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<document id="4FC924BCEA37C4A2BF1F5D421EF8B62E" ID-DOI="10.5281/zenodo.277513" ID-GBIF-Dataset="c499f937-b2bb-4333-8504-958183e30f73" ID-ISSN="1175-5326" ID-Zenodo-Dep="277513" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460307345794" checkinUser="plazi" docAuthor="Sturaro, Marcelo José &amp; Avila-Pires, Teresa C. S." docDate="2011" docId="C708878CFFFD995FFF75FE4FFA0DEC26" docLanguage="en" docName="zt02869p036.pdf" docOrigin="Zootaxa 2869" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gonatodes tapajonicus Rodrigues 1980" docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="3B31FFF4FFEC9943FFE2FF9EFF90EA46" masterDocTitle="Taxonomic revision of the geckos of the Gonatodes concinnatus complex (Squamata: Sphaerodactylidae), with description of two new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="18" updateTime="1698253556106" updateUser="plazi">
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<mods:title id="475243B784035FBB7D0D2D360C06E7F1">Taxonomic revision of the geckos of the Gonatodes concinnatus complex (Squamata: Sphaerodactylidae), with description of two new species</mods:title>
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<mods:namePart id="6BA66AA3F6EF5028231D56057B2B6A88">Sturaro, Marcelo José</mods:namePart>
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<mods:namePart id="97512022AB446023003F0AB3ADC843FD">Avila-Pires, Teresa C. S.</mods:namePart>
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<mods:date id="010D85C91B53B75B8A2744692B424279">2011</mods:date>
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<treatment id="C708878CFFFD995FFF75FE4FFA0DEC26" ID-DOI="http://doi.org/10.5281/zenodo.5618486" ID-GBIF-Taxon="119400476" ID-Zenodo-Dep="5618486" LSID="urn:lsid:plazi:treatment:C708878CFFFD995FFF75FE4FFA0DEC26" httpUri="http://treatment.plazi.org/id/C708878CFFFD995FFF75FE4FFA0DEC26" lastPageId="28" lastPageNumber="29" pageId="17" pageNumber="18">
<subSubSection id="07BB6511FFFD9952FF75FE4FFEDAE84A" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="4F1E369AFFFD9952FF75FE4FFDE1EBAC" blockId="17.[151,625,464,524]" box="[151,625,464,491]" pageId="17" pageNumber="18">
<heading id="145681F6FFFD9952FF75FE4FFDE1EBAC" bold="true" box="[151,625,464,491]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="88A14D19FFFD9952FF75FE4FFDE1EBAC" authority="Rodrigues, 1980" authorityName="Rodrigues" authorityYear="1980" box="[151,625,464,491]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFFD9952FF75FE4FFDE1EBAC" bold="true" box="[151,625,464,491]" pageId="17" pageNumber="18">
<emphasis id="7DD5EA88FFFD9952FF75FE4FFE35EBAD" bold="true" box="[151,421,465,491]" italics="true" pageId="17" pageNumber="18">Gonatodes tapajonicus</emphasis>
<bibRefCitation id="2B304B6BFFFD9952FE4EFE4EFDE1EBAC" author="Rodrigues" box="[428,625,464,490]" pageId="17" pageNumber="18" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues, 1980</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4F1E369AFFFD9952FF75FE6AFEDAE84A" blockId="17.[151,625,464,524]" box="[151,330,500,524]" pageId="17" pageNumber="18">
(
<figureCitation id="D79A2A1FFFFD9952FF7DFE6AFF77E84A" box="[159,231,500,524]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="17" pageNumber="18">Figs. 1</figureCitation>
C, 9, 10)
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFFD9952FF75FDA2FD5EE836" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="4F1E369AFFFD9952FF75FDA2FD5EE836" blockId="17.[151,1436,571,624]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="6FB111B4FFFD9952FF75FDA2FD5EE836" pageId="17" pageNumber="18">
<taxonomicName id="88A14D19FFFD9952FF75FDA2FDF1E817" authority="Rodrigues, 1980: 309" authorityName="Rodrigues" authorityPageNumber="309" authorityYear="1980" box="[151,609,571,594]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFFD9952FF75FDA2FEEFE814" box="[151,383,572,594]" italics="true" pageId="17" pageNumber="18">Gonatodes tapajonicus</emphasis>
<treatmentCitation id="CE00108BFFFD9952FE65FDA5FDF1E817" author="Rodrigues" box="[391,609,571,593]" page="309" pageId="17" pageNumber="18" year="1980">
<bibRefCitation id="2B304B6BFFFD9952FE65FDA5FDA0E817" author="Rodrigues" box="[391,560,571,593]" pageId="17" pageNumber="18" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues, 1980</bibRefCitation>
: 309
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="901A8838FFFD9952FD92FDA5FD58E817" box="[624,712,571,593]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
MZUSP 53676, type-locality: Cachoeira do Limão, Rio Tapajós, Pará,
<collectingCountry id="37B6760AFFFD9952FF25FDC4FE98E836" box="[199,264,602,624]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
);
<treatmentCitation id="CE00108BFFFD9952FEFAFDC4FE4DE836" author="Vanzolini" box="[280,477,602,624]" page="10" pageId="17" pageNumber="18" year="1986">
<bibRefCitation id="2B304B6BFFFD9952FEFAFDC4FE28E836" author="Vanzolini" box="[280,440,602,624]" pageId="17" pageNumber="18" refString="Vanzolini, P. E. (1986) Addenda and corrigenda to the Catalogue of Neotropical Squamata. Smithsonian Herpetological Information Service, 70, 1 - 25." type="journal article" year="1986">Vanzolini, 1986</bibRefCitation>
: 10
</treatmentCitation>
;
<treatmentCitation id="CE00108BFFFD9952FE0AFDC4FD5AE836" author="Avila-Pires" box="[488,714,602,624]" page="283" pageId="17" pageNumber="18" year="1995">
<bibRefCitation id="2B304B6BFFFD9952FE0AFDC4FD08E836" author="Avila-Pires" box="[488,664,602,624]" pageId="17" pageNumber="18" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Avila-Pires, 1995</bibRefCitation>
: 283
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFFD9952FF75FD03FCEDE9D3" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="4F1E369AFFFD9952FF75FD03FCEDE9D3" blockId="17.[151,1437,669,2034]" pageId="17" pageNumber="18">
<emphasis id="7DD5EA88FFFD9952FF75FD03FEE4E8F0" bold="true" box="[151,372,669,694]" pageId="17" pageNumber="18">Material examined</emphasis>
(numbers in bold between parentheses refer to localities in
<figureCitation id="D79A2A1FFFFD9952FC1EFD03FBAEE8F3" box="[1020,1086,669,693]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="17" pageNumber="18">Fig. 3</figureCitation>
):
<emphasis id="7DD5EA88FFFD9952FBBAFD03FB59E8F0" bold="true" box="[1112,1225,669,694]" pageId="17" pageNumber="18">
<collectingCountry id="37B6760AFFFD9952FBBAFD03FB54E8F0" box="[1112,1220,669,694]" name="Brazil" pageId="17" pageNumber="18">BRAZIL</collectingCountry>
.
</emphasis>
<collectionCode id="29B0AE5FFFFD9952FB2DFD03FA84E8F3" box="[1231,1300,669,693]" pageId="17" pageNumber="18">PARÁ</collectionCode>
.
<emphasis id="7DD5EA88FFFD9952FAC1FD03FF65E89C" bold="true" pageId="17" pageNumber="18">Município Itaituba</emphasis>
. Comunidade Aldeia Nova (&quot;Aldêia Akay Muuybu&quot;), anteriorly known as Comunidade Boa Fé, right margen of Rio Tapajós (
<geoCoordinate id="2A95505DFFFD9952FE45FD76FDD7E946" box="[423,583,741,768]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="-4.6995">04o 41' 58.2&quot;S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFFD9952FD62FD76FCBCE946" box="[640,812,741,768]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-56.37694">56o 22' 37.0&quot;W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFFD9952FCA7FD79FCF0E946" bold="true" box="[837,864,743,768]" pageId="17" pageNumber="18">45</emphasis>
):
<collectionCode id="29B0AE5FFFFD9952FC98FD76FC57E946" box="[890,967,744,768]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="17" pageNumber="18">MPEG</collectionCode>
2770827718,
<specimenCount id="59A7FD13FFFD9952FB97FD76FB61E946" box="[1141,1265,744,768]" pageId="17" pageNumber="18" type="juvenile">2 juveniles</specimenCount>
,
<specimenCount id="59A7FD13FFFD9952FB1CFD76FAC5E946" box="[1278,1365,744,768]" pageId="17" pageNumber="18" type="male">5 males</specimenCount>
and
<specimenCount id="59A7FD13FFFD9952FA6CFD76FF7FE961" pageId="17" pageNumber="18" type="female">4 females</specimenCount>
, leg. M.J. Sturaro, P.L.V. Peloso and J.
<collectingCountry id="37B6760AFFFD9952FD75FC91FD3EE961" box="[663,686,783,807]" name="Iceland" pageId="17" pageNumber="18">O</collectingCountry>
. Gomes. Cachoeira do Limão, right margin of Rio Tapajós (~
<geoCoordinate id="2A95505DFFFD9952FAB0FC91FA0CE961" box="[1362,1436,780,807]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="-4.6833334">4o41S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFFD9952FF2AFCABFEBAE90B" box="[200,298,818,845]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-56.35">56o21W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFFD9952FEA7FCABFEF1E908" bold="true" box="[325,353,821,846]" pageId="17" pageNumber="18">46</emphasis>
):
<collectionCode id="29B0AE5FFFFD9952FE99FCABFE47E90B" box="[379,471,821,845]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="18">MZUSP</collectionCode>
53676 (
<typeStatus id="901A8838FFFD9952FDD4FCABFD0BE90B" box="[566,667,821,845]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
), male;
<collectionCode id="29B0AE5FFFFD9952FD14FCABFCC2E90B" box="[758,850,821,845]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="18">MZUSP</collectionCode>
53669, 5367174, 53677 (
<typeStatus id="901A8838FFFD9952FB67FCABFB64E90B" box="[1157,1268,821,845]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
), a male and
<specimenCount id="59A7FD13FFFD9952FA6CFCABFF75E937" pageId="17" pageNumber="18" type="female">5 female</specimenCount>
, leg. M.T. Rodrigues. Floresta Nacional de Altamira, Distrito de Moraes de Almeida (
<geoCoordinate id="2A95505DFFFD9952FB28FCC7FAAAE937" box="[1226,1338,857,881]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="-6.018889">06º 18”S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFFD9952FA9AFCC7FF6EE9D3" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-55.30472">55º 1817”W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFFD9952FEF4FCE3FEA2E9D0" bold="true" box="[278,306,893,918]" pageId="17" pageNumber="18">47</emphasis>
):
<collectionCode id="29B0AE5FFFFD9952FEABFCE3FE06E9D3" box="[329,406,893,917]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="17" pageNumber="18">MPEG</collectionCode>
25095, female, leg. C. Lima and R.R. Silva.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFFD9952FF25FC3FFF78EEBB" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="4F1E369AFFFD9952FF25FC3FFF78EEBB" blockId="17.[151,1437,669,2034]" pageId="17" pageNumber="18">
<emphasis id="7DD5EA88FFFD9952FF25FC3FFEAFE9FC" bold="true" box="[199,319,929,954]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
A relatively large
<taxonomicName id="88A14D19FFFD9952FDFEFC3CFD03E9FC" box="[540,659,930,954]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="7DD5EA88FFFD9952FDFEFC3CFD03E9FC" box="[540,659,930,954]" italics="true" pageId="17" pageNumber="18">Gonatodes</emphasis>
</taxonomicName>
, with maximum SVL of
<quantity id="88599B7FFFFD9952FC5FFC3FFB82E9FF" box="[957,1042,929,953]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="18" unit="mm" value="55.0">55 mm</quantity>
. Scales around midbody 118133. Ventral scales in a longitudinal row 5363. Proximal subdigital lamellae as wide as digits, in total 1822 under fourth finger, 2227 under fourth toe. Three, occasionally four, lateral rows of scales on distal portion of fingers and toes. Tail with midventral scales distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral in contact laterodistally with two laterodistal scales per side (respectively 112” and 111”). A white suprahumeral ocellus, bordered by black, present both in males and females, although more conspicuous in males. Males with a vermiculated pattern of light and dark (vivid yellow and reddish brown in life) spots both on head and body dorsally; gular region with dark oblique streaks (yellow with reddish brown streaks in life).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFFD9951FF25FA97FC9DED9A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="4F1E369AFFFD9952FF25FA97FBE8EF93" blockId="17.[151,1437,669,2034]" pageId="17" pageNumber="18">
<emphasis id="7DD5EA88FFFD9952FF25FA97FEC6EF64" bold="true" box="[199,342,1289,1314]" pageId="17" pageNumber="18">Description.</emphasis>
Maximum SVL in males of
<quantity id="88599B7FFFFD9952FD4FFA97FD6FEF67" box="[685,767,1289,1313]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="17" pageNumber="18" unit="mm" value="53.0">53 mm</quantity>
(MZUSP 53676), in females of
<quantity id="88599B7FFFFD9952FB62FA97FB42EF67" box="[1152,1234,1289,1313]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="18" unit="mm" value="55.0">55 mm</quantity>
(MZUSP 53671) (
<bibRefCitation id="2B304B6BFFFD9952FF7CFAB3FEC2EF03" author="Rodrigues" box="[158,338,1325,1349]" pageId="17" pageNumber="18" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues 1980</bibRefCitation>
). Tail round in cross section, tapering toward tip, 1.11.3 (1.21 ± 0.10,
<emphasis id="7DD5EA88FFFD9952FBB8FAB1FBFCEF00" box="[1114,1132,1327,1350]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=6) times SLV. Head length 0.220.27 (0.23 ± 0.01,
<emphasis id="7DD5EA88FFFD9952FE46FACDFE27EF2C" box="[420,439,1363,1386]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times SLV, 1.41.6 (1.47 ± 0.06,
<emphasis id="7DD5EA88FFFD9952FC87FACDFCE8EF2C" box="[869,888,1363,1386]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times as long as wide, 1.31.5 (1.41 ± 0.06,
<emphasis id="7DD5EA88FFFD9952FF75FAE9FF3AEFC8" box="[151,170,1399,1422]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times as wide as high. Snout round, moderately elongate (
<figureCitation id="D79A2A1FFFFD9952FC92FAEBFC3CEFCB" box="[880,940,1397,1421]" captionStart="FIGURE 9" captionStartId="19.[151,250,1982,2005]" captionTargetBox="[184,1431,190,1981]" captionTargetId="figure@19.[151,1436,147,2037]" captionTargetPageId="19" captionText="FIGURE 9. Gonatodes tapajonicus (paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="17" pageNumber="18">Fig 9</figureCitation>
A), gently sloping toward top of head. Neck slightly narrower than head and anterior portion of body. Body cylindrical. Limbs well developed, lower arm 0.13 0.15 (0.14 ± 0.01,
<emphasis id="7DD5EA88FFFD9952FE83FA21FEE3EF90" box="[353,371,1471,1494]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times SVL, lower leg 0.120.16 (0.15, ± 0.01,
<emphasis id="7DD5EA88FFFD9952FC4FFA21FC50EF90" box="[941,960,1471,1494]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times SVL.
