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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. &amp; Wesselingh, Frank P." docDate="2023" docId="DB4FB853CD0056BBBBFFBFD893962178" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Esperiana esperi" docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:title id="AC9D17D1109879F48FB357FE07E10A6A">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:namePart id="AB0234B3DB6A346121788839A3CE7341">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="395E90621B5677CDB81E231187010282">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="A6BA2DC3FCBE4C47AD99ACA1CF220E99">Wesselingh, Frank P.</mods:namePart>
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<mods:affiliation id="E25100074272CF7AF66DEEA5F36E733E">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands &amp; Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<treatment id="DB4FB853CD0056BBBBFFBFD893962178" LSID="urn:lsid:plazi:treatment:DB4FB853CD0056BBBBFFBFD893962178" httpUri="http://treatment.plazi.org/id/DB4FB853CD0056BBBBFFBFD893962178" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection id="6C3A5AF5F2D6CD1EA9929D9D7DC0D945" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="0028D4521AEA42C5B8783D9B69936AE7" pageId="0" pageNumber="53">
<taxonomicName id="CE2A8C7776D07DB30D626F428D7C234D" ID-CoL="3BJDB" LSID="DB4FB853-CD00-56BB-BBFF-BFD893962178" authority="(Ferussac, 1823)" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi">
Esperiana esperi (
<normalizedToken id="C3919E32ABFC2A137E0B0A0DD81385E6" originalValue="Férussac">Ferussac</normalizedToken>
, 1823)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="AFF455F14F6FEF23C91D7C4DF4E8075A" pageId="0" pageNumber="53" type="description">
<paragraph id="FA48F0A427678B34BFC98BCF3EDDB765" pageId="0" pageNumber="53">
<figureCitation id="DDA88CC8BF3E31C886C4967CB7B50481" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Melanopsidae of the Kolankaya I fauna. A-C. Esperiana esperi (Ferussac, 1823), RGM 962691, sample 1. D, E. E. esperi, RGM 962692, sample 1. F. E. esperi, RGM 962693, sample 1. Scale bars: 5 mm." figureDoi="10.3897/zitteliana.97.115682.figure5" httpUri="https://binary.pensoft.net/fig/950742" pageId="0" pageNumber="53">Fig. 5A-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6EE90218ACE94228AF1D7A521D236211" pageId="0" pageNumber="53" type="reference_group">
<paragraph id="12A97E3C9854AD830C3C7B15173CA0B8" pageId="0" pageNumber="53">
<taxonomicName id="CD367E094E15E4DBA8673D558ABB7EBB" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis Esperi" order="Neogastropoda" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="Esperi">*1823 Melanopsis Esperi</taxonomicName>
, nobis -
<normalizedToken id="566F5EAC660277143A3EA920FD2105D2" originalValue="Férussac">Ferussac</normalizedToken>
: 160.
</paragraph>
<paragraph id="128CB81519782563917F58457A3F3521" pageId="0" pageNumber="53">
<taxonomicName id="61BA887AC4D9CF76854892573A5CD158" authorityName="Stefanescu" authorityYear="1896" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis (Melanopsis) bergeroni" order="Neogastropoda" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="bergeroni" subGenus="Melanopsis">1974a Melanopsis (Melanopsis) cf. bergeroni</taxonomicName>
Stefanescu - Taner: 116, pl. 9, figs 16-21.
</paragraph>
<paragraph id="61F2CA565F4B8A79B75298F97C9AEFEC" pageId="0" pageNumber="53">
<taxonomicName id="22D91C15FFD2625C97944DF670210E42" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi">2018 Esperiana esperi</taxonomicName>
(
<normalizedToken id="2AA57B01357888167BEF1D42DAF3A591" originalValue="Férussac">Ferussac</normalizedToken>
, 1823) - Neubauer et al.: 89. fig. 17N.
</paragraph>
<paragraph id="5E99DDC193E5A38F9A66BFE159AD1A48" pageId="0" pageNumber="53">
<taxonomicName id="6FBAA3E88B6DCB1293A776A8D351D8AC" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi">2012 Esperiana esperi</taxonomicName>
(
<normalizedToken id="915688A22416BA770219618AEE106753" originalValue="Férussac">Ferussac</normalizedToken>
, 1823) - Welter-Schultes: 35, unnumbered textfig.
</paragraph>
<paragraph id="ACBDCF77E5976A4B109D19A183DC2699" pageId="0" pageNumber="53">
<taxonomicName id="854B54C343085F88658E88C4AFB0307A" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana (Esperiana) esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi" subGenus="Esperiana">2019 Esperiana (Esperiana) esperi</taxonomicName>
(A.
