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<document id="C4E729198678B393E3F26D9723A866B3" ID-CLB-Dataset="21648" ID-DOI="10.5852/ejt.2020.669" ID-GBIF-Dataset="47b43e12-37bc-4b2c-b654-0e668351cc97" ID-ISSN="2118-9773" ID-Zenodo-Dep="3900298" ID-ZooBank="06FEC792-67DE-46F1-A5D3-4B0F17AF3B73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1592505667257" checkinUser="valdenar" docAuthor="Haran, Julien M., Hansen, Steffan, Benoit, Laure &amp; Cbgp, Pia Addison" docDate="2020" docId="BD7C0F46FFCD5306FDDAFC6CFC0D0BD4" docLanguage="en" docName="ejt-669_haran_hansen_addison.pdf.imf" docOrigin="European Journal of Taxonomy 669" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phlyctinus aloevorus Haran &amp; Hansen &amp; Benoit &amp; Cbgp 2020, sp. nov." docType="treatment" docUuid="28603A47-8AF4-4676-A2E2-BC82092AD72A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="24" masterDocId="4145773EFFD95311FF99FF97FFB50A6A" masterDocTitle="Description of five new species in the genus Phlyctinus Schoenherr (Coleoptera, Curculionidae): a first step in deciphering the P. callosus complex" masterLastPageNumber="29" masterPageNumber="1" pageNumber="21" updateTime="1698842622188" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="335645E98518E30ACC363609DEF95762">Description of five new species in the genus Phlyctinus Schoenherr (Coleoptera, Curculionidae): a first step in deciphering the P. callosus complex</mods:title>
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<mods:namePart id="4B0F3CDA516F4F2F9B3EDF87781AB143">Haran, Julien M.</mods:namePart>
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<mods:namePart id="DFEB010EDB1E69D7167FFE034ADB7BE1">Hansen, Steffan</mods:namePart>
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<mods:affiliation id="B0C46FF82D0A88288F37B30B3AE89987">Department of Conservation Ecology and Entomology, Stellenbosch University, Stellenbosch, South Africa. &amp; Email: shansen. rsa @ gmail. com &amp; urn: lsid: zoobank. org: author: 52 F 0 FC 34 - 03 F 6 - 4 F 04 - A 1 AA-FE 7607 B 0 EFBC</mods:affiliation>
<mods:nameIdentifier id="B05F55AFD1BD8EDBB0C1C45B4C3B8EF7" type="email">shansen.rsa@gmail.com</mods:nameIdentifier>
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<mods:namePart id="45965F7354844844F563315704426CC3">Benoit, Laure</mods:namePart>
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<mods:affiliation id="C65E4E6C6727775B99138595364D4A1C">CBGP, CIRAD, Montpellier SupAgro, INRA, IRD, Montpellier University, Montpellier, France. &amp; Email: laure. benoit @ cirad. fr &amp; urn: lsid: zoobank. org: author: 61963 F 74 - 724 B- 4174 - 9 E 9 A- 8817 A 3516 B 0 E</mods:affiliation>
<mods:nameIdentifier id="E88F5901736C17855B8494CD873A39F7" type="email">laure.benoit@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="B5A136A85FECAB9B8FFD4E2CC16F8062">Cbgp, Pia Addison</mods:namePart>
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<mods:title id="B260C4688DC5B35FA0323A10DB508785">European Journal of Taxonomy</mods:title>
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<mods:date id="1316D8D24E8F0C18C363F99AA3B0D8B3">2020</mods:date>
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<treatment id="BD7C0F46FFCD5306FDDAFC6CFC0D0BD4" ID-DOI="http://doi.org/10.5281/zenodo.5662138" ID-GBIF-Taxon="164615712" ID-Zenodo-Dep="5662138" LSID="urn:lsid:zoobank.org:act:28603A47-8AF4-4676-A2E2-BC82092AD72A" httpUri="http://treatment.plazi.org/id/BD7C0F46FFCD5306FDDAFC6CFC0D0BD4" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="7DCFEDDBFFCD5305FDDAFC6CFC440E7C" box="[579,1009,1019,1046]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="356ABE50FFCD5305FDDAFC6CFC440E7C" blockId="20.[367,1223,1019,1116]" box="[579,1009,1019,1046]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FDDAFC6CFC440E7C" box="[579,1009,1019,1046]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="F2D5C5D3FFCD5305FDDAFC6CFC250E7C" ID-CoL="8KBN2" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[579,912,1019,1046]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCD5305FDDAFC6CFCF50E7C" bold="true" box="[579,832,1019,1046]" italics="true" pageId="20" pageNumber="21">Phlyctinus aloevorus</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FC0EFC6BFC440E7C" box="[919,1009,1020,1046]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="356ABE50FFCD5305FD21FBD6FCCE0E36" blockId="20.[367,1223,1019,1116]" box="[696,891,1089,1116]" pageId="20" pageNumber="21">
<figureCitation id="ADEEA2D5FFCD5305FD21FBD6FCA40E36" box="[696,785,1089,1116]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[192,1395,350,1949]" captionTargetId="figure@7.[189,1398,319,1964]" captionTargetPageId="7" captionText="Fig. 1 (opposite page). Habitus of species of Phlyctinus Schoenherr, 1826 males (left) and females (right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3900300" httpUri="https://zenodo.org/record/3900300/files/figure.png" pageId="20" pageNumber="21">Figs 1F</figureCitation>
