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<document id="A0BD45C1C38095383BEAB7C59D712DFC" ID-DOI="10.11646/zootaxa.3670.2.11" ID-GBIF-Dataset="bc516484-c1c5-4ee3-8912-7ff89f906a8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="219876" ID-ZooBank="67402B31-9D14-4116-8752-2927A1084240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633918393" checkinUser="plazi" docAuthor="Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando &amp; Oromí, Pedro" docDate="2013" docId="015C614EF409FF9D5C94F895061FFCBA" docLanguage="en" docName="zt03670p299.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Roncocreagris blothroides Beier 1962" docType="treatment" docVersion="8" lastPageNumber="289" masterDocId="FD651936F40BFF9B5C03FFB30766FFAE" masterDocTitle="On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species" masterLastPageNumber="299" masterPageNumber="283" pageNumber="285" updateTime="1698334986200" updateUser="plazi">
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<mods:title id="BAE787AD7A8CD9E86BEC087988584A8D">On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species</mods:title>
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<mods:namePart id="6BE13890998404D37C2F4C1FCE8B48B0">Reboleira, Ana Sofia P. S.</mods:namePart>
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<mods:namePart id="696DE8D5FE0A592C6D9AB280FBD89032">Zaragoza, Juan A.</mods:namePart>
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<mods:namePart id="BCF8B17136B94EB7BBFA0DC70E54C171">Oromí, Pedro</mods:namePart>
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<treatment id="015C614EF409FF9D5C94F895061FFCBA" ID-DOI="http://doi.org/10.5281/zenodo.6146628" ID-GBIF-Taxon="119570118" ID-Zenodo-Dep="6146628" LSID="urn:lsid:plazi:treatment:015C614EF409FF9D5C94F895061FFCBA" httpUri="http://treatment.plazi.org/id/015C614EF409FF9D5C94F895061FFCBA" lastPageId="6" lastPageNumber="289" pageId="2" pageNumber="285">
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<paragraph id="894AD058F409FF995C94F895051FF8EE" blockId="2.[151,633,1830,1889]" box="[151,633,1830,1856]" pageId="2" pageNumber="285">
<heading id="D2026734F409FF995C94F895051FF8EE" bold="true" box="[151,633,1830,1856]" fontSize="11" level="1" pageId="2" pageNumber="285" reason="1">
<taxonomicName id="4EF5ABDBF409FF995C94F895051FF8EE" ID-CoL="4TB43" authority="Beier, 1962" authorityName="Beier" authorityYear="1962" box="[151,633,1830,1856]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF409FF995C94F895051FF8EE" bold="true" box="[151,633,1830,1856]" pageId="2" pageNumber="285">
<emphasis id="BB810C4AF409FF995C94F89506B3F8EE" bold="true" box="[151,469,1830,1856]" italics="true" pageId="2" pageNumber="285">Roncocreagris blothroides</emphasis>
(
<bibRefCitation id="ED64ADA9F409FF995DE7F8950516F8EE" author="Beier" box="[484,624,1830,1856]" pageId="2" pageNumber="285" refString="Beier, M. (1962) Eine neue Microcreagris aus Portugal. Voyage au Portugal du Dr. K. Lindberg. Boletim da Sociedade Portuguesa de Ciencias Naturais, (2) 9, 25 - 26." type="journal article" year="1962">Beier, 1962</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="894AD058F409FF995C94F8FA0671F8CF" blockId="2.[151,633,1830,1889]" box="[151,279,1865,1889]" pageId="2" pageNumber="285">
(
<figureCitation id="11CECCDDF409FF995C9CF8FA0669F8CF" box="[159,271,1865,1889]" captionStart-0="FIGURE 2" captionStart-1="FIGURES 3 12" captionStartId-0="4.[151,255,853,877]" captionStartId-1="5.[151,269,1863,1887]" captionTargetBox-0="[151,1436,194,832]" captionTargetBox-1="[168,1411,187,1831]" captionTargetId-0="figure@4.[151,1436,194,832]" captionTargetId-1="figure@5.[151,1436,166,1851]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, Sicó massif, habitus of living specimen, dorsal view." captionText-1="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri-0="https://zenodo.org/record/219878/files/figure.png" httpUri-1="https://zenodo.org/record/219879/files/figure.png" pageId="2" pageNumber="285">Figs 211</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F409FF995C94F82105AEF86B" pageId="2" pageNumber="285" type="reference_group">
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<taxonomicName id="4EF5ABDBF409FF995C94F821055AF809" authority="Beier 1962: 25" authorityName="Beier" authorityPageNumber="25" authorityYear="1962" box="[151,572,1937,1959]" class="Arachnida" family="Neobisiidae" genus="Microcreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF409FF995C94F82106C6F809" box="[151,416,1937,1959]" italics="true" pageId="2" pageNumber="285">Microcreagris blothroides</emphasis>
<treatmentCitation id="0854F649F409FF995DA5F822055AF809" author="Beier" box="[422,572,1937,1959]" page="25" pageId="2" pageNumber="285" year="1962">
<bibRefCitation id="ED64ADA9F409FF995DA5F822057CF809" author="Beier" box="[422,538,1937,1959]" pageId="2" pageNumber="285" refString="Beier, M. (1962) Eine neue Microcreagris aus Portugal. Voyage au Portugal du Dr. K. Lindberg. Boletim da Sociedade Portuguesa de Ciencias Naturais, (2) 9, 25 - 26." type="journal article" year="1962">Beier 1962</bibRefCitation>
: 25
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26, fig. 1.
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</paragraph>
<paragraph id="894AD058F409FF995C94F80205AEF86B" blockId="2.[151,712,1937,1990]" box="[151,712,1967,1990]" pageId="2" pageNumber="285">
<treatmentCitationGroup id="A9E5F776F409FF995C94F80205AEF86B" box="[151,712,1967,1990]" pageId="2" pageNumber="285">
<taxonomicName id="4EF5ABDBF409FF995C94F8020693F86B" ID-CoL="4TB43" authority="Beier" authorityName="Beier" box="[151,501,1967,1990]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF409FF995C94F80206C5F86B" box="[151,419,1967,1990]" italics="true" pageId="2" pageNumber="285">Roncocreagris blothroides</emphasis>
(Beier)
</taxonomicName>
:
<treatmentCitation id="0854F649F409FF995DFCF81C05A2F86B" author="Mahnert" box="[511,708,1967,1989]" page="212" pageId="2" pageNumber="285" year="1976">
<bibRefCitation id="ED64ADA9F409FF995DFCF81C05F4F86B" author="Mahnert" box="[511,658,1967,1989]" pageId="2" pageNumber="285" refString="Mahnert, V. (1976) Zur Kenntnis der Gattungen Acanthocreagris und Roncocreagris (Arachnida, Pseudoscorpiones, Neobisiidae). Revue suisse de Zoologie, 83, 193 - 214." type="journal article" year="1976">Mahnert 1976</bibRefCitation>
: 212
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="BB810C4AF408FF985C94FF240660FF1E" bold="true" box="[151,262,151,176]" pageId="3" pageNumber="286">Material.</emphasis>
<collectingCountry id="F1E290C8F408FF985D15FF24061EFF01" box="[278,376,151,175]" name="Portugal" pageId="3" pageNumber="286">Portugal</collectingCountry>
, Sicó massif, district of Leiria, three caves: Santiago da Guarda, Gruta da Cerâmica (39º55ʹ36.57ʺN, 8º31ʹ0 3.63ʺW;
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.
