7" blockId="22.[151,1437,582,1363]" box="[199,501,654,679]" pageId="22" pageNumber="173">
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<emphasis id="0EFDFD60536BFFE4FF0FFD69FEA26FE7" bold="true" box="[199,352,654,679]" pageId="22" pageNumber="173">Distribution.</emphasis>
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<collectingCountry id="449E61E2536BFFE4FEAFFD69FE336FE7" box="[359,497,654,679]" name="Madagascar" pageId="22" pageNumber="173">Madagascar</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF0FFD55FF2669DF" pageId="22" pageNumber="173" type="diagnosis">
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<paragraph id="3C362172536BFFE4FF0FFD55FF2669DF" blockId="22.[151,1437,582,1363]" pageId="22" pageNumber="173">
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<emphasis id="0EFDFD60536BFFE4FF0FFD55FE836F8B" bold="true" box="[199,321,690,715]" pageId="22" pageNumber="173">Diagnosis.</emphasis>
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Dorsoventrally compressed, pronotum weakly declivous in front, crown sloping abruptly to deep groove separating it from convex tylus (Fig. 19A); crown distinctly pointed but a third length of pronotum, tylus exceeding antennal ledges by half its length, width of tylus twice distance between ocelli, posterior border arcuate, more strongly curved than adjacent hind margin of head; antennal ledge high (Fig. 19B), armed with 3 preantennal bristles in vertical row before antennal pit (Fig. 19C); antennal postpedicel with tongue-shaped basiconic sensillum well below flattened base of arista (Fig. 19C), surrounded by 10 or more septate pits, 2 of which bear reduced coeloconic sensilla, the latter lying close to a deep, circular pit containing 4 coeloconic sensilla (Fig. 19D); rostrum extending to hind coxae. Lateral margin of pronotum as long as eye, less than half as long (0.45×) as interocular distance; width of pronotum 0.7 × as great as head. Pronotum and tegmina sparsely punctate; tegmina with veins obscure, a broad but low bulla across base of central anteapical cell, a tiny black bulla just beyond tip of clavus; pecten of 2nd tarsomere of hind leg with 7–8 black-tipped spines.
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<collectingCountry id="449E61E2536BFFE4FC42FBFDFC006973" box="[906,962,1050,1075]" name="India" pageId="22" pageNumber="173">Hind</collectingCountry>
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wing with 3 fine hooks on costal margin. Fore femora 3 × as long as wide; hind tibia slender, 4 × as long as wide, bearing 2 distinct lateral spines; pectens of hind tibia with 7–8 black-tipped spines; those of basitarsomere with 5 black-tipped spines, of 2nd tarsomere with 6 spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF0FFB4DFDE96880" pageId="22" pageNumber="173" type="discussion">
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<paragraph id="3C362172536BFFE4FF0FFB4DFBBF6812" blockId="22.[151,1437,582,1363]" pageId="22" pageNumber="173">
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<emphasis id="0EFDFD60536BFFE4FF0FFB4DFEF96983" bold="true" box="[199,315,1194,1219]" pageId="22" pageNumber="173">Remarks.</emphasis>
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The short head resembles that of
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<taxonomicName id="FB895AF1536BFFE4FD04FB4DFCEA6983" box="[716,808,1194,1219]" class="Insecta" family="Aphrophoridae" genus="Sepullia" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="genus">
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<emphasis id="0EFDFD60536BFFE4FD04FB4DFCEA6983" box="[716,808,1194,1219]" italics="true" pageId="22" pageNumber="173">Sepullia</emphasis>
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</taxonomicName>
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but the tylus bounded by a distinct groove across the crown is unique. The well-developed coeloconic sensilla lying in a deep pit just behind the tongue-shaped basiconic sensillum resemble both sensilla