,
<figureCitation id="ADEEA2D5FFCD5305FC87FBD5FC880E36" box="[798,829,1090,1116]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[252,1379,330,1800]" captionTargetId="figure@10.[189,1398,308,1806]" captionTargetPageId="10" captionText="Fig. 2. Penis of Phlyctinus Schoenherr, 1826, in dorsal (left) and lateral (right) view, and copulatory sclerite (top right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 500 µm. Copulatory sclerite not to scale." figureDoi="http://doi.org/10.5281/zenodo.3900302" httpUri="https://zenodo.org/record/3900302/files/figure.png" pageId="20" pageNumber="21">2F</figureCitation>
,
<figureCitation id="ADEEA2D5FFCD5305FCD0FBD5FCE20E36" box="[841,855,1090,1116]" captionStart="Fig" captionStartId="18.[189,232,1817,1843]" captionTargetBox="[264,1322,286,2013]" captionTargetId="figure@17.[244,1344,265,2019]" captionTargetPageId="17" captionText="Fig. 4 (opposite page). Habitus in natura, host plants and biotopes of Phlyctinus Schoenherr, 1826 species. A. P. callosus (Schoenherr, 1826). B. P. xerophilus Haran sp. nov. CD. P. grootbosensis Haran sp. nov. and its host plant, Osteospermum moniliferum L. EF. P. planithorax Haran sp. nov. and its host plant, Hymenolepis crithmifolia (L.) Greuter. GH. Biotope and host plant (Arctotis angustifolia L.) of P. littoralis Haran sp. nov. IJ. P. aloevorus Haran sp. nov. on its host plant, Aloe arborescens Mill., with details of feeding damage on floral peduncle" figureDoi="http://doi.org/10.5281/zenodo.3900306" httpUri="https://zenodo.org/record/3900306/files/figure.png" pageId="20" pageNumber="21">4</figureCitation>
IJ
</paragraph>
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<subSubSection id="7DCFEDDBFFCD5305FF24FB14FD2D0E85" pageId="20" pageNumber="21" type="etymology">
<paragraph id="356ABE50FFCD5305FF24FB14FEF10EF7" blockId="20.[189,324,1155,1181]" box="[189,324,1155,1181]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24FB14FEF10EF7" bold="true" box="[189,324,1155,1181]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24FB14FEF10EF7" bold="true" box="[189,324,1155,1181]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FB26FD2D0E85" blockId="20.[189,1398,1201,1263]" pageId="20" pageNumber="21">
This species name is formed from the genus name of its main host plant,
<taxonomicName id="F2D5C5D3FFCD5305FBAEFB26FAFE0EA1" authority="Mill." authorityName="Mill." box="[1079,1355,1201,1227]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCD5305FBAEFB26FAB10EA1" box="[1079,1284,1201,1227]" italics="true" pageId="20" pageNumber="21">Aloe arborescens</emphasis>
Mill.
</taxonomicName>
, on which it causes distinct feeding damage.
</paragraph>
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<emphasis id="07A16242FFCD5305FF24FA81FE1E0F5A" bold="true" box="[189,427,1302,1328]" pageId="20" pageNumber="21">Material examined</emphasis>
</heading>
</paragraph>
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<emphasis id="07A16242FFCD5305FF73FAD2FEE90F35" bold="true" box="[234,348,1349,1375]" pageId="20" pageNumber="21">
<typeStatus id="EA6E00F2FFCD5305FF73FAD2FEE90F35" box="[234,348,1349,1375]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FAFFFB290FCF" blockId="20.[189,1399,1349,1446]" pageId="20" pageNumber="21">
<materialsCitation id="85BDB40DFFCD5305FF24FAFFFB290FCF" ID-GBIF-Occurrence="2640284354" collectingDate="2019-03-23" collectionCode="CBGP, SAMC" collectorName="J. Haran" country="South Africa" latitude="-34.352417" location="Betty's Bay" longLatPrecision="1" longitude="18.91275" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<collectingCountry id="4DC2FEC0FFCD5305FF24FAFFFDF90FE8" box="[189,588,1384,1410]" name="South Africa" pageId="20" pageNumber="21">REPUBLIC OF SOUTH AFRICA</collectingCountry>
• ♂;
<collectingRegion id="F71170B2FFCD5305FD1FFAFFFC2D0FE8" box="[646,920,1384,1410]" country="South Africa" name="Western Cape" pageId="20" pageNumber="21">Western Cape province</collectingRegion>
,
<location id="300AE88BFFCD5305FC3BFAFFFB9B0FE8" LSID="urn:lsid:plazi:treatment:BD7C0F46FFCD5306FDDAFC6CFC0D0BD4:300AE88BFFCD5305FC3BFAFFFB9B0FE8" box="[930,1070,1384,1410]" country="South Africa" latitude="-34.352417" longLatPrecision="1" longitude="18.91275" name="Betty's Bay" pageId="20" pageNumber="21" stateProvince="Western Cape">Bettys Bay</location>
;
<geoCoordinate id="50E1D897FFCD5305FBA1FAF1FB780FE8" box="[1080,1229,1382,1410]" degrees="34" direction="south" minutes="21.145" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-34.352417">34º21.145ʹ S</geoCoordinate>