</paragraph>
<paragraph id="4F1E369AFFFD9952FF27FA7FFB68EDB7" blockId="17.[151,1437,669,2034]" pageId="17" pageNumber="18">
Rostral convex, 1.82.4 (2.11 ± 0.17,
<emphasis id="7DD5EA88FFFD9952FD8EFA7DFDEFEFBC" box="[620,639,1507,1530]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times as wide as high; posterior margin with a shallow depression medially and slightly indented by 13 medial postrostrals, with or without a medial cleft extending anteriorly. Postrostrals 3 (79%), 4 (10.5%), or 5 (10.5%), laterals ones (supranasals) distinctly larger than medial ones (
<emphasis id="7DD5EA88FFFD9952FAB7F9B5FAF7EC04" box="[1365,1383,1579,1602]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) (
<figureCitation id="D79A2A1FFFFD9952FF7CF9D3FF4DEC23" box="[158,221,1613,1637]" captionStart="FIGURE 9" captionStartId="19.[151,250,1982,2005]" captionTargetBox="[184,1431,190,1981]" captionTargetId="figure@19.[151,1436,147,2037]" captionTargetPageId="19" captionText="FIGURE 9. Gonatodes tapajonicus (paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="17" pageNumber="18">Fig 9</figureCitation>
A). Nasal bordered by rostral, first supralabial (only a narrow contact in some specimens), 3 (68.4%) or 4 (31.6%) postnasals (
<emphasis id="7DD5EA88FFFD9952FE63F9EDFE04ECCC" box="[385,404,1651,1674]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19), and supranasal; internostril distance 0.20.3 (0.25 ± 0.02,
<emphasis id="7DD5EA88FFFD9952FB9FF9EDFB00ECCC" box="[1149,1168,1651,1674]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times head width. Supranasal roughly oval or circular, 1.01.5 (1.20 ± 0.14;
<emphasis id="7DD5EA88FFFD9952FCECF909FCB0ECE8" box="[782,800,1687,1710]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times as wide as long. Postnasals slightly larger than, or similar in size to, adjacent loreals. Scales on snout convex, hexagonal to round, juxtaposed, relatively uniform in size. Canthus rostralis rounded. Loreal region with scales slightly more elongate than those on snout, largest on row adjacent to supralabials, 1013 (11.1 ± 0.8,
<emphasis id="7DD5EA88FFFD9952FD90F89DFD15ED5C" box="[626,645,1795,1818]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) loreals on a line between postnasals and anterior corner of eye. Top and posterior portion of head, as well as supraorbital region, with granular scales. A short supraciliary flap present, anteriorly with 710 (8.6 ± 1.0,
<emphasis id="7DD5EA88FFFD9952FDEBF8D5FD8CED24" box="[521,540,1867,1890]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) enlarged and flattened scales, among which 04 (2.6 ± 1.1;
<emphasis id="7DD5EA88FFFD9952FAEBF8D5FA8CED24" box="[1289,1308,1867,1890]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) small, conical spines. Pupil round, eye diameter 0.200.27 (0.23 ± 0.01,
<emphasis id="7DD5EA88FFFD9952FC87F8F1FCE7EDC0" box="[869,887,1903,1926]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19) times head length. Scales on temporal region similar to those on top of head. Ear-opening much smaller than eye, oval, posterior to, and at same level of, commissure of mouth. Supralabials 58 (6.7 ± 0.7;
<emphasis id="7DD5EA88FFFD9952FD10F829FC95ED88" box="[754,773,1975,1998]" italics="true" pageId="17" pageNumber="18">N</emphasis>
=19), distinctly enlarged anteriorly and decreasing in size posteriorly, one or two of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FF09FD1AEB79" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
Mental large, distinctly wider anteriorly than posteriorly, with posterior margin forming a wide angle, 1.11.4 (1.25 ± 0.11;
<emphasis id="7DD5EA88FFFE9951FECEFF23FEAFEA92" box="[300,319,189,212]" italics="true" pageId="18" pageNumber="19">N</emphasis>
=19) times as wide as long; 1 (10.5%), 2 (63.2%) or 3 (26.3%) postmentals (
<figureCitation id="D79A2A1FFFFE9951FB72FF22FB43EA92" box="[1168,1235,188,212]" captionStart="FIGURE 9" captionStartId="19.[151,250,1982,2005]" captionTargetBox="[184,1431,190,1981]" captionTargetId="figure@19.[151,1436,147,2037]" captionTargetPageId="19" captionText="FIGURE 9. Gonatodes tapajonicus (paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
B). Scales on chin flat, smooth, polygonal, juxtaposted, larger anteriorly, decreasing in size posteriorly. Infralabials 57 (5.8 ± 0.7;
<emphasis id="7DD5EA88FFFE9951FF75FE9BFF3AEB5A" box="[151,170,261,284]" italics="true" pageId="18" pageNumber="19">N</emphasis>
=19), distinctly enlarged anteriorly and decreasing in size posteriorly; 13 of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FED2FC8BEBEA" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">Scales on nape small and granular, becoming slightly larger on sides of neck. Scales on throat anteriorly granular, posteriorly flat, smooth, hexagonal or round, increasing in size toward the posterior region, with a short transitional zone between the anterior and posterior parts.</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FE29FA00E8C2" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
Dorsals granular, increasing in size toward the flanks. Ventrals larger than dorsals, roughly hexagonal, flat, smooth, imbricate, in oblique rows; 5363 (57.4 ± 3.3;
<emphasis id="7DD5EA88FFFE9951FCFAFE43FCBBEBB2" box="[792,811,477,500]" italics="true" pageId="18" pageNumber="19">N</emphasis>
=19) scales along the midventral line between anterior margin of forelimbs and vent; 1821 (19.2 ± 0.9;
<emphasis id="7DD5EA88FFFE9951FD0EFD9FFD6FE85E" box="[748,767,513,536]" italics="true" pageId="18" pageNumber="19">N</emphasis>
=18) scales in a transverse line at midbody, with a short transitional zone between ventrals and scales on flanks. Scales around midbody 118133 (124.2 ± 3.2;
<emphasis id="7DD5EA88FFFE9951FAAFFDBBFAF0E87A" box="[1357,1376,549,572]" italics="true" pageId="18" pageNumber="19">N</emphasis>
=18). Scales on preanal plate similar to ventrals, except for those bordering vent, which are very small. Escutcheon present in males on posterior portion of belly and on four rows (body-knee direction) on ventral surface of thighs.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FD11FD98E971" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">Scales on anterodorsal surface of forelimbs flat, smooth, roundish or rhomboidal, imbricate, largest close to the wrist; on posterodorsal and ventral surface of forelimbs convex, smooth, rhomboidal, juxtaposed, relatively small. Scales on anterodorsal surface of thighs and ventral surface of hind limbs flat, smooth, roundish or rhomboidal, imbricate and relatively larger; on posterodorsal surface of thighs and dorsal surface of lower legs smaller, convex, smooth, rhomboidal, juxtaposed.</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FCDAFBBBE9AA" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
Lamellae under second (
<emphasis id="7DD5EA88FFFE9951FE3DFCDAFE65E91A" bold="true" box="[479,501,836,860]" pageId="18" pageNumber="19">II</emphasis>
) through fourth (
<emphasis id="7DD5EA88FFFE9951FD5CFCDAFD4DE91A" bold="true" box="[702,733,836,860]" pageId="18" pageNumber="19">IV</emphasis>
) fingers (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFFE9951FB00FCDAFB68E91A" bold="true" box="[1250,1272,836,860]" pageId="18" pageNumber="19">II</emphasis>
: 1518 (56),
<emphasis id="7DD5EA88FFFE9951FF75FCF6FF28E9C6" bold="true" box="[151,184,872,896]" pageId="18" pageNumber="19">III</emphasis>
: 1820 (57), and
<emphasis id="7DD5EA88FFFE9951FE73FCF6FE3EE9C6" bold="true" box="[401,430,872,896]" pageId="18" pageNumber="19">IV</emphasis>
: 1822 (68) (fig 9C). Lamellae under second through fourth toes (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFFE9951FED1FC12FED9E9E2" bold="true" box="[307,329,908,932]" pageId="18" pageNumber="19">II</emphasis>
: 1418 (57),
<emphasis id="7DD5EA88FFFE9951FE11FC12FD84E9E2" bold="true" box="[499,532,908,932]" pageId="18" pageNumber="19">III</emphasis>
: 1622 (58), and
<emphasis id="7DD5EA88FFFE9951FD0DFC12FC9CE9E2" bold="true" box="[751,780,908,932]" pageId="18" pageNumber="19">IV</emphasis>
: 2227 (1012) (
<figureCitation id="D79A2A1FFFFE9951FC36FC12FB88E9E2" box="[980,1048,908,932]" captionStart="FIGURE 9" captionStartId="19.[151,250,1982,2005]" captionTargetBox="[184,1431,190,1981]" captionTargetId="figure@19.[151,1436,147,2037]" captionTargetPageId="19" captionText="FIGURE 9. Gonatodes tapajonicus (paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
D). Claws exposed, non-retractile, between two basal scales. Fingers and toes with three, occasionally four, lateral scales distally, between the fourth (counted from the claw towards the hand) subdigital lamella, and the dorsal scale.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF27FC69FC26EE82" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
Scales on tail dorsally and laterally relatively small, rhomboidal, flat, smooth, imbricate. On ventral surface of tail scales smooth, flat, imbricate, increasing in size toward midventral line; midventral scales, except close to the base of the tail, distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral scale in contact laterodistally with two scales per side respectively 112” and 111” in the codification of
<bibRefCitation id="2B304B6BFFFE9951FDE5FB32FCD3EE82" author="Avila-Pires" box="[519,835,1196,1220]" pageId="18" pageNumber="19" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">
Avila-Pires (1995:
<figureCitation id="D79A2A1FFFFE9951FD3EFB32FCABEE82" box="[732,827,1196,1220]" captionStart="FIGURE 2" captionStartId="6.[151,250,1440,1463]" captionTargetBox="[190,1402,193,1415]" captionTargetId="figure@6.[185,1402,193,1419]" captionTargetPageId="6" captionText="FIGURE 2. Bivariate scattergrams from stepwise discriminant function analysis using scale counts and measurements. Red = Pattern A. Cyan-green = Pattern B. Lime-green = Pattern C. Yellow = Pattern D. Blue = Pattern E." httpUri="https://zenodo.org/record/277515/files/figure.png" pageId="18" pageNumber="19">Figure 2</figureCitation>
)
</bibRefCitation>
(
<figureCitation id="D79A2A1FFFFE9951FCB0FB32FC06EE82" box="[850,918,1196,1220]" captionStart="FIGURE 9" captionStartId="19.[151,250,1982,2005]" captionTargetBox="[184,1431,190,1981]" captionTargetId="figure@19.[151,1436,147,2037]" captionTargetPageId="19" captionText="FIGURE 9. Gonatodes tapajonicus (paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
E).
</paragraph>
<paragraph id="4F1E369AFFFE9951FF25FB51FECEEF87" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="7DD5EA88FFFE9951FF25FB51FE5AEEAE" bold="true" box="[199,458,1231,1256]" pageId="18" pageNumber="19">Color in preservative.</emphasis>
In males (
<figureCitation id="D79A2A1FFFFE9951FDDCFB4EFDE7EEAE" box="[574,631,1232,1256]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="18" pageNumber="19">Fig 1</figureCitation>
C), dorsal surface of head with relatively large reddish-brown and beige vermiculations. Sides of head with oblique and/or longitudinal, beige and brown bands. Back and flanks, base of tail and limbs with beige and brown vermiculations. A large, conspicuous, white suprahumeral ocellus with black margin. Ventral surface of head and gular region beige with oblique black streaks, in contact or not at midventral line; chest beige; belly dark-brown; underside of limbs and base of tail brown. Dorsal surface of tail brown, ventral surface reddish-brown; distal part in one specimen with a beige ring around the tail. Escutcheon area (belly and thighs) light gray.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF25FA53FA83ECDE" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">In females, dorsal surface of head and limbs with brown and beige irregular spots. Sides of head with beige and brown oblique and/or longitudinal bands. Back gray with dorsolateral pairs of black and white spots; flanks gray with black and brown irregular spots. A moderately large, conspicuous, white suprahumeral ocellus, with black margin, smaller than that of males. Ventral surface of head and gular region white with dark oblique streaks, in contact or not at midventral line; belly and underside of limbs light gray. Tail brown and/or black dorsally, white and/or brown ventrally; original tail distally with white bands that form complete rings around the tail.</paragraph>
<paragraph id="4F1E369AFFFE9951FF25F93DFC0CED0A" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="7DD5EA88FFFE9951FF25F93DFEE2ECFA" bold="true" box="[199,370,1699,1724]" pageId="18" pageNumber="19">Color in life.</emphasis>
In males (
<figureCitation id="D79A2A1FFFFE9951FDE5F93AFDF1ECFA" box="[519,609,1700,1724]" captionStart="FIGURE 10" captionStartId="20.[151,250,1040,1063]" captionTargetBox="[151,1436,161,1018]" captionTargetId="figure@20.[151,1436,161,1018]" captionTargetPageId="20" captionText="FIGURE 10. Gonatodes tapajonicus from Aldeia Nova, righ margin of Rio Tapajós, Pará, Brazil. Adult male (A) in dorsal (MPEG 27714, SVL = 52.5 mm) and (B) ventral (MPEG 27718, SVL = 48.7 mm) views. Adult female (C) in dorsal (MPEG 27715, SVL = 39.5 mm) and (D) ventral (MPEG 27716, SVL = 51.1 mm) views (Photo C by Pedro L. Peloso)." httpUri="https://zenodo.org/record/277523/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
AB), head dorsally and laterally reddish-brown and vivid yellow. Suprahumeral ocellus white with black margin. Vermiculations on back brown and yellow, smaller vermiculations on flanks bluish-white and black. Head ventrally and gular region vivid yellow with reddish-brown streaks, chest vivid yellow. Remaining ventral region dark gray, lighter on escutcheon areas (belly and thighs). Tail dark gray to black all around, except near the base, where it is similar to the back.
</paragraph>
<paragraph id="4F1E369AFFFE9951FF25F8C7FC9DED9A" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
In females (
<figureCitation id="D79A2A1FFFFE9951FEADF8C6FE31ED36" box="[335,417,1880,1904]" captionStart="FIGURE 10" captionStartId="20.[151,250,1040,1063]" captionTargetBox="[151,1436,161,1018]" captionTargetId="figure@20.[151,1436,161,1018]" captionTargetPageId="20" captionText="FIGURE 10. Gonatodes tapajonicus from Aldeia Nova, righ margin of Rio Tapajós, Pará, Brazil. Adult male (A) in dorsal (MPEG 27714, SVL = 52.5 mm) and (B) ventral (MPEG 27718, SVL = 48.7 mm) views. Adult female (C) in dorsal (MPEG 27715, SVL = 39.5 mm) and (D) ventral (MPEG 27716, SVL = 51.1 mm) views (Photo C by Pedro L. Peloso)." httpUri="https://zenodo.org/record/277523/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
CD), head dorsally and lateraly gray and brown. Suprahumeral ocellus white with black margin. Back brown with pairs of black and gray spots. Head ventrally and gular region beige with light-yellow and gray streaks, chest white. Remaining ventral region white. Tail anteriorly brown with black spots dorsally, orange ventrally; posteriorly with black and white rings.