<normalizedToken id="83065F9A2EC73A048787FAE65C025C8B" originalValue="Férussac">Ferussac</normalizedToken>
, 1823) -
<normalizedToken id="988FA21F10CCE4657B0191F6BE66E638" originalValue="Glöer">Gloeer</normalizedToken>
: 83, textfig. 83.
</paragraph>
</subSubSection>
<subSubSection id="3F402F7DAE678214A9E032186DB685BA" pageId="0" pageNumber="53" type="materials_examined">
<paragraph id="3A24EC700319387F0B95CC3A8AA90E85" pageId="0" pageNumber="53">Material.</paragraph>
<paragraph id="90B830B4CAC87D3EF560CA1E32EB1CA2" pageId="0" pageNumber="53">
Sample 1:
<specimenCount id="3A08CD5D485448C00A9C25F1D89EBE1C" type="generic">3 specimens</specimenCount>
(RGM 962691-962693),
<specimenCount id="66CA39540392C35F4E48A2368751CCED" type="generic">10 specimens</specimenCount>
(SNSB-BSPG
<date id="B9A9C7600220D07FE309FC7616895B01" value="2023-12-04">2023 XII 4</date>
), ca.
<specimenCount id="D29D2A960904C66EA7032805123CFF5A" type="generic">300 specimens</specimenCount>
(RGM 1310843); sample 3:
<specimenCount id="EDE78CB1434ED3E5C386F316DBE6B955" type="generic">13 specimens</specimenCount>
(RGM 1310844),
<specimenCount id="1E1EE0C31B9024BCFF03908C20E0707F" type="generic">1 specimen</specimenCount>
(RGM 1365348).
</paragraph>
</subSubSection>
<subSubSection id="5C3A286FC7802DB3AA66109869225C44" pageId="0" pageNumber="53" type="description">
<paragraph id="7CA683E3E2DE08BBF6FD10819F7B8474" pageId="0" pageNumber="53">Dimensions.</paragraph>
<paragraph id="52413E544345030C20BA32EA16CD0A6A" pageId="0" pageNumber="53">
16.25
<normalizedToken id="9C3F1EFD683BA89FDF749CA5F118EB93" originalValue="×">x</normalizedToken>
7.78 mm (RGM 962691; Fig.
<figureCitation id="14A8FC53C9CF7D00CE6017A3ABDFF3C5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Melanopsidae of the Kolankaya I fauna. A-C. Esperiana esperi (Ferussac, 1823), RGM 962691, sample 1. D, E. E. esperi, RGM 962692, sample 1. F. E. esperi, RGM 962693, sample 1. Scale bars: 5 mm." figureDoi="10.3897/zitteliana.97.115682.figure5" httpUri="https://binary.pensoft.net/fig/950742" pageId="0" pageNumber="53">5A-C</figureCitation>
), 14.44
<normalizedToken id="E9FB45BC374412E0DE86A97373915768" originalValue="×">x</normalizedToken>
7.47 mm (RGM 962692; Fig.
<figureCitation id="9D6B8679CD9BA234C55F2A5B4813F416" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Melanopsidae of the Kolankaya I fauna. A-C. Esperiana esperi (Ferussac, 1823), RGM 962691, sample 1. D, E. E. esperi, RGM 962692, sample 1. F. E. esperi, RGM 962693, sample 1. Scale bars: 5 mm." figureDoi="10.3897/zitteliana.97.115682.figure5" httpUri="https://binary.pensoft.net/fig/950742" pageId="0" pageNumber="53">5D, E</figureCitation>
).
</paragraph>
<caption id="46DFC0265B727EE70428008E821DDAD8" doi="10.3897/zitteliana.97.115682.figure5" httpUri="https://binary.pensoft.net/fig/950742" pageId="0" pageNumber="53" start="Figure 5" startId="F5">
<paragraph id="659BB5E91460259D97FAA6CBCA105091" pageId="0" pageNumber="53">
<emphasis id="49F430173FA3C85ED0C7C7AF4D53B220" bold="true" pageId="0" pageNumber="53">Figure 5.</emphasis>
<taxonomicName id="6D9F0242F5AF5E9E6BD58F0CF1D86045" baseAuthorityName="Adams &amp; Adams" baseAuthorityYear="1854" class="Bivalvia" family="Melanopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Sorbeoconcha" pageId="0" pageNumber="53" phylum="Mollusca" rank="family">Melanopsidae</taxonomicName>
of the Kolankaya I fauna.