,
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;
<date id="416B9890FFCD5305FF24FA1CFEEB0FCF" box="[189,350,1419,1446]" pageId="20" pageNumber="21" value="2019-03-23">
<collectingDate id="512F6178FFCD5305FF24FA1CFEEB0FCF" box="[189,350,1419,1446]" pageId="20" pageNumber="21" value="2019-03-23">23 Mar. 2019</collectingDate>
</date>
;
<collectorName id="9820DB86FFCD5305FEF3FA1BFE790FCC" box="[362,460,1420,1446]" pageId="20" pageNumber="21">J. Haran</collectorName>
leg.;
<collectedFrom id="86C9FDADFFCD5305FD97FA1BFD480FCF" box="[526,765,1419,1446]" pageId="20" pageNumber="21">
on
<taxonomicName id="F2D5C5D3FFCD5305FDABFA1CFD480FCF" box="[562,765,1419,1445]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCD5305FDABFA1CFD480FCF" box="[562,765,1419,1445]" italics="true" pageId="20" pageNumber="21">Aloe arborescens</emphasis>
</taxonomicName>
</collectedFrom>
;
<collectionCode id="53C42695FFCD5305FC94FA1CFCEC0FCF" box="[781,857,1419,1445]" pageId="20" pageNumber="21">CBGP</collectionCode>
code JHAR02252;
<collectionCode id="53C42695FFCD5305FBD8FA1CFB2D0FCF" box="[1089,1176,1419,1445]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="20" pageNumber="21" type="Museum">SAMC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="356ABE50FFCD5305FF73FA5AFED20F8D" blockId="20.[189,1399,1485,1723]" box="[234,359,1485,1511]" pageId="20" pageNumber="21">
<emphasis id="07A16242FFCD5305FF73FA5AFED20F8D" bold="true" box="[234,359,1485,1511]" pageId="20" pageNumber="21">
<typeStatus id="EA6E00F2FFCD5305FF73FA5AFED20F8D" box="[234,359,1485,1511]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FA67FEDC0CD1" blockId="20.[189,1399,1485,1723]" pageId="20" pageNumber="21">
<materialsCitation id="85BDB40DFFCD5305FF24FA67FEC80C47" ID-GBIF-Occurrence="2640284341" collectionCode="CBGP" country="South Africa" location="Western Cape province" pageId="20" pageNumber="21" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Western Cape province" typeStatus="holotype">
<collectingCountry id="4DC2FEC0FFCD5305FF24FA67FDFB0C60" box="[189,590,1520,1546]" name="South Africa" pageId="20" pageNumber="21">REPUBLIC OF SOUTH AFRICA</collectingCountry>
<emphasis id="07A16242FFCD5305FDF0FA67FC390C60" bold="true" box="[617,908,1520,1546]" pageId="20" pageNumber="21">
<collectingRegion id="F71170B2FFCD5305FDF0FA67FC390C60" box="[617,908,1520,1546]" country="South Africa" name="Western Cape" pageId="20" pageNumber="21">Western Cape province</collectingRegion>
</emphasis>
<specimenCount id="23D375D9FFCD5305FC3AFA67FC530C60" box="[931,998,1520,1546]" pageId="20" pageNumber="21" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="23D375D9FFCD5305FC6BFA67FB810C60" box="[1010,1076,1520,1546]" pageId="20" pageNumber="21" type="female">3 ♀♀</specimenCount>
; same collection data as for
<typeStatus id="EA6E00F2FFCD5305FF24F984FE920C47" box="[189,295,1555,1581]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
;
<collectionCode id="53C42695FFCD5305FEA8F984FEC80C47" box="[305,381,1555,1581]" pageId="20" pageNumber="21">CBGP</collectionCode>
</materialsCitation>
<materialsCitation id="85BDB40DFFCD5305FE0BF984FC480C47" ID-GBIF-Occurrence="2640284358" box="[402,1021,1555,1581]" collectionCode="SAMC" country="South Africa" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Western Cape province" typeStatus="holotype">
<specimenCount id="23D375D9FFCD5305FE0BF984FE080C47" box="[402,445,1555,1581]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
,
<specimenCount id="23D375D9FFCD5305FE50F984FE410C47" box="[457,500,1555,1581]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
; same collection data as for
<typeStatus id="EA6E00F2FFCD5305FCAFF984FC150C47" box="[822,928,1555,1581]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
;
<collectionCode id="53C42695FFCD5305FC33F984FC480C47" box="[938,1021,1555,1581]" collectionName="South Africa, Cape Town, Iziko Museum of Capetown (formerly South African Museum)" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="20" pageNumber="21" type="Museum">SAMC</collectionCode>
</materialsCitation>
<materialsCitation id="85BDB40DFFCD5305FB8AF984FDE30C1E" ID-GBIF-Occurrence="2640284321" collectingDate="2019-11-24" collectionCode="CBGP" collectorName="J. Haran" latitude="-34.351166" location="Botanical Garden" longLatPrecision="1" longitude="18.927147" municipality="Bay" pageId="20" pageNumber="21" specimenCode="JHAR03063" specimenCount="3">
<specimenCount id="23D375D9FFCD5305FB8AF984FBDF0C47" box="[1043,1130,1555,1581]" pageId="20" pageNumber="21" type="generic">3 specs</specimenCount>
; Bettys
<collectingMunicipality id="D50E242AFFCD5305FB55F984FB490C47" box="[1228,1276,1555,1581]" pageId="20" pageNumber="21">Bay</collectingMunicipality>
