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<document id="655DCA0530CFF40DC06539E15753CD6F" ID-DOI="10.11646/zootaxa.3946.2.1" ID-GBIF-Dataset="f56bb3a6-9852-4218-9c33-b1b3c79670b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="233167" ID-ZooBank="9EE92E94-8743-49E9-B96E-A057C77D9BC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032553967" checkinUser="plazi" docAuthor="Andrew Hamilton, K. G." docDate="2015" docId="B42090645361FFEEFF5FFD8DFE2968F2" docLanguage="en" docName="zt03946p189.pdf" docOrigin="Zootaxa 3946 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tremapterus (Nyanja) viduus Stal, comb. nov." docType="treatment" docVersion="7" lastPageNumber="179" masterDocId="4819E81C537DFFF2FFC8FFE7FFC26D40" masterDocTitle="A new tribe and species of Clastopterinae (Hemiptera: Cercopoidea: Clastopteridae) from Africa, Asia and North America" masterLastPageNumber="189" masterPageNumber="151" pageNumber="179" updateTime="1698573836090" updateUser="plazi">
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<mods:title id="F00C37D3B10DE130C66FCEAC69E8B1D6">A new tribe and species of Clastopterinae (Hemiptera: Cercopoidea: Clastopteridae) from Africa, Asia and North America</mods:title>
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<mods:namePart id="3F7F58A5CFC3B56614CE40FEB70C06DB">Andrew Hamilton, K. G.</mods:namePart>
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<mods:date id="8C68CBE55732C00ABCDF4AD7C1747E19">2015</mods:date>
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<treatment id="B42090645361FFEEFF5FFD8DFE2968F2" ID-DOI="http://doi.org/10.5281/zenodo.6106188" ID-GBIF-Taxon="119608449" ID-Zenodo-Dep="6106188" LSID="urn:lsid:plazi:treatment:B42090645361FFEEFF5FFD8DFE2968F2" httpUri="http://treatment.plazi.org/id/B42090645361FFEEFF5FFD8DFE2968F2" lastPageNumber="179" pageId="28" pageNumber="179">
<subSubSection id="749372F95361FFEEFF5FFD8DFCDE6FA6" pageId="28" pageNumber="179" type="nomenclature">
<paragraph id="3C3621725361FFEEFF5FFD8DFD1B6FC4" blockId="28.[151,729,618,644]" box="[151,729,618,644]" pageId="28" pageNumber="179">
<heading id="677E961E5361FFEEFF5FFD8DFD1B6FC4" bold="true" box="[151,729,618,644]" fontSize="11" level="1" pageId="28" pageNumber="179" reason="1">
<emphasis id="0EFDFD605361FFEEFF5FFD8DFD1B6FC4" bold="true" box="[151,729,618,644]" pageId="28" pageNumber="179">
<taxonomicName id="FB895AF15361FFEEFF5FFD8DFD846FC4" authority="Stal" authorityName="Stal" box="[151,582,618,644]" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="viduus" status="comb. nov." subGenus="Nyanja">
<emphasis id="0EFDFD605361FFEEFF5FFD8DFEF36FC4" bold="true" box="[151,305,618,644]" italics="true" pageId="28" pageNumber="179">Tremapterus</emphasis>
(
<emphasis id="0EFDFD605361FFEEFE89FD8DFE5B6FC4" bold="true" box="[321,409,618,644]" italics="true" pageId="28" pageNumber="179">Nyanja</emphasis>
)
<emphasis id="0EFDFD605361FFEEFE62FD8DFE386FC4" bold="true" box="[426,506,618,644]" italics="true" pageId="28" pageNumber="179">viduus</emphasis>
(Stål)
</taxonomicName>
,
<taxonomicNameLabel id="15CE401B5361FFEEFD9BFD8DFD1B6FC4" box="[595,729,618,644]" pageId="28" pageNumber="179">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3C3621725361FFEEFF5FFD55FDC76F87" blockId="28.[151,796,689,742]" box="[151,517,689,711]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF5FFD55FF646F87" box="[151,166,690,711]" italics="true" pageId="28" pageNumber="179">A</emphasis>
[
<emphasis id="0EFDFD605361FFEEFF67FD56FEE36F87" box="[175,289,689,711]" italics="true" pageId="28" pageNumber="179">cocephalus</emphasis>
]
<taxonomicName id="FB895AF15361FFEEFEE7FD56FDC76F87" authority="Stal, 1855: 98" authorityName="Stal" authorityPageNumber="98" authorityYear="1855" box="[303,517,689,711]" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="0EFDFD605361FFEEFEE7FD56FEB36F87" box="[303,369,689,711]" italics="true" pageId="28" pageNumber="179">viduus</emphasis>