<emphasis id="694B82508268AA224A18B3375131C3AC" bold="true" pageId="0" pageNumber="53">A-C.</emphasis>
<taxonomicName id="5EB54AA586098A07829EA1BFCF83B4E7" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi">
<emphasis id="BA611CDA2E0A8C0A971222168C543CA8" italics="true" pageId="0" pageNumber="53">Esperiana esperi</emphasis>
</taxonomicName>
(
<normalizedToken id="16972B7A08DDD6E3AD526AA424C83299" originalValue="Férussac">Ferussac</normalizedToken>
, 1823), RGM 962691, sample 1.
<emphasis id="98316D8729B029E0849757F15B92746A" bold="true" pageId="0" pageNumber="53">D, E.</emphasis>
<taxonomicName id="BF3EEFA0CD56F16D63EB2E8F0F0988F9" lsidName="E. esperi" pageId="0" pageNumber="53" rank="species" species="esperi">
<emphasis id="8C5ADDFBD210E734CF24BE86A2AA8031" italics="true" pageId="0" pageNumber="53">E. esperi</emphasis>
</taxonomicName>
, RGM 962692, sample 1.
<emphasis id="96647E706D2EBE31FD5D61E1BCEC6C38" bold="true" pageId="0" pageNumber="53">F.</emphasis>
<taxonomicName id="13E4FE95E11293D01F0EC1434646D510" lsidName="E. esperi" pageId="0" pageNumber="53" rank="species" species="esperi">
<emphasis id="9E59AD67549675C66FE4BE125D460BF5" italics="true" pageId="0" pageNumber="53">E. esperi</emphasis>
</taxonomicName>
, RGM 962693, sample 1. Scale bars: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1414B2C85AA0C7F8AE8A5351A0594E9C" pageId="0" pageNumber="53" type="remarks">
<paragraph id="F03B77CE498EA89870B523A490155F6D" pageId="0" pageNumber="53">Remarks.</paragraph>
<paragraph id="B22D57E9E252925339216E8B9B831391" pageId="0" pageNumber="53">
The present material matches well the extant
<taxonomicName id="33EDC69353F5CE20D8C0449C9852C5FC" baseAuthorityName="Ferussac" baseAuthorityYear="1823" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana esperi" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperi">
<emphasis id="24D6736EFAE6DD1C7661A42585B663CF" italics="true" pageId="0" pageNumber="53">Esperiana esperi</emphasis>
</taxonomicName>
concerning the elongate, smooth shell with adpressed whorls, lanceolate aperture with thickened inner lip and the slight concavity between base of the last whorl and base of the aperture (
<bibRefCitation id="30D9A44C74A1245215A30F37518F7CC8" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="53" refId="B170" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
;
<bibRefCitation id="94335DDB26D815A17AEB97CA5292BDE3" author="Gloeer, P" journalOrPublisher="Identification key, Anatomy, Ecology, Distribution. Privately published" pageId="0" pageNumber="53" refId="B61" refString="Gloeer, P, 2019. The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails. Identification key, Anatomy, Ecology, Distribution. Privately published" title="The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken id="7F9CAFEB6AD255A2E05173C600114089" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
). Even the color pattern, consisting of dissected zigzag lines to a chess-like arrangement of orange spots, and the corroded apex are typical of the modern species.