,
<location id="300AE88BFFCD5305FA9FF984FEA20C3A" LSID="urn:lsid:plazi:treatment:BD7C0F46FFCD5306FDDAFC6CFC0D0BD4:300AE88BFFCD5305FA9FF984FEA20C3A" latitude="-34.351166" longLatPrecision="1" longitude="18.927147" municipality="Bay" name="Botanical Garden" pageId="20" pageNumber="21">Botanical Garden</location>
;
<geoCoordinate id="50E1D897FFCD5305FEBAF9A3FE670C3A" box="[291,466,1588,1616]" degrees="34" direction="south" minutes="21" orientation="latitude" pageId="20" pageNumber="21" precision="1" seconds="04.20" value="-34.351166">34º21ʹ04.20ʺ S</geoCoordinate>
,
<geoCoordinate id="50E1D897FFCD5305FE46F9A3FD270C3B" box="[479,658,1588,1617]" degrees="18" direction="east" minutes="55" orientation="longitude" pageId="20" pageNumber="21" precision="1" seconds="37.73" value="18.927147">18º55ʹ37.73ʺ E</geoCoordinate>
;
<date id="416B9890FFCD5305FD07F9A1FC8A0C3A" box="[670,831,1590,1617]" pageId="20" pageNumber="21" value="2019-11-24">
<collectingDate id="512F6178FFCD5305FD07F9A1FC8A0C3A" box="[670,831,1590,1617]" pageId="20" pageNumber="21" value="2019-11-24">24 Nov. 2019</collectingDate>
</date>
;
<collectorName id="9820DB86FFCD5305FCD2F9A0FC180C3B" box="[843,941,1591,1617]" pageId="20" pageNumber="21">J. Haran</collectorName>
leg.; on
<taxonomicName id="F2D5C5D3FFCD5305FB8BF9A1FB680C3A" box="[1042,1245,1590,1616]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCD5305FB8BF9A1FB680C3A" box="[1042,1245,1590,1616]" italics="true" pageId="20" pageNumber="21">Aloe arborescens</emphasis>
</taxonomicName>
;
<collectionCode id="53C42695FFCD5305FB74F9A1FA8C0C3A" box="[1261,1337,1590,1616]" pageId="20" pageNumber="21">CBGP</collectionCode>
code
<specimenCode id="6573162BFFCD5305FF24F9CEFEE60C19" box="[189,339,1625,1651]" pageId="20" pageNumber="21">JHAR03063</specimenCode>
;
<collectionCode id="53C42695FFCD5305FEC6F9CEFE1E0C19" box="[351,427,1625,1651]" pageId="20" pageNumber="21">CBGP</collectionCode>
(ethanol coll.)
</materialsCitation>
<materialsCitation id="85BDB40DFFCD5305FDF7F9CEFEDC0CD1" ID-GBIF-Occurrence="2640284318" collectingDate="2019-11-24" collectionCode="CBGP" collectorName="J. Haran" latitude="-34.342693" location="Kleinmond" longLatPrecision="1" longitude="19.014853" municipality="Kleinmond" pageId="20" pageNumber="21" specimenCode="JHAR03064" specimenCount="7" specimenCount-male="1">
<specimenCount id="23D375D9FFCD5305FDF7F9CEFD2F0C1E" box="[622,666,1625,1652]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
,
<specimenCount id="23D375D9FFCD5305FD31F9CEFCB50C1E" box="[680,768,1625,1652]" pageId="20" pageNumber="21" type="generic">6 specs</specimenCount>
;
<collectingMunicipality id="D50E242AFFCD5305FC95F9CEFC210C19" box="[780,916,1625,1651]" pageId="20" pageNumber="21">Kleinmond</collectingMunicipality>
;
<geoCoordinate id="50E1D897FFCD5305FC06F9CFFBFA0C19" box="[927,1103,1624,1652]" degrees="34" direction="south" minutes="20" orientation="latitude" pageId="20" pageNumber="21" precision="1" seconds="33.70" value="-34.342693">34º20ʹ33.70ʺ S</geoCoordinate>
,
<geoCoordinate id="50E1D897FFCD5305FBC2F9CFFABB0C1E" box="[1115,1294,1624,1652]" degrees="19" direction="east" minutes="00" orientation="longitude" pageId="20" pageNumber="21" precision="1" seconds="53.47" value="19.014853">19º00ʹ53.47ʺ E</geoCoordinate>
;
<date id="416B9890FFCD5305FA83F9CEFF4E0CFD" pageId="20" pageNumber="21" value="2019-11-24">
<collectingDate id="512F6178FFCD5305FA83F9CEFF4E0CFD" pageId="20" pageNumber="21" value="2019-11-24">24 Nov. 2019</collectingDate>
</date>
;
<collectorName id="9820DB86FFCD5305FE92F9EAFEC40CFD" box="[267,369,1661,1687]" pageId="20" pageNumber="21">J. Haran</collectorName>
leg.; on
<taxonomicName id="F2D5C5D3FFCD5305FE7AF9EAFD070CFD" box="[483,690,1661,1687]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCD5305FE7AF9EAFD070CFD" box="[483,690,1661,1687]" italics="true" pageId="20" pageNumber="21">Aloe arborescens</emphasis>
</taxonomicName>
and
<taxonomicName id="F2D5C5D3FFCD5305FD6AF9EAFC6C0CFD" box="[755,985,1661,1687]" class="Magnoliopsida" family="Aizoaceae" genus="Carpobrotus" kingdom="Plantae" order="Caryophyllales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="edulis">
<emphasis id="07A16242FFCD5305FD6AF9EAFC6C0CFD" box="[755,985,1661,1687]" italics="true" pageId="20" pageNumber="21">Carpobrotus edulis</emphasis>
</taxonomicName>
;
<collectionCode id="53C42695FFCD5305FC75F9EAFB8D0CFD" box="[1004,1080,1661,1687]" pageId="20" pageNumber="21">CBGP</collectionCode>
code
<specimenCode id="6573162BFFCD5305FB1CF9EAFAAE0CFC" box="[1157,1307,1660,1687]" pageId="20" pageNumber="21">JHAR03064</specimenCode>
;
<collectionCode id="53C42695FFCD5305FAB2F9EAFAC20CFD" box="[1323,1399,1661,1687]" pageId="20" pageNumber="21">CBGP</collectionCode>