</paragraph>
</subSubSection>
<caption id="1BDE6612FFFF9950FF75F820FA0CED93" box="[151,1436,1982,2005]" httpUri="https://zenodo.org/record/277522/files/figure.png" pageId="19" pageNumber="20" targetBox="[184,1431,190,1981]" targetPageId="19">
<paragraph id="4F1E369AFFFF9950FF75F820FA0CED93" blockId="19.[151,1436,1982,2005]" box="[151,1436,1982,2005]" pageId="19" pageNumber="20">
<emphasis id="7DD5EA88FFFF9950FF75F820FE82ED92" bold="true" box="[151,274,1982,2005]" pageId="19" pageNumber="20">FIGURE 9.</emphasis>
<taxonomicName id="88A14D19FFFF9950FEFBF821FD93ED93" box="[281,515,1983,2005]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFFF9950FEFBF821FD93ED93" box="[281,515,1983,2005]" italics="true" pageId="19" pageNumber="20">Gonatodes tapajonicus</emphasis>
</taxonomicName>
(paratype, MZUSP 53674). (A) Dorsal and (B) ventral views of head; ventral views of (C)
</paragraph>
</caption>
<caption id="1BDE6612FFF89957FF75FB8EFB6EEE25" httpUri="https://zenodo.org/record/277523/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,161,1018]" targetPageId="20">
<paragraph id="4F1E369AFFF89957FF75FB8EFB6EEE25" blockId="20.[151,1436,1040,1124]" pageId="20" pageNumber="21">
<emphasis id="7DD5EA88FFF89957FF75FB8EFEB1EE63" bold="true" box="[151,289,1040,1063]" pageId="20" pageNumber="21">FIGURE 10.</emphasis>
<taxonomicName id="88A14D19FFF89957FEC8FB8EFD86EE60" box="[298,534,1040,1062]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFF89957FEC8FB8EFD86EE60" box="[298,534,1040,1062]" italics="true" pageId="20" pageNumber="21">Gonatodes tapajonicus</emphasis>
</taxonomicName>
from Aldeia Nova, righ margin of Rio Tapajós, Pará, Brazil. Adult male (A) in dorsal (MPEG 27714, SVL = 52.5 mm) and (B) ventral (MPEG 27718, SVL = 48.7 mm) views. Adult female (C) in dorsal (MPEG 27715, SVL = 39.5 mm) and (D) ventral (MPEG 27716, SVL = 51.1 mm) views (Photo C by Pedro L. Peloso).
</paragraph>
</caption>
<subSubSection id="07BB6511FFF89957FF25FB11FB98EEEE" box="[199,1032,1167,1192]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="4F1E369AFFF89957FF25FB11FB98EEEE" blockId="20.[151,1436,1167,1519]" box="[199,1032,1167,1192]" pageId="20" pageNumber="21">
<emphasis id="7DD5EA88FFF89957FF25FB11FEF0EEEE" bold="true" box="[199,352,1167,1192]" pageId="20" pageNumber="21">Distribution.</emphasis>
Interfluvium TapajósXingu, state of Pará,
<collectingCountry id="37B6760AFFF89957FCAFFB0EFC01EEEE" box="[845,913,1168,1192]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
(
<figureCitation id="D79A2A1FFFF89957FC42FB0EFC73EEEE" box="[928,995,1168,1192]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF89957FF25FB2BFEC2EC3B" pageId="20" pageNumber="21" type="discussion">
<paragraph id="4F1E369AFFF89957FF25FB2BFAF3EFA9" blockId="20.[151,1436,1167,1519]" pageId="20" pageNumber="21">
<emphasis id="7DD5EA88FFF89957FF25FB2BFEABEE8B" bold="true" box="[199,315,1205,1229]" pageId="20" pageNumber="21">Remarks.</emphasis>
<bibRefCitation id="2B304B6BFFF89957FEA0FB2AFD96EE8A" author="Rodrigues" box="[322,518,1204,1228]" pageId="20" pageNumber="21" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues (1980)</bibRefCitation>
pointed out that the type-series was collected in an area with predominance of the palm
<taxonomicName id="88A14D19FFF89957FF34FB47FE08EEB6" box="[214,408,1240,1264]" class="Liliopsida" family="Arecaceae" genus="Euterpe" kingdom="Animalia" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="oleracea">
<emphasis id="7DD5EA88FFF89957FF34FB47FE08EEB6" box="[214,408,1240,1264]" italics="true" pageId="20" pageNumber="21">Euterpe oleracea</emphasis>
</taxonomicName>
(“açaizal”), along a trail known as Boa Fé, in front of Cachoeira do Limão, Tapajós river, at approximately
<geoCoordinate id="2A95505DFFF89957FEB9FB61FEEFEF4C" box="[347,383,1276,1303]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="0.06666667">
0
<geoCoordinate id="2A95505DFFF89957FE8BFB61FEEFEF4C" box="[361,383,1276,1303]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-4.6833334">4o</geoCoordinate>
</geoCoordinate>
41´S
<geoCoordinate id="2A95505DFFF89957FE5EFB61FDB0EF51" box="[444,544,1276,1303]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-45.35">45o21´W</geoCoordinate>
. It is possible that it is the same locality where MPEG 2770827718 have been collected, community Boa Fé being the old name of this locality. Similar to the type-series, these specimens were in an “açaizal” in a flooded depression within the forest; they were found mostly on the palm
<taxonomicName id="88A14D19FFF89957FB47FAD6FAF6EF19" box="[1189,1382,1351,1375]" class="Liliopsida" family="Arecaceae" genus="Euterpe" kingdom="Animalia" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="oleracea">
<emphasis id="7DD5EA88FFF89957FB47FAD6FAF6EF19" box="[1189,1382,1351,1375]" italics="true" pageId="20" pageNumber="21">Euterpe oleracea</emphasis>
</taxonomicName>
, one specimen on a tree trunk, between 30 and
<quantity id="88599B7FFFF89957FD91FAF5FD56EFC5" box="[627,710,1387,1411]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="cm" value="200.0">200 cm</quantity>
above the ground. Contrary to what was observed by
<bibRefCitation id="2B304B6BFFF89957FACBFAF5FF73EFE1" author="Rodrigues" pageId="20" pageNumber="21" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues (1980)</bibRefCitation>
, we also found
<taxonomicName id="88A14D19FFF89957FE7BFA0EFDB9EFE1" box="[409,553,1423,1447]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="humeralis">
<emphasis id="7DD5EA88FFF89957FE7BFA0EFDB9EFE1" box="[409,553,1423,1447]" italics="true" pageId="20" pageNumber="21">G. humeralis</emphasis>
</taxonomicName>
in the “açaizal”, in syntopy with
<taxonomicName id="88A14D19FFF89957FC48FA0EFBDDEFE1" box="[938,1101,1424,1447]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFF89957FC48FA0EFBDDEFE1" box="[938,1101,1424,1447]" italics="true" pageId="20" pageNumber="21">G. tapajonicus</emphasis>
</taxonomicName>
. The
<taxonomicName id="88A14D19FFF89957FB73FA0EFAC4EFE1" box="[1169,1364,1423,1447]" class="Liliopsida" family="Arecaceae" genus="Euterpe" kingdom="Animalia" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="oleracea">
<emphasis id="7DD5EA88FFF89957FB73FA0EFAC4EFE1" box="[1169,1364,1423,1447]" italics="true" pageId="20" pageNumber="21">Euterpe oleracea</emphasis>
</taxonomicName>
forest was one of the few in the area still well-preserved, due to the exploitation in other similar environments of the palm heart for the food market. The species was not found in the more disturbed açaizais, nor outside this habitat.
</paragraph>
<paragraph id="4F1E369AFFF89957FF75F9DDFEC2EC3B" blockId="20.[151,531,1603,1661]" pageId="20" pageNumber="21">
<taxonomicName id="88A14D19FFF89957FF75F9DDFE22EC1B" box="[151,434,1603,1629]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="nascimentoi" status="sp. nov.">
<emphasis id="7DD5EA88FFF89957FF75F9DDFE22EC1B" bold="true" box="[151,434,1603,1629]" italics="true" pageId="20" pageNumber="21">Gonatodes nascimentoi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFF89957FE5BF9DAFD83EC1D" bold="true" box="[441,531,1604,1628]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="66E657F3FFF89957FE5BF9DAFD83EC1D" box="[441,531,1604,1628]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D79A2A1FFFF89957FF7DF9FBFF4FEC3B" box="[159,223,1637,1661]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
D, 11, 12)
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF89956FF75F930FEB6EB21" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="4F1E369AFFF89957FF75F930FE3DED71" blockId="20.[151,1436,1709,2031]" pageId="20" pageNumber="21">
<emphasis id="7DD5EA88FFF89957FF75F930FE94EC80" bold="true" box="[151,260,1710,1734]" pageId="20" pageNumber="21">
<typeStatus id="901A8838FFF89957FF75F930FE94EC80" box="[151,260,1710,1734]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
(number in bold between parentheses refer to localities in
<figureCitation id="D79A2A1FFFF89957FC50F933FC69EC83" box="[946,1017,1709,1733]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
):
<collectionCode id="29B0AE5FFFF89957FBF0F933FBF0EC83" box="[1042,1120,1709,1733]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="20" pageNumber="21">MPEG</collectionCode>
25596 (field number BML 446), an adult male from Fazenda Caracol, right margin of Rio Xingu, Município Anapu, Pará,
<collectingCountry id="37B6760AFFF89957FB31F94AFA87ECAA" box="[1235,1303,1748,1772]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
(
<geoCoordinate id="2A95505DFFF89957FACBF94AFA0CECAA" box="[1321,1436,1745,1772]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-3.4583335">3o2730”S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFF89957FF2BF965FEC5ED55" box="[201,341,1784,1811]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-51.675835">51o4033”W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFF89957FE92F965FE1CED52" bold="true" box="[368,396,1787,1812]" pageId="20" pageNumber="21">50</emphasis>
), collected in pitfall trap inside primary forest, by A. A. Lima, M. J. Sturaro and R. A. T. Rocha,
<date id="3B1F105AFFF89957FF0EF881FE3AED71" box="[236,426,1823,1848]" pageId="20" pageNumber="21" value="2008-01-12">12 January 2008</date>
.
</paragraph>
<paragraph id="4F1E369AFFF89956FF25F8DAFEB6EB21" blockId="20.[151,1436,1709,2031]" lastBlockId="21.[151,1437,151,2026]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="7DD5EA88FFF89957FF25F8DAFEAFED1A" bold="true" box="[199,319,1860,1884]" pageId="20" pageNumber="21">
<typeStatus id="901A8838FFF89957FF25F8DAFEAFED1A" box="[199,319,1860,1884]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
</emphasis>
(numbers in bold between parentheses refer to localities in
<figureCitation id="D79A2A1FFFF89957FC3FF8DDFBB1ED1D" box="[989,1057,1859,1883]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
):
<emphasis id="7DD5EA88FFF89957FBD5F8DDFB36ED1A" bold="true" box="[1079,1190,1859,1884]" pageId="20" pageNumber="21">
<collectingCountry id="37B6760AFFF89957FBD5F8DDFB31ED1A" box="[1079,1185,1859,1884]" name="Brazil" pageId="20" pageNumber="21">BRASIL</collectingCountry>
.
</emphasis>
<collectionCode id="29B0AE5FFFF89957FB4CF8DDFA83ED1D" box="[1198,1299,1859,1883]" pageId="20" pageNumber="21">AMAPÁ</collectionCode>
.
<emphasis id="7DD5EA88FFF89957FAC3F8DDFEC6ED39" bold="true" pageId="20" pageNumber="21">Município Ferreira Gomes</emphasis>
. Floresta Nacional do Amapá (
<geoCoordinate id="2A95505DFFF89957FD22F8F9FCDCED39" box="[704,844,1895,1919]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="1.1102778">01º0637N</geoCoordinate>
and
<geoCoordinate id="2A95505DFFF89957FC65F8F9FB88ED39" box="[903,1048,1895,1919]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-51.893612">51º5337W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFF89957FBD1F8F8FBDFED39" bold="true" box="[1075,1103,1894,1919]" pageId="20" pageNumber="21">48</emphasis>
):
<collectionCode id="29B0AE5FFFF89957FB88F8F9FB37ED39" box="[1130,1191,1895,1919]" httpUri="http://grbio.org/cool/qq2e-yp5c" name="Istituto di Entomologia Agraria dell'Universita" pageId="20" pageNumber="21">IEPA</collectionCode>
/FL 361, male, leg. J. Lima,
<date id="3B1F105AFFF89957FF02F811FEC6EDE0" box="[224,342,1934,1958]" pageId="20" pageNumber="21" value="2005-06">June 2005</date>
.
<collectionCode id="29B0AE5FFFF89957FEBDF810FE37EDE0" box="[351,423,1934,1958]" pageId="20" pageNumber="21">PARÁ</collectionCode>
.
<emphasis id="7DD5EA88FFF89957FE57F813FD3AEDE0" bold="true" box="[437,682,1933,1958]" pageId="20" pageNumber="21">Município Almeirim</emphasis>
. Monte Dourado, forest near Estação Ecológica do Jari (
<geoCoordinate id="2A95505DFFF89957FAC6F810FA0CEDE0" box="[1316,1436,1931,1958]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-0.5908333">0o3527S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFF89957FF2AF82CFECAED8C" box="[200,346,1970,1994]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-52.735832">52º4409W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFF89957FE96F82FFE00ED8C" bold="true" box="[372,400,1969,1994]" pageId="20" pageNumber="21">49</emphasis>
):
<collectionCode id="29B0AE5FFFF89957FE48F82CFE68ED8C" box="[426,504,1970,1994]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="20" pageNumber="21">MPEG</collectionCode>
2382227,
<specimenCount id="59A7FD13FFF89957FD62F82CFD49ED8C" box="[640,729,1970,1994]" pageId="20" pageNumber="21" type="male">5 males</specimenCount>
and a female, leg. T. A. Gardner and M. A. Ribeiro-Jr,
<date id="3B1F105AFFF89957FAB9F82DFE9FEDA8" pageId="20" pageNumber="21" value="2005-05" valueMax="2005-06-31" valueMin="2005-05-00">May June 2005</date>
;
<collectionCode id="29B0AE5FFFF89957FEFFF848FEFBEDA8" box="[285,363,2006,2030]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="20" pageNumber="21">MPEG</collectionCode>
27719, female, leg. M. J. Sturaro and J.
<collectingCountry id="37B6760AFFF89957FCAAF848FCCFEDA8" box="[840,863,2006,2030]" name="Iceland" pageId="20" pageNumber="21">O</collectingCountry>
. Gomes,
<date id="3B1F105AFFF89957FC2EF849FBF8EDA8" box="[972,1128,2006,2031]" pageId="20" pageNumber="21" value="2009-01">January 2009</date>
.
<emphasis id="7DD5EA88FFF89957FB97F848FADBEDA9" bold="true" box="[1141,1355,2006,2031]" pageId="20" pageNumber="21">Município Anapu</emphasis>
. Typelocality (
<emphasis id="7DD5EA88FFF99956FF1EFF09FE88EAF6" bold="true" box="[252,280,151,176]" pageId="21" pageNumber="22">50</emphasis>
):
<collectionCode id="29B0AE5FFFF99956FED0FF09FE10EAE9" box="[306,384,151,175]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="22">MPEG</collectionCode>
25164, male, leg. A. A. Lima and R. A. T. Rocha,
<date id="3B1F105AFFF99956FC2BFF09FB14EAE9" box="[969,1156,151,175]" pageId="21" pageNumber="22" value="2007-11">November 2007</date>
;
<collectionCode id="29B0AE5FFFF99956FB6DFF09FB4DEAE9" box="[1167,1245,151,175]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="22">MPEG</collectionCode>
25595, 25597, a juvenile, a female and a male, leg. A. A. Lima, M. J. Sturaro and R. A. T. Rocha,
<date id="3B1F105AFFF99956FBA8FF23FB74EA92" box="[1098,1252,188,213]" pageId="21" pageNumber="22" value="2008-01">January 2008</date>
;
<collectionCode id="29B0AE5FFFF99956FB10FF22FAD0EA92" box="[1266,1344,188,212]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="22">MPEG</collectionCode>
25598 25601, a male and
<specimenCount id="59A7FD13FFF99956FE8DFF41FE4FEAB1" box="[367,479,223,247]" pageId="21" pageNumber="22" type="female">3 females</specimenCount>
, leg. A. A. Lima, M. J. Sturaro, R. A. T. Rocha, P. L. V. Peloso and F. S. Rodrigues,
<date id="3B1F105AFFF99956FF75FE99FEB6EB59" box="[151,294,263,287]" pageId="21" pageNumber="22" value="2008-03">March 2008</date>
.