</paragraph>
<paragraph id="027046CEEBDCB28D15795E02DD6FD230" pageId="0" pageNumber="53">
Shape, apertural characteristics and color pattern are also found in the Plio-Pleistocene species
<taxonomicName id="1026FA965C89862D6E7CCF146938D3D4" authorityName="Stefanescu" authorityYear="1896" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis bergeroni" order="Neogastropoda" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="bergeroni">
<emphasis id="56D7702C630997D2A56E47C9DDA4B2B0" italics="true" pageId="0" pageNumber="53">Melanopsis bergeroni</emphasis>
</taxonomicName>
Stefanescu, 1896 from the Dacian Basin (Romania) (
<bibRefCitation id="411AF0DECE863ACAD64561CACDBF1CED" author="Stefanescu, S" journalOrPublisher="Paleontologique" pageId="0" pageNumber="53" pagination="1 - 147" refId="B147" refString="Stefanescu, S, 1896. Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15. Paleontologique 6: 1 - 147" title="Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15." volume="6" year="1896">Stefanescu 1896</bibRefCitation>
;
<bibRefCitation id="619DAF5FD8D0A0B3B647D50FF69ED3D1" author="Pană, I" journalOrPublisher="Institutul de cercetǎri, inginerie tehnologicǎ și proiectǎri miniere pentru lignit, Craiova" pageId="0" pageNumber="53" refId="B105" refString="Pană, I, Enache, C, Andreescu, I, 1981. Fauna de moluste a depozitelor cu ligniti din Oltenia. Institutul de cercetǎri, inginerie tehnologicǎ și proiectǎri miniere pentru lignit, Craiova" title="Fauna de moluste a depozitelor cu ligniti din Oltenia." year="1981">
<normalizedToken id="763629F202494D243096B83B08C8187D" originalValue="Pană">Pană</normalizedToken>
et al. 1981
</bibRefCitation>
;
<bibRefCitation id="D00D26EDA52E3AC35A96180663D1A14F" author="Pană, I" editor="Papaianopol, I" journalOrPublisher="Neogen der Zentrale Paratethys, Bd. X, Pl 2. Romanien. Editura Academiei Romane, Bucuresti" pageId="0" pageNumber="53" pagination="296 - 349" refId="B104" refString="Pană, I, 2003. Les nannogastropodes. In: Papaianopol, I, Marinescu, F, Krstic, N, Macalet, R, Eds., Chronostratigraphie und Neostratotypen. Neogen der Zentrale Paratethys, Bd. X, Pl2. Romanien. Editura Academiei Romane, Bucuresti: 296 - 349" title="Les nannogastropodes." volumeTitle="Chronostratigraphie und Neostratotypen." year="2003">
<normalizedToken id="B10CA3BB3B3A1B9401EF3DC4C1DE552A" originalValue="Pană">Pană</normalizedToken>
2003
</bibRefCitation>
). The same is true for
<taxonomicName id="D9054FFCB3094F8DF0DD7D35CD7182F4" authorityName="Stefanescu" authorityYear="1896" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis esperioides" order="Neogastropoda" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="esperioides">
<emphasis id="5F733DE9E860F77ED9DEF8DBB13FE87A" italics="true" pageId="0" pageNumber="53">Melanopsis esperioides</emphasis>
</taxonomicName>
Stefanescu, 1896 from the same region, which seems to be only slightly more slender and larger on average than
<taxonomicName id="607CAF96545F7F087D999C6B6B0758D2" lsidName="M. bergeroni" pageId="0" pageNumber="53" rank="species" species="bergeroni">
<emphasis id="88073539ABB18252550C4A0B2F2FAA57" italics="true" pageId="0" pageNumber="53">M. bergeroni</emphasis>
</taxonomicName>
(
<bibRefCitation id="587155A5166C0A40B70093017CBFCEEF" author="Stefanescu, S" journalOrPublisher="Paleontologique" pageId="0" pageNumber="53" pagination="1 - 147" refId="B147" refString="Stefanescu, S, 1896. Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15. Paleontologique 6: 1 - 147" title="Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15." volume="6" year="1896">Stefanescu 1896</bibRefCitation>
). It might well be that both species range within the morphological variability of
<taxonomicName id="F5DB9D946BD812B6FAFFD04776E5D716" lsidName="E. esperi" pageId="0" pageNumber="53" rank="species" species="esperi">
<emphasis id="EFCD5472823845C2B0431BD6010FE242" italics="true" pageId="0" pageNumber="53">E. esperi</emphasis>
</taxonomicName>
, but a final verdict on their status requires comparison with topotypic material from the Dacian Basin. In case of synonymy, the subgenus
<taxonomicName id="63C6631E44CD343FDDC2EE1943A2D727" authority="" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis (Calodiona)" order="Neogastropoda" pageId="0" pageNumber="53" phylum="Mollusca" rank="subGenus" subGenus="Calodiona">Melanopsis (Calodiona)</taxonomicName>
Stefanescu, 1896, of which
<taxonomicName id="60D68E01268F086256DAD94A3AAC7113" lsidName="M. bergeroni" pageId="0" pageNumber="53" rank="species" species="bergeroni">