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a.s.l.),
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, 1 3 (
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),
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, 2 Ƥ (1
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), 1 3 (
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); Redinha, Gruta de Santa Maria da Estrela (39º55ʹ41.15ʺN, 8º32ʹ59.4ʺW;
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.
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a.s.l.),
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, 1 3 (
<collectionCode id="EFE4489DF408FF985E36FEFF05EFFECA" box="[565,649,332,356]" pageId="3" pageNumber="286">DEUA</collectionCode>
); Abiúl, Algar da Confraria I (39°55ʹ51.01&quot;N, 8°31ʹ35.64&quot;W;
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.
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a.s.l.),
<date id="FD4BF698F408FF985CFCFEDC06ECFE29" box="[255,394,367,391]" pageId="3" pageNumber="286" value="2007-07-22">22.VII.2007</date>
, 1 Ƥ (
<collectionCode id="EFE4489DF408FF985DCCFEDC0545FE29" box="[463,547,367,391]" pageId="3" pageNumber="286">DEUA</collectionCode>
). All specimens lgt. A.S.P.S. Reboleira.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F408FF985CC4FE2006E9FDCB" pageId="3" pageNumber="286" type="diagnosis">
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<emphasis id="BB810C4AF408FF985CC4FE200627FE02" bold="true" box="[199,321,403,428]" pageId="3" pageNumber="286">Diagnosis.</emphasis>
No eyes or eye-spots. Extreme troglomorphic adaptations. Carapace usually with 18 setae, of which 4 on posterior margin. Tergite I with 4 setae. Galeal chelicera short in both sexes, with 35 tiny apical rami. Pedipalp: femur ratio about 8.0, femur roughly same length as the patella and almost as long as the movable chelal finger; chela+ ratio about 8.0 in males, 7.0
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females; chelal hand maximum width distal of middle, ratio movable finger/hand+ about 1.7; trichobothrium
<emphasis id="BB810C4AF408FF985EB2FD9905AAFDEF" box="[689,716,554,577]" italics="true" pageId="3" pageNumber="286">ist</emphasis>
about halfway between
<emphasis id="BB810C4AF408FF985FDEFD9A0495FDEF" box="[989,1011,553,577]" italics="true" pageId="3" pageNumber="286">ib</emphasis>
and
<emphasis id="BB810C4AF408FF98582AFD99035EFDEF" box="[1065,1080,554,577]" italics="true" pageId="3" pageNumber="286">it</emphasis>
, ratio
<emphasis id="BB810C4AF408FF985882FD9A03D9FDEF" box="[1153,1215,553,577]" italics="true" pageId="3" pageNumber="286">ibist</emphasis>
/
<emphasis id="BB810C4AF408FF9858C4FD990398FDEF" box="[1223,1278,554,577]" italics="true" pageId="3" pageNumber="286">istit</emphasis>
0.91.2; ratio
<emphasis id="BB810C4AF408FF985C94FDFD07B9FDC8" box="[151,223,590,614]" italics="true" pageId="3" pageNumber="286">isbist</emphasis>
/
<emphasis id="BB810C4AF408FF985CEBFDFD064AFDC8" box="[232,300,590,614]" italics="true" pageId="3" pageNumber="286">ibisb</emphasis>
0.81.0.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F408FF9F5CC4FDC204F8F9B2" lastPageId="4" lastPageNumber="287" pageId="3" pageNumber="286" type="description">
<paragraph id="894AD058F408FF985CC4FDC20441FD03" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
<emphasis id="BB810C4AF408FF985CC4FDC2063FFD24" bold="true" box="[199,345,625,650]" pageId="3" pageNumber="286">Description.</emphasis>
<emphasis id="BB810C4AF408FF985D60FDC2041BFD27" box="[355,893,625,649]" italics="true" pageId="3" pageNumber="286">Males, followed by females in square brackets</emphasis>
. Large species (
<figureCitation id="11CECCDDF408FF98583EFDC203E0FD27" box="[1085,1158,625,649]" captionStart="FIGURE 2" captionStartId="4.[151,255,853,877]" captionTargetBox="[151,1436,194,832]" captionTargetId="figure@4.[151,1436,194,832]" captionTargetPageId="4" captionText="FIGURE 2. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, Sicó massif, habitus of living specimen, dorsal view." httpUri="https://zenodo.org/record/219878/files/figure.png" pageId="3" pageNumber="286">Fig. 2</figureCitation>
). Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps pale brown.
</paragraph>
<paragraph id="894AD058F408FF985CC4FD0A03EBFCB7" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Carapace markedly longer than broad (
<figureCitation id="11CECCDDF408FF985E9CFD0A058CFD7F" box="[671,746,697,721]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Fig. 3</figureCitation>
). Without eyes or eye-spots. Anterior margin moderately prominent medially, with blunt, low epistome with some tiny denticles (
<figureCitation id="11CECCDDF408FF985FC7FD6E036DFD5B" box="[964,1035,733,757]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Fig. 4</figureCitation>
). Chaetotaxy: 1819 setae, formula 4:4:6:45. Four microlyrifissures in ocular zone, two between median and posterior zones.
</paragraph>
<paragraph id="894AD058F408FF985CC4FC9607B0FC2B" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Coxal area. Manducatory process with 3 setae. Anterior process of coxa I long and apically pointed; medial process straight, with some low tubercles (
<figureCitation id="11CECCDDF408FF985E76FCFA05DCFCCF" box="[629,698,841,865]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Fig. 5</figureCitation>
). Pedipalpal coxa with 7 setae, pedal coxa
<date id="FD4BF698F408FF98589EFCF903BCFCCF" box="[1181,1242,841,865]" pageId="3" pageNumber="286">I 57</date>
, II: 56, III: 34, IV: 6.
</paragraph>
<paragraph id="894AD058F408FF985CC4FC2203DCFBF3" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Tergal chaetotaxy IX: 35:6:67:78:7:7:7:7:7:7 [46:67:67:78:78:710:78:78:68:7]. Male genital area with 1011 setae on sternite II; sternite III with 1116 setae, 6 of them along posterior margin of genital opening. Female genital area with 56 microsetae on sternite II, sternite III with 810 setae. Sternal chaetotaxy IV X: 78:11:1112:1112:1011:910:9 [67:1011:1011:1113:1011:10:9], each sternite VI and VII with 2 discal setae (included in sternal formula). Segment XI: 910 setae, including 6 TS. Chaetotaxy of stigmata of sternites
<date id="FD4BF698F408FF985CF8FBF506FBFBF3" box="[251,413,1093,1117]" pageId="3" pageNumber="286">III and IV 13</date>
and 13 respectively. Anal cone with two dorsal and two ventral setae.
</paragraph>
<paragraph id="894AD058F408FF985CC4FBDA05DDFA9B" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Chelicerae (
<figureCitation id="11CECCDDF408FF985D4CFBDA06D1FB2F" box="[335,439,1129,1153]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Figs 67</figureCitation>
). Palm with 6 setae, subgaleal seta 0.630.68 from base of movable finger. Galea short (length
<quantity id="4E0D7DBDF408FF985CEDFB3E0630FB0B" box="[238,342,1165,1189]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="286" unit="mm" value="0.04">0.04 mm</quantity>
), apically with 35 short rami, about same length and shape in both sexes. Fixed finger with 10 15 medium or small teeth, two medial teeth of them larger than the others; movable finger with 813 teeth, one large and blunt medial tooth, the others medium or small, dental row ending proximad to subgaleal seta. Rallum with 78 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 3336 blades, serrula interior 2731 blades.
</paragraph>
<paragraph id="894AD058F408FF985CC4FAF20384F89E" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Pedipalps (
<figureCitation id="11CECCDDF408FF985D49FAF206A0FAF7" box="[330,454,1345,1369]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Figs 810</figureCitation>
). Trochanter, femur, distal third portion of the patella and chelal hand with low granulation, more pronounced on paraxial faces. Lyrifissures as in
<figureCitation id="11CECCDDF408FF985F89FAD6049BFAD3" box="[906,1021,1381,1405]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Figs 810</figureCitation>
. Femur with one tiny tubercle distad of middle on antiaxial face, 23 distal glandular pores present. Patella dorsally with one micropore at base of pedicel and 12 distal glandular pores. Chelal hand dorsally more or less parallel-sided, maximum width distad of middle; pedicel dorsal bears 23 dorsal micropores; antiaxial face with an irregular row of 35 glandular pores close to base of finger. Fixed finger with 139148 teeth, most of them apically cusped, dental row reaching to level of trichobothrium
<emphasis id="BB810C4AF408FF985D64F9AF06EAF99A" box="[359,396,1564,1588]" italics="true" pageId="3" pageNumber="286">esb</emphasis>
;
<emphasis id="BB810C4AF408FF985D9FF9AF0520F99A" box="[412,582,1564,1588]" italics="true" pageId="3" pageNumber="286">nodus ramosus</emphasis>
short, reaching level of 35th distal teeth; trichobothrium
<emphasis id="BB810C4AF408FF9858D4F9AE0381F99A" box="[1239,1255,1565,1588]" italics="true" pageId="3" pageNumber="286">it</emphasis>
proximad to
<emphasis id="BB810C4AF408FF985982F9AE02F3F99A" box="[1409,1429,1565,1588]" italics="true" pageId="3" pageNumber="286">et</emphasis>
, about half way between
<emphasis id="BB810C4AF408FF985DB3F9F206A9F9F6" box="[432,463,1601,1624]" italics="true" pageId="3" pageNumber="286">est</emphasis>
and
<emphasis id="BB810C4AF408FF985E0AF9F2057BF9F6" box="[521,541,1601,1624]" italics="true" pageId="3" pageNumber="286">et</emphasis>
; trichobothrium
<emphasis id="BB810C4AF408FF985EDDF9F2059EF9F6" box="[734,760,1601,1624]" italics="true" pageId="3" pageNumber="286">ist</emphasis>
closer to the base of finger than to apex; distance between trichobothria
<emphasis id="BB810C4AF408FF985D32F9D70621F9D2" box="[305,327,1636,1660]" italics="true" pageId="3" pageNumber="286">ib</emphasis>
and
<emphasis id="BB810C4AF408FF985D80F9D606F8F9D2" box="[387,414,1637,1660]" italics="true" pageId="3" pageNumber="286">ist</emphasis>
0.91.2 times that between
<emphasis id="BB810C4AF408FF985EE4F9D60467F9D2" box="[743,769,1637,1660]" italics="true" pageId="3" pageNumber="286">ist</emphasis>
and
<emphasis id="BB810C4AF408FF985F3DF9D6042BF9D2" box="[830,845,1637,1660]" italics="true" pageId="3" pageNumber="286">it</emphasis>
; distance between trichobothria
<emphasis id="BB810C4AF408FF9858C5F9D70381F9D2" box="[1222,1255,1636,1660]" italics="true" pageId="3" pageNumber="286">isb</emphasis>
and
<emphasis id="BB810C4AF408FF985927F9D60259F9D2" box="[1316,1343,1637,1660]" italics="true" pageId="3" pageNumber="286">ist</emphasis>
0.81.0 times that between
<emphasis id="BB810C4AF408FF985D73F93B06F7F90E" box="[368,401,1672,1696]" italics="true" pageId="3" pageNumber="286">isb</emphasis>
and
<emphasis id="BB810C4AF408FF985DC4F93B06BBF90E" box="[455,477,1672,1696]" italics="true" pageId="3" pageNumber="286">ib</emphasis>
. Chelal microsetae pattern (as defined by
<bibRefCitation id="ED64ADA9F408FF985FB4F93A0305F90E" author="Zaragoza" box="[951,1123,1672,1697]" pageId="3" pageNumber="286" refString="Zaragoza, J. A. (2008) On the status of the subspecies of Roncocreagris galeonuda (Pseudoscorpiones: Neobisiidae): importance of the chelal microsetae pattern. Remarks on the genus Roncocreagris Mahnert. Revista Iberica de Aracnologia, 15, 35 - 46." type="journal article" year="2008">Zaragoza 2008</bibRefCitation>
): all groups present,
<emphasis id="BB810C4AF408FF98594FF93A0217F90E" box="[1356,1393,1673,1696]" italics="true" pageId="3" pageNumber="286">Em</emphasis>
: 4 6 curved setae (same length as normal setae),
<emphasis id="BB810C4AF408FF985EA0F91E05B1F96A" box="[675,727,1709,1732]" italics="true" pageId="3" pageNumber="286">Mm:</emphasis>
23,
<emphasis id="BB810C4AF408FF985F19F91E0451F96A" box="[794,823,1709,1732]" italics="true" pageId="3" pageNumber="286">Im</emphasis>
: 1. Movable finger with 132141 teeth, most of them apically cusped, dental row shorter than on fixed finger, ending just distad of trichobothrium
<emphasis id="BB810C4AF408FF9858BFF96303ACF946" box="[1212,1226,1744,1768]" italics="true" pageId="3" pageNumber="286">b</emphasis>
; distance between trichobothria
<emphasis id="BB810C4AF408FF985D32F947062CF8A2" box="[305,330,1780,1804]" italics="true" pageId="3" pageNumber="286">sb</emphasis>
and
<emphasis id="BB810C4AF408FF985D86F94606FEF8A2" box="[389,408,1781,1804]" italics="true" pageId="3" pageNumber="286">st</emphasis>
1.31.8 times that between
<emphasis id="BB810C4AF408FF985EDCF947059EF8A2" box="[735,760,1780,1804]" italics="true" pageId="3" pageNumber="286">sb</emphasis>
and
<emphasis id="BB810C4AF408FF985F37F9470424F8A2" box="[820,834,1780,1804]" italics="true" pageId="3" pageNumber="286">b</emphasis>
. One sensillum near tip of both fingers; one diploid sensillum
<emphasis id="BB810C4AF408FF985D0AF8AA0645F89E" box="[265,291,1817,1840]" italics="true" pageId="3" pageNumber="286">pc</emphasis>
close to dental margin and not raised, close to or slightly distad of trichobothrium
<emphasis id="BB810C4AF408FF9858C1F8AB03BDF89E" box="[1218,1243,1816,1840]" italics="true" pageId="3" pageNumber="286">sb</emphasis>
.
</paragraph>
<paragraph id="894AD058F408FF985CC4F88E06D7F832" blockId="3.[151,1437,151,1948]" pageId="3" pageNumber="286">
Legs. Claws of legs I and IV with a tiny dorsal tooth proximad of middle (
<figureCitation id="11CECCDDF408FF98580BF88F033FF8FA" box="[1032,1113,1852,1876]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Fig. 11</figureCitation>
), subterminal setae with 34 rami (
<figureCitation id="11CECCDDF408FF985CDCF8D3065EF8D6" box="[223,312,1888,1912]" captionStart="FIGURES 3 12" captionStartId="5.[151,269,1863,1887]" captionTargetBox="[168,1411,187,1831]" captionTargetId="figure@5.[151,1436,166,1851]" captionTargetPageId="5" captionText="FIGURES 3 12. Roncocreagris blothroides (Beier, 1962), Gruta da Cerâmica, male. 3. carapace; 4. anterior margin of carapace; 5. anterior and medial processes of left coxa I; 6. left chelicera; 7. apex of movable finger of left chelicera, showing galea; 8. right pedipalp minus chela, dorsal view; 9. right chela, dorsal view; 10. right chela, lateral view; 11. claw of right leg IV; 12. subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 8 10)." httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="3" pageNumber="286">Fig. 12</figureCitation>
). Leg IV tibia with four long setae, TS ratio 0.330.49 for longest, basitarsus TS: 0.120.14, telotarsus TS: 0.210.38.
</paragraph>
<caption id="DD8A80D0F40FFF9F5C94FCE607B7FC23" httpUri="https://zenodo.org/record/219878/files/figure.png" pageId="4" pageNumber="287" targetBox="[151,1436,194,832]" targetPageId="4">
<paragraph id="894AD058F40FFF9F5C94FCE607B7FC23" blockId="4.[151,1436,853,909]" pageId="4" pageNumber="287">
<emphasis id="BB810C4AF40FFF9F5C94FCE6067EFCC3" bold="true" box="[151,280,853,877]" pageId="4" pageNumber="287">FIGURE 2.</emphasis>
<taxonomicName id="4EF5ABDBF40FFF9F5D1CFCE505A8FCC3" authority="Beier, 1962" authorityName="Beier" authorityYear="1962" box="[287,718,854,877]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40FFF9F5D1CFCE5055CFCC3" box="[287,570,854,877]" italics="true" pageId="4" pageNumber="287">Roncocreagris blothroides</emphasis>
(Beier, 1962)
</taxonomicName>
, Gruta da Cerâmica, Sicó massif, habitus of living specimen, dorsal view.
</paragraph>
</caption>
<paragraph id="894AD058F40FFF9F5CC4FC0504F8F9B2" blockId="4.[151,1437,949,2033]" pageId="4" pageNumber="287">
<emphasis id="BB810C4AF40FFF9F5CC4FC050698FC60" bold="true" box="[199,510,949,974]" pageId="4" pageNumber="287">Measurements and ratios.</emphasis>
<emphasis id="BB810C4AF40FFF9F5E06FC050368FC60" box="[517,1038,950,974]" italics="true" pageId="4" pageNumber="287">Males, followed by females in square brackets</emphasis>
. Body 3.083.1. Carapace 1.01.2/ 0.70.79 (1.41.5) [1.161.22/0.790.87 (1.41.5)]. Chelicera: palm 0.650.72/0.30.4 (2.1) [0.70.73/0.350.37 (2.0)], movable finger 0.410.45 [0.460.49]. Pedipalp: trochanter 0.931.02/0.230.24 (4.04.3) [1.021.04/ 0.240.26 (4.04.3)]; femur 1.842.12/0.230.26 (8.08.2) [2.02.09/0.290.3 (7.68.1)]; patella 1.752.1/0.26 0.3 (6.87.1) [2.022.09/0.290.3 (6.87.2)], pedicel 0.440.5 [0.490.51], club 1.311.6/0.260.3 (5.15.4) [1.531.59/0.290.3 (5.15.5)], ratio club/pedicel: 3.03.2 [3.13.2]; chela+ 2.983.38/0.380.42 (7.98.1) [3.35 3.44/0.440.48 (7.17.8)], chela - 2.823.16/0.380.42 (7.57.6) [3.153.23/0.440.48 (6.67.3)]; hand+ 1.151.26 (3.03.1) [1.281.31 (2.72.9)], hand - 0.991.04 (2.52.6) [1.071.1 (2.32.4)]; movable finger 1.852.15 [2.11 2.19]; ratio finger/hand+ 1.61.7 [1.61.7)]; ratio chela+/carapace 2.83.0 [2.83.0]; femur/carapace 1.8 [1.61.8]; finger/femur 1.0 [1.01.1]; femur/patella 1.011.05 [1.0]; patella/hand+ 1.51.7 [1.51.6]. Leg I: femur 0.91.05/ 0.140.15 (6.46.9) [1.01.07/0.150.16 (6.66.7)]; patella 0.700.84/0.120.13 (6.06.5) [0.740.84/0.130.14 (5.76.2)]; tibia 0.861.02/.090.1 (9.410.7) [0.981.02/0.1 (10.210.4)]; basitarsus 0.470.56/0.080.09 (6.0 6.6) [0.520.53/0.08 (6.5/6.7)]; telotarsus 0.600.71/0.07 (9.09.9) [0.660.7/0.07 (9.19.4)]; ratio femur/patella 1.3 [1.31.4]; telotarsus/basitarsus 1.3 [1.3]. Leg IV: femur+patella 1.581.78/0.210.22 (7.68.1) [1.71.8/0.21 0.23 (7.58.8)]; tibia 1.551.79/0.130.14 (11.913.0) [1.681.74/0.140.15 (11.812.1)]; basitarsus 0.590.69/ 0.10.11 (6.16.5) [0.650.7/0.11 (6.06.4)]; telotarsus 0.710.84/0.080.09 (8.49.3) [0.820.87/0.09 (9.09.3)]; ratio femur+patella/tibia 1.0 [1.0]; telotarsus/basitarsus 1.2 [1.21.3].
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F40FFF9D5CC4F99B02FAFE02" lastPageId="6" lastPageNumber="289" pageId="4" pageNumber="287" type="discussion">
<paragraph id="894AD058F40FFF9F5CC4F99B0306F95A" blockId="4.[151,1437,949,2033]" pageId="4" pageNumber="287">
<emphasis id="BB810C4AF40FFF9F5CC4F99B065DF9EE" bold="true" box="[199,315,1576,1600]" pageId="4" pageNumber="287">Remarks.</emphasis>
<bibRefCitation id="ED64ADA9F40FFF9F5D41F99B06B6F9EE" author="Beier" box="[322,464,1576,1600]" pageId="4" pageNumber="287" refString="Beier, M. (1962) Eine neue Microcreagris aus Portugal. Voyage au Portugal du Dr. K. Lindberg. Boletim da Sociedade Portuguesa de Ciencias Naturais, (2) 9, 25 - 26." type="journal article" year="1962">Beier (1962)</bibRefCitation>
described the tarsal claws of
<taxonomicName id="4EF5ABDBF40FFF9F5F1FF99A04DCF9EE" box="[796,954,1576,1600]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40FFF9F5F1FF99A04DCF9EE" box="[796,954,1576,1600]" italics="true" pageId="4" pageNumber="287">R. blothroides</emphasis>
</taxonomicName>
as lacking a dorsal tooth and used this as a characteristic to separate this species from
<taxonomicName id="4EF5ABDBF40FFF9F5E84F9FE044AF9CA" box="[647,812,1612,1636]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="BB810C4AF40FFF9F5E84F9FE044AF9CA" box="[647,812,1612,1636]" italics="true" pageId="4" pageNumber="287">R. cavernicola</emphasis>
</taxonomicName>
; as did
<bibRefCitation id="ED64ADA9F40FFF9F5F8FF9FF0327F9CA" author="Mahnert" box="[908,1089,1612,1636]" pageId="4" pageNumber="287" refString="Mahnert, V. (1976) Zur Kenntnis der Gattungen Acanthocreagris und Roncocreagris (Arachnida, Pseudoscorpiones, Neobisiidae). Revue suisse de Zoologie, 83, 193 - 214." type="journal article" year="1976">Mahnert (1976)</bibRefCitation>
, based on Beiers description.
<bibRefCitation id="ED64ADA9F40FFF9F5C94F9C20637F926" author="Zaragoza" box="[151,337,1648,1673]" pageId="4" pageNumber="287" refString="Zaragoza, J. A. (2002) Dos nuevos Roncocreagris cavernicolas de Galicia (Arachnida, Pseudoscorpiones, Neobisiidae). Revista Iberica de Aracnologia, 5, 91 - 98." type="journal article" year="2002">Zaragoza (2002)</bibRefCitation>
suggested that, due to the tiny size of the teeth in some
<taxonomicName id="4EF5ABDBF40FFF9F5FC7F9C2030EF926" box="[964,1128,1649,1672]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF40FFF9F5FC7F9C2030EF926" box="[964,1128,1649,1672]" italics="true" pageId="4" pageNumber="287">Roncocreagris</emphasis>
</taxonomicName>
species, it was not difficult to imagine that Beier might have overlooked them in
<taxonomicName id="4EF5ABDBF40FFF9F5F13F92604D4F902" box="[784,946,1684,1708]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40FFF9F5F13F92604D4F902" box="[784,946,1684,1708]" italics="true" pageId="4" pageNumber="287">R. blothroides</emphasis>
</taxonomicName>
. Given the agreement in all other studied characters, we have no doubt that the specimens described above belong to
<taxonomicName id="4EF5ABDBF40FFF9F5813F90A03D7F97E" box="[1040,1201,1720,1744]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40FFF9F5813F90A03D7F97E" box="[1040,1201,1720,1744]" italics="true" pageId="4" pageNumber="287">R. blothroides</emphasis>
</taxonomicName>
, hence the absence/ presence of teeth on the tarsal claws is not a valid diagnostic character for this species.
</paragraph>
<paragraph id="894AD058F40FFF9F5CC4F8B30624F806" blockId="4.[151,1437,949,2033]" pageId="4" pageNumber="287">
Beiers original diagnosis (1962) of
<taxonomicName id="4EF5ABDBF40FFF9F5E6BF8B20461F8B6" box="[616,775,1792,1816]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40FFF9F5E6BF8B20461F8B6" box="[616,775,1792,1816]" italics="true" pageId="4" pageNumber="287">R. blothroides</emphasis>
</taxonomicName>
stated that it bears 4 setae on the posterior margin of the carapace and 4 setae on tergite I. Of the
<specimenCount id="9FF31BD1F40FFF9F5E55F897058BF892" box="[598,749,1828,1852]" pageId="4" pageNumber="287" type="generic">24 specimens</specimenCount>
checked, 16 coincide with this (4+4), two bear 4+3, two 4+5, one 4+6, one 5+4, and two 5+5. No specimens bear 6 or more setae on posterior margin of carapace, and this number of setae is very rare on tergite I. This pattern is important for separating taxonomic groups within the genus
<taxonomicName id="4EF5ABDBF40FFF9F5C94F822065DF806" box="[151,315,1937,1960]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="BB810C4AF40FFF9F5C94F822065DF806" box="[151,315,1937,1960]" italics="true" pageId="4" pageNumber="287">Roncocreagris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="894AD058F40FFF9F5CC4F8070427F85E" blockId="4.[151,1437,949,2033]" pageId="4" pageNumber="287">
The discal setae of the sternites are glandular with a visible duct, as first described and illustrated by
<bibRefCitation id="ED64ADA9F40FFF9F594DF8070787F85E" author="Judson" pageId="4" pageNumber="287" refString="Judson, M. L. I. (1992) Roncocreagris murphyorum n. sp. and Occitanobisium nanum (Beier) n. comb. (Neobisiidae) from Iberia, with notes on the sternal glands of pseudoscorpions (Chelonethi). Bulletin of the British Arachnological Society, 9, 26 - 30." type="journal article" year="1992">Judson (1992)</bibRefCitation>
and also mentioned and illustrated by
<bibRefCitation id="ED64ADA9F40FFF9F5E90F86A0458F85E" author="Zaragoza" box="[659,830,2008,2033]" pageId="4" pageNumber="287" refString="Zaragoza, J. A. (2002) Dos nuevos Roncocreagris cavernicolas de Galicia (Arachnida, Pseudoscorpiones, Neobisiidae). Revista Iberica de Aracnologia, 5, 91 - 98." type="journal article" year="2002">Zaragoza 2002</bibRefCitation>
.
</paragraph>
<caption id="DD8A80D0F40EFF9E5C94F8F40563F870" httpUri="https://zenodo.org/record/219879/files/figure.png" pageId="5" pageNumber="288" targetBox="[168,1411,187,1831]" targetPageId="5">
<paragraph id="894AD058F40EFF9E5C94F8F40563F870" blockId="5.[151,1436,1863,2014]" pageId="5" pageNumber="288">
<emphasis id="BB810C4AF40EFF9E5C94F8F40637F8F1" bold="true" box="[151,337,1863,1887]" pageId="5" pageNumber="288">FIGURES 312.</emphasis>
<taxonomicName id="4EF5ABDBF40EFF9E5D5AF8F4046BF8F0" authority="Beier, 1962" authorityName="Beier" authorityYear="1962" box="[345,781,1863,1886]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40EFF9E5D5AF8F40513F8F0" box="[345,629,1863,1886]" italics="true" pageId="5" pageNumber="288">Roncocreagris blothroides</emphasis>
(Beier, 1962)
</taxonomicName>
, Gruta da Cerâmica, male.
<emphasis id="BB810C4AF40EFF9E5836F8F4032EF8F1" bold="true" box="[1077,1096,1863,1887]" pageId="5" pageNumber="288">3.</emphasis>
carapace;
<emphasis id="BB810C4AF40EFF9E58BFF8F403B6F8F1" bold="true" box="[1212,1232,1863,1887]" pageId="5" pageNumber="288">4.</emphasis>
anterior margin of carapace;
<emphasis id="BB810C4AF40EFF9E5D07F8D4067FF8D1" bold="true" box="[260,281,1895,1919]" pageId="5" pageNumber="288">5.</emphasis>
anterior and medial processes of left coxa I;
<emphasis id="BB810C4AF40EFF9E5F0FF8D40479F8D1" bold="true" box="[780,799,1895,1919]" pageId="5" pageNumber="288">6.</emphasis>
left chelicera;
<emphasis id="BB810C4AF40EFF9E5FC6F8D404BFF8D1" bold="true" box="[965,985,1895,1919]" pageId="5" pageNumber="288">7.</emphasis>
apex of movable finger of left chelicera, showing galea;
<emphasis id="BB810C4AF40EFF9E5D42F8340630F831" bold="true" box="[321,342,1927,1951]" pageId="5" pageNumber="288">8.</emphasis>
right pedipalp minus chela, dorsal view;
<emphasis id="BB810C4AF40EFF9E5F1AF834044AF831" bold="true" box="[793,812,1927,1951]" pageId="5" pageNumber="288">9.</emphasis>
right chela, dorsal view;
<emphasis id="BB810C4AF40EFF9E5840F8340302F831" bold="true" box="[1091,1124,1927,1951]" pageId="5" pageNumber="288">10.</emphasis>
right chela, lateral view;
<emphasis id="BB810C4AF40EFF9E597FF83402FCF831" bold="true" box="[1404,1434,1927,1951]" pageId="5" pageNumber="288">11.</emphasis>
claw of right leg IV;
<emphasis id="BB810C4AF40EFF9E5D74F81406FEF811" bold="true" box="[375,408,1959,1983]" pageId="5" pageNumber="288">12.</emphasis>
subterminal tarsal seta of right leg IV. Scale bars (in mm): 0.10 (Figs 5, 7, 11, 12), 0.20 (Figs 4, 6), 0.30 (Fig. 3), 0.50 (Figs 810).
</paragraph>
</caption>
<paragraph id="894AD058F40DFF9D5CC4FF240789FF59" blockId="6.[151,1436,151,788]" pageId="6" pageNumber="289">
The sensillum of the movable finger, near dental margin and close to the trichobothrium
<emphasis id="BB810C4AF40DFF9D58E5FF2A039FFF1E" box="[1254,1273,153,176]" italics="true" pageId="6" pageNumber="289">st</emphasis>
, is diploid as mentioned and illustrated for other species by
<bibRefCitation id="ED64ADA9F40DFF9D5EB8FF0F0439FF7A" author="Judson" box="[699,863,188,212]" pageId="6" pageNumber="289" refString="Judson, M. L. I. (1992) Roncocreagris murphyorum n. sp. and Occitanobisium nanum (Beier) n. comb. (Neobisiidae) from Iberia, with notes on the sternal glands of pseudoscorpions (Chelonethi). Bulletin of the British Arachnological Society, 9, 26 - 30." type="journal article" year="1992">Judson (1992)</bibRefCitation>
and
<bibRefCitation id="ED64ADA9F40DFF9D5F9DFF0E0338FF7A" author="Zaragoza" box="[926,1118,188,213]" pageId="6" pageNumber="289" refString="Zaragoza, J. A. (2003) Roncocreagris iglesiasae, nueva especie cavernicola de Asturias (Arachnida, Pseudoscorpiones, Neobisiidae). Revista Iberica de Aracnologia, 7, 89 - 94." type="journal article" year="2003">Zaragoza (2003)</bibRefCitation>
, but it is not raised in this species.
</paragraph>
<paragraph id="894AD058F40DFF9D5CC4FEB602FAFE02" blockId="6.[151,1436,151,788]" pageId="6" pageNumber="289">
<taxonomicName id="4EF5ABDBF40DFF9D5CC4FEB60695FEB2" box="[199,499,260,284]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5CC4FEB60695FEB2" box="[199,499,260,284]" italics="true" pageId="6" pageNumber="289">Roncocreagris blothroides</emphasis>
</taxonomicName>
and
<taxonomicName id="4EF5ABDBF40DFF9D5E2CFEB605B0FEB2" box="[559,726,260,284]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="BB810C4AF40DFF9D5E2CFEB605B0FEB2" box="[559,726,260,284]" italics="true" pageId="6" pageNumber="289">R. cavernicola</emphasis>
</taxonomicName>
share the feature of being the only species to usually have 4 setae on the posterior margin of the carapace and on tergite I. They differ as follows: the chelal measurements and ratios are distinctly larger in
<taxonomicName id="4EF5ABDBF40DFF9D5DD4FEFE0512FECA" box="[471,628,332,356]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5DD4FEFE0512FECA" box="[471,628,332,356]" italics="true" pageId="6" pageNumber="289">R. blothroides</emphasis>
</taxonomicName>
than in
<taxonomicName id="4EF5ABDBF40DFF9D5ECCFEFE0414FECA" box="[719,882,332,356]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="BB810C4AF40DFF9D5ECCFEFE0414FECA" box="[719,882,332,356]" italics="true" pageId="6" pageNumber="289">R. cavernicola</emphasis>
</taxonomicName>
; trichobothrium
<emphasis id="BB810C4AF40DFF9D582CFEFE032FFECA" box="[1071,1097,333,356]" italics="true" pageId="6" pageNumber="289">ist</emphasis>
is medial with respect to
<emphasis id="BB810C4AF40DFF9D596BFEFF0218FECA" box="[1384,1406,332,356]" italics="true" pageId="6" pageNumber="289">ib</emphasis>
/
<emphasis id="BB810C4AF40DFF9D5986FEFE02F2FECA" box="[1413,1428,333,356]" italics="true" pageId="6" pageNumber="289">it</emphasis>
, and
<emphasis id="BB810C4AF40DFF9D5CC5FEC207B0FE26" box="[198,214,369,392]" italics="true" pageId="6" pageNumber="289">it</emphasis>
is medial with respect to
<emphasis id="BB810C4AF40DFF9D5DF0FEC20574FE26" box="[499,530,369,392]" italics="true" pageId="6" pageNumber="289">est</emphasis>
/
<emphasis id="BB810C4AF40DFF9D5E19FEC20548FE26" box="[538,558,369,392]" italics="true" pageId="6" pageNumber="289">et</emphasis>
in
<taxonomicName id="4EF5ABDBF40DFF9D5E52FEC20588FE26" box="[593,750,368,392]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5E52FEC20588FE26" box="[593,750,368,392]" italics="true" pageId="6" pageNumber="289">R. blothroides</emphasis>
</taxonomicName>
, whereas in
<emphasis id="BB810C4AF40DFF9D5F79FEC20359FE26" box="[890,1087,368,392]" italics="true" pageId="6" pageNumber="289">
<taxonomicName id="4EF5ABDBF40DFF9D5F79FEC2037BFE26" box="[890,1053,368,392]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="cavernicola">R. cavernicola</taxonomicName>
ist
</emphasis>
is placed more basally and
<emphasis id="BB810C4AF40DFF9D5970FEC202E5FE26" box="[1395,1411,369,392]" italics="true" pageId="6" pageNumber="289">it</emphasis>
is closer to
<emphasis id="BB810C4AF40DFF9D5CFEFE26067AFE02" box="[253,284,405,428]" italics="true" pageId="6" pageNumber="289">est</emphasis>
than to
<emphasis id="BB810C4AF40DFF9D5D74FE2606EDFE02" box="[375,395,405,428]" italics="true" pageId="6" pageNumber="289">et</emphasis>
; the male galea bears some rami apically in
<taxonomicName id="4EF5ABDBF40DFF9D5F7EFE26037CFE02" box="[893,1050,404,428]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5F7EFE26037CFE02" box="[893,1050,404,428]" italics="true" pageId="6" pageNumber="289">R. blothroides</emphasis>
</taxonomicName>
, but it is simple in
<taxonomicName id="4EF5ABDBF40DFF9D58F1FE2602F3FE02" box="[1266,1429,404,428]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="BB810C4AF40DFF9D58F1FE2602F3FE02" box="[1266,1429,404,428]" italics="true" pageId="6" pageNumber="289">R. cavernicola</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F40DFF9D5CC4FE040530FD92" pageId="6" pageNumber="289" type="distribution">
<paragraph id="894AD058F40DFF9D5CC4FE040530FD92" blockId="6.[151,1436,151,788]" pageId="6" pageNumber="289">
<emphasis id="BB810C4AF40DFF9D5CC4FE040695FE7E" bold="true" box="[199,499,439,464]" pageId="6" pageNumber="289">Distribution and habitat.</emphasis>
All specimens of
<taxonomicName id="4EF5ABDBF40DFF9D5EC0FE0A0488FE7E" box="[707,1006,440,464]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5EC0FE0A0488FE7E" box="[707,1006,440,464]" italics="true" pageId="6" pageNumber="289">Roncocreagris blothroides</emphasis>
</taxonomicName>
were collected in the deepest parts of three caves, the two farthest separated by over
<quantity id="4E0D7DBDF40DFF9D5EACFE6F059DFE5A" box="[687,763,476,500]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="289" unit="km" value="25.0">25 km</quantity>
. In addition to the
<typeStatus id="564E6EFAF40DFF9D5FD7FE6E0362FE5B" box="[980,1028,477,501]" pageId="6" pageNumber="289">type</typeStatus>
localities of Santa Maria da Estrela and Moura (= Arrifana) Caves, this species is newly recorded from Cerâmica and Confraria Caves, all of which are located in the Sicó karst massif (