<emphasis id="7DD5EA88FFF99956FED0FE98FE6FEB59" bold="true" box="[306,511,262,287]" pageId="21" pageNumber="22">Município Portel</emphasis>
. Fazenda Riacho Monte Verde (~
<geoCoordinate id="2A95505DFFF99956FC72FE99FC4BEB59" box="[912,987,260,287]" direction="south" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="-3.25">3o15S</geoCoordinate>
<geoCoordinate id="2A95505DFFF99956FC06FE99FBD8EB59" box="[996,1096,260,287]" direction="west" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-50.316666">50o19W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFF99956FB86FE98FB10EB59" bold="true" box="[1124,1152,262,287]" pageId="21" pageNumber="22">51</emphasis>
):
<collectionCode id="29B0AE5FFFF99956FB79FE99FB79EB59" box="[1179,1257,263,287]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="22">MPEG</collectionCode>
24643, female, leg. T.C.S. Avila Pires and J.
<collectingCountry id="37B6760AFFF99956FE3AFEB4FE60EB04" box="[472,496,298,322]" name="Iceland" pageId="21" pageNumber="22">O</collectingCountry>
. Gomes,
<date id="3B1F105AFFF99956FDBBFEB4FC90EB04" box="[601,768,298,322]" pageId="21" pageNumber="22" value="2007-02">February 2007</date>
;
<collectionCode id="29B0AE5FFFF99956FCE9FEB4FCC9EB04" box="[779,857,298,322]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="22">MPEG</collectionCode>
2464450,
<specimenCount id="59A7FD13FFF99956FC3CFEB4FBA4EB04" box="[990,1076,298,322]" pageId="21" pageNumber="22" type="male">6 males</specimenCount>
and a female, leg. J.
<collectingCountry id="37B6760AFFF99956FAC1FEB4FAAAEB04" box="[1315,1338,298,322]" name="Iceland" pageId="21" pageNumber="22">O</collectingCountry>
. Gomes,
<date id="3B1F105AFFF99956FF75FED1FEB3EB21" box="[151,291,335,359]" pageId="21" pageNumber="22" value="2007-03">March 2007</date>
.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF99956FF25FEECFDB0E8AB" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="4F1E369AFFF99956FF25FEECFDB0E8AB" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">
<emphasis id="7DD5EA88FFF99956FF25FEECFED1EBCD" bold="true" box="[199,321,370,395]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
A relatively large
<taxonomicName id="88A14D19FFF99956FDF5FEEDFD00EBCD" box="[535,656,371,395]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="7DD5EA88FFF99956FDF5FEEDFD00EBCD" box="[535,656,371,395]" italics="true" pageId="21" pageNumber="22">Gonatodes</emphasis>
</taxonomicName>
, with maximum SLV of
<quantity id="88599B7FFFF99956FC4CFEECFB86EBCC" box="[942,1046,370,394]" metricMagnitude="-2" metricUnit="m" metricValue="5.57" pageId="21" pageNumber="22" unit="mm" value="55.7">55.7 mm</quantity>
. Scales around midbody 118143. Ventral scales in a longitudinal row 4960. Proximal subdigital lamellae as wide as digits, in total 1823 under fourth finger, 2328 under fourth toe. Three, occasionally four, lateral rows of scales on distal portion of fingers and toes. Tail with midventral scales distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral in contact laterodistally with two scales per side (respectively 112” and 111”). A white suprahumeral bar, bordered by black, both in males and females (but thinner in the latter); from very short (hardly higher than long) to almost reaching the middorsal region. Males with a vermiculated pattern of light and dark (in life respectively drab or orange-yellow, and reddish-brown) spots on head and body dorsally; gular region light with dark oblique streaks or redddish with white oblique streaks (orange with orange-yellow streaks in life).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF99955FF25FD66FE3EEDA2" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="4F1E369AFFF99956FF25FD66FBDFE984" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">
<emphasis id="7DD5EA88FFF99956FF25FD66FEC9E957" bold="true" box="[199,345,760,785]" pageId="21" pageNumber="22">Description.</emphasis>
Maximum SVL in males of
<quantity id="88599B7FFFF99956FD4AFD66FC9EE956" box="[680,782,760,784]" metricMagnitude="-2" metricUnit="m" metricValue="5.57" pageId="21" pageNumber="22" unit="mm" value="55.7">55.7 mm</quantity>
(MPEG 23822), in females of 54.0 mm (MJS 035). Tail round in cross section, tapering toward tip, 1.11.3 (1.15 ± 0.09,
<emphasis id="7DD5EA88FFF99956FC9EFC83FC1FE972" box="[892,911,797,820]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=8) times SLV. Head length 0.210.28 (0.24 ± 0.02,
<emphasis id="7DD5EA88FFF99956FF35FCDFFF7AE91E" box="[215,234,833,856]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times SLV, 1.21.6 (1.40 ± 0.08,
<emphasis id="7DD5EA88FFF99956FD40FCDFFD25E91E" box="[674,693,833,856]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times as long as wide, 1.21.5 (1.40 ± 0.10,
<emphasis id="7DD5EA88FFF99956FB0DFCDFFA92E91E" box="[1263,1282,833,856]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times as wide as high. Snout round, moderately elongate (
<figureCitation id="D79A2A1FFFF99956FD3CFCFDFCBCE93D" box="[734,812,867,891]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="21" pageNumber="22">Fig 11</figureCitation>
A), gently sloping toward top of head. Neck slightly narrower than head and anterior portion of body. Body cylindrical. Limbs well developed, lower arm 0.140.17 (0.15 ± 0.01,
<emphasis id="7DD5EA88FFF99956FECFFC35FED0E984" box="[301,320,939,962]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times SVL, lower leg 0.130.17 (0.15 ± 0.01,
<emphasis id="7DD5EA88FFF99956FC62FC35FC03E984" box="[896,915,939,962]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times SVL.
</paragraph>
<paragraph id="4F1E369AFFF99956FF27FC53FD93EFBF" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">
Rostral convex, 1.83.0 (2.01 ± 0.28,
<emphasis id="7DD5EA88FFF99956FD8FFC51FD10E9A0" box="[621,640,975,998]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times as wide as high; posterior margin with a shallow depression medially and slightly indented by 02 medial postrostrals, with or without a medial cleft extending anteriorly. Postrostrals 2 (12.5%), 3 (83.3%), or 4 (4.2%), laterals ones (supranasals) distinctly larger than medial one(s), when these are present (
<emphasis id="7DD5EA88FFF99956FE53FBA7FE54EE16" box="[433,452,1081,1104]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) (
<figureCitation id="D79A2A1FFFF99956FDE9FBA6FDC6EE16" box="[523,598,1080,1104]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="21" pageNumber="22">Fig 11</figureCitation>
A). Nasal bordered by rostral, first supralabial (only a narrow contact in some specimens), 3 postnasals (N=24), and supranasal; internostril distance 0.14.0.21 (0.17 ± 0.01,
<emphasis id="7DD5EA88FFF99956FAEDFBC3FAB2EE32" box="[1295,1314,1117,1140]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times head width. Supranasal roughly oval, circular or semicircular, 0.91.4 (1.13 ± 0.13;
<emphasis id="7DD5EA88FFF99956FBB3FB1EFBF4EED1" box="[1105,1124,1152,1175]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times as wide as long. Postnasals slightly larger than, or similar in size to, adjacent loreals. Scales on snout convex, hexagonal to round, juxtaposed, relatively uniform in size. Canthus rostralis rounded. Loreal region with scales slightly more elongate than those on snout, largest on row adjacent to supralabials, 1013 (10.2 ± 0.8,
<emphasis id="7DD5EA88FFF99956FBCEFB74FBAFEF47" box="[1068,1087,1258,1281]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) loreals on a line between postnasals and anterior corner of eye. Top and posterior portion of head, as well as supraorbital region, with granular scales. A short supraciliary flap present, anteriorly with 712 (9.5 ± 1.4,
<emphasis id="7DD5EA88FFF99956FBAEFAAFFBCFEF0E" box="[1100,1119,1329,1352]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) enlarged and flattened scales, among which 03 (1.2 ± 1.3;
<emphasis id="7DD5EA88FFF99956FDCCFACBFDD1EF2A" box="[558,577,1365,1388]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) small, conical spines. Pupil round, eye diameter 0.200.27 (0.22 ± 0.02,
<emphasis id="7DD5EA88FFF99956FF75FAE6FF3AEFC9" box="[151,170,1400,1423]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times head length. Scales on temporal region similar to those on top of head. Ear-opening much smaller than eye, oval, posterior to, and at same level of, commissure of mouth. Supralabials 57 (6.2 ± 0.5;
<emphasis id="7DD5EA88FFF99956FB3FFA02FB60EFF5" box="[1245,1264,1436,1459]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24), distinctly enlarged anteriorly and decreasing in size posteriorly, one or two of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFF99956FF27F99BFD1BECED" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">
Mental large, distinctly wider anteriorly than posteriorly, with posterior margin forming a wide angle or a right angle, 1.21.4 (1.28 ± 0.7;
<emphasis id="7DD5EA88FFF99956FE2BF9B4FE4CEC07" box="[457,476,1578,1601]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24) times as wide as long; 2 (95.8%), or 3 (4.2%) postmentals (
<figureCitation id="D79A2A1FFFF99956FB54F9B6FA97EC06" box="[1206,1287,1576,1600]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="21" pageNumber="22">Fig. 11</figureCitation>
B). Scales on chin flat, smooth, polygonal, juxtaposted, larger anteriorly, decreasing in size posteriorly. Infralabials 58 (6.4 ± 0.9;
<emphasis id="7DD5EA88FFF99956FF2BF9EFFF4CECCE" box="[201,220,1649,1672]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=24), distinctly enlarged anteriorly and decreasing in size posteriorly; 13 of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFF99956FF27F928FC8BED53" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">Scales on nape small and granular, becoming slightly larger on sides of neck. Scales on throat anteriorly granular, posteriorly flat, smooth, hexagonal or round, increasing in size toward the posterior region, with a short transitional zone between the anterior and posterior parts.</paragraph>
<paragraph id="4F1E369AFFF99956FF27F8BEFA1CEDAC" blockId="21.[151,1437,151,2026]" pageId="21" pageNumber="22">
Dorsals granular, increasing in size toward the flanks. Ventrals larger than dorsals, roughly hexagonal, flat, smooth, imbricate, in oblique rows; 5260 (56.3 ± 2.4;
<emphasis id="7DD5EA88FFF99956FCFAF8DBFCBBED1A" box="[792,811,1861,1884]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=22) scales along the midventral line between anterior margin of forelimbs and vent; 1824 (21.2 ± 1.3;
<emphasis id="7DD5EA88FFF99956FD0EF8F7FD6FEDC6" box="[748,767,1897,1920]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=22) scales in a transverse line at midbody, with a short transitional zone between ventrals and scales on flanks. Scales around midbody 118143 (129.9 ± 6.4;
<emphasis id="7DD5EA88FFF99956FAAFF812FAF0EDE5" box="[1357,1376,1932,1955]" italics="true" pageId="21" pageNumber="22">N</emphasis>
=22). Scales on preanal plate similar to ventrals, except for those bordering vent, which are very small. Escutcheon present in males on posterior portion of belly and on 45 rows (body-knee direction) on ventral surface of thighs.
</paragraph>
<paragraph id="4F1E369AFFFA9955FF27FF09FD98EB78" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">Scales on anterodorsal surface of forelimbs flat, smooth, roundish or rhomboidal, imbricate, largest close to the wrist; on posterodorsal and ventral surface of forelimbs convex, smooth, rhomboidal, juxtaposed, relatively small. Scales on anterodorsal surface of thighs and ventral surface of hind limbs flat, smooth, roundish or rhomboidal, imbricate and relatively larger; on posterodorsal surface of thighs and dorsal surface of lower legs smaller, convex, smooth, rhomboidal, juxtaposed.</paragraph>
<paragraph id="4F1E369AFFFA9955FF27FED7FBBBEBA9" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
Lamellae under second (
<emphasis id="7DD5EA88FFFA9955FE3DFED7FE65EB27" bold="true" box="[479,501,329,353]" pageId="22" pageNumber="23">II</emphasis>
) through fourth (
<emphasis id="7DD5EA88FFFA9955FD5CFED7FD4DEB27" bold="true" box="[702,733,329,353]" pageId="22" pageNumber="23">IV</emphasis>
) fingers (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFFA9955FB00FED7FB68EB27" bold="true" box="[1250,1272,329,353]" pageId="22" pageNumber="23">II</emphasis>
: 1519 (46),
<emphasis id="7DD5EA88FFFA9955FF75FEF3FF28EBC3" bold="true" box="[151,184,365,389]" pageId="22" pageNumber="23">III</emphasis>
: 1822 (56), and
<emphasis id="7DD5EA88FFFA9955FE6FFEF3FE3AEBC3" bold="true" box="[397,426,365,389]" pageId="22" pageNumber="23">IV</emphasis>
: 1823 (68) (
<figureCitation id="D79A2A1FFFFA9955FDB3FEF2FD0AEBC2" box="[593,666,364,388]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="22" pageNumber="23">Fig 11</figureCitation>
C). Lamellae under second through fourth toes (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFFA9955FED1FE0EFED9EBEE" bold="true" box="[307,329,400,424]" pageId="22" pageNumber="23">II</emphasis>
: 1519 (56),
<emphasis id="7DD5EA88FFFA9955FE16FE0EFD85EBEE" bold="true" box="[500,533,400,424]" pageId="22" pageNumber="23">III</emphasis>
: 1823 (57), and
<emphasis id="7DD5EA88FFFA9955FD12FE0EFC9DEBEE" bold="true" box="[752,781,400,424]" pageId="22" pageNumber="23">IV</emphasis>
: 2328 (912) (
<figureCitation id="D79A2A1FFFFA9955FC25FE0EFB87EBEE" box="[967,1047,400,424]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
D). Claws exposed, non-retractile, between two basal scales. Fingers and toes with three, occasionally four, lateral scales distally, between the fourth (counted from the claw towards the hand) subdigital lamella, and the dorsal scale.
</paragraph>
<paragraph id="4F1E369AFFFA9955FF27FE64FC53E882" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
Scales on tail dorsally and laterally relatively small, rhomboidal, flat, smooth, imbricate. On ventral surface of tail scales smooth, flat, imbricate, increasing in size toward midventral line; midventral scales, except close to the base of the tail, distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral scale in contact laterodistally with two scales per side respectively 112” and 111” in the codification of
<bibRefCitation id="2B304B6BFFFA9955FDE5FD32FCD3E882" author="Avila-Pires" box="[519,835,684,708]" pageId="22" pageNumber="23" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">
Avila-Pires (1995:
<figureCitation id="D79A2A1FFFFA9955FD3EFD32FCABE882" box="[732,827,684,708]" captionStart="FIGURE 2" captionStartId="6.[151,250,1440,1463]" captionTargetBox="[190,1402,193,1415]" captionTargetId="figure@6.[185,1402,193,1419]" captionTargetPageId="6" captionText="FIGURE 2. Bivariate scattergrams from stepwise discriminant function analysis using scale counts and measurements. Red = Pattern A. Cyan-green = Pattern B. Lime-green = Pattern C. Yellow = Pattern D. Blue = Pattern E." httpUri="https://zenodo.org/record/277515/files/figure.png" pageId="22" pageNumber="23">Figure 2</figureCitation>
)
</bibRefCitation>
(
<figureCitation id="D79A2A1FFFFA9955FCB0FD32FC32E882" box="[850,930,684,708]" captionStart="FIGURE 11" captionStartId="23.[151,250,1960,1983]" captionTargetBox="[183,1396,220,1959]" captionTargetId="figure@23.[182,1426,161,1990]" captionTargetPageId="23" captionText="FIGURE 11. Gonatodes nascimentoi sp. nov. (holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
E).
</paragraph>
<paragraph id="4F1E369AFFFA9955FF25FD4EFA99E94D" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
<emphasis id="7DD5EA88FFFA9955FF25FD4EFD92E8AE" bold="true" box="[199,514,719,744]" pageId="22" pageNumber="23">
Measurements of
<typeStatus id="901A8838FFFA9955FE7EFD4EFD92E8AE" box="[412,514,720,744]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
</emphasis>
(in millimeters): SVL 48.7, TL 54.0, HL 11.7, HW 7.9, HD 6.8, ED 2.6,
<collectingCountry id="37B6760AFFFA9955FADBFD4EFAFBE8A1" box="[1337,1387,720,743]" name="India" pageId="22" pageNumber="23">IND</collectingCountry>
2.0, SSL 0.8, SSW 1.12, RSL 1.3, RSW 2.5, MSL 2.6, MSW 3.2, LAL 8.1, LLL 7.8, HSH 5.34, HSL 2.6.