(ethanol coll.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCD5304FF24F976FDE10DBC" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="356ABE50FFCD5305FF24F976FEFB0C91" blockId="20.[189,334,1761,1787]" box="[189,334,1761,1787]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24F976FEFB0C91" bold="true" box="[189,334,1761,1787]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24F976FEFB0C91" bold="true" box="[189,334,1761,1787]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24F887FE4F0D40" blockId="20.[189,506,1807,1834]" box="[189,506,1807,1834]" pageId="20" pageNumber="21">
BODY LENGTH.
<quantity id="F22D13B5FFCD5305FEFCF898FE430D40" box="[357,502,1807,1834]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="20" pageNumber="21" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.56.5 mm</quantity>
.
</paragraph>
<paragraph id="356ABE50FFCD5305FF24F8C6FB450DD8" blockId="20.[189,1398,1873,1970]" pageId="20" pageNumber="21">COLOUR. Body integument dark red, head and prothorax darker, usually black; vestiture of elytra with elliptic elongate scales, twice as long as wide, not concealing integument; scales brown with scattered ill-defined paler spots, in females with a pale transverse band on apical ¼ of interstriae 13.</paragraph>
<paragraph id="356ABE50FFCC5304FF24FE9BFBE90885" blockId="21.[189,1399,267,751]" pageId="21" pageNumber="22">HEAD. Rostrum longer than wide (w/l ratio: 0.72) in both sexes, at base as wide as at apex, sides slightly concave near middle of length; epifrons slightly narrower in middle of length than width of eye in dorsal view, at base slightly narrower than distance between eyes, sides subparallel, slightly widening apicad, upper face with a longitudinal carina extending from basal transverse groove to nasal plate, visible through scales; nasal plate V-shaped, reaching level of antennal insertion basally, bare of scales, with only minute setae inserted in scattered punctures and 3 pairs of long setae apically near insertion of mandibles (2 short and 1 very long); antennal scrobes slightly curved in lateral view, directed to middle of eye, separated from it by a strip of scales half to ½ narrower than diameter of eye; mandibles trisetose, bare of scales. Forehead with a short longitudinal furrow between eyes, scales and setae arranged centripetally towards a point below base of eyes; eyes protruding, margin rounded, surface moderately conical in dorsal view. Antennae slender, scape 0.7 × as long as funicle, bisinuate, subclavate at apex, exceeding anterior margin of prothorax in repose; funicle with segments 12 elongate, 1 slightly shorter (0.87 ×) than 2, segments 36 longer than wide, conical, shortening apicad, segment 7 longer and slightly wider than 6; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous.</paragraph>
<paragraph id="356ABE50FFCC5304FF24FC80FEF409F0" blockId="21.[189,1399,790,922]" pageId="21" pageNumber="22">PROTHORAX. Slightly wider than long (w/l ratio: 1.18), widest before middle of length, apical margin 0.7 × as wide as basal margin, sides rounded, apical half slightly concave; integument densely punctate, with shiny elongate concentric ridges and a median carina; upper face regularly convex, lacking cuticular depression.</paragraph>
<paragraph id="356ABE50FFCC5304FF24FC55FBED0E2F" blockId="21.[189,1398,961,1094]" pageId="21" pageNumber="22">ELYTRA. Sides convex, widest near middle of length (ratio w/l: 0.790.83), integument finely punctate and reticulate, shiny, interstriae raised, 34× as wide as width of striae in basal half; large declivital callosities absent or very reduced in apical ¼ of interstriae 3, 5 and 7; each interstria with a series of semi-erect, elongate pale brown scales, slightly more condensed on callosities.</paragraph>
<paragraph id="356ABE50FFCC5304FF24FBFAFC3D0F5D" blockId="21.[189,1399,1132,1335]" pageId="21" pageNumber="22">
ABDOMEN. Underside with scattered scales similar to those on elytra, not concealing integument; ventrite 1 shorter than 2+3+4, slightly convex in middle (
<specimenCount id="23D375D9FFCC5304FC98FB07FCA10EC0" box="[769,788,1168,1194]" pageId="21" pageNumber="22" type="female"></specimenCount>
) or concave with sub-erect setae shorter than claws (♂), ventrite 5 longer than 3+4, with recumbent elongate setae in middle and near apex and a weak longitudinal carina near apex (
<specimenCount id="23D375D9FFCC5304FDB0FB41FD890E9A" box="[553,572,1238,1264]" pageId="21" pageNumber="22" type="female"></specimenCount>
) or with a large cavity extending over apical ¾, with long erect setae mainly condensed at middle of basal margin of cavity and a brush on each side of apical margin; bottom of cavity with a few short recumbent or semi-erect setae (♂).