Stål, 1855: 98
</taxonomicName>
</paragraph>
<paragraph id="3C3621725361FFEEFF5FFD36FCDE6FA6" blockId="28.[151,796,689,742]" box="[151,796,720,742]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF5FFD36FF646FA6" box="[151,166,721,742]" italics="true" pageId="28" pageNumber="179">A</emphasis>
[
<emphasis id="0EFDFD605361FFEEFF67FD37FEE36FA6" box="[175,289,720,742]" italics="true" pageId="28" pageNumber="179">cocephalus</emphasis>
]
<taxonomicName id="FB895AF15361FFEEFEE7FD37FDD76FA6" authority="Stal, 1855: 98" authorityName="Stal" authorityPageNumber="98" authorityYear="1855" box="[303,533,720,742]" class="Insecta" family="Aphrophoridae" genus="Acocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="0EFDFD605361FFEEFEE7FD37FE406FA6" box="[303,386,720,742]" italics="true" pageId="28" pageNumber="179">funebris</emphasis>
Stål, 1855: 98
</taxonomicName>
(synonymy by
<bibRefCitation id="58185C835361FFEEFD7CFD37FCD76FA6" author="Stal" box="[692,789,720,742]" pageId="28" pageNumber="179" refString="Stal, C. (1866) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276." type="journal article" year="1866">Stål 1866</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF5FFCF5FE866E33" pageId="28" pageNumber="179" type="diagnosis">
<paragraph id="3C3621725361FFEEFF5FFCF5FE866E33" blockId="28.[151,1436,786,955]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF5FFCF5FED36E6B" bold="true" box="[151,273,786,811]" pageId="28" pageNumber="179">Diagnosis.</emphasis>
A sexually dimorphic species from
<collectingCountry id="449E61E25361FFEEFD16FCF5FCB86E6B" box="[734,890,786,811]" name="South Africa" pageId="28" pageNumber="179">South Africa</collectingCountry>
, males (form
<taxonomicName id="FB895AF15361FFEEFBF8FCF5FB4E6E6B" box="[1072,1164,786,811]" class="Insecta" family="Aphrophoridae" genus="Acocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="0EFDFD605361FFEEFBF8FCF5FB4E6E6B" box="[1072,1164,786,811]" italics="true" pageId="28" pageNumber="179">funebris</emphasis>
</taxonomicName>
) dark with prominent crossbands and a shorter head (
<figureCitation id="A4B23DF75361FFEEFE33FCD1FDFC6E0F" box="[507,574,822,847]" captionStart="FIGURE 7" captionStartId="16.[151,250,1839,1861]" captionTargetBox="[167,1409,513,1788]" captionTargetId="figure@16.[157,1421,510,1807]" captionTargetPageId="16" captionText="FIGURE 7. Dorsal aspects of Grellaphia costalis (A), G. robusta sp. nov. (B) and (at slightly larger scale) Tremapterus subgenus Abbalomba (C D) and T. subgenus Nyanja (E Q): C, Tremapterus schoutedeni; D, T. facialis; E, T. viduus, female lectotype; F, same, male paralectotype; G, T. perfasciatus sp. nov., male holotype; H, same, female; J, T. virididorsum sp. nov., holotype; K, T. indecisus sp. nov., holotype; L, T. scutellatus sp. nov., holotype; M, T. humilis sp. nov., holotype; N, T. muiri; O, T. bukamensis, lectotype; P, T. johannae, lectotype; Q, T. bimaculatus, lectotype. Scale line: 1 mm." httpUri="https://zenodo.org/record/233171/files/figure.png" pageId="28" pageNumber="179">Fig. 7</figureCitation>
F) and females (form
<taxonomicName id="FB895AF15361FFEEFCF0FCD1FC436E0F" box="[824,897,822,847]" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="0EFDFD605361FFEEFCF0FCD1FC436E0F" box="[824,897,822,847]" italics="true" pageId="28" pageNumber="179">viduus</emphasis>
</taxonomicName>
) pale, with tegminal pale markings confined to costa (
<figureCitation id="A4B23DF75361FFEEFF28FCBDFEE16E33" box="[224,291,858,883]" captionStart="FIGURE 7" captionStartId="16.[151,250,1839,1861]" captionTargetBox="[167,1409,513,1788]" captionTargetId="figure@16.[157,1421,510,1807]" captionTargetPageId="16" captionText="FIGURE 7. Dorsal aspects of Grellaphia costalis (A), G. robusta sp. nov. (B) and (at slightly larger scale) Tremapterus subgenus Abbalomba (C D) and T. subgenus Nyanja (E Q): C, Tremapterus schoutedeni; D, T. facialis; E, T. viduus, female lectotype; F, same, male paralectotype; G, T. perfasciatus sp. nov., male holotype; H, same, female; J, T. virididorsum sp. nov., holotype; K, T. indecisus sp. nov., holotype; L, T. scutellatus sp. nov., holotype; M, T. humilis sp. nov., holotype; N, T. muiri; O, T. bukamensis, lectotype; P, T. johannae, lectotype; Q, T. bimaculatus, lectotype. Scale line: 1 mm." httpUri="https://zenodo.org/record/233171/files/figure.png" pageId="28" pageNumber="179">Fig. 7</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF0FFC99FD646968" pageId="28" pageNumber="179" type="discussion">
<paragraph id="3C3621725361FFEEFF0FFC99FCC26EFB" blockId="28.[151,1436,786,955]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF0FFC99FEF96ED7" bold="true" box="[199,315,894,919]" pageId="28" pageNumber="179">Remarks.</emphasis>
Metcalf and Wade (1962) list
<taxonomicName id="FB895AF15361FFEEFD56FC99FD256ED7" box="[670,743,894,919]" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="0EFDFD605361FFEEFD56FC99FD256ED7" box="[670,743,894,919]" italics="true" pageId="28" pageNumber="179">viduus</emphasis>
</taxonomicName>
as junior synonym of
<taxonomicName id="FB895AF15361FFEEFC27FC99FB896ED7" box="[1007,1099,894,919]" class="Insecta" family="Aphrophoridae" genus="Acocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="0EFDFD605361FFEEFC27FC99FB896ED7" box="[1007,1099,894,919]" italics="true" pageId="28" pageNumber="179">funebris</emphasis>
</taxonomicName>
, but this is incorrect because priority was determined by the first reviser (
<bibRefCitation id="58185C835361FFEEFD4DFC45FD336EFB" author="Stal" box="[645,753,930,955]" pageId="28" pageNumber="179" refString="Stal, C. (1866) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276." type="journal article" year="1866">Stål 1866</bibRefCitation>
).
</paragraph>
<paragraph id="3C3621725361FFEEFF5FFBE9FD646968" blockId="28.[151,678,1038,1064]" box="[151,678,1038,1064]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF5FFBE9FD646968" bold="true" box="[151,678,1038,1064]" pageId="28" pageNumber="179">
<taxonomicName id="FB895AF15361FFEEFF5FFBE9FD846968" box="[151,582,1038,1064]" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="virididorsum" status="sp. nov." subGenus="Nyanja">
<emphasis id="0EFDFD605361FFEEFF5FFBE9FEF36968" bold="true" box="[151,305,1038,1064]" italics="true" pageId="28" pageNumber="179">Tremapterus</emphasis>
(
<emphasis id="0EFDFD605361FFEEFE89FBE9FE5B6968" bold="true" box="[321,409,1038,1064]" italics="true" pageId="28" pageNumber="179">Nyanja</emphasis>
)
<emphasis id="0EFDFD605361FFEEFE62FBE9FD846968" bold="true" box="[426,582,1038,1064]" italics="true" pageId="28" pageNumber="179">virididorsum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15CE401B5361FFEEFD84FBE8FD646968" box="[588,678,1039,1064]" pageId="28" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF5FFBB2FCD1692E" box="[151,787,1109,1134]" pageId="28" pageNumber="179" type="etymology">
<paragraph id="3C3621725361FFEEFF5FFBB2FCD1692E" blockId="28.[151,1436,1109,1458]" box="[151,787,1109,1134]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF5FFBB2FEDC692E" bold="true" box="[151,286,1109,1134]" pageId="28" pageNumber="179">Etymology.</emphasis>
<emphasis id="0EFDFD605361FFEEFEECFBB2FEA9692E" box="[292,363,1109,1134]" italics="true" pageId="28" pageNumber="179">viridis</emphasis>
(adjective), green;
<emphasis id="0EFDFD605361FFEEFD8DFBB2FD5B692E" box="[581,665,1109,1134]" italics="true" pageId="28" pageNumber="179">dorsum</emphasis>
(n.), back.
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF0FFB9EFCB669BD" pageId="28" pageNumber="179" type="diagnosis">
<paragraph id="3C3621725361FFEEFF0FFB9EFCB669BD" blockId="28.[151,1436,1109,1458]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF0FFB9EFE8369D2" bold="true" box="[199,321,1145,1170]" pageId="28" pageNumber="179">Diagnosis.</emphasis>
Pronotum greenish, contrasting with orange scutellum; otherwise, form and colour as in
<taxonomicName id="FB895AF15361FFEEFAD5FB9DFEE269F6" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="perfasciatus" subGenus="Nyanja">
<emphasis id="0EFDFD605361FFEEFAD5FB9DFAF069D1" box="[1309,1330,1146,1169]" italics="true" pageId="28" pageNumber="179">T.</emphasis>
(
<emphasis id="0EFDFD605361FFEEFA8AFB9DFA5169D1" box="[1346,1427,1146,1169]" italics="true" pageId="28" pageNumber="179">Nyanja</emphasis>
)
<emphasis id="0EFDFD605361FFEEFF5FFB79FEE269F6" box="[151,288,1182,1206]" italics="true" pageId="28" pageNumber="179">perfasciatus</emphasis>
</taxonomicName>
, but lower half of face and propleura ivory, dorsum with 2nd pronotal band nearly effaced, tegmina darker brown, with irregular pale irrorations (
<figureCitation id="A4B23DF75361FFEEFD5CFB26FD15699A" box="[660,727,1217,1242]" captionStart="FIGURE 7" captionStartId="16.[151,250,1839,1861]" captionTargetBox="[167,1409,513,1788]" captionTargetId="figure@16.[157,1421,510,1807]" captionTargetPageId="16" captionText="FIGURE 7. Dorsal aspects of Grellaphia costalis (A), G. robusta sp. nov. (B) and (at slightly larger scale) Tremapterus subgenus Abbalomba (C D) and T. subgenus Nyanja (E Q): C, Tremapterus schoutedeni; D, T. facialis; E, T. viduus, female lectotype; F, same, male paralectotype; G, T. perfasciatus sp. nov., male holotype; H, same, female; J, T. virididorsum sp. nov., holotype; K, T. indecisus sp. nov., holotype; L, T. scutellatus sp. nov., holotype; M, T. humilis sp. nov., holotype; N, T. muiri; O, T. bukamensis, lectotype; P, T. johannae, lectotype; Q, T. bimaculatus, lectotype. Scale line: 1 mm." httpUri="https://zenodo.org/record/233171/files/figure.png" pageId="28" pageNumber="179">Fig. 7</figureCitation>
J).
<collectingCountry id="449E61E25361FFEEFD30FB26FCF3699A" box="[760,817,1217,1242]" name="Malta" pageId="28" pageNumber="179">Male</collectingCountry>
unknown. Ovipositor with 2nd valvulae fused on basal half, very broad, tapered to sharp tip (
<figureCitation id="A4B23DF75361FFEEFDF6FB02FD4F69BE" box="[574,653,1253,1278]" captionStart="FIGURE 11" captionStartId="21.[151,250,1653,1675]" captionTargetBox="[295,1293,523,1626]" captionTargetId="figure@21.[288,1299,519,1631]" captionTargetPageId="21" captionText="FIGURE 11. Second valvulae of Taphrotylus (A), Sepullia (M P) and Tremapterus subgenera Nyanja (B G), Selenion (H), Tremiziana (J L), and subgenus Abbalomba (Q R). A, Taphrotylus insignificans, holotype; B, Tremapterus bimaculatus; C, T. humilis; D, T. perfasciatus; E, T. indecisus; F, T. scutellatus; G, T. virididorsum; H, T. longicaudatus, holotype; J, T. araneosus, paratype; K, T. scapulatus, paratype; L, T. oxythymus, paratype; M, Sepullia fuscolimbata; N, S. viridicollis; O, S. abyssinia; P, S. murrayi; Q, T. typicus; R, T. facialis. Scale line: 1 mm." httpUri="https://zenodo.org/record/233175/files/figure.png" pageId="28" pageNumber="179">Fig. 11</figureCitation>
G). Length:
<quantity id="FB718C975361FFEEFCDFFB01FCAD69BD" box="[791,879,1254,1278]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="28" pageNumber="179" unit="mm" value="4.1">4.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF0FFAEEFE2B6805" pageId="28" pageNumber="179" type="materials_examined">
<paragraph id="3C3621725361FFEEFF0FFAEEFE2B6805" blockId="28.[151,1436,1109,1458]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF0FFAEEFECA6862" bold="true" box="[199,264,1289,1314]" pageId="28" pageNumber="179">
<typeStatus id="E3329FD05361FFEEFF0FFAEEFEC66862" box="[199,260,1289,1314]" pageId="28" pageNumber="179">Type</typeStatus>
.
</emphasis>
<typeStatus id="E3329FD05361FFEEFED8FAEEFEBA6862" box="[272,376,1289,1314]" pageId="28" pageNumber="179" type="holotype">Holotype</typeStatus>
♀,
<emphasis id="0EFDFD605361FFEEFE6CFAEEFDDD6862" bold="true" box="[420,543,1289,1314]" pageId="28" pageNumber="179">
<collectingCountry id="449E61E25361FFEEFE6CFAEEFDDD6862" box="[420,543,1289,1314]" name="Angola" pageId="28" pageNumber="179">ANGOLA</collectingCountry>
</emphasis>
: Bruco,
<date id="483707B25361FFEEFD4AFAEEFC446862" box="[642,902,1289,1314]" pageId="28" pageNumber="179" value="1972-02-26" valueMax="1972-03-02" valueMin="1972-02-26">26 Feb.2 March 1972</date>
, beaten from Seanchocarpus; Southern African Exp. B.M. 1972-1; in
<collectionCode id="5A98B9B75361FFEEFE45FAC9FE276805" box="[397,485,1326,1349]" pageId="28" pageNumber="179">BMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="749372F95361FFEEFF0FFAB6FE2968F2" pageId="28" pageNumber="179" type="discussion">
<paragraph id="3C3621725361FFEEFF0FFAB6FE2968F2" blockId="28.[151,1436,1109,1458]" pageId="28" pageNumber="179">
<emphasis id="0EFDFD605361FFEEFF0FFAB6FEF9682A" bold="true" box="[199,315,1361,1386]" pageId="28" pageNumber="179">Remarks.</emphasis>
Although this species is known only from the female, it comes from the desert of southwestern
<collectingCountry id="449E61E25361FFEEFF5FFA92FF2868CE" box="[151,234,1397,1422]" name="Angola" pageId="28" pageNumber="179">Angola</collectingCountry>
and is therefore unlikely to be conspecific with the only congeneric species known only from the male:
<taxonomicName id="FB895AF15361FFEEFA4FFA91FE8A68F2" class="Insecta" family="Aphrophoridae" genus="Tremapterus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="perfasciata" subGenus="Nyanja">
<emphasis id="0EFDFD605361FFEEFA4FFA91FA5968CD" box="[1415,1435,1398,1421]" italics="true" pageId="28" pageNumber="179">T.</emphasis>
(
<emphasis id="0EFDFD605361FFEEFF68FA7DFF7868F1" box="[160,186,1434,1457]" italics="true" pageId="28" pageNumber="179">N.</emphasis>
)
<emphasis id="0EFDFD605361FFEEFF02FA7DFE8A68F2" box="[202,328,1434,1458]" italics="true" pageId="28" pageNumber="179">perfasciata</emphasis>
</taxonomicName>
from
<collectingCountry id="449E61E25361FFEEFE44FA7EFE2568F2" box="[396,487,1433,1458]" name="Uganda" pageId="28" pageNumber="179">Uganda</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>