</paragraph>
<paragraph id="4F1E369AFFFA9955FF25FC88FC5CE933" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
<emphasis id="7DD5EA88FFFA9955FF25FC88FE7DE968" bold="true" box="[199,493,790,815]" pageId="22" pageNumber="23">
Scale counts of
<typeStatus id="901A8838FFFA9955FE9FFC88FE78E968" box="[381,488,790,814]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
:
</emphasis>
SAM 127 scales, VLR 55, VTR 21, SL 6,
<collectingCountry id="37B6760AFFFA9955FC2BFC89FC74E968" box="[969,996,791,814]" name="Iceland" pageId="22" pageNumber="23">IL</collectingCountry>
5, PR 3, SP 1, PN 3, LS 12, PM 2, SCS 8, SSC 3, PL2F 5, DL2F 12, LS2F 3, PL3F 4, DL3F 14, LS3F 4, PL4F 6, DL4F 13, LS4F 4, PL2T 5, DL2T 12, LS2T 4, PL3T 5, DL3T 15, LS3T 3, PL4T 10, DL4T 13, LS4T 3, RSE 4.
</paragraph>
<paragraph id="4F1E369AFFFA9955FF25FC1EFE99EEF3" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
<emphasis id="7DD5EA88FFFA9955FF25FC1EFE72E9DF" bold="true" box="[199,482,896,921]" pageId="22" pageNumber="23">Color in preservative.</emphasis>
In males (
<figureCitation id="D79A2A1FFFFA9955FD98FC1EFD55E9DE" box="[634,709,896,920]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="22" pageNumber="23">Fig. 1</figureCitation>
D), dorsal surface of head with reddish-brown and beige vermiculations. Sides of head, in some specimens, with oblique and/or longitudinal, beige and brown bands. Back and flanks, base of tail and hind limbs with relatively large, beige color and dark-brown vermiculations. A conspicuous, white suprahumeral vertical bar with black margin, which varies from only slightly to distinctly higher than long (see under “Remarks” below). Ventral surface of head and gular region beige with black streaks, in some specimens dark-brown with beige streaks, in contact or not at midvental line; chest beige; belly darkbrown; underside of limbs and base of tail brown. Dorsal surface of tail brown, underside dark-brown or black; a white band may be present distally, forming a complete ring around the tail. Escutcheon area (belly and thighs) light gray.
</paragraph>
<paragraph id="4F1E369AFFFA9955FF25FB5FFD33EFCF" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">In females, dorsal surface of head and limbs with brown and beige irregular spots. Back gray with dorsolateral pairs of black and white spots; flanks gray with black and brown irregular spots. Suprahumeral bar conspicuous, white with black margins, thinner than that of males; in some cases they almost reach the middorsal region. Ventral surface of head and gular region white with black oblique streaks, in contact or not at midventral line; belly and underside of limbs light-gray. Tail gray dorsally and white ventrally; original tail distally with white and black bands that form complete rings around the tail.</paragraph>
<paragraph id="4F1E369AFFFA9955FF25FA0AFDBDED10" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
<emphasis id="7DD5EA88FFFA9955FF25FA0AFECCEFEB" bold="true" box="[199,348,1428,1453]" pageId="22" pageNumber="23">Color in life</emphasis>
(Photos of MPEG 23822, 2382425, 2559699; field notes of MPEG 24643): In males (
<figureCitation id="D79A2A1FFFFA9955FA8CFA0AFF23EF96" captionStart="FIGURE 12" captionStartId="24.[151,250,1063,1086]" captionTargetBox="[151,1436,193,1041]" captionTargetId="figure@24.[151,1436,193,1042]" captionTargetPageId="24" captionText="FIGURE 12. Gonatodes nascimentoi sp. nov. from Anapu, Rio Xingu, Pará, Brazil. Adult male (paratype, MPEG 25598, SVL = 46.7 mm) in dorsal (A) and ventral (B) views; adult female (paratype, MPEG 25599, SVL = 39.7 mm) in dorsal (C) and ventral (D) views. (Photos by Pedro L. Peloso)." httpUri="https://zenodo.org/record/277525/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
AB), head dorsally and laterally with orange and orange-yellow, or orange and drab, vermiculations. On back vermiculations may be smaller than, or similar in size to, those on head and the lighter spots are frequently bordered by black (occasionally also on head). Suprahumeral bar or ocellus white with black margins. Flanks with smaller, bluish-white and black vermiculations. Head ventrally and gular region orange with orange-yellow streaks, chest orange. Remaining ventral region plumbeous, lighter on escutcheon areas (belly and thighs). Tail dark gray to black all around, except near the base, where it is similar to the back; distal portion with white spots. Color in life of MPEG 24643, a male, was described based on the color guide of
<bibRefCitation id="2B304B6BFFFA9955FBABF913FB60ECE3" author="Smithe" box="[1097,1264,1677,1701]" pageId="22" pageNumber="23" refString="Smithe, F. B. (1975) Naturalist's Color Guide. The American Museum of Natural History, New York. 8 pp." type="book" year="1975">Smithe (1975)</bibRefCitation>
: Head dorsally “buff” (24) and “mahogany red” (132B); body “brick red” (132A) with “chamois” (123D) vermiculation; a white bar with black margins; lower flanks “blackish neutral gray” (82) with “pearl gray” (81) spots. Ventrally, head and chest (until forearm) “buff” (24) with convergent “scarlet” (14) streaks; otherwise ventral region and tail mainly “blackish neutral gray” (82), midventrally medium (84) to light (85) neutral gray; tail dorsally with a darker and lighter pattern (TCSAP, field notes).
</paragraph>
<paragraph id="4F1E369AFFFA9955FF25F8FFFE3EEDA2" blockId="22.[151,1437,151,2020]" pageId="22" pageNumber="23">
Females (
<figureCitation id="D79A2A1FFFFA9955FED6F8FFFE14ED3F" box="[308,388,1889,1913]" captionStart="FIGURE 12" captionStartId="24.[151,250,1063,1086]" captionTargetBox="[151,1436,193,1041]" captionTargetId="figure@24.[151,1436,193,1042]" captionTargetPageId="24" captionText="FIGURE 12. Gonatodes nascimentoi sp. nov. from Anapu, Rio Xingu, Pará, Brazil. Adult male (paratype, MPEG 25598, SVL = 46.7 mm) in dorsal (A) and ventral (B) views; adult female (paratype, MPEG 25599, SVL = 39.7 mm) in dorsal (C) and ventral (D) views. (Photos by Pedro L. Peloso)." httpUri="https://zenodo.org/record/277525/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
CD) dorsally brown or grayish-brown, with a paired series of dark brown to black irregular, transversely elongate, spots. Suprahumeral bar white with black margins, thinner than those in males. Head ventrally and gular region white with black streaks, chest light-gray. Tail gray with black spots all around; posterior portion with white spots.
</paragraph>
</subSubSection>
<caption id="1BDE6612FFFB9954FF75F836FC67ED9B" httpUri="https://zenodo.org/record/277524/files/figure.png" pageId="23" pageNumber="24" targetBox="[183,1396,220,1959]" targetPageId="23">
<paragraph id="4F1E369AFFFB9954FF75F836FC67ED9B" blockId="23.[151,1435,1960,2014]" pageId="23" pageNumber="24">
<emphasis id="7DD5EA88FFFB9954FF75F836FEB0EDF8" bold="true" box="[151,288,1960,1983]" pageId="23" pageNumber="24">FIGURE 11.</emphasis>
<taxonomicName id="88A14D19FFFB9954FECBF836FD8AEDF9" box="[297,538,1960,1983]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="nascimentoi" status="sp. nov.">
<emphasis id="7DD5EA88FFFB9954FECBF836FD8AEDF9" box="[297,538,1960,1983]" italics="true" pageId="23" pageNumber="24">Gonatodes nascimentoi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFFB9954FDC1F837FDE6EDF8" bold="true" box="[547,630,1961,1982]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="66E657F3FFFB9954FDC1F837FDE6EDF8" box="[547,630,1961,1982]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, MPEG 25596). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="1BDE6612FFF4995BFF75FBB9FDE8EE3C" httpUri="https://zenodo.org/record/277525/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1436,193,1041]" targetPageId="24">
<paragraph id="4F1E369AFFF4995BFF75FBB9FDE8EE3C" blockId="24.[151,1436,1063,1147]" pageId="24" pageNumber="25">
<emphasis id="7DD5EA88FFF4995BFF75FBB9FE8FEE7A" bold="true" box="[151,287,1063,1086]" pageId="24" pageNumber="25">FIGURE 12.</emphasis>
<taxonomicName id="88A14D19FFF4995BFEC7FBB9FD83EE7B" box="[293,531,1063,1085]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="nascimentoi" status="sp. nov.">
<emphasis id="7DD5EA88FFF4995BFEC7FBB9FD83EE7B" box="[293,531,1063,1085]" italics="true" pageId="24" pageNumber="25">Gonatodes nascimentoi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFF4995BFDFBFBB6FDFAEE7B" bold="true" box="[537,618,1064,1085]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="66E657F3FFF4995BFDFBFBB6FDFAEE7B" box="[537,618,1064,1085]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Anapu, Rio Xingu, Pará, Brazil. Adult male (paratype, MPEG 25598, SVL = 46.7 mm) in dorsal (A) and ventral (B) views; adult female (paratype, MPEG 25599, SVL = 39.7 mm) in dorsal (C) and ventral (D) views. (Photos by Pedro L. Peloso).
</paragraph>
</caption>
<subSubSection id="07BB6511FFF4995BFF25FB38FD21EF41" pageId="24" pageNumber="25" type="distribution">
<paragraph id="4F1E369AFFF4995BFF25FB38FD21EF41" blockId="24.[151,1436,1190,1827]" pageId="24" pageNumber="25">
<emphasis id="7DD5EA88FFF4995BFF25FB38FEF0EEF9" bold="true" box="[199,352,1190,1215]" pageId="24" pageNumber="25">Distribution.</emphasis>
Eastern Brazilian Amazonia, in the states of Pará and Amapá. Until now it is only known south of the Amazon river in the interfluvium XinguTocantins, Pará state, and north of the Amazon river east of Paru river, in the states of Pará and Amapá (
<figureCitation id="D79A2A1FFFF4995BFDA8FB71FD1DEF41" box="[586,653,1263,1287]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="24" pageNumber="25">Fig. 3</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF4995BFF25FA8DFEF8EF34" pageId="24" pageNumber="25" type="etymology">
<paragraph id="4F1E369AFFF4995BFF25FA8DFEF8EF34" blockId="24.[151,1436,1190,1827]" pageId="24" pageNumber="25">
<emphasis id="7DD5EA88FFF4995BFF25FA8DFEDEEF6D" bold="true" box="[199,334,1299,1323]" pageId="24" pageNumber="25">Etymology.</emphasis>
The species is named after Francisco Paiva do Nascimento, who worked for many years in Museu Paraense Emílio Goeldi and contributed to our knowledge of Amazonian reptiles, most of all about the snakes from eastern Amazonia.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF4995AFF25FAE1FECBEA94" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="4F1E369AFFF4995BFF25FAE1FC28EC29" blockId="24.[151,1436,1190,1827]" pageId="24" pageNumber="25">
<emphasis id="7DD5EA88FFF4995BFF25FAE1FEABEFD1" bold="true" box="[199,315,1407,1431]" pageId="24" pageNumber="25">Remarks.</emphasis>
The suprahumeral spot varies among populations and, to a smaller degree, within populations. In the only specimen from Amapá examined, the spot has the shape of a short bar, almost an ocellus. Specimens from Pará, north of the Amazon River, show a vertical bar reaching dorsally at least the dorsolateral region, in some cases almost the middorsal region. South of the Amazon, all specimens from Monte Verde present a short vertical bar (not reaching the dorsolateral region, but always with dorso-ventral axis longer than antero-posterior axis), while in the specimens from Anapu the vertical bar is similar to those from northern Pará (although one specimen presents a vertical bar and a small ocellus above the bar, on both sides).
</paragraph>
<paragraph id="4F1E369AFFF4995BFF36F9E4FC92ED65" blockId="24.[151,1436,1190,1827]" pageId="24" pageNumber="25">
Specimens were collected by active search or in pitfall traps, in primary or little-disturbed forest. Six specimens were found on the buttresses or trunks of live trees, between
<quantity id="88599B7FFFF4995BFC57F901FBB3ECF1" box="[949,1059,1695,1719]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="24" pageNumber="25" unit="cm" value="32.5" valueMax="50.0" valueMin="15.0">1550 cm</quantity>
above ground; one was inside the loose bark of a dead tree,
<quantity id="88599B7FFFF4995BFE5AF95CFD9BEC9C" box="[440,523,1730,1754]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="cm" value="120.0">120 cm</quantity>
above ground; a male and a female were on a fallen trunk; and at the Xingu river a pair was found on a rock,
<quantity id="88599B7FFFF4995BFE34F979FD8CECB9" box="[470,540,1767,1791]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="cm" value="50.0">50 cm</quantity>
above ground. At this locality specimens hid inside rock crevices or tree trunk holes when disturbed. Weights vary between
<quantity id="88599B7FFFF4995BFD70F894FD6EED65" box="[658,766,1802,1827]" metricMagnitude="-3" metricUnit="kg" metricValue="2.65" metricValueMax="5.1" metricValueMin="0.2" pageId="24" pageNumber="25" unit="g" value="2.65" valueMax="5.1" valueMin="0.2">0.25.1 g</quantity>
.