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<typeStatus id="E3329FD0536BFFE4FDB6FB14FD7B684B" box="[638,697,1267,1291]" pageId="22" pageNumber="173">types</typeStatus>
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of
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<taxonomicName id="FB895AF1536BFFE4FD17FB15FCA0684B" box="[735,866,1266,1291]" class="Insecta" family="Aphrophoridae" genus="Beesoniella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="genus">
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<emphasis id="0EFDFD60536BFFE4FD17FB15FCA0684B" box="[735,866,1266,1291]" italics="true" pageId="22" pageNumber="173">Beesoniella</emphasis>
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</taxonomicName>
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(Fig. 21D), a genus with entirely different habitus (
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<figureCitation id="A4B23DF7536BFFE4FF57FAF1FF2D686F" box="[159,239,1302,1327]" captionStart="FIGURES 3 – 6" captionStartId="10.[151,264,1759,1782]" captionTargetBox="[217,1368,199,1734]" captionTargetId="figure@10.[217,1369,193,1738]" captionTargetPageId="10" captionText="FIGURES 3 – 6. Comparison of face (3 – 4 A), tegmen (3 – 5 B), hind wing (3 – 5 C), lateral aspect of head, pronotum and scutellum (6 A – M) and dorsal aspect (N – O) in genera of Clastopterini (3 – 4), Machaerotinae (5), Sepullini (6 A – L, O) and Aphrophorinae (6 M – N): 3 A, Clastoptera atrapicata; 3 B – C, Clastoptera globosa Fowler; 6 A, Clastoptera achatina; 4 B – C, Iba (s. s.) venosa; 5 B – C, Aphrosiphon bauhiniae, from China (1935), 6 B, Iba (Parahindoloides) lumuana, lectotype, 6 C, Beesoniella sylvestris; 6 D, Sepullia fuscolimbatus; 6 E, Tremapterus (Patriziana) somalicus; 6 F, Tremapterus (s. s.) nigropunctatus, lectotype; 6 G, Tremapterus (s. s.) palimpsestus sp. nov.; 6 H, Tremapterus (Abbalomba) schoutedeni, holotype; 6 J, Tremapterus (Abbalomba) facialis, lectotype; 6 K, Tremapterus (Abbalomba) typicus; 6 L, Grellaphia costalis; 6 O, Sepullia viridicollis var. curtula; 6 M – N, Witteella major (M – O from Jacobi 1910, with hind legs of Sepullia corrected to place spines on outer surface)." httpUri="https://zenodo.org/record/233170/files/figure.png" pageId="22" pageNumber="173">Fig. 6</figureCitation>
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C). Although the male genitalia are unknown, and the overall appearance is that of a small
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<taxonomicName id="FB895AF1536BFFE4FF5FFADDFEAC6813" box="[151,366,1338,1363]" class="Insecta" family="Cercopidae" genus="Pseudomachaerota" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="genus">
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<emphasis id="0EFDFD60536BFFE4FF5FFADDFEAC6813" box="[151,366,1338,1363]" italics="true" pageId="22" pageNumber="173">Pseudomachaerota</emphasis>
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</taxonomicName>
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, the balance of head characters shows a clear relationship to
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<taxonomicName id="FB895AF1536BFFE4FBD2FADDFBB46813" box="[1050,1142,1338,1363]" class="Insecta" family="Aphrophoridae" genus="Sepullia" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="genus">
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<emphasis id="0EFDFD60536BFFE4FBD2FADDFBB46813" box="[1050,1142,1338,1363]" italics="true" pageId="22" pageNumber="173">Sepullia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="3C362172536BFFE4FF5FFA41FDE96880" blockId="22.[151,555,1446,1472]" box="[151,555,1446,1472]" pageId="22" pageNumber="173">
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<taxonomicName id="FB895AF1536BFFE4FF5FFA41FE086880" box="[151,458,1446,1472]" class="Insecta" family="Clastopteridae" genus="Taphrotylus" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="insignificans" status="sp. nov.">