<emphasis id="9944301B193EAC5179F5959A888AF944" italics="true" pageId="0" pageNumber="53">M. bergeroni</emphasis>
</taxonomicName>
is the type species, would become a synonym of
<taxonomicName id="EB20346D9ACB1FD027E2351B293861C8" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Melanopsidae" genus="Esperiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Esperiana" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="7D6FC7F63B79C13D148FCE7227CE7960" italics="true" pageId="0" pageNumber="53">Esperiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="518E3A7C8AEB41BF062E6331F75F8F21" pageId="0" pageNumber="53">
The specimens from the Denizli Basin studied by
<bibRefCitation id="13ABC3325D3D221B2315425F85D97A7A" author="Taner, G" journalOrPublisher="Bulletin of Mineral Research and Exploration Institute of Turkey (MTA)" pageId="0" pageNumber="53" pagination="89 - 126" refId="B148" refString="Taner, G, 1974a. Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue. Bulletin of Mineral Research and Exploration Institute of Turkey (MTA) 82: 89 - 126" title="Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue." volume="82" year="1974 a">Taner (1974a)</bibRefCitation>
and identified as
<taxonomicName id="139A72B30159C3503D81AA10E1BA32B8" lsidName="M. bergeroni" pageId="0" pageNumber="53" rank="species" species="bergeroni">M. cf. bergeroni</taxonomicName>
falls well into the range of
<taxonomicName id="D821060FAB3668B550A7D6D6A9E93574" lsidName="E. esperi" pageId="0" pageNumber="53" rank="species" species="esperi">
<emphasis id="3A23A332C745CB2F6E6571B307598D89" italics="true" pageId="0" pageNumber="53">E. esperi</emphasis>
</taxonomicName>
and considered conspecific herein.
</paragraph>
</subSubSection>
<subSubSection id="3CE2F363FDE31F923D7D6B351BE8EB07" pageId="0" pageNumber="53" type="distribution">
<paragraph id="E922C01107BAD2679B2407C5A932BCD9" pageId="0" pageNumber="53">Distribution.</paragraph>
<paragraph id="759736C0C9A2B977EBC0FDECE9CB2F27" pageId="0" pageNumber="53">
Today in central to southeastern Europe (western Black Sea region to Hungary, Slovakia, Austria, and Slovenia) and northwestern Anatolia (
<bibRefCitation id="48642CC19E2394D6A96FC1A243EF78E5" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="53" refId="B170" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
;
<bibRefCitation id="2B1B9814F65E80228C3ABC623CBB3A74" author="Gloeer, P" journalOrPublisher="Identification key, Anatomy, Ecology, Distribution. Privately published" pageId="0" pageNumber="53" refId="B61" refString="Gloeer, P, 2019. The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails. Identification key, Anatomy, Ecology, Distribution. Privately published" title="The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken id="9227165D767F4109AE8B2230658199F0" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
). Also found in Pleistocene strata of the North Caspian Basin (
<bibRefCitation id="A99AE00AB9DA548090DB794727A8AD81" DOI="https://doi.org/10.3897/zookeys.770.25365" author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="53" pagination="43 - 103" refId="B91" refString="Neubauer, TA, van de Velde, S, Yanina, T, Wesselingh, FP, 2018. A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys 770: 43 - 103, DOI: https://doi.org/10.3897/zookeys.770.25365" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2452B289EBF5399C374C28FDD42AC98B" pageId="0" pageNumber="53" type="superfamily truncatelloidea gray, 1840">
<paragraph id="1D4B4B77F7BBE28F24D9F1BE4C85BB99" pageId="0" pageNumber="53">
Superfamily
<taxonomicName id="B8CBFF25379A50DC963BD754E7E1CB6D" lsidName="" pageId="0" pageNumber="53" rank="superFamily" superFamily="Truncatelloidea">Truncatelloidea</taxonomicName>
Gray, 1840
</paragraph>
<paragraph id="3AAEAE9FABB4953113FCC9DE018318E4" pageId="0" pageNumber="53">
<emphasis id="3FBB44481076CD53A8EA6408DB602CFA" bold="true" pageId="0" pageNumber="53">
Family
<taxonomicName id="6242B6F294A885024C157C23347C5408" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
Stimpson, 1865
</emphasis>
</paragraph>
<paragraph id="DFC0AAB4EE1701E831110EE569A2E359" pageId="0" pageNumber="53">
<emphasis id="B2D54BC67C23BADE579BB2F11EB2584C" bold="true" pageId="0" pageNumber="53">
Subfamily
<taxonomicName id="508053349D898F3AF7871F581385398C" authorityName="B.Dybowski" authorityYear="1913" lsidName="" pageId="0" pageNumber="53" rank="subFamily" subFamily="Caspiinae">Caspiinae</taxonomicName>
B. Dybowski, 1913
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>