</paragraph>
<paragraph id="4F1E369AFFF5995AFF75FF09FECBEA94" blockId="25.[151,464,151,210]" pageId="25" pageNumber="26">
<taxonomicName id="88A14D19FFF5995AFF75FF09FEFFEAF7" box="[151,367,151,177]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="riveroi" status="sp. nov.">
<emphasis id="7DD5EA88FFF5995AFF75FF09FEFFEAF7" bold="true" box="[151,367,151,177]" italics="true" pageId="25" pageNumber="26">Gonatodes riveroi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFF5995AFE94FF07FE40EAF6" bold="true" box="[374,464,153,177]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="66E657F3FFF5995AFE94FF07FE40EAF6" box="[374,464,153,177]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D79A2A1FFFF5995AFF7DFF24FF79EA94" box="[159,233,186,210]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="25" pageNumber="26">Figs. 1</figureCitation>
E, 13, 14)
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF5995AFF75FE9CFAE5EB7E" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="4F1E369AFFF5995AFF75FE9CFAE5EB7E" blockId="25.[151,1436,258,312]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="6FB111B4FFF5995AFF75FE9CFAE5EB7E" pageId="25" pageNumber="26">
<taxonomicName id="88A14D19FFF5995AFF75FE9CFDF4EB5E" authority="Vanzolini, 1955: 126" authorityName="Vanzolini" authorityPageNumber="126" authorityYear="1955" box="[151,612,258,280]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFF5995AFF75FE9CFE15EB5E" box="[151,389,258,280]" italics="true" pageId="25" pageNumber="26">Gonatodes concinnatus</emphasis>
: Vanzolini, 1955: 126
</taxonomicName>
(part); 1968: 26 (part);
<treatmentCitation id="CE00108BFFF5995AFCB6FE9CFB89EB5E" author="Mechler" box="[852,1049,258,280]" page="331" pageId="25" pageNumber="26" year="1968">
<bibRefCitation id="2B304B6BFFF5995AFCB6FE9CFC79EB5E" author="Mechler" box="[852,1001,258,280]" pageId="25" pageNumber="26" refString="Mechler, B. (1968) Les geckonides de la Colombie. Revue Suisse de Zoologie 75, 305 - 371." type="journal article" year="1968">Mechler, 1968</bibRefCitation>
: 331
</treatmentCitation>
(part);
<bibRefCitation id="2B304B6BFFF5995AFB86FE9DFA0CEB5E" author="Peters" box="[1124,1436,259,280]" pageId="25" pageNumber="26" refString="Peters, J. A. &amp; Donoso-Barros, R. (1970) Catalogue of the Neotropical Squamata: part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293." type="journal article" year="1970">Peters &amp; Donoso-Barros, 1970</bibRefCitation>
(part);
<treatmentCitation id="CE00108BFFF5995AFEEFFEBFFD97EB71" author="Rivero-Blanco" box="[269,519,289,311]" page="94" pageId="25" pageNumber="26" year="1979">
<bibRefCitation id="2B304B6BFFF5995AFEEFFEBFFE74EB71" author="Rivero-Blanco" box="[269,484,289,311]" pageId="25" pageNumber="26" refString="Rivero-Blanco, C. (1979) The Neotropical lizard genus Gonatodes Fitzinger (Sauria: Sphaerodactylinae). PhD Thesis, Texas A &amp; M University. 233 pp." type="book" year="1979">Rivero-Blanco, 1979</bibRefCitation>
: 94
</treatmentCitation>
(part);
<treatmentCitation id="CE00108BFFF5995AFDB6FEBFFC6AEB71" author="Sanchez" box="[596,1018,289,311]" page="317" pageId="25" pageNumber="26" year="1995">
<bibRefCitation id="2B304B6BFFF5995AFDB6FEBFFC57EB71" author="Sanchez" box="[596,967,289,311]" pageId="25" pageNumber="26" refString="Sanchez, H.; Castano, O. &amp; Cardenas, G. (1995) Diversidad de los reptiles en Colombia. In: Rangel-Ch, J. O. (Ed.), Colombia Diversidad biotica I. Guadalupe, Santafe de Bogota, Colombia, pp. 277 - 325." type="book chapter" year="1995">Sanchez, Castaño &amp; Cardenas, 1995</bibRefCitation>
: 317
</treatmentCitation>
;
<treatmentCitation id="CE00108BFFF5995AFBE6FEBFFABFEB71" author="Bartlett" box="[1028,1327,289,311]" page="158" pageId="25" pageNumber="26" year="2003">
<bibRefCitation id="2B304B6BFFF5995AFBE6FEBFFB6FEB71" author="Bartlett" box="[1028,1279,289,311]" pageId="25" pageNumber="26" refString="Bartlett, R. D. &amp; Bartlett, P. (2003) Reptiles and Amphibians of the Amazon: An Ecotourist's Guide. University Press of Florida, Gainesville, 291 pp." type="book" year="2003">Bartlett &amp; Bartlett, 2003</bibRefCitation>
: 158
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF5995AFF75FEC1FBB9EED7" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="4F1E369AFFF5995AFF75FEC1FD58EB87" blockId="25.[151,1437,350,2033]" pageId="25" pageNumber="26">
<emphasis id="7DD5EA88FFF5995AFF75FEC1FE94EB31" bold="true" box="[151,260,351,375]" pageId="25" pageNumber="26">
<typeStatus id="901A8838FFF5995AFF75FEC1FE94EB31" box="[151,260,351,375]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
(number in bold between parentheses refer to localities in
<figureCitation id="D79A2A1FFFF5995AFC35FEC0FBB1EB30" box="[983,1057,350,374]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="25" pageNumber="26">Fig. 3</figureCitation>
):
<collectionCode id="29B0AE5FFFF5995AFBDCFEC0FB0BEB30" box="[1086,1179,350,374]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
49153 (field number FMEDEM 1465), adult male, from Villavicencio, State of Meta,
<collectingCountry id="37B6760AFFF5995AFC96FE1BFC72EBDB" box="[884,994,389,413]" name="Colombia" pageId="25" pageNumber="26">Colombia</collectingCountry>
(~
<geoCoordinate id="2A95505DFFF5995AFBE3FE1BFBDDEBDB" box="[1025,1101,386,413]" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-4.15">4o09S</geoCoordinate>
and
<geoCoordinate id="2A95505DFFF5995AFB66FE1BFB79EBDB" box="[1156,1257,386,413]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-73.61667">73o37W</geoCoordinate>
) (
<emphasis id="7DD5EA88FFF5995AFAE1FE1BFA8FEBD8" bold="true" box="[1283,1311,389,414]" pageId="25" pageNumber="26">37</emphasis>
), collected by W. W. Lamar and F. Medem,
<date id="3B1F105AFFF5995AFDE6FE37FD53EB87" box="[516,707,425,449]" pageId="25" pageNumber="26" value="1977-10-22">22 October 1977</date>
.
</paragraph>
<paragraph id="4F1E369AFFF5995AFF25FE53FBB9EED7" blockId="25.[151,1437,350,2033]" pageId="25" pageNumber="26">
<emphasis id="7DD5EA88FFF5995AFF25FE53FEAFEBA3" bold="true" box="[199,319,461,485]" pageId="25" pageNumber="26">
<typeStatus id="901A8838FFF5995AFF25FE53FEAFEBA3" box="[199,319,461,485]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(numbers in bold between parentheses refer to localities in
<figureCitation id="D79A2A1FFFF5995AFBEFFE53FBC5EBA3" box="[1037,1109,461,485]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="25" pageNumber="26">Fig. 3</figureCitation>
):
<emphasis id="7DD5EA88FFF5995AFB92FE53FA86EBA0" bold="true" box="[1136,1302,461,486]" pageId="25" pageNumber="26">
<collectingCountry id="37B6760AFFF5995AFB92FE53FA81EBA0" box="[1136,1297,461,486]" name="Colombia" pageId="25" pageNumber="26">COLOMBIA</collectingCountry>
.
</emphasis>
BOYACÁ. Garagoa (
<emphasis id="7DD5EA88FFF5995AFEEBFE6FFEB5E84C" bold="true" box="[265,293,497,522]" pageId="25" pageNumber="26">34</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFEDDFE6FFEE5E84F" LSID="urn:lsid:biocol.org:col:34700" box="[319,373,497,521]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="25" pageNumber="26">CAS</collectionCode>
71238,
<collectionCode id="29B0AE5FFFF5995AFE34FE6FFDB7E84F" LSID="urn:lsid:biocol.org:col:34871" box="[470,551,497,521]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
84968,
<collectionCode id="29B0AE5FFFF5995AFD65FE6FFD48E84F" LSID="urn:lsid:biocol.org:col:34871" box="[647,728,497,521]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
9249092, a male and
<specimenCount id="59A7FD13FFF5995AFC08FE6FFBCCE84F" box="[1002,1116,497,521]" pageId="25" pageNumber="26" type="female">4 females</specimenCount>
, leg. Nicéforo Maria, 1931. BOYACÁCUNDINAMARCA
<collectionCode id="29B0AE5FFFF5995AFDECFD88FDCBE868" box="[526,603,534,558]" pageId="25" pageNumber="26">META</collectionCode>
border. Guaicáramo (according to
<bibRefCitation id="2B304B6BFFF5995AFBE6FD88FB5EE86B" author="Paynter" box="[1028,1230,533,558]" pageId="25" pageNumber="26" refString="Paynter Jr., R. A. (1997) Ornithological gazetteer of Colombia. Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, 537 pp." type="book" year="1997">
Paynter Jr.,
<quantity id="88599B7FFFF5995AFB74FD8BFB5EE86B" box="[1174,1230,533,557]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="25" pageNumber="26" unit="in" value="1997.0">1997</quantity>
</bibRefCitation>
in the hills near confluence of Rio Guavio with Rio Upía, at eastern base of the eastern Andes) (
<emphasis id="7DD5EA88FFF5995AFBD3FDA7FBDDE814" bold="true" box="[1073,1101,569,594]" pageId="25" pageNumber="26">35</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFB85FDA7FB28E817" LSID="urn:lsid:biocol.org:col:34871" box="[1127,1208,569,593]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
84972, female, leg. Nicéforo Maria, 1931.
<collectionCode id="29B0AE5FFFF5995AFE75FDC0FE74E830" box="[407,484,606,630]" pageId="25" pageNumber="26">META</collectionCode>
. San Juan de Arama (
<emphasis id="7DD5EA88FFF5995AFD3AFDC3FD64E830" bold="true" box="[728,756,605,630]" pageId="25" pageNumber="26">36</emphasis>
): KU 19263536, a male and a female, leg. M.
<collectingCountry id="37B6760AFFF5995AFAE5FDC3FAD0E833" box="[1287,1344,605,629]" name="China" pageId="25" pageNumber="26">Chin</collectingCountry>
,
<date id="3B1F105AFFF5995AFAA9FDC0FF43E8DF" pageId="25" pageNumber="26" value="1967-08">August 1967</date>
. Type-locality (
<emphasis id="7DD5EA88FFF5995AFE60FD1FFE0EE8DC" bold="true" box="[386,414,641,666]" pageId="25" pageNumber="26">37</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFE57FD1FFE62E8DF" LSID="urn:lsid:biocol.org:col:33791" box="[437,498,641,665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
13279697, a juvenile and a female,
<date id="3B1F105AFFF5995AFC77FD1FFB96E8DF" box="[917,1030,641,665]" pageId="25" pageNumber="26" value="1972-07">July 1972</date>
;
<collectionCode id="29B0AE5FFFF5995AFBF3FD1CFBF7E8DC" LSID="urn:lsid:biocol.org:col:34925" box="[1041,1127,642,666]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
3528087,
<collectionCode id="29B0AE5FFFF5995AFB09FD1CFAD1E8DC" LSID="urn:lsid:biocol.org:col:34925" box="[1259,1345,642,666]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
35290 91,
<specimenCount id="59A7FD13FFF5995AFF23FD3BFE87E8FB" box="[193,279,677,701]" pageId="25" pageNumber="26" type="male">5 males</specimenCount>
and
<specimenCount id="59A7FD13FFF5995AFEACFD3BFE2DE8FB" box="[334,445,677,701]" pageId="25" pageNumber="26" type="female">5 females</specimenCount>
, leg. Nicéforo Maria,
<date id="3B1F105AFFF5995AFD51FD38FCDDE8FB" box="[691,845,677,702]" pageId="25" pageNumber="26" value="1927-01">January 1927</date>
; KU
<date id="3B1F105AFFF5995AFC6AFD3BFC4DE8FB" box="[904,989,677,701]" pageId="25" pageNumber="26" value="79-11-05">110579</date>
80, a juvenile and a male, collector not specified,
<date id="3B1F105AFFF5995AFEECFD57FE13E8A7" box="[270,387,713,737]" pageId="25" pageNumber="26" value="1967-07">July 1967</date>
; KU 15193335,
<specimenCount id="59A7FD13FFF5995AFDB4FD57FD22E8A7" box="[598,690,713,737]" pageId="25" pageNumber="26" type="male">3 males</specimenCount>
, collector not specified,
<date id="3B1F105AFFF5995AFC36FD54FBDDE8A7" box="[980,1101,713,738]" pageId="25" pageNumber="26" value="1967-05">May 1967</date>
;
<collectionCode id="29B0AE5FFFF5995AFBBFFD57FB09E8A7" LSID="urn:lsid:biocol.org:col:33791" box="[1117,1177,713,737]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
19213, male,
<date id="3B1F105AFFF5995AFAA7FD54FF42E943" pageId="25" pageNumber="26" value="1924-01">January 1924</date>
;
<collectionCode id="29B0AE5FFFF5995AFF3CFD73FE8AE943" LSID="urn:lsid:biocol.org:col:33791" box="[222,282,749,773]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
7738889, a male and a female, collector and date not specified;
<collectionCode id="29B0AE5FFFF5995AFC19FD73FBA8E943" LSID="urn:lsid:biocol.org:col:33791" box="[1019,1080,749,773]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
11000914, a juvenile,
<specimenCount id="59A7FD13FFF5995AFAA7FD73FA0BE943" box="[1349,1435,749,773]" pageId="25" pageNumber="26" type="male">3 males</specimenCount>
and
<specimenCount id="59A7FD13FFF5995AFF29FC8FFEAEE96F" box="[203,318,785,809]" pageId="25" pageNumber="26" type="female">2 females</specimenCount>
,
<date id="3B1F105AFFF5995AFEADFC8CFE59E96F" box="[335,457,785,809]" pageId="25" pageNumber="26" value="1968-06">June 1968</date>
;
<collectionCode id="29B0AE5FFFF5995AFE3BFC8FFD86E96F" LSID="urn:lsid:biocol.org:col:33791" box="[473,534,785,809]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
<date id="3B1F105AFFF5995AFDC3FC8FFDE5E96F" box="[545,629,785,809]" pageId="25" pageNumber="26" value="41-11-07">110741</date>
42,
<specimenCount id="59A7FD13FFF5995AFD53FC8FFC9FE96F" box="[689,783,785,809]" pageId="25" pageNumber="26" type="male">2 males</specimenCount>
, leg. G. Gorman,
<date id="3B1F105AFFF5995AFC04FC8CFBF0E96F" box="[998,1120,785,809]" pageId="25" pageNumber="26" value="1968-06">June 1968</date>
;
<collectionCode id="29B0AE5FFFF5995AFB92FC8FFB5DE96F" box="[1136,1229,785,809]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
<date id="3B1F105AFFF5995AFB3BFC8FFAABE96F" box="[1241,1339,785,809]" pageId="25" pageNumber="26" value="2145" valueMax="2146">214546</date>
(former
<collectionCode id="29B0AE5FFFF5995AFF75FCA8FF7DE908" LSID="urn:lsid:biocol.org:col:34925" box="[151,237,822,846]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
3529293),
<specimenCount id="59A7FD13FFF5995AFE99FCABFE79E90B" box="[379,489,821,845]" pageId="25" pageNumber="26" type="female">2 females</specimenCount>
, leg. Nicéforo Maria, no date;
<collectionCode id="29B0AE5FFFF5995AFCA2FCABFC0DE90B" box="[832,925,821,845]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
49152, female, leg. M. Lugor and F. Medem,
<date id="3B1F105AFFF5995AFF75FCC7FEA4E937" box="[151,308,857,881]" pageId="25" pageNumber="26" value="1977-10">October 1977</date>
;
<collectionCode id="29B0AE5FFFF5995AFEA3FCC7FE0EE937" box="[321,414,857,881]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
49154, female, leg. W. W. Lamar and F. Medem,
<date id="3B1F105AFFF5995AFC34FCC7FBE3E937" box="[982,1139,857,881]" pageId="25" pageNumber="26" value="1977-10">October 1977</date>
;
<collectionCode id="29B0AE5FFFF5995AFB62FCC7FB41E937" LSID="urn:lsid:biocol.org:col:34871" box="[1152,1233,857,881]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
84969, male, leg. Nicéforo Maria, 1931. Villavicencio, Finca “El Buque” (
<emphasis id="7DD5EA88FFF5995AFCD0FCE3FCDEE9D0" bold="true" box="[818,846,893,918]" pageId="25" pageNumber="26">37</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFC88FCE3FC57E9D3" box="[874,967,893,917]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
4915562,
<specimenCount id="59A7FD13FFF5995AFBB4FCE3FB20E9D3" box="[1110,1200,893,917]" pageId="25" pageNumber="26" type="male">3 males</specimenCount>
and
<specimenCount id="59A7FD13FFF5995AFB12FCE3FAF3E9D3" box="[1264,1379,893,917]" pageId="25" pageNumber="26" type="female">5 females</specimenCount>
, leg. E.Thierry and F. Medem,
<date id="3B1F105AFFF5995AFE3AFC3FFDECE9FF" box="[472,636,929,953]" pageId="25" pageNumber="26" value="1977-10">October 1977</date>
. Villavicencio, Instituto 'Roberto Franco' (
<emphasis id="7DD5EA88FFF5995AFB62FC3FFB0CE9FC" bold="true" box="[1152,1180,929,954]" pageId="25" pageNumber="26">37</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFB5EFC3FFB69E9FF" LSID="urn:lsid:biocol.org:col:33791" box="[1212,1273,929,953]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
7739197,
<specimenCount id="59A7FD13FFF5995AFA6CFC3FFF6FE99B" pageId="25" pageNumber="26" type="juvenile">2 juveniles</specimenCount>
,
<specimenCount id="59A7FD13FFF5995AFEEEFC5BFEF4E99B" box="[268,356,965,989]" pageId="25" pageNumber="26" type="male">2 males</specimenCount>
and
<specimenCount id="59A7FD13FFF5995AFE7CFC5BFD9FE99B" box="[414,527,965,989]" pageId="25" pageNumber="26" type="female">3 females</specimenCount>
, collector not specified,
<date id="3B1F105AFFF5995AFCCBFC5BFC0BE99B" box="[809,923,965,989]" pageId="25" pageNumber="26" value="1963-07">July 1963</date>
;
<collectionCode id="29B0AE5FFFF5995AFC4BFC5BFC75E99B" LSID="urn:lsid:biocol.org:col:33791" box="[937,997,965,989]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
146137, male, collector not specified,
<date id="3B1F105AFFF5995AFF75FC77FEA4EE47" box="[151,308,1001,1025]" pageId="25" pageNumber="26" value="1974-10">
October
<quantity id="88599B7FFFF5995AFF1AFC77FEA4EE47" box="[248,308,1001,1025]" metricMagnitude="6" metricUnit="m" metricValue="1.9744999999999997" pageId="25" pageNumber="26" unit="km" value="1974.5">1974</quantity>
</date>
. 5 km NE of Villavicencio (
<emphasis id="7DD5EA88FFF5995AFD8FFC77FD19EE44" bold="true" box="[621,649,1001,1026]" pageId="25" pageNumber="26">37</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFD40FC74FD69EE47" box="[674,761,1002,1025]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="25" pageNumber="26">UMMZ</collectionCode>
127206, male, leg. W. Mobely and K. Adler,
<date id="3B1F105AFFF5995AFB1AFC77FAF8EE47" box="[1272,1384,1001,1025]" pageId="25" pageNumber="26" value="1965-07">July 1965</date>
. Rio Negro, c.
