401 lines
43 KiB
XML
401 lines
43 KiB
XML
<document id="FF81F0A867531188237779A07DE46A61" ID-CLB-Dataset="46924" ID-DOI="10.5281/zenodo.199951" ID-GBIF-Dataset="9581ddd0-efed-4ff4-a955-4f6db2dad6c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="199951" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460350412969" checkinUser="plazi" docAuthor="Pinto, Roberta Richard, Passos, Paulo, Portilla, José Rances Caicedo, Arredondo, Juan Camilo & Fernandes, Ronaldo" docDate="2010" docId="864C87F9FF8B145EFF7B59F1FE5DAA0A" docLanguage="en" docName="zt02724p028.pdf" docOrigin="Zootaxa 2724" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tricheilostoma joshuai Dunn 1944" docType="treatment" docVersion="10" lastPageNumber="10" masterDocId="7A75FF81FF8D1457FFEC5806FF86A855" masterDocTitle="Taxonomy of the Threadsnakes of the tribe Epictini (Squamata: Serpentes: Leptotyphlopidae) in Colombia" masterLastPageNumber="28" masterPageNumber="1" pageNumber="7" updateTime="1698293673783" updateUser="ExternalLinkService">
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<mods:titleInfo id="0B6BD711A24ACBD304DA419840E7C891">
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<mods:title id="998BFD2C44F636DC0B5281D5B5506DF4">Taxonomy of the Threadsnakes of the tribe Epictini (Squamata: Serpentes: Leptotyphlopidae) in Colombia</mods:title>
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<mods:namePart id="573E99D40C63A3C844D89292C0E1C7E4">Pinto, Roberta Richard</mods:namePart>
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<mods:roleTerm id="42391D3612C6298332A3E310CB91705C">Author</mods:roleTerm>
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<mods:namePart id="206E2207A531EF18BEACC1BDB87F73D0">Passos, Paulo</mods:namePart>
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<mods:roleTerm id="99792DFE47026FBCDD5089397084FCDC">Author</mods:roleTerm>
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<mods:namePart id="6CCCE80289019B4E16A6CF58A9164CFE">Portilla, José Rances Caicedo</mods:namePart>
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<mods:namePart id="0B49EC865464C815CC32A9CACAB7FEB4">Arredondo, Juan Camilo</mods:namePart>
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<mods:name id="8BDA438B7B3704AD853CDA09CA3D03EE" type="personal">
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<mods:namePart id="121C2DDE61653F3E9FAC0482A2A9AD1F">Fernandes, Ronaldo</mods:namePart>
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<mods:title id="66FED6E67096E2D618578F0D663D35E6">Zootaxa</mods:title>
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<mods:date id="3BAEACF81D9B285579A55445A2A1B32C">2010</mods:date>
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<mods:detail id="74A77EE41B3D9939DECA7F55A297A77B" type="volume">
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<mods:number id="AEB8268F771298077BE62525A4E4A16F">2724</mods:number>
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<mods:identifier id="7169FC2884E6EE50E266BFB0A5038AE1" type="DOI">10.5281/zenodo.199951</mods:identifier>
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<treatment id="864C87F9FF8B145EFF7B59F1FE5DAA0A" ID-DOI="http://doi.org/10.5281/zenodo.6204209" ID-GBIF-Taxon="119407853" ID-Zenodo-Dep="6204209" LSID="urn:lsid:plazi:treatment:864C87F9FF8B145EFF7B59F1FE5DAA0A" httpUri="http://treatment.plazi.org/id/864C87F9FF8B145EFF7B59F1FE5DAA0A" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
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<subSubSection id="46FF6564FF8B1451FF7B59F1FF79AA67" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="0E5A36EFFF8B1451FF7B59F1FDC9AA44" blockId="6.[151,591,503,562]" box="[151,591,503,529]" pageId="6" pageNumber="7">
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<heading id="55128183FF8B1451FF7B59F1FDC9AA44" bold="true" box="[151,591,503,529]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<taxonomicName id="C9E54D6CFF8B1451FF7B59F1FDC9AA44" authority="Dunn 1944" authorityName="Dunn" authorityYear="1944" box="[151,591,503,529]" class="Reptilia" family="Leptotyphlopidae" genus="Tricheilostoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="joshuai">
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<emphasis id="3C91EAFDFF8B1451FF7B59F1FDC9AA44" bold="true" box="[151,591,503,529]" pageId="6" pageNumber="7">
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<emphasis id="3C91EAFDFF8B1451FF7B59F1FE2BAA44" bold="true" box="[151,429,503,529]" italics="true" pageId="6" pageNumber="7">Tricheilostoma joshuai</emphasis>
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||
(
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<bibRefCitation id="6A744B1EFF8B1451FE5059F1FDC1AA44" author="Dunn" box="[444,583,503,529]" pageId="6" pageNumber="7" refString="Dunn, E. R. (1944) A review of the Colombian snakes of the families Typhlopidae and Leptotyphlopidae. Caldasia, 3, 47 - 55." type="journal article" year="1944">Dunn 1944</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="0E5A36EFFF8B1451FF7B5A1CFF79AA67" blockId="6.[151,591,503,562]" box="[151,255,538,562]" pageId="6" pageNumber="7">
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<figureCitation id="96DE2A6AFF8B1451FF7B5A1CFF63AA67" box="[151,229,538,562]" captionStart="FIGURE 4" captionStartId="7.[151,250,1390,1413]" captionText="FIGURE 4. General view of the paratype of Tricheilostoma joshuai (MLS 11)." pageId="6" pageNumber="7">Figs. 4</figureCitation>
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||
,