<figureCitation id="11CECCDDF40DFF9D5E00FD970523FD92" box="[515,581,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,255,1630,1654]" captionTargetBox="[342,1243,854,1606]" captionTargetId="figure@2.[340,1247,829,1616]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of subterranean species of Roncocreagris Mahnert, 1974 in Portuguese caves. Cave numbers: 1, Arrifana (R. blothroides); 2, Santa Maria da Estrela (R. blothroides); 3, Confraria I (R. blothroides); 4, Algarinho (R. gespei); 5, Lapa (R. cavernicola); 6, Cerâmica (R. blothroides, R. borgesi); 7, Corujeiras (R. cavernicola); 8, Bolhos (R. occidentalis); 9, Javali (R. occidentalis)." httpUri="https://zenodo.org/record/219877/files/figure.png" pageId="6" pageNumber="289">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C1EF83D3F40DFF9D5CC4FDFA061FFCBA" pageId="6" pageNumber="289" type="reference_group">
<paragraph id="894AD058F40DFF9D5CC4FDFA061FFCBA" blockId="6.[151,1436,151,788]" pageId="6" pageNumber="289">
<taxonomicName id="4EF5ABDBF40DFF9D5CC4FDFA0696FDCE" ID-CoL="4TB43" box="[199,496,584,608]" class="Arachnida" family="Neobisiidae" genus="Roncocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="blothroides">
<emphasis id="BB810C4AF40DFF9D5CC4FDFA0696FDCE" box="[199,496,584,608]" italics="true" pageId="6" pageNumber="289">Roncocreagris blothroides</emphasis>
</taxonomicName>
is an abundant pseudoscorpion in the studied caves, with up to ten individuals being found during a single cave visit. It was found inhabiting a temperature range of 13.616.6ºC at soil level. This species shares its habitat with some frequent troglophile springtails, such as
<taxonomicName id="4EF5ABDBF40DFF9D5831FD2302FEFD09" authority="Folsom, 1896" authorityName="Folsom" authorityYear="1896" box="[1074,1432,655,680]" class="Entognatha" family="Neelidae" genus="Neelus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="murinus">
<emphasis id="BB810C4AF40DFF9D5831FD230381FD06" box="[1074,1255,656,680]" italics="true" pageId="6" pageNumber="289">Neelus murinus</emphasis>
Folsom, 1896
</taxonomicName>
,
<taxonomicName id="4EF5ABDBF40DFF9D5C94FD07052EFD62" authority="Huther, 1959" authorityName="Huther" authorityYear="1959" box="[151,584,692,716]" class="Entognatha" family="Hypogastruridae" genus="Willemia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="buddenbrocki">
<emphasis id="BB810C4AF40DFF9D5C94FD0706C6FD62" box="[151,416,692,716]" italics="true" pageId="6" pageNumber="289">Willemia buddenbrocki</emphasis>
Hüther, 1959
</taxonomicName>
,
<taxonomicName id="4EF5ABDBF40DFF9D5E5AFD070367FD62" authority="Gisin, 1952" authorityName="Gisin" authorityYear="1952" box="[601,1025,692,716]" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="insubraruis">
<emphasis id="BB810C4AF40DFF9D5E5AFD07040FFD62" box="[601,873,692,716]" italics="true" pageId="6" pageNumber="289">Onychiurus insubraruis</emphasis>
Gisin, 1952
</taxonomicName>
,
<taxonomicName id="4EF5ABDBF40DFF9D5810FD0707B4FD41" authority="Gama, 1964" authorityName="Gama" authorityYear="1964" class="Entognatha" family="Onychiuridae" genus="Onychiurus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="subgranulosus">
<emphasis id="BB810C4AF40DFF9D5810FD070223FD62" box="[1043,1349,692,716]" italics="true" pageId="6" pageNumber="289">Onychiurus subgranulosus</emphasis>
Gama, 1964
</taxonomicName>
,
<taxonomicName id="4EF5ABDBF40DFF9D5CDFFD6B053CFD41" authority="Willem, 1902" authorityName="Willem" authorityYear="1902" box="[220,602,727,752]" class="Entognatha" family="Isotomidae" genus="Folsomia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="candida">
<emphasis id="BB810C4AF40DFF9D5CDFFD6B06C0FD5E" box="[220,422,728,752]" italics="true" pageId="6" pageNumber="289">Folsomia candida</emphasis>
(Willem, 1902)
</taxonomicName>
,
<taxonomicName id="4EF5ABDBF40DFF9D5E66FD6A04DCFD41" authority="Gama, 1964" authorityName="Gama" authorityYear="1964" box="[613,954,727,752]" class="Entognatha" family="Isotomidae" genus="Proisotoma" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="gisini">
<emphasis id="BB810C4AF40DFF9D5E66FD6A044DFD5E" box="[613,811,729,752]" italics="true" pageId="6" pageNumber="289">Proisotoma gisini</emphasis>
Gama, 1964
</taxonomicName>
and
<taxonomicName id="4EF5ABDBF40DFF9D5FECFD6B02FAFD41" authority="Borner, 1901" authorityName="Borner" authorityYear="1901" box="[1007,1436,727,752]" class="Entognatha" family="Tullbergiidae" genus="Tullbergia" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="krausbaueri">
<emphasis id="BB810C4AF40DFF9D5FECFD6B0397FD5E" box="[1007,1265,728,752]" italics="true" pageId="6" pageNumber="289">Tullbergia krausbaueri</emphasis>
(Börner, 1901)
</taxonomicName>
(
<bibRefCitation id="ED64ADA9F40DFF9D5C9DFD4F0643FCBA" author="Gama" box="[158,293,764,788]" pageId="6" pageNumber="289" refString="Gama, M. M. (1962) Collemboles des grottes du Portugal. Voyage au Portugal du Dr. K. Lindberg. Resultats zoologiques 5. Boletim da Sociedade Portuguesa de Ciencias Naturais, 2 ª serie, 9, 100 - 108." type="journal article" year="1962">Gama 1962</bibRefCitation>
,
<bibRefCitation id="ED64ADA9F40DFF9D5D33FD4F060AFCBA" author="Gama" box="[304,364,764,788]" pageId="6" pageNumber="289" refString="Gama, M. M. (1965) Colembolos de Portugal Continental. Memorias e estudos do Museu Zoologico da Universidade de Coimbra, 292 pp." type="book" year="1965">1965</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>