<quantity id="88599B7FFFF5995AFEE5FB93FEDFEE63" box="[263,335,1037,1061]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="km" value="12.0">12 km</quantity>
from Villavicencio, Finca 'Las Orquideas' (“below” [south of?] road BogotáVillavicencio) (
<emphasis id="7DD5EA88FFF5995AFA8DFB93FA1BEE60" bold="true" box="[1391,1419,1037,1062]" pageId="25" pageNumber="26">37</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFF75FBAFFF44EE0F" LSID="urn:lsid:biocol.org:col:33791" box="[151,212,1073,1097]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26">MCZ</collectionCode>
150022, male, leg. F. Medem, 1975. Güejar River, Finca Guadualito (
<emphasis id="7DD5EA88FFF5995AFBE3FBAFFB8DEE0C" bold="true" box="[1025,1053,1073,1098]" pageId="25" pageNumber="26">38</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFBD5FBAFFB04EE0F" box="[1079,1172,1073,1097]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
44777, female, leg. L. Klein and F. Medem,
<date id="3B1F105AFFF5995AFE7DFBCBFDCEEE2B" box="[415,606,1109,1133]" pageId="25" pageNumber="26" value="1967-09">September 1967</date>
. Puerto Lopez (
<emphasis id="7DD5EA88FFF5995AFCFCFBCBFCAAEE28" bold="true" box="[798,826,1109,1134]" pageId="25" pageNumber="26">39</emphasis>
):
<collectionCode id="29B0AE5FFFF5995AFCBAFBC8FC3EEE28" LSID="urn:lsid:biocol.org:col:34925" box="[856,942,1110,1134]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
97350, juvenile, leg. A.V. Hutchison and Mahoney,
<date id="3B1F105AFFF5995AFEEFFBE4FE36EED7" box="[269,422,1145,1170]" pageId="25" pageNumber="26" value="1966-01">January 1966</date>
;
<collectionCode id="29B0AE5FFFF5995AFE50FBE4FD98EED4" LSID="urn:lsid:biocol.org:col:34925" box="[434,520,1146,1170]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH</collectionCode>
98406, female, leg. V.H. Hutchison,
<date id="3B1F105AFFF5995AFC45FBE7FBB5EED7" box="[935,1061,1145,1169]" pageId="25" pageNumber="26" value="1966-04">April 1966</date>
.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF5995AFF25FB03FCAFEFBF" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="4F1E369AFFF5995AFF25FB03FCAFEFBF" blockId="25.[151,1437,350,2033]" pageId="25" pageNumber="26">
<emphasis id="7DD5EA88FFF5995AFF25FB03FED1EEF0" bold="true" box="[199,321,1181,1206]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
A moderately large
<taxonomicName id="88A14D19FFF5995AFDC8FB03FD33EEF3" box="[554,675,1181,1205]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="7DD5EA88FFF5995AFDC8FB03FD33EEF3" box="[554,675,1181,1205]" italics="true" pageId="25" pageNumber="26">Gonatodes</emphasis>
</taxonomicName>
, with maximum SLV of
<quantity id="88599B7FFFF5995AFC5CFB03FBB5EEF3" box="[958,1061,1181,1205]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="25" pageNumber="26" unit="mm" value="47.6">47.6 mm</quantity>
. Scales around midbody 96123. Ventral scales in a longitudinal row 4455. Proximal subdigital lamellae as wide as digits, in total 1421 under fourth finger, 1725 under fourth toe. Two or three lateral rows of scales on distal portion of fingers and toes. Tail with midventral scales distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact with one laterodistal scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral in contact with two laterodistal scales per side (respectively 112” and 111”). A white suprahumeral bar bordered by black, present both in males and females, although more conspicuous in males; it reaches dorsally at least up to the dorsolateral region, in some females it almost reaches the middorsal region; never in the form of an ocellus. Males with head dorsally without vermiculation; back and limbs finely vermiculated; gular region without dark streaks.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF5995FFF25F99BFBC0EF22" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26" type="description">
<paragraph id="4F1E369AFFF5995AFF25F99BFB17EC97" blockId="25.[151,1437,350,2033]" pageId="25" pageNumber="26">
<emphasis id="7DD5EA88FFF5995AFF25F99BFEC8EC58" bold="true" box="[199,344,1541,1566]" pageId="25" pageNumber="26">Description.</emphasis>
Maximum SVL in males of
<quantity id="88599B7FFFF5995AFD47F99BFC9BEC5B" box="[677,779,1541,1565]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="25" pageNumber="26" unit="mm" value="47.6">47.6 mm</quantity>
(USNM 84968), in females of
<quantity id="88599B7FFFF5995AFB96F99BFB4AEC5B" box="[1140,1242,1541,1565]" metricMagnitude="-2" metricUnit="m" metricValue="4.71" pageId="25" pageNumber="26" unit="mm" value="47.1">47.1 mm</quantity>
(USNM 92490). Tail round in cross section, tapering toward tip, 1.01.3 (1.17 ± 0.11,
<emphasis id="7DD5EA88FFF5995AFC23F9B4FC44EC07" box="[961,980,1578,1601]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=11) times SLV. Head length 0.220.28 (0.24 ± 0.01,
<emphasis id="7DD5EA88FFF5995AFED6F9D1FED7EC20" box="[308,327,1615,1638]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=61) times SLV, 1.31.6 (1.46 ± 0.05,
<emphasis id="7DD5EA88FFF5995AFCE0F9D1FC85EC20" box="[770,789,1615,1638]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=57) times as long as wide, 1.21.8 (1.38 ± 0.10,
<emphasis id="7DD5EA88FFF5995AFAB6F9D1FAF7EC20" box="[1364,1383,1615,1638]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=56) times as wide as high. Snout round, moderately elongate (
<figureCitation id="D79A2A1FFFF5995AFCA2F9EFFC04ECCF" box="[832,916,1649,1673]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
A), gently sloping toward top of head. Neck slightly narrower than head and anterior portion of body. Body cylindrical. Limbs well developed, lower arm 0.11 0.16 (0.13 ± 0.01,
<emphasis id="7DD5EA88FFF5995AFE87F924FEE8EC97" box="[357,376,1722,1745]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=62) times SVL, lower leg 0.120.16 (0.14 ± 0.01,
<emphasis id="7DD5EA88FFF5995AFC5AF924FC5BEC97" box="[952,971,1722,1745]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=62) times SVL.
</paragraph>
<paragraph id="4F1E369AFFF59959FF27F943FD2FEBEA" blockId="25.[151,1437,350,2033]" lastBlockId="26.[151,1437,151,2013]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Rostral convex, 1.62.3 (1.96 ± 0.13,
<emphasis id="7DD5EA88FFF5995AFD8FF941FD10ECB0" box="[621,640,1759,1782]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=61) times as wide as high; posterior margin with a shallow depression medially and slightly indented by 03 medial postrostrals, with or without a median cleft extending anteriorly. Postrostrals 2 (4.9%), 3 (82%), 4 (4.9%) or 5 (8.2%), lateral ones (supranasals) distinctly larger than medial one(s), when these are present (
<emphasis id="7DD5EA88FFF5995AFE53F8D4FE54ED27" box="[433,452,1866,1889]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=61) (
<figureCitation id="D79A2A1FFFF5995AFDE8F8D7FDC7ED27" box="[522,599,1865,1889]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="25" pageNumber="26">Fig 13</figureCitation>
A). Nasal bordered by rostral, first supralabial (only a narrow contact in some specimens), 2 (96.8%) or 3 (3.2%) postnasals (N=63), and supranasal; internostril distance 0.20.0.27 (0.24 ± 0.02,
<emphasis id="7DD5EA88FFF5995AFF35F80CFF7AEDEF" box="[215,234,1938,1961]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=55) times head width. Supranasal roughly oval or circular, 0.91.6 (1.23 ± 0.15;
<emphasis id="7DD5EA88FFF5995AFB73F80CFB34EDEF" box="[1169,1188,1938,1961]" italics="true" pageId="25" pageNumber="26">N</emphasis>
=62) times as wide as long. Postnasals slightly larger than, or similar in size to, adjacent loreals. Scales on snout convex, hexagonal to round, juxtaposed, relatively uniform in size. Canthus rostralis rounded. Loreal region with scales slightly more elongate than those on snout, largest on row adjacent to supralabials, 911 (9.8 ± 0.7,
<emphasis id="7DD5EA88FFF69959FB6FFF07FB30EAF6" box="[1165,1184,153,176]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=61) loreals on a line between postnasals and anterior corner of eye. Top and posterior portion of head, as well as supraorbital region, with granular scales. A short supraciliary flap present, anteriorly with 612 (8.3 ± 1.0,
<emphasis id="7DD5EA88FFF69959FB50FF7FFB55EABE" box="[1202,1221,225,248]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=60) enlarged and flattened scales, among which 05 (1.7 ± 1.4;
<emphasis id="7DD5EA88FFF69959FD4FFE9BFD50EB5A" box="[685,704,261,284]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=60) small, conical spines. Pupil round, eye diameter 0.190.26 (0.22 ± 0.01,
<emphasis id="7DD5EA88FFF69959FED4FEB7FED9EB06" box="[310,329,297,320]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=56) times head length. Scales on temporal region similar to those on top of head. Ear-opening much smaller than eye, oval, posterior to, and at same level of, commissure of mouth. Supralabials 58 (6.1 ± 0.6;
<emphasis id="7DD5EA88FFF69959FF75FEEFFF3AEBCE" box="[151,170,369,392]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=62), distinctly enlarged anteriorly and decreasing in size posteriorly, one or two of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFF69959FF27FE29FC7FE819" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
Mental large, distinctly wider anteriorly than posteriorly, with posterior margin forming a moderate or wide angle, 1.11.5 (1.27 ± 0.08;
<emphasis id="7DD5EA88FFF69959FE36FE43FE77EBB2" box="[468,487,477,500]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=59) times as wide as long; 1 (1.6%), 2 (76.2%), 3 (19.0%), or 4 (3.2%) postmentals (
<figureCitation id="D79A2A1FFFF69959FF7DFE61FF64E851" box="[159,244,511,535]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="26" pageNumber="27">Fig. 13</figureCitation>
B). Scales on chin flat, smooth, polygonal, juxtaposted, larger anteriorly, decreasing in size posteriorly. Infralabials 47 (5.4 ± 0.7;
<emphasis id="7DD5EA88FFF69959FE32FDBBFE73E87A" box="[464,483,549,572]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=63), distinctly enlarged anteriorly and decreasing in size posteriorly; one or two of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFF69959FF27FDF2FC8BE88A" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">Scales on nape small and granular, becoming slightly larger on sides of neck. Scales on throat anteriorly granular, posteriorly flat, smooth, hexagonal or round, increasing in size toward the posterior region, with a short transitional zone between the anterior and posterior parts.</paragraph>
<paragraph id="4F1E369AFFF69959FF27FD49FA87E9E2" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
Dorsals granular, increasing in size toward the flanks. Ventrals larger than dorsals, roughly hexagonal, flat, smooth, imbricate, in oblique rows; 4456 (50.8 ± 2.3;
<emphasis id="7DD5EA88FFF69959FCFAFD63FCBBE952" box="[792,811,765,788]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=58) scales along the midventral line between anterior margin of forelimbs and vent; 1521 (18.0 ± 1.2;
<emphasis id="7DD5EA88FFF69959FD0EFCBFFD6FE97E" box="[748,767,801,824]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=62) scales in a transverse line at midbody, with a short transitional zone between ventrals and scales on flanks. Scales around midbody 96123 (110.1 ± 5.6;
<emphasis id="7DD5EA88FFF69959FB1DFCDBFA82E91A" box="[1279,1298,837,860]" italics="true" pageId="26" pageNumber="27">N</emphasis>
=60). Scales on preanal plate similar to ventrals, except for those bordering vent, which are very small. Escutcheon present in males on posterior portion of belly and on 35 rows (body-knee direction) on ventral surface of thighs.
</paragraph>
<paragraph id="4F1E369AFFF69959FF27FC31FD98EE1E" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">Scales on anterodorsal surface of forelimbs flat, smooth, roundish or rhomboidal, imbricate, largest close to the wrist; on posterodorsal and ventral surface of forelimbs convex, smooth, rhomboidal, juxtaposed, relatively small. Scales on anterodorsal surface of thighs and ventral surface of hindlimbs flat, smooth, roundish or rhomboidal, imbricate and relatively larger; on posterodorsal surface of thighs and dorsal surface of lower legs smaller, convex, smooth, rhomboidal, juxtaposed.</paragraph>
<paragraph id="4F1E369AFFF69959FF27FBFAFBBBEF4A" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
Lamellae under second (
<emphasis id="7DD5EA88FFF69959FE3DFBFAFE65EE3A" bold="true" box="[479,501,1124,1148]" pageId="26" pageNumber="27">II</emphasis>
) through fourth (
<emphasis id="7DD5EA88FFF69959FD5CFBFAFD4DEE3A" bold="true" box="[702,733,1124,1148]" pageId="26" pageNumber="27">IV</emphasis>
) fingers (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFF69959FB00FBFAFB68EE3A" bold="true" box="[1250,1272,1124,1148]" pageId="26" pageNumber="27">II</emphasis>
: 1317 (56),
<emphasis id="7DD5EA88FFF69959FF75FB16FF28EEE6" bold="true" box="[151,184,1160,1184]" pageId="26" pageNumber="27">III</emphasis>
: 1519 (46), and
<emphasis id="7DD5EA88FFF69959FE6EFB16FE39EEE6" bold="true" box="[396,425,1160,1184]" pageId="26" pageNumber="27">IV</emphasis>
: 1421 (58) (
<figureCitation id="D79A2A1FFFF69959FDB2FB16FD0BEEE6" box="[592,667,1160,1184]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="26" pageNumber="27">Fig 13</figureCitation>
C). Lamellae under second through fourth toes (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFF69959FED1FB32FED9EE82" bold="true" box="[307,329,1196,1220]" pageId="26" pageNumber="27">II</emphasis>
: 1218 (57),
<emphasis id="7DD5EA88FFF69959FE11FB32FD84EE82" bold="true" box="[499,532,1196,1220]" pageId="26" pageNumber="27">III</emphasis>
: 1621 (58), and
<emphasis id="7DD5EA88FFF69959FD0DFB32FC9CEE82" bold="true" box="[751,780,1196,1220]" pageId="26" pageNumber="27">IV</emphasis>
: 1725 (812) (
<figureCitation id="D79A2A1FFFF69959FC24FB32FB88EE82" box="[966,1048,1196,1220]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="26" pageNumber="27">Fig. 13</figureCitation>
D). Claws exposed, non-retractile, between two basal scales. Fingers and toes with three, occasionally two, lateral scales distally, between the fourth (counted from the claw towards the hand) subdigital lamella, and the dorsal scale.