</paragraph>
<paragraph id="356ABE50FFCC5304FF24FAC9FB0A0FF1" blockId="21.[189,1398,1374,1436]" pageId="21" pageNumber="22">LEGS. Femora clavate, unarmed, with double vestiture of recumbent and semi-erect greyish scales; tibiae straight, unarmed, slightly bisinuate on ventral side; claws simple, free, equal in length.</paragraph>
<paragraph id="356ABE50FFCC5304FF24FA54FBB40C2D" blockId="21.[189,1399,1474,1607]" pageId="21" pageNumber="22">
GENITALIA. Body of penis elongate, 0.75 × as long as apodemes, sides convex, widest slightly before middle of length (w/l ratio: 0.36), in lateral view curvature weak, stronger near base (
<figureCitation id="ADEEA2D5FFCC5304FB14FA71FB500C6A" box="[1165,1253,1510,1536]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[252,1379,330,1800]" captionTargetId="figure@10.[189,1398,308,1806]" captionTargetPageId="10" captionText="Fig. 2. Penis of Phlyctinus Schoenherr, 1826, in dorsal (left) and lateral (right) view, and copulatory sclerite (top right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 500 µm. Copulatory sclerite not to scale." figureDoi="http://doi.org/10.5281/zenodo.3900302" httpUri="https://zenodo.org/record/3900302/files/figure.png" pageId="21" pageNumber="22">Fig. 2F</figureCitation>
); copulatory sclerite with left arm longer than body, bearing setae at base; comb of right arm with setae converging, curved, straight at apex; right area of body slightly wider than left area.
</paragraph>
<paragraph id="356ABE50FFCC5304FF24F9FAFE050CED" blockId="21.[189,432,1645,1671]" box="[189,432,1645,1671]" pageId="21" pageNumber="22">
<heading id="6E22093CFFCC5304FF24F9FAFE050CED" bold="true" box="[189,432,1645,1671]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="07A16242FFCC5304FF24F9FAFE050CED" bold="true" box="[189,432,1645,1671]" pageId="21" pageNumber="22">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCC5304FF24F90BFB7F0CB0" blockId="21.[189,1398,1692,1754]" pageId="21" pageNumber="22">Males may be distinguished externally from females by the homogenous elytral vestiture (with a transverse pale band in apical ½ in females), and the cavity and erect setae on ventrite 5.</paragraph>
<paragraph id="356ABE50FFCC5304FF24F896FEFA0D71" blockId="21.[189,335,1793,1819]" box="[189,335,1793,1819]" pageId="21" pageNumber="22">
<heading id="6E22093CFFCC5304FF24F896FEFA0D71" bold="true" box="[189,335,1793,1819]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="07A16242FFCC5304FF24F896FEFA0D71" bold="true" box="[189,335,1793,1819]" pageId="21" pageNumber="22">Life history</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCC5304FF24F8B8FDE10DBC" blockId="21.[189,1399,1839,2006]" pageId="21" pageNumber="22">
<taxonomicName id="F2D5C5D3FFCC5304FF24F8B8FE030D23" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,438,1839,1865]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCC5304FF24F8B8FE030D23" box="[189,438,1839,1865]" italics="true" pageId="21" pageNumber="22">Phlyctinus aloevorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCC5304FE26F8A7FDAE0D20" box="[447,539,1840,1866]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="ADEEA2D5FFCC5304FDB4F8B8FDCA0D23" box="[557,639,1839,1865]" captionStart="Fig" captionStartId="18.[189,232,1817,1843]" captionTargetBox="[264,1322,286,2013]" captionTargetId="figure@17.[244,1344,265,2019]" captionTargetPageId="17" captionText="Fig. 4 (opposite page). Habitus in natura, host plants and biotopes of Phlyctinus Schoenherr, 1826 species. A. P. callosus (Schoenherr, 1826). B. P. xerophilus Haran sp. nov. CD. P. grootbosensis Haran sp. nov. and its host plant, Osteospermum moniliferum L. EF. P. planithorax Haran sp. nov. and its host plant, Hymenolepis crithmifolia (L.) Greuter. GH. Biotope and host plant (Arctotis angustifolia L.) of P. littoralis Haran sp. nov. IJ. P. aloevorus Haran sp. nov. on its host plant, Aloe arborescens Mill., with details of feeding damage on floral peduncle" figureDoi="http://doi.org/10.5281/zenodo.3900306" httpUri="https://zenodo.org/record/3900306/files/figure.png" pageId="21" pageNumber="22">Fig. 4I</figureCitation>
) forms large populations on
<taxonomicName id="F2D5C5D3FFCC5304FC42F8B8FAC20D20" box="[987,1399,1839,1866]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCC5304FC42F8B8FB1D0D23" box="[987,1192,1839,1865]" italics="true" pageId="21" pageNumber="22">Aloe arborescens</emphasis>
(Asphodelaceae)
</taxonomicName>
and, to a lesser extent, on
<taxonomicName id="F2D5C5D3FFCC5304FE7EF8C5FCD50D07" box="[487,864,1874,1901]" class="Magnoliopsida" family="Aizoaceae" genus="Carpobrotus" kingdom="Plantae" order="Caryophyllales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="edulis">
<emphasis id="07A16242FFCC5304FE7EF8C5FD7C0D06" box="[487,713,1874,1900]" italics="true" pageId="21" pageNumber="22">Carpobrotus edulis</emphasis>
(Aizoaceae)
</taxonomicName>
. Contrary to other species of the genus, which spend the day sheltering in the leaf litter,
<taxonomicName id="F2D5C5D3FFCC5304FD2DF8E0FCF20DFA" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[692,839,1910,1936]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCC5304FD2DF8E0FCF20DFA" box="[692,839,1910,1936]" italics="true" pageId="21" pageNumber="22">P. aloevorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCC5304FCC9F8E1FC180DFA" box="[848,941,1910,1936]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
was found hidden between the young leaves of
<taxonomicName id="F2D5C5D3FFCC5304FEA9F80EFED30DD9" box="[304,358,1945,1971]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="07A16242FFCC5304FEA9F80EFED30DD9" box="[304,358,1945,1971]" italics="true" pageId="21" pageNumber="22">Aloe</emphasis>
</taxonomicName>
L. during the day (
<figureCitation id="ADEEA2D5FFCC5304FDC9F80EFD130DD9" box="[592,678,1945,1971]" captionStart="Fig" captionStartId="18.[189,232,1817,1843]" captionTargetBox="[264,1322,286,2013]" captionTargetId="figure@17.[244,1344,265,2019]" captionTargetPageId="17" captionText="Fig. 4 (opposite page). Habitus in natura, host plants and biotopes of Phlyctinus Schoenherr, 1826 species. A. P. callosus (Schoenherr, 1826). B. P. xerophilus Haran sp. nov. CD. P. grootbosensis Haran sp. nov. and its host plant, Osteospermum moniliferum L. EF. P. planithorax Haran sp. nov. and its host plant, Hymenolepis crithmifolia (L.) Greuter. GH. Biotope and host plant (Arctotis angustifolia L.) of P. littoralis Haran sp. nov. IJ. P. aloevorus Haran sp. nov. on its host plant, Aloe arborescens Mill., with details of feeding damage on floral peduncle" figureDoi="http://doi.org/10.5281/zenodo.3900306" httpUri="https://zenodo.org/record/3900306/files/figure.png" pageId="21" pageNumber="22">Fig. 4J</figureCitation>
), feeding mainly on flower peduncles (
<figureCitation id="ADEEA2D5FFCC5304FBE2F80EFB640DD9" box="[1147,1233,1945,1971]" captionStart="Fig" captionStartId="18.[189,232,1817,1843]" captionTargetBox="[264,1322,286,2013]" captionTargetId="figure@17.[244,1344,265,2019]" captionTargetPageId="17" captionText="Fig. 4 (opposite page). Habitus in natura, host plants and biotopes of Phlyctinus Schoenherr, 1826 species. A. P. callosus (Schoenherr, 1826). B. P. xerophilus Haran sp. nov. CD. P. grootbosensis Haran sp. nov. and its host plant, Osteospermum moniliferum L. EF. P. planithorax Haran sp. nov. and its host plant, Hymenolepis crithmifolia (L.) Greuter. GH. Biotope and host plant (Arctotis angustifolia L.) of P. littoralis Haran sp. nov. IJ. P. aloevorus Haran sp. nov. on its host plant, Aloe arborescens Mill., with details of feeding damage on floral peduncle" figureDoi="http://doi.org/10.5281/zenodo.3900306" httpUri="https://zenodo.org/record/3900306/files/figure.png" pageId="21" pageNumber="22">Fig. 4J</figureCitation>
). Adults were collected in March and November.