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<emphasis id="0EFDFD60536BFFE4FF5FFA41FE086880" bold="true" box="[151,458,1446,1472]" italics="true" pageId="22" pageNumber="173">Taphrotylus insignificans</emphasis>
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</taxonomicName>
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<emphasis id="0EFDFD60536BFFE4FE1AFA40FDE96880" bold="true" box="[466,555,1447,1472]" pageId="22" pageNumber="173">
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<taxonomicNameLabel id="15CE401B536BFFE4FE1AFA40FDE96880" box="[466,555,1447,1472]" pageId="22" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF5FFA0AFD386B46" box="[151,762,1517,1542]" pageId="22" pageNumber="173" type="etymology">
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<paragraph id="3C362172536BFFE4FF5FFA0AFD386B46" blockId="22.[151,1436,1517,1944]" box="[151,762,1517,1542]" pageId="22" pageNumber="173">
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||
<emphasis id="0EFDFD60536BFFE4FF5FFA0AFEDC6B46" bold="true" box="[151,286,1517,1542]" pageId="22" pageNumber="173">Etymology.</emphasis>
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<emphasis id="0EFDFD60536BFFE4FEECFA09FE5F6B46" box="[292,413,1518,1542]" italics="true" pageId="22" pageNumber="173">significans</emphasis>
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(adjective) marked, important.
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</paragraph>
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</subSubSection>
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<subSubSection id="749372F9536BFFE4FF0FF9F6FE0A6B9D" pageId="22" pageNumber="173" type="diagnosis">
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||
<paragraph id="3C362172536BFFE4FF0FF9F6FE0A6B9D" blockId="22.[151,1436,1517,1944]" pageId="22" pageNumber="173">
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||
<emphasis id="0EFDFD60536BFFE4FF0FF9F6FE836B6A" bold="true" box="[199,321,1553,1578]" pageId="22" pageNumber="173">Diagnosis.</emphasis>
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Yellow ochre with extensive patch on lower half of frons and faint brown spots across fore margin of pronotum embrowned; fore and middle femora and tibiae with brown longitudinal stripe and a paler, indefinite area below this dotted with brown setal bases; basal two-thirds of hind femur, inner surface of hind tibia and tarsal claws brown; lateral spines of hind tibia black. Ovipositor 2nd valvulae
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<quantity id="FB718C97536BFFE4FC2CF99AFB896BD5" box="[996,1099,1661,1686]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="173" unit="mm" value="0.75">0.75 mm</quantity>
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long, wedge-shaped, tapered throughout length to pointed tip, with a single tooth on dorsal margin at one-third length from tip (
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<figureCitation id="A4B23DF7536BFFE4FB2FF946FAF56BFA" box="[1255,1335,1697,1722]" captionStart="FIGURE 11" captionStartId="21.[151,250,1653,1675]" captionTargetBox="[295,1293,523,1626]" captionTargetId="figure@21.[288,1299,519,1631]" captionTargetPageId="21" captionText="FIGURE 11. Second valvulae of Taphrotylus (A), Sepullia (M – P) and Tremapterus subgenera Nyanja (B – G), Selenion (H), Tremiziana (J – L), and subgenus Abbalomba (Q – R). A, Taphrotylus insignificans, holotype; B, Tremapterus bimaculatus; C, T. humilis; D, T. perfasciatus; E, T. indecisus; F, T. scutellatus; G, T. virididorsum; H, T. longicaudatus, holotype; J, T. araneosus, paratype; K, T. scapulatus, paratype; L, T. oxythymus, paratype; M, Sepullia fuscolimbata; N, S. viridicollis; O, S. abyssinia; P, S. murrayi; Q, T. typicus; R, T. facialis. Scale line: 1 mm." httpUri="https://zenodo.org/record/233175/files/figure.png" pageId="22" pageNumber="173">Fig. 11</figureCitation>
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A).