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<figureCitation id="96DE2A6AFF8B1451FF1D5A1CFF79AA67" box="[241,255,538,562]" captionStart="FIGURE 5" captionStartId="8.[151,255,1912,1936]" captionTargetBox="[195,1384,194,1886]" captionTargetId="figure@8.[195,1386,194,1886]" captionTargetPageId="8" captionText="FIGURE 5. Dorsal (A), lateral (B), and ventral (C) views of head of the Tricheilostoma joshuai (MVZ 68688)." httpUri="https://zenodo.org/record/199955/files/figure.png" pageId="6" pageNumber="7">5</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="46FF6564FF8B1451FF7B5A64FB39AB1A" pageId="6" pageNumber="7" type="reference_group">
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||
<paragraph id="0E5A36EFFF8B1451FF7B5A64FD0AAA2D" blockId="6.[151,1357,610,847]" box="[151,652,610,632]" pageId="6" pageNumber="7">
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<taxonomicName id="C9E54D6CFF8B1451FF7B5A64FE77AA2D" authority="Dunn 1944" authorityName="Dunn" authorityYear="1944" box="[151,497,610,632]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="joshuai">
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<emphasis id="3C91EAFDFF8B1451FF7B5A64FEF1AA2D" box="[151,375,610,632]" italics="true" pageId="6" pageNumber="7">Leptotyphlops joshuai</emphasis>
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<bibRefCitation id="6A744B1EFF8B1451FE915A65FE77AA2D" author="Dunn" box="[381,497,611,632]" pageId="6" pageNumber="7" refString="Dunn, E. R. (1944) A review of the Colombian snakes of the families Typhlopidae and Leptotyphlopidae. Caldasia, 3, 47 - 55." type="journal article" year="1944">Dunn 1944</bibRefCitation>
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</taxonomicName>
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, Caldasia 3:53.
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</paragraph>
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<paragraph id="0E5A36EFFF8B1451FF7B5A87FD36AAE0" blockId="6.[151,1357,610,847]" pageId="6" pageNumber="7">
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||
<emphasis id="3C91EAFDFF8B1451FF7B5A87FE0BAAC2" box="[151,397,641,663]" italics="true" pageId="6" pageNumber="7">
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||
<taxonomicName id="C9E54D6CFF8B1451FF7B5A87FEA0AAC2" box="[151,294,641,663]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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joshuai—
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</emphasis>
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Bailey 1946, Occasional Papers of the Museum of Zoology the University of Michigan, 492:4.
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<emphasis id="3C91EAFDFF8B1451FF7B5AA6FE0BAAE3" box="[151,397,672,694]" italics="true" pageId="6" pageNumber="7">
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||
<taxonomicName id="C9E54D6CFF8B1451FF7B5AA6FEA0AAE3" box="[151,294,672,694]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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joshuai—
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</emphasis>
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||
<bibRefCitation id="6A744B1EFF8B1451FE615AA7FD87AAE3" author="Dunn" box="[397,513,672,694]" pageId="6" pageNumber="7" refString="Dunn, E. R. (1946) A new snake from the eastern Andes of Colombia. Caldasia, 4, 121 - 122." type="journal article" year="1946">Dunn 1946</bibRefCitation>
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, Caldasia, 4:122.
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||
</paragraph>
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||
<paragraph id="0E5A36EFFF8B1451FF7B5AB9FD6BAAA7" blockId="6.[151,1357,610,847]" pageId="6" pageNumber="7">
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||
<emphasis id="3C91EAFDFF8B1451FF7B5AB9FE0BAA80" box="[151,397,703,725]" italics="true" pageId="6" pageNumber="7">
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||
<taxonomicName id="C9E54D6CFF8B1451FF7B5AB9FEA0AA80" box="[151,294,703,725]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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||
joshuai—
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||
</emphasis>
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||
<bibRefCitation id="6A744B1EFF8B1451FE615AB9FD4EAA81" author="Peters" box="[397,712,702,724]" pageId="6" pageNumber="7" refString="Peters, J. A., & Orejas-Miranda, B. R. (1970) Catalogue of the Neotropical Squamata: Part 1. Snakes. Bulletin of the United States of National Museum, 297, 1 - 347." type="journal article" year="1970">Peters & Orejas-Miranda 1970</bibRefCitation>
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||
, Bulletin of the
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<collectingCountry id="76F2767FFF8B1451FC875AB8FC74AA81" box="[875,1010,702,724]" name="United States of America" pageId="6" pageNumber="7">United States</collectingCountry>
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||
National Museum, 297:170.
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||
<emphasis id="3C91EAFDFF8B1451FF7B5ADBFE0BAAA6" box="[151,397,733,755]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="C9E54D6CFF8B1451FF7B5ADBFEA0AAA6" box="[151,294,733,755]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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joshuai—
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||
</emphasis>
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||
<bibRefCitation id="6A744B1EFF8B1451FE615ADBFD86AAA7" author="Hahn" box="[397,512,733,755]" pageId="6" pageNumber="7" refString="Hahn, D. E. (1980) Liste der rezenten Amphibien und Reptilien Anomalepididae, Leptotyphlopidae, Typhlopidae. Das Tierreich, 101, 1 - 93." type="journal article" year="1980">Hahn 1980</bibRefCitation>
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, Das Tierreich, 101:19.
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||
</paragraph>
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||
<paragraph id="0E5A36EFFF8B1451FF7B5AFAFB39AB1A" blockId="6.[151,1357,610,847]" pageId="6" pageNumber="7">
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||
<emphasis id="3C91EAFDFF8B1451FF7B5AFAFE0BAB47" box="[151,397,764,786]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="C9E54D6CFF8B1451FF7B5AFAFEA0AB47" box="[151,294,764,786]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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joshuai—
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</emphasis>
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||
<bibRefCitation id="6A744B1EFF8B1451FE615AFDFC8FAB44" author="McDiarmid" box="[397,777,763,785]" pageId="6" pageNumber="7" refString="McDiarmid, R. W., Campbell, J. A. & Toure, T. (1999) Snake species of the world. A taxonomic and geographic reference, vol. 1. Washington DC: The Herpetologists' League, 512 pp." type="book" year="1999">McDiarmid, Campbell & Touré 1999</bibRefCitation>
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||
, Snakes Species of the World, 1:33–34.
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||
<emphasis id="3C91EAFDFF8B1451FF7B5B1CFE0BAB65" box="[151,397,794,816]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="C9E54D6CFF8B1451FF7B5B1CFEA0AB65" box="[151,294,794,816]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Leptotyphlops</taxonomicName>
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joshuai—
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||
</emphasis>
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||
<bibRefCitation id="6A744B1EFF8B1451FE615B1DFD60AB65" author="Passos" box="[397,742,794,816]" pageId="6" pageNumber="7" refString="Passos, P., Caramaschi, U. & Pinto, R. R. (2006) Redescription of Leptotyphlops koppesi Amaral, 1954, and description of a new species of the Leptotyphlops dulcis group from Central Brazil (Serpentes: Leptotyphlopidae). Amphibia- Reptilia, 27, 347 - 357." type="journal article" year="2006">Passos, Caramaschi & Pinto 2006</bibRefCitation>
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||
, Amphibia-Reptilia, 27:349.
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||
<emphasis id="3C91EAFDFF8B1451FF7B5B3FFE10AB1A" box="[151,406,825,847]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="C9E54D6CFF8B1451FF7B5B3FFEA9AB1A" box="[151,303,825,847]" class="Reptilia" family="Leptotyphlopidae" genus="Tricheilostoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Tricheilostoma</taxonomicName>
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||
joshuai—
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||
</emphasis>
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||
Hedges, Adalsteinsson & Branch
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||
<emphasis id="3C91EAFDFF8B1451FD005B3CFC86AB1A" box="[748,768,826,847]" italics="true" pageId="6" pageNumber="7">in</emphasis>
|
||
Adalsteisson
|
||
<emphasis id="3C91EAFDFF8B1451FC625B3CFC3CAB1A" box="[910,954,825,847]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
. 2009, Zootaxa, 2244:11.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46FF6564FF8B1451FF7B5B79FB9DAC7C" pageId="6" pageNumber="7" type="materials_examined">
|
||
<paragraph id="0E5A36EFFF8B1451FF7B5B79FE2AABEF" blockId="6.[151,1436,892,2037]" pageId="6" pageNumber="7">
|
||
<emphasis id="3C91EAFDFF8B1451FF7B5B79FE8DABC2" bold="true" box="[151,267,895,919]" pageId="6" pageNumber="7">
|
||
<typeStatus id="D15E884DFF8B1451FF7B5B79FE80ABC2" box="[151,262,895,919]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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||
.
|
||
</emphasis>
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||
<collectionCode id="68F4AE2AFF8B1451FEF95B79FEC8ABC2" box="[277,334,895,919]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
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||
13, from municipality of Jericó (
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||
<geoCoordinate id="6BD15028FF8B1451FD395B79FCC4ABC2" box="[725,834,892,919]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="5.7833333">05o 47' N</geoCoordinate>
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||
,
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||
<geoCoordinate id="6BD15028FF8B1451FCBC5B79FC04ABDF" box="[848,898,892,919]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="1.25">
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||
0
|
||
<geoCoordinate id="6BD15028FF8B1451FCB25B79FC04ABDF" box="[862,898,892,919]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-75.78333">75o</geoCoordinate>
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||
</geoCoordinate>
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||
47' W; ca.
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||
<quantity id="C91D9B0AFF8B1451FBE55B79FBE2ABC2" box="[1033,1124,895,919]" metricMagnitude="3" metricUnit="m" metricValue="1.9669999999999999" pageId="6" pageNumber="7" unit="m" value="1967.0">1967 m</quantity>
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||
), department of Antioquia,
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||
<collectingCountry id="76F2767FFF8B1451FF7B5BA4FE83ABEF" box="[151,261,930,954]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
|
||
(see remarks).
|
||
</paragraph>
|
||
<paragraph id="0E5A36EFFF8B1451FF255BC1FB9DAC7C" blockId="6.[151,1436,892,2037]" pageId="6" pageNumber="7">
|
||
<emphasis id="3C91EAFDFF8B1451FF255BC1FECCAB8A" bold="true" box="[201,330,967,991]" pageId="6" pageNumber="7">
|
||
<typeStatus id="D15E884DFF8B1451FF255BC1FEC3AB8A" box="[201,325,967,991]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
|
||
:
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||
</emphasis>
|
||
<collectionCode id="68F4AE2AFF8B1451FEBF5BC1FE0BAB8A" box="[339,397,967,991]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
|
||
11,
|
||
<collectionCode id="68F4AE2AFF8B1451FE2D5BC1FE7CAB8A" box="[449,506,967,991]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
|
||
<date id="7A5B102FFF8B1451FDE85BC1FD07AB8A" box="[516,641,967,991]" pageId="6" pageNumber="7" value="2646" valueMax="2647">2646–2647</date>
|
||
from municipality of Jericó, department of Antioquia.
|
||
<collectionCode id="68F4AE2AFF8B1451FB125BC1FAB1AB8A" box="[1278,1335,967,991]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
|
||
12 from “Río Cauca”, department of Antioquia.
|
||
<collectionCode id="68F4AE2AFF8B1451FDB65BECFD15AC57" box="[602,659,1002,1026]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
|
||
14 lacking specific locality, department of Antioquia.
|
||
<collectionCode id="68F4AE2AFF8B1451FAEC5BECFABCAC57" box="[1280,1338,1002,1026]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="6" pageNumber="7">MLS</collectionCode>
|
||
15 from Villamaría (
|
||
<geoCoordinate id="6BD15028FF8B1451FEF05C17FE02AC7C" box="[284,388,1038,1065]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="6.0333333">05o 62' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BD15028FF8B1451FE635C17FE47AC49" box="[399,449,1038,1065]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="1.25">
|
||
0
|
||
<geoCoordinate id="6BD15028FF8B1451FE715C17FE47AC49" box="[413,449,1038,1065]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-75.51667">75o</geoCoordinate>
|
||
</geoCoordinate>
|
||
31' W; ca.
|
||
<quantity id="C91D9B0AFF8B1451FDD05C17FD12AC7C" box="[572,660,1041,1065]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="6" pageNumber="7" unit="m" value="1840.0">1840 m</quantity>
|
||
), department of Caldas,
|
||
<collectingCountry id="76F2767FFF8B1451FC495C17FB91AC7C" box="[933,1047,1041,1065]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46FF6564FF8B1451FF255C33FD06AD70" pageId="6" pageNumber="7" type="diagnosis">
|
||
<paragraph id="0E5A36EFFF8B1451FF255C33FD06AD70" blockId="6.[151,1436,892,2037]" pageId="6" pageNumber="7">
|
||
<emphasis id="3C91EAFDFF8B1451FF255C33FEC5AC1B" bold="true" box="[201,323,1077,1102]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
|
||
<taxonomicName id="C9E54D6CFF8B1451FEBD5C33FDDEAC18" box="[337,600,1077,1101]" class="Reptilia" family="Leptotyphlopidae" genus="Tricheilostoma" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="joshuai">
|
||
<emphasis id="3C91EAFDFF8B1451FEBD5C33FDDEAC18" box="[337,600,1077,1101]" italics="true" pageId="6" pageNumber="7">Tricheilostoma joshuai</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all congeners by the following combination of characters: snout rounded in dorsal view, truncate in lateral view; supraocular present; rostral triangular in dorsal view; ocular subhexagonal, with rounded shape in the eye level; temporal distinct; fused caudals; eye in anterior portion of expanded area of ocular scale; three supralabials (2+1); four infralabials; 174–193 middorsal scales in males and
|
||
<quantity id="C91D9B0AFF8B1451FEFC5CC3FE09AC88" box="[272,399,1221,1245]" metricMagnitude="0" metricUnit="m" metricValue="4.8641" metricValueMax="5.0546" metricValueMin="4.6736" pageId="6" pageNumber="7" unit="in" value="191.5" valueMax="199.0" valueMin="184.0">184–199 in</quantity>
|
||
females; 169–181 midventral scales in males and
|
||
<quantity id="C91D9B0AFF8B1451FC255CC3FBCEAC88" box="[969,1096,1221,1245]" metricMagnitude="0" metricUnit="m" metricValue="4.5593" metricValueMax="4.7498" metricValueMin="4.3688" pageId="6" pageNumber="7" unit="in" value="179.5" valueMax="187.0" valueMin="172.0">172–187 in</quantity>
|
||
females; 13–17 subcaudals in males and
|
||
<quantity id="C91D9B0AFF8B1451FEF85CEFFEFFAD54" box="[276,377,1257,1281]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="6" pageNumber="7" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
|
||
females; 12 scales around the middle of tail; seven dark brown dorsal scale rows with pale brown edge, venter and labial scales cream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46FF6564FF8B1450FF255D37FAFEAFA6" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="0E5A36EFFF8B1451FF255D37FBBCADC4" blockId="6.[151,1436,892,2037]" pageId="6" pageNumber="7">
|
||
<emphasis id="3C91EAFDFF8B1451FF255D37FDA8AD1F" bold="true" box="[201,558,1329,1354]" pageId="6" pageNumber="7">
|
||
Redescription of the
|
||
<typeStatus id="D15E884DFF8B1451FE2C5D34FDAFAD1F" box="[448,553,1330,1354]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female,
|
||
<quantity id="C91D9B0AFF8B1451FD395D37FCB4AD1C" box="[725,818,1329,1353]" metricMagnitude="-1" metricUnit="m" metricValue="2.59" pageId="6" pageNumber="7" unit="mm" value="259.0">259 mm</quantity>
|
||
TL,
|
||
<quantity id="C91D9B0AFF8B1451FC845D37FC31AD1C" box="[872,951,1329,1353]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="mm" value="16.0">16 mm</quantity>
|
||
TAL;
|
||
<quantity id="C91D9B0AFF8B1451FBEC5D37FBD3AD1C" box="[1024,1109,1329,1353]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="6" pageNumber="7" unit="mm" value="8.3">8.3 mm</quantity>
|
||
MB; 16.2 TL/TAL; 31.2 TL/ MB;
|
||
<quantity id="C91D9B0AFF8B1451FF395D53FEA9AD38" box="[213,303,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="7" unit="mm" value="4.9">4.9 mm</quantity>
|
||
HL; 3.0 mm relative eye diameter;
|
||
<quantity id="C91D9B0AFF8B1451FD375D53FCB0AD38" box="[731,822,1365,1389]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="0.3">0.3 mm</quantity>
|
||
relative rostral width; head slightly depressed; body subcylindrical, slightly enlarged on head and slightly tapered caudally near the tail.
|
||
</paragraph>
|
||
<paragraph id="0E5A36EFFF8B1450FF255D9BFAEBAA99" blockId="6.[151,1436,892,2037]" lastBlockId="7.[151,1436,151,716]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
Head subcylindrical, almost twice as long as wide; cervical constriction barely distinct; snout slightly truncate in dorsal and ventral views, slightly obtuse in lateral view; rostral straight in frontal and ventral views, dorsal apex triangular, reaching an imaginary transverse line between anterior margins of ocular scales; rostral contacting supranasal and infranasal laterally and frontal dorsally; nasal completely divided horizontally by oblique suture crossing nostril; nostril roughly elliptic, obliquely oriented and placed in the middle of the nasal suture; supranasal about twice as high as long, bordering rostral anteriorly, infranasal inferiorly, first supralabial and ocular posteriorly, and frontal and supraocular dorsally; supranasal as long as upper border of infranasal scale; infranasal about twice as high as long; upper lip border formed by rostral, infranasal, two anterior supralabials, ocular and posterior supralabial scales; temporal scale distinct from dorsal scales of lateral rows; three supralabials (five according to
|
||
<bibRefCitation id="6A744B1EFF8B1451FEC65EE4FE2DAEAC" author="Dunn" box="[298,427,1761,1785]" pageId="6" pageNumber="7" refString="Dunn, E. R. (1944) A review of the Colombian snakes of the families Typhlopidae and Leptotyphlopidae. Caldasia, 3, 47 - 55." type="journal article" year="1944">Dunn 1944</bibRefCitation>
|
||
), two anterior and one posterior to ocular (2+1); first supralabial small, not reaching level of nostril and eye; second supralabial about twice as high as long, exceeding nostril and the lower portion of eye level; third supralabial with trapezoidal shape, lower than second one, slightly longer than high, reaching eye level, its posterior margin in broad contact with temporal; ocular enlarged, subhexagonal, rounded in the eye level, twice as high as long, contacting the posterior margins of supranasal and second supralabial anteriorly, parietal and third supralabial posteriorly, and supraocular dorsally, its dorsal apex straight; eye distinct (
|
||
<quantity id="C91D9B0AFF8B1451FB605F93FB6AAFF8" box="[1164,1260,1941,1965]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="7" unit="mm" value="0.6">0.6 mm</quantity>
|
||
), placed in the anterior area of the expanded upper part of ocular, displaced above the nostril level; supraocular three times longer than wide, subtly longer and slender than frontal scale, contacting supranasal anteriorly, frontal, postfrontal and ocular scales laterally, and parietal posteriorly; midsaggital head scales (postfrontal, interparietal and interoccipital) subequal in size, hexagonal in dorsal view, weakly imbricate; frontal not enlarged, smaller than other midsaggital scales, as wide as long, contacting rostral and supranasals anteriorly, supraoculars laterally, and postfrontal posteriorly; postfrontal as wide as long, contacting frontal, supraoculars, parietals and interparietal; interparietal wider than long, contacting postfrontal, parietals, occipitals, and interoccipital; interoccipital contacting interparietal, occipitals, and the first dorsal scale of the vertebral row; parietal and occipital similar in shape, irregularly pentagonal; parietal twice as wide as long, lower margin contacting the upper border of third supralabial, posterior margin contacting temporal, occipital and interparietal, anterior border in broad contact with ocular, supraocular and postfrontal; occipital twice as wide as long, its lower limit not attaining the level of the upper margin of third supralabial, though separated of the latter by temporal; symphysial trapezoidal, anterior and posterior borders respectively straight and slightly convex, three times as wide as long; four infralabials behind symphysial on both sides; first three infralabials subequal, slightly higher than long; fourth longer than other infralabials, approximately as long as second one; dorsal scales homogeneous, cycloid, smooth, slightly imbricate, and almost two times as wide as long; 195 (191 according to
|
||
<bibRefCitation id="6A744B1EFF8A1450FCA45A6BFC41AAD1" author="Dunn" box="[840,967,620,644]" pageId="7" pageNumber="8" refString="Dunn, E. R. (1944) A review of the Colombian snakes of the families Typhlopidae and Leptotyphlopidae. Caldasia, 3, 47 - 55." type="journal article" year="1944">Dunn 1944</bibRefCitation>
|
||
) middorsal scales; 181 midventral scales; 14 scale rows around midbody, reducing to 12 rows in the middle of the tail; cloacal shield short and semicircular, wider than long; 15 subcaudals; caudals fused; terminal spine short, conical, with stout base, longer than wide.
|
||
</paragraph>
|
||
<caption id="5A9A6667FF8A1450FF7B5D68FC3CADD0" box="[151,954,1390,1413]" pageId="7" pageNumber="8">
|
||
<paragraph id="0E5A36EFFF8A1450FF7B5D68FC3CADD0" blockId="7.[151,954,1390,1413]" box="[151,954,1390,1413]" pageId="7" pageNumber="8">
|
||
<emphasis id="3C91EAFDFF8A1450FF7B5D68FE95ADD6" bold="true" box="[151,275,1390,1413]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
|
||
General view of the paratype of
|
||
<taxonomicName id="C9E54D6CFF8A1450FD8C5D68FCCCADD1" box="[608,842,1390,1412]" class="Reptilia" family="Leptotyphlopidae" genus="Tricheilostoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="joshuai">
|
||
<emphasis id="3C91EAFDFF8A1450FD8C5D68FCCCADD1" box="[608,842,1390,1412]" italics="true" pageId="7" pageNumber="8">Tricheilostoma joshuai</emphasis>
|
||
</taxonomicName>
|
||
(MLS 11).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0E5A36EFFF8A1450FF255DA1FE60AE1F" blockId="7.[151,1436,1447,2035]" pageId="7" pageNumber="8">
|
||
<emphasis id="3C91EAFDFF8A1450FF255DA1FD22AD95" bold="true" box="[201,676,1447,1472]" pageId="7" pageNumber="8">
|
||
Colour of the
|
||
<typeStatus id="D15E884DFF8A1450FE9A5DAEFE5BAD95" box="[374,477,1448,1472]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
in preservative:
|
||
</emphasis>
|
||
Seven dorsal scale rows uniformly dark brown and seven ventral scale rows cream; head dorsally paler than dorsal scales of body, with some brown pigmentation concentrated on centre portion of cephalic scales; paler colour extending from rostral to interoccipital; lower margins of scales forming the creamish upper lip border; cloacal shield pale brown, slightly darker than general ventral coloration; terminal spine not pigmented.
|
||
</paragraph>
|
||
<paragraph id="0E5A36EFFF8A1450FF255E53FC89AEED" blockId="7.[151,1436,1447,2035]" pageId="7" pageNumber="8">
|
||
<emphasis id="3C91EAFDFF8A1450FF255E53FE3DAE3B" bold="true" box="[201,443,1621,1646]" pageId="7" pageNumber="8">Sexual dimorphism:</emphasis>
|
||
Females were significantly longer (F(1,13) = 11.8; p <0.01) and showed a higher TL/TAL ratio (F(1, 13) = 7.1; p <0.05) than males. However, males have significantly more subcaudal scales (F(1,13) = 11.2; p <0.01) and largest tail F(1, 13) = 8.1; p <0.01) than females.
|
||
</paragraph>
|
||
<paragraph id="0E5A36EFFF8A1450FF255EC3FAFEAFA6" blockId="7.[151,1436,1447,2035]" pageId="7" pageNumber="8">
|
||
<emphasis id="3C91EAFDFF8A1450FF255EC3FEC6AE8B" bold="true" box="[201,320,1733,1758]" pageId="7" pageNumber="8">Variation:</emphasis>
|
||
Middorsal scales 174–193 (
|
||
<emphasis id="3C91EAFDFF8A1450FD6C5EC1FD21AE8B" box="[640,679,1735,1758]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
187.4 ± 5.2,
|
||
<emphasis id="3C91EAFDFF8A1450FCD15EC1FCCDAE8B" box="[829,843,1735,1758]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) in males and 184–199 (
|
||
<emphasis id="3C91EAFDFF8A1450FB705EC1FB45AE8B" box="[1180,1219,1735,1758]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
193.8 ± 6.7,
|
||
<emphasis id="3C91EAFDFF8A1450FAB55EC1FAE1AE8B" box="[1369,1383,1735,1758]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 4) in females; midventral scales 169–181 (
|
||
<emphasis id="3C91EAFDFF8A1450FDB75EEFFD02AF55" box="[603,644,1769,1792]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
173.9 ± 4.2,
|
||
<emphasis id="3C91EAFDFF8A1450FCF25EEFFCAAAF55" box="[798,812,1769,1792]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 8) in males and 172–187 (
|
||
<emphasis id="3C91EAFDFF8A1450FB9A5EEFFB19AF55" box="[1142,1183,1769,1792]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
180.0 ± 7.6,
|
||
<emphasis id="3C91EAFDFF8A1450FAD55EEFFAC1AF55" box="[1337,1351,1769,1792]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 3) in females; subcaudal scales
|
||
<quantity id="C91D9B0AFF8A1450FE515F0DFDA6AF76" box="[445,544,1803,1827]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.318" metricValueMin="3.302" pageId="7" pageNumber="8" unit="in" value="15.0" valueMax="17.0" valueMin="13.0">13–17 in</quantity>
|
||
males (
|
||
<emphasis id="3C91EAFDFF8A1450FD945F0AFD02AF76" box="[632,644,1804,1827]" italics="true" pageId="7" pageNumber="8">x¯</emphasis>
|
||
= 16.0 ± 1.2,
|
||
<emphasis id="3C91EAFDFF8A1450FCCF5F0AFCB7AF76" box="[803,817,1804,1827]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) and 13–15 (
|
||
<emphasis id="3C91EAFDFF8A1450FBEC5F0AFBA0AF76" box="[1024,1062,1804,1827]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
13.8 ± 1.1,
|
||
<emphasis id="3C91EAFDFF8A1450FB425F0AFB3AAF76" box="[1198,1212,1804,1827]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 5) in females; TL
|
||
<quantity id="C91D9B0AFF8A1450FF7B5F2BFE99AF10" box="[151,287,1837,1861]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" metricValueMax="2.18" metricValueMin="0.9" pageId="7" pageNumber="8" unit="mm" value="154.0" valueMax="218.0" valueMin="90.0">90–218 mm</quantity>
|
||
(
|
||
<emphasis id="3C91EAFDFF8A1450FEDC5F29FEBAAF13" box="[304,316,1839,1862]" italics="true" pageId="7" pageNumber="8">x¯</emphasis>
|
||
= 149.3 ± 49.5,
|
||
<emphasis id="3C91EAFDFF8A1450FE135F29FD8BAF13" box="[511,525,1839,1862]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) in males and
|
||
<quantity id="C91D9B0AFF8A1450FD1E5F2BFC0EAF10" box="[754,904,1837,1861]" metricMagnitude="-1" metricUnit="m" metricValue="2.315" metricValueMax="3.0" metricValueMin="1.63" pageId="7" pageNumber="8" unit="mm" value="231.5" valueMax="300.0" valueMin="163.0">163–300 mm</quantity>
|
||
(
|
||
<emphasis id="3C91EAFDFF8A1450FC755F29FC44AF13" box="[921,962,1839,1862]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
246.2 ± 55.6,
|
||
<emphasis id="3C91EAFDFF8A1450FB875F29FBFFAF13" box="[1131,1145,1839,1862]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 5) in females; TL/TAL ratio 10.8–17.0 (
|
||
<emphasis id="3C91EAFDFF8A1450FEBC5F57FEDBAF3D" box="[336,349,1873,1896]" italics="true" pageId="7" pageNumber="8">x¯</emphasis>
|
||
= 13.1 ± 1.9,
|
||
<emphasis id="3C91EAFDFF8A1450FE155F57FD81AF3D" box="[505,519,1873,1896]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) in males and 13.6–24.6 (
|
||
<emphasis id="3C91EAFDFF8A1450FC8E5F57FC0EAF3D" box="[866,904,1873,1896]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
17.3 ± 4.3,
|
||
<emphasis id="3C91EAFDFF8A1450FBE25F57FB9AAF3D" box="[1038,1052,1873,1896]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 5) in females; TAL 5.9–9.3% of TL in males (
|
||
<emphasis id="3C91EAFDFF8A1450FEDD5F72FEDCAFDE" box="[305,346,1908,1931]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
7.8 ± 0.0,
|
||
<emphasis id="3C91EAFDFF8A1450FE345F72FE60AFDE" box="[472,486,1908,1931]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) and 4.1–7.4% in females (
|
||
<emphasis id="3C91EAFDFF8A1450FCB05F72FC03AFDE" box="[860,901,1908,1931]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
6.0 ± 0.0,
|
||
<emphasis id="3C91EAFDFF8A1450FBEF5F72FB97AFDE" box="[1027,1041,1908,1931]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 5); TL/MB ratio 37.8–50.0 (
|
||
<emphasis id="3C91EAFDFF8A1450FA9D5F72FA1AAFDE" box="[1393,1436,1908,1931]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
42.3 ± 4.2,
|
||
<emphasis id="3C91EAFDFF8A1450FEF05F91FEACAFFB" box="[284,298,1943,1966]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 10) in males and 34.0–55.2 (
|
||
<emphasis id="3C91EAFDFF8A1450FD795F91FD46AFFB" box="[661,704,1943,1966]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
43.9 ± 9.4,
|
||
<emphasis id="3C91EAFDFF8A1450FCA35F91FCDBAFFB" box="[847,861,1943,1966]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 5) in females; TAL/MT ratio 2.8–3.6 (
|
||
<emphasis id="3C91EAFDFF8A1450FADA5F91FAC2AFFB" box="[1334,1348,1943,1966]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 2) in males and 2.9–3.5 (
|
||
<emphasis id="3C91EAFDFF8A1450FE9F5FBFFEF9AF85" box="[371,383,1977,2000]" italics="true" pageId="7" pageNumber="8">x¯</emphasis>
|
||
= 3.2 ± 0.3,
|
||
<emphasis id="3C91EAFDFF8A1450FDFC5FBFFD98AF85" box="[528,542,1977,2000]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 3) in females; relative eye diameter 0.3–0.4 (
|
||
<emphasis id="3C91EAFDFF8A1450FBDD5FBFFBB9AF85" box="[1073,1087,1977,2000]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 2) in males and 0.4–0.5 (
|
||
<emphasis id="3C91EAFDFF8A1450FA985FBFFA1CAF85" box="[1396,1434,1977,2000]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
0.5 ± 0.0,
|
||
<emphasis id="3C91EAFDFF8A1450FEE45FDAFE90AFA6" box="[264,278,2012,2035]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 3) in females; rostral width 2.4–3.7 (
|
||
<emphasis id="3C91EAFDFF8A1450FD225FDAFD5AAFA6" box="[718,732,2012,2035]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 2) in males and 2.1–2.3 (
|
||
<emphasis id="3C91EAFDFF8A1450FBFD5FDAFBBEAFA6" box="[1041,1080,2012,2035]" italics="true" pageId="7" pageNumber="8">x¯ =</emphasis>
|
||
2.2 ± 0.1,
|
||
<emphasis id="3C91EAFDFF8A1450FB5E5FDAFB46AFA6" box="[1202,1216,2012,2035]" italics="true" pageId="7" pageNumber="8">n</emphasis>
|
||
= 3) in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="5A9A6667FF85145FFF7B5F7EFABCAFDA" box="[151,1338,1912,1936]" httpUri="https://zenodo.org/record/199955/files/figure.png" pageId="8" pageNumber="9" targetBox="[195,1384,194,1886]" targetPageId="8">
|
||
<paragraph id="0E5A36EFFF85145FFF7B5F7EFABCAFDA" blockId="8.[151,1338,1912,1936]" box="[151,1338,1912,1936]" pageId="8" pageNumber="9">
|
||
<emphasis id="3C91EAFDFF85145FFF7B5F7EFE9FAFC5" bold="true" box="[151,281,1912,1936]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
|
||
Dorsal (A), lateral (B), and ventral (C) views of head of the
|
||
<taxonomicName id="C9E54D6CFF85145FFC4D5F7FFB13AFC5" box="[929,1173,1913,1936]" class="Reptilia" family="Leptotyphlopidae" genus="Tricheilostoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="joshuai">
|
||
<emphasis id="3C91EAFDFF85145FFC4D5F7FFB13AFC5" box="[929,1173,1913,1936]" italics="true" pageId="8" pageNumber="9">Tricheilostoma joshuai</emphasis>
|
||
</taxonomicName>
|
||
(MVZ 68688).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="46FF6564FF84145EFF255891FCF5A8A2" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="0E5A36EFFF84145EFF255891FCF5A8A2" blockId="9.[151,1437,151,607]" pageId="9" pageNumber="10">
|
||
<emphasis id="3C91EAFDFF84145EFF255891FEE4A8E5" bold="true" box="[201,354,151,176]" pageId="9" pageNumber="10">Distribution.</emphasis>
|
||
Cauca valley between the Cordilleras Occidental and Central of
|
||
<collectingCountry id="76F2767FFF84145EFB625891FB79A8FA" box="[1166,1279,151,175]" name="Colombia" pageId="9" pageNumber="10">Colombia</collectingCountry>
|
||
, from Jericó (
|
||
<geoCoordinate id="6BD15028FF84145EFF7358BAFEACA881" box="[159,298,188,212]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="5.7941666">05º47’39”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BD15028FF84145EFED458BAFE4EA881" box="[312,456,188,212]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-75.78972">75º47’23”W</geoCoordinate>
|
||
), department of Antioquia south to San Antonio (
|
||
<geoCoordinate id="6BD15028FF84145EFBE158BAFBE9A881" box="[1037,1135,188,212]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="3.2166667">03º13’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BD15028FF84145EFB9158BAFB60A881" box="[1149,1254,188,212]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="-76.65">76º39’W</geoCoordinate>
|
||
), department of Valle del Cauca; between altitudes of 1600 up to
|
||
<quantity id="C91D9B0AFF84145EFD5058D9FC97A8A2" box="[700,785,223,247]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="10" unit="m" value="2200.0">2200 m</quantity>
|
||
(
|
||
<figureCitation id="96DE2A6AFF84145EFCCC58D9FCE3A8A2" box="[800,869,223,247]" captionStart="FIGURE 3" captionStartId="5.[151,255,1139,1163]" captionTargetBox="[151,1436,194,1117]" captionTargetId="figure@5.[151,1436,194,1118]" captionTargetPageId="5" captionText="FIGURE 3. Geographic distribution of species of the tribe Epictini in Colombia. Type localities correspond to white symbols." httpUri="https://zenodo.org/record/199954/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46FF6564FF84145EFF255902FE5DAA0A" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="0E5A36EFFF84145EFF255902FE5DAA0A" blockId="9.[151,1437,151,607]" pageId="9" pageNumber="10">
|
||
<emphasis id="3C91EAFDFF84145EFF255902FEBBA949" bold="true" box="[201,317,260,284]" pageId="9" pageNumber="10">Remarks.</emphasis>
|
||
<bibRefCitation id="6A744B1EFF84145EFEA45903FE5DA949" author="Dunn" box="[328,475,260,284]" pageId="9" pageNumber="10" refString="Dunn, E. R. (1944) A review of the Colombian snakes of the families Typhlopidae and Leptotyphlopidae. Caldasia, 3, 47 - 55." type="journal article" year="1944">Dunn (1944)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="C9E54D6CFF84145EFDB05902FCDEA949" box="[604,856,260,284]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="joshuai">
|
||
<emphasis id="3C91EAFDFF84145EFDB05902FCDEA949" box="[604,856,260,284]" italics="true" pageId="9" pageNumber="10">Leptotyphlops joshuai</emphasis>
|
||
</taxonomicName>
|
||
on the basis of seven specimens from the Cauca Valley. However, the author did not provide the institutional number for all specimens in the
|
||
<typeStatus id="D15E884DFF84145EFB40592EFB5AA915" box="[1196,1244,296,320]" pageId="9" pageNumber="10">type</typeStatus>
|
||
series. Although La Salle
|
||
<typeStatus id="D15E884DFF84145EFF12594BFEBFA930" box="[254,313,333,357]" pageId="9" pageNumber="10">types</typeStatus>
|
||
actually are kept together in a separate cabinet from remaining collection, the specimen currently labelled as
|
||
<typeStatus id="D15E884DFF84145EFEFA5969FEFEA9D2" box="[278,376,367,391]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
did not match with morphometric and meristic data given in the original description. As most of the bottles and labels in the La Salle collection were replaced after the fire accident in 1948 (frequently without the maintenance of the original species label), many
|
||
<typeStatus id="D15E884DFF84145EFD0B59BEFCA4A985" box="[743,802,440,464]" pageId="9" pageNumber="10">types</typeStatus>
|
||
were mixed with specimens from the main collection or lost (P. Passos pers. obs.). Besides, the specimen separated in the type’s cabinet (MLS 11) was not labelled as such, while other leptotyphlopid
|
||
<typeStatus id="D15E884DFF84145EFE205A06FD81AA4D" box="[460,519,512,536]" pageId="9" pageNumber="10">types</typeStatus>
|
||
were properly labelled (e.g.,
|
||
<taxonomicName id="C9E54D6CFF84145EFCB75A07FC7BAA4D" box="[859,1021,512,536]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="brevissimus">
|
||
<emphasis id="3C91EAFDFF84145EFCB75A07FC7BAA4D" box="[859,1021,512,536]" italics="true" pageId="9" pageNumber="10">L. brevissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C9E54D6CFF84145EFBD45A07FB28AA4D" box="[1080,1198,512,536]" class="Reptilia" family="Leptotyphlopidae" genus="Rena" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nicefori">
|
||
<emphasis id="3C91EAFDFF84145EFBD45A07FB28AA4D" box="[1080,1198,512,536]" italics="true" pageId="9" pageNumber="10">L. nicefori</emphasis>
|
||
</taxonomicName>