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCF5307FF24FE9CFE880B1D" pageId="22" pageNumber="23" type="distribution">
<paragraph id="356ABE50FFCF5307FF24FE9CFEE30B4F" blockId="22.[189,342,267,293]" box="[189,342,267,293]" pageId="22" pageNumber="23">
<heading id="6E22093CFFCF5307FF24FE9CFEE30B4F" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="07A16242FFCF5307FF24FE9CFEE30B4F" bold="true" box="[189,342,267,293]" pageId="22" pageNumber="23">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCF5307FF24FEAEFE880B1D" blockId="22.[189,1398,313,375]" pageId="22" pageNumber="23">
This species was only found at various limited locations on the
<collectingRegion id="F71170B2FFCF5307FC26FEADFC4E0B3E" box="[959,1019,314,340]" country="South Africa" name="Western Cape" pageId="22" pageNumber="23">Cape</collectingRegion>
coast between Bettys Bay and Hermanus.
</paragraph>
</subSubSection>
<caption id="61AAEED8FFCF5307FF24F814FAA70D89" ID-DOI="http://doi.org/10.5281/zenodo.3900308" ID-Zenodo-Dep="3900308" httpUri="https://zenodo.org/record/3900308/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1923,1949]" targetBox="[249,1378,546,1865]" targetPageId="22">
<paragraph id="356ABE50FFCF5307FF24F814FAA70D89" blockId="22.[189,1399,1923,2020]" pageId="22" pageNumber="23">
Fig. 5. PhyML tree of
<taxonomicName id="F2D5C5D3FFCF5307FE56F814FC930DF7" authority="Schoenherr, 1826" authorityName="Schoenherr" authorityYear="1826" box="[463,806,1923,1949]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="07A16242FFCF5307FE56F814FDFE0DF7" box="[463,587,1923,1949]" italics="true" pageId="22" pageNumber="23">Phlyctinus</emphasis>
<bibRefCitation id="5144C3A1FFCF5307FDCCF814FC930DF7" author="Schoenherr C. J." box="[597,806,1923,1949]" pageId="22" pageNumber="23" refId="ref16598" refString="Schoenherr C. J. 1826. Curculionidum Dispositio Methodica, cum Generum Characteribus, Descriptionibus atque Observationibus variis, seu Prodromus ad Synonymiae Insectorum, Partem IV. Fleischer, Leipzig [Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 9327" type="book" year="1826">Schoenherr, 1826</bibRefCitation>
</taxonomicName>
species inferred from mitochondrial Cytochrome oxidase I (
<emphasis id="07A16242FFCF5307FEA0F831FED90DAA" box="[313,364,1958,1984]" italics="true" pageId="22" pageNumber="23">COI</emphasis>
, 658 base pairs). Bootstrap support values as obtained from 1000 replicates (*above 80, ** above 95). Specimens codes were shortened for representation: JHAR0 = J and _0101 = -1.
</paragraph>
</caption>
<subSubSection id="7DCFEDDBFFCE5306FF24FE9CFC0D0BD4" pageId="23" pageNumber="24" type="discussion">
<paragraph id="356ABE50FFCE5306FF24FE9CFE9A0B4F" blockId="23.[189,303,267,293]" box="[189,303,267,293]" pageId="23" pageNumber="24">
<heading id="6E22093CFFCE5306FF24FE9CFE9A0B4F" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="07A16242FFCE5306FF24FE9CFE9A0B4F" bold="true" box="[189,303,267,293]" pageId="23" pageNumber="24">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCE5306FF24FEAEFC0D0BD4" blockId="23.[189,1399,313,446]" pageId="23" pageNumber="24">
<taxonomicName id="F2D5C5D3FFCE5306FF24FEAEFE010B39" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,436,313,339]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCE5306FF24FEAEFE010B39" box="[189,436,313,339]" italics="true" pageId="23" pageNumber="24">Phlyctinus aloevorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCE5306FE25FEADFDA20B3E" box="[444,535,314,340]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="F2D5C5D3FFCE5306FD52FEADFCF80B39" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[715,845,313,339]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCE5306FD52FEADFCF80B39" box="[715,845,313,339]" italics="true" pageId="23" pageNumber="24">P. littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCE5306FCCCFEADFC050B3E" box="[853,944,314,340]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
species group, in which it is closest to
<taxonomicName id="F2D5C5D3FFCE5306FF24FECAFED60B1D" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,355,349,375]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="planithorax" status="sp. nov.">
<emphasis id="07A16242FFCE5306FF24FECAFED60B1D" box="[189,355,349,375]" italics="true" pageId="23" pageNumber="24">P. planithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCE5306FEF3FECAFE700B1D" box="[362,453,349,375]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
(see discussion under that species for distinguishing characters). It can easily be distinguished from specimens of
<taxonomicName id="F2D5C5D3FFCE5306FDDAFE16FD730BF0" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[579,710,384,410]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCE5306FDDAFE16FD730BF0" box="[579,710,384,410]" italics="true" pageId="23" pageNumber="24">P. littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCE5306FD54FE17FC9D0BF0" box="[717,808,384,410]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
by the presence of a distinct median carina on the prothorax. Both species show an uncorrected p-distance of 5.3%.
</paragraph>
</subSubSection>
</treatment>
</document>