</paragraph>
<paragraph id="4F1E369AFFF69959FF27FA86FC05EFA2" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
Scales on tail dorsally and laterally relatively small, rhomboidal, flat, smooth, imbricate. On ventral surface of tail scales smooth, flat, imbricate, increasing in size toward midventral line; midventral scales, except close to the base of the tail, distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (on proximal portion of tail, when present) or single midventral scale in contact laterodistally with two scales per side respectively 112” and 111” in the codification of
<bibRefCitation id="2B304B6BFFF69959FE3AFA52FC84EFA2" author="Avila-Pires" box="[472,788,1484,1508]" pageId="26" pageNumber="27" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">
Avila-Pires (1995:
<figureCitation id="D79A2A1FFFF69959FD4EFA52FC9BEFA2" box="[684,779,1484,1508]" captionStart="FIGURE 2" captionStartId="6.[151,250,1440,1463]" captionTargetBox="[190,1402,193,1415]" captionTargetId="figure@6.[185,1402,193,1419]" captionTargetPageId="6" captionText="FIGURE 2. Bivariate scattergrams from stepwise discriminant function analysis using scale counts and measurements. Red = Pattern A. Cyan-green = Pattern B. Lime-green = Pattern C. Yellow = Pattern D. Blue = Pattern E." httpUri="https://zenodo.org/record/277515/files/figure.png" pageId="26" pageNumber="27">Figure 2</figureCitation>
)
</bibRefCitation>
(
<figureCitation id="D79A2A1FFFF69959FCC1FA52FCE4EFA2" box="[803,884,1484,1508]" captionStart="FIGURE 13" captionStartId="27.[151,250,1966,1989]" captionTargetBox="[225,1312,246,1965]" captionTargetId="figure@27.[194,1411,193,1982]" captionTargetPageId="27" captionText="FIGURE 13. Gonatodes riveroi sp. nov. (holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm." httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="26" pageNumber="27">Fig. 13</figureCitation>
E).
</paragraph>
<paragraph id="4F1E369AFFF69959FF25FA6EFE9BEC16" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
<emphasis id="7DD5EA88FFF69959FF25FA6EFD95EC4E" bold="true" box="[199,517,1519,1544]" pageId="26" pageNumber="27">
Measurements of
<typeStatus id="901A8838FFF69959FE7CFA6EFD95EC4E" box="[414,517,1520,1544]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
</emphasis>
(in millimeters): SVL 42.5, TL 46.0 (distally regenerated), HL 9.8, HW 6.7, HD 4.7, ED 2.0,
<collectingCountry id="37B6760AFFF69959FEC8F98BFECCEC6A" box="[298,348,1557,1580]" name="India" pageId="26" pageNumber="27">IND</collectingCountry>
1.7, SSL 0.7, SSW 0.8, RSL 1.1, RSW 2.1, MSL 2.2, MSW 2.6, LAL 5.3, LLL 6.2, HSH 2.8, HSL 1.57.
</paragraph>
<paragraph id="4F1E369AFFF69959FF25F9C5FC5BECFA" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
<emphasis id="7DD5EA88FFF69959FF25F9C5FE7FEC32" bold="true" box="[199,495,1627,1652]" pageId="26" pageNumber="27">
Scale counts of
<typeStatus id="901A8838FFF69959FE9DF9C2FE7AEC32" box="[383,490,1628,1652]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
:
</emphasis>
SAM 116 scales, VLR 51, VTR 17, SL 6,
<collectingCountry id="37B6760AFFF69959FC32F9C3FC7BEC32" box="[976,1003,1629,1652]" name="Iceland" pageId="26" pageNumber="27">IL</collectingCountry>
6, PR 3, SP 1, PN 3, LS 9, PM 2, SCS 8, SSC 2, PL2F 6, DL2F 11, LS2F 3, PL3F 6, DL3F 13, LS3F 3, PL4F 7, DL4F 11, LS4F 3, PL2T 6, DL2T 11, LS2T 3, PL3T 6, DL3T 13, LS3T 3, PL4T 11, DL4T 12, LS4T 3, RSE 4.
</paragraph>
<paragraph id="4F1E369AFFF69959FF25F959FB76ED37" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="27">
<emphasis id="7DD5EA88FFF69959FF25F959FE5FECA6" bold="true" box="[199,463,1735,1760]" pageId="26" pageNumber="27">Color in preservative.</emphasis>
In males (
<figureCitation id="D79A2A1FFFF69959FDA9F956FD1DECA6" box="[587,653,1736,1760]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="26" pageNumber="27">Fig. 1</figureCitation>
E), dorsal surface of head reddish-brown, spotless. Back and flanks, base of tail and hind limbs with relatively small, beige and brown vermiculations. A large, conspicuous, white suprahumeral bar with black margins, extending dorsally at least up to the dorsolateral region. Ventral surface of head, gular region and chest brown, without oblique streaks; belly dark-brown or dark-gray; underside of limbs beige. Tail dark brown dorsally and brown ventrally. Escutcheon area (belly and thighs) light gray.
</paragraph>
<paragraph id="4F1E369AFFF6995FFF27F8E3FD06EF5A" blockId="26.[151,1437,151,2013]" lastBlockId="28.[151,1437,1176,1632]" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">In females, dorsal surface of head and limbs light-brown or gray, with black irregular spots; brown and black oblique and/or longitudinal bands may be present on sides of head. Back gray with dorsolateral pairs of black spots; flanks gray with black and brown irregular spots. A moderately large, conspicuous, white suprahumeral bar, with black margins, but less defined than that on males; in some specimens it almost reaches the middorsal region. Ventral surface of head and gular region white with dark oblique streaks, in contact or not at midventral line; belly and underside of limbs white or beige. Tail gray dorsally and white ventrally; original tail distally with white and black bands that form complete rings around the tail.</paragraph>
<caption id="1BDE6612FFF79958FF75F830FC26EDA5" httpUri="https://zenodo.org/record/277526/files/figure.png" pageId="27" pageNumber="28" targetBox="[225,1312,246,1965]" targetPageId="27">
<paragraph id="4F1E369AFFF79958FF75F830FC26EDA5" blockId="27.[151,1436,1966,2019]" pageId="27" pageNumber="28">
<emphasis id="7DD5EA88FFF79958FF75F830FEB0ED82" bold="true" box="[151,288,1966,1989]" pageId="27" pageNumber="28">FIGURE 13.</emphasis>
<taxonomicName id="88A14D19FFF79958FEC5F831FE70ED83" box="[295,480,1967,1989]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="riveroi" status="sp. nov.">
<emphasis id="7DD5EA88FFF79958FEC5F831FE70ED83" box="[295,480,1967,1989]" italics="true" pageId="27" pageNumber="28">Gonatodes riveroi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFF79958FE05F831FDA9ED82" bold="true" box="[487,569,1967,1988]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="66E657F3FFF79958FE05F831FDA9ED82" box="[487,569,1967,1988]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, MZUSP 49153). (A) Dorsal and (B) ventral views of head; ventral views of (C) right hand and (D) right foot; (E) ventral view of tail. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="1BDE6612FFF0995FFF75FBB0FEB0EE2E" httpUri="https://zenodo.org/record/277527/files/figure.png" pageId="28" pageNumber="29" targetBox="[219,1361,392,1010]" targetPageId="28">
<paragraph id="4F1E369AFFF0995FFF75FBB0FEB0EE2E" blockId="28.[151,1436,1070,1128]" pageId="28" pageNumber="29">
<emphasis id="7DD5EA88FFF0995FFF75FBB0FEB1EE02" bold="true" box="[151,289,1070,1093]" pageId="28" pageNumber="29">FIGURE 14.</emphasis>
<taxonomicName id="88A14D19FFF0995FFECAFBB1FE72EE03" box="[296,482,1071,1093]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="riveroi" status="sp. nov.">
<emphasis id="7DD5EA88FFF0995FFECAFBB1FE72EE03" box="[296,482,1071,1093]" italics="true" pageId="28" pageNumber="29">Gonatodes riveroi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFF0995FFE0BFBB1FDACEE02" bold="true" box="[489,572,1071,1092]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="66E657F3FFF0995FFE0BFBB1FDACEE02" box="[489,572,1071,1092]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Villavicencio, Meta, Colombia (UMMZ 127206, SVL = 41.9 mm). (Photo by Kraig Adler).
</paragraph>
</caption>
<paragraph id="4F1E369AFFF0995FFF27FAB9FBC0EF22" blockId="28.[151,1437,1176,1632]" pageId="28" pageNumber="29">
<emphasis id="7DD5EA88FFF0995FFF27FAB9FEC2EF06" bold="true" box="[197,338,1319,1344]" pageId="28" pageNumber="29">Color in life</emphasis>
(photo of UMMZ 127206, male): Head dorsally and laterally brown. Back with brown and yellow vermiculations. Suprahumeral bar white with black margins. Tail dark-brown (
<figureCitation id="D79A2A1FFFF0995FFC10FAD2FBD3EF22" box="[1010,1091,1356,1380]" captionStart="FIGURE 14" captionStartId="28.[151,250,1070,1093]" captionTargetBox="[219,1361,392,1010]" captionTargetId="figure@28.[151,1436,193,1049]" captionTargetPageId="28" captionText="FIGURE 14. Gonatodes riveroi sp. nov. from Villavicencio, Meta, Colombia (UMMZ 127206, SVL = 41.9 mm). (Photo by Kraig Adler)." httpUri="https://zenodo.org/record/277527/files/figure.png" pageId="28" pageNumber="29">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF0995FFF26FAF1FC66EF96" pageId="28" pageNumber="29" type="distribution">
<paragraph id="4F1E369AFFF0995FFF26FAF1FC66EF96" blockId="28.[151,1437,1176,1632]" pageId="28" pageNumber="29">
<emphasis id="7DD5EA88FFF0995FFF26FAF1FECBEFCE" bold="true" box="[196,347,1391,1416]" pageId="28" pageNumber="29">Distribution.</emphasis>
Central
<collectingCountry id="37B6760AFFF0995FFE5FFAEEFDB9EFCE" box="[445,553,1392,1416]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
east of the Andes (
<figureCitation id="D79A2A1FFFF0995FFCEBFAEEFCDCEFCE" box="[777,844,1392,1416]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="28" pageNumber="29">Fig. 3</figureCitation>
A).
<bibRefCitation id="2B304B6BFFF0995FFC98FAEEFBFBEFCE" box="[890,1131,1392,1416]" pageId="28" pageNumber="29" refString="Sanchez, H.; Castano, O. &amp; Cardenas, G. (1995) Diversidad de los reptiles en Colombia. In: Rangel-Ch, J. O. (Ed.), Colombia Diversidad biotica I. Guadalupe, Santafe de Bogota, Colombia, pp. 277 - 325." type="book chapter">
Sanchez
<emphasis id="7DD5EA88FFF0995FFC3DFAEFFB81EFCE" box="[991,1041,1392,1416]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. (1995)
</bibRefCitation>
reported
<taxonomicName id="88A14D19FFF0995FFB3AFAEFFAEDEFCE" box="[1240,1405,1393,1416]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFF0995FFB3AFAEFFAEDEFCE" box="[1240,1405,1393,1416]" italics="true" pageId="28" pageNumber="29">G. concinnatus</emphasis>
</taxonomicName>
in
<collectingCountry id="37B6760AFFF0995FFF75FA0AFE93EFEA" box="[151,259,1428,1452]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
from “Amazonas, Casanare, Boyacá, Cundinamarca, Meta, Pasto, Orinoco, Guaviare.” It is necessary, however, to verify whether all these records refer to the species here described.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF0995FFF26FA43FCF8EC5E" pageId="28" pageNumber="29" type="etymology">
<paragraph id="4F1E369AFFF0995FFF26FA43FCF8EC5E" blockId="28.[151,1437,1176,1632]" pageId="28" pageNumber="29">
<emphasis id="7DD5EA88FFF0995FFF26FA43FED9EFB3" bold="true" box="[196,329,1501,1525]" pageId="28" pageNumber="29">Etymology.</emphasis>
The species is named after Dr. Carlos Rivero-Blanco, who made an extensive study on
<taxonomicName id="88A14D19FFF0995FFAE3FA42FAE8EFB2" box="[1281,1400,1500,1524]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="7DD5EA88FFF0995FFAE3FA42FAE8EFB2" box="[1281,1400,1500,1524]" italics="true" pageId="28" pageNumber="29">Gonatodes</emphasis>
</taxonomicName>
, as well as contributions to environmental conservation in
<collectingCountry id="37B6760AFFF0995FFD12F99EFCF4EC5E" box="[752,868,1536,1560]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFF0995FFF26F9BBFA0DEC26" pageId="28" pageNumber="29" type="discussion">
<paragraph id="4F1E369AFFF0995FFF26F9BBFA0DEC26" blockId="28.[151,1437,1176,1632]" pageId="28" pageNumber="29">
<emphasis id="7DD5EA88FFF0995FFF26F9BBFEA6EC7B" bold="true" box="[196,310,1573,1597]" pageId="28" pageNumber="29">Remarks.</emphasis>
<bibRefCitation id="2B304B6BFFF0995FFEDFF9BAFDBFEC7A" author="Rivero-Blanco" box="[317,559,1572,1596]" pageId="28" pageNumber="29" refString="Rivero-Blanco, C. (1979) The Neotropical lizard genus Gonatodes Fitzinger (Sauria: Sphaerodactylinae). PhD Thesis, Texas A &amp; M University. 233 pp." type="book" year="1979">Rivero-Blanco (1979)</bibRefCitation>
considered the specimens from Villavincencio and vicinity, in Meta,
<collectingCountry id="37B6760AFFF0995FFACBF9BAFA08EC7A" box="[1321,1432,1572,1596]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
, as intergrades between
<taxonomicName id="88A14D19FFF0995FFE77F9D6FCB1EC26" box="[405,801,1608,1632]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="subSpecies" species="concinnatus" subSpecies="concinnatus">
<emphasis id="7DD5EA88FFF0995FFE77F9D6FCB1EC26" box="[405,801,1608,1632]" italics="true" pageId="28" pageNumber="29">Gonatodes concinnatus concinnatus</emphasis>
</taxonomicName>
, from
<collectingCountry id="37B6760AFFF0995FFC89F9D7FC0DEC26" box="[875,925,1609,1632]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
and
<collectingCountry id="37B6760AFFF0995FFC30F9D6FBBFEC26" box="[978,1071,1608,1632]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
, and
<emphasis id="7DD5EA88FFF0995FFB85F9D7FB73EC26" box="[1127,1251,1608,1632]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="88A14D19FFF0995FFB85F9D7FB4EEC26" box="[1127,1246,1608,1632]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="subSpecies" species="concinnatus" subSpecies="ligiae">G. c. ligiae</taxonomicName>
,
</emphasis>
from
<collectingCountry id="37B6760AFFF0995FFAC7F9D6FA09EC26" box="[1317,1433,1608,1632]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>