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<collectingCountry id="449E61E2536BFFE4FAAAF946FA5E6BFA" box="[1378,1436,1697,1722]" name="Malta" pageId="22" pageNumber="173">Male</collectingCountry>
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unknown. Length: 4.0 mm.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="749372F9536BFFE4FF0FF90BFC3D6A0F" pageId="22" pageNumber="173" type="materials_examined">
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<paragraph id="3C362172536BFFE4FF0FF90BFC3D6A0F" blockId="22.[151,1436,1517,1944]" pageId="22" pageNumber="173">
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||
<emphasis id="0EFDFD60536BFFE4FF0FF90BFECA6A45" bold="true" box="[199,264,1772,1797]" pageId="22" pageNumber="173">
|
||
<typeStatus id="E3329FD0536BFFE4FF0FF90BFEC66A45" box="[199,260,1772,1797]" pageId="22" pageNumber="173">Type</typeStatus>
|
||
.
|
||
</emphasis>
|
||
♀,
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||
<emphasis id="0EFDFD60536BFFE4FEF1F90BFDC46A45" bold="true" box="[313,518,1772,1797]" pageId="22" pageNumber="173">
|
||
<collectingCountry id="449E61E2536BFFE4FEF1F90BFDC46A45" box="[313,518,1772,1797]" name="Madagascar" pageId="22" pageNumber="173">MADAGASCAR</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="0EFDFD60536BFFE4FDD0F90BFD776A44" box="[536,693,1772,1797]" italics="true" pageId="22" pageNumber="173">Toliara Prov.,</emphasis>
|
||
Fôret Classée d’Analevelona,
|
||
<quantity id="FB718C97536BFFE4FBD3F90BFBB86A45" box="[1051,1146,1772,1797]" metricMagnitude="4" metricUnit="m" metricValue="2.92" pageId="22" pageNumber="173" unit="km" value="29.2">29.2 km</quantity>
|
||
343o NNW Mahaboboka,
|
||
<geoCoordinate id="59BD47B5536BFFE4FF5FF8F4FED16A6B" box="[151,275,1807,1835]" direction="south" orientation="latitude" pageId="22" pageNumber="173" precision="15" value="-22.675">22o40'30"S</geoCoordinate>
|
||
<geoCoordinate id="59BD47B5536BFFE4FED3F8F4FE586A6B" box="[283,410,1807,1835]" direction="east" orientation="longitude" pageId="22" pageNumber="173" precision="15" value="44.190002">44o11'24"E</geoCoordinate>
|
||
,
|
||
<quantity id="FB718C97536BFFE4FE6EF8F4FE3E6A6B" box="[422,508,1811,1835]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="173" unit="m" value="1100.0">1100 m</quantity>
|
||
el.,
|
||
<date id="483707B2536BFFE4FDE6F8F4FD2C6A6B" box="[558,750,1811,1836]" pageId="22" pageNumber="173" value="2003-02-18" valueMax="2003-02-22" valueMin="2003-02-18">18–22 Feb. 2003</date>
|
||
(Fisher, Griswold et al.) Malaise trap, in montane rainforest BLF7818.
|
||
<typeStatus id="E3329FD0536BFFE4FED8F8D1FEBA6A0F" box="[272,376,1846,1871]" pageId="22" pageNumber="173" type="holotype">Holotype</typeStatus>
|
||
in California Academy of Sciences (CASENT 3008099).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="749372F9536BFFE4FF0FF8BCFAC16AD7" pageId="22" pageNumber="173" type="discussion">
|
||
<paragraph id="3C362172536BFFE4FF0FF8BCFAC16AD7" blockId="22.[151,1436,1517,1944]" pageId="22" pageNumber="173">
|
||
<emphasis id="0EFDFD60536BFFE4FF0FF8BCFEF96A34" bold="true" box="[199,315,1883,1908]" pageId="22" pageNumber="173">Remarks.</emphasis>
|
||
Although this was a recently collected specimen, it failed to barcode. The condition of the specimen, with head displaced on the thorax, suggests that it may have begun to rot while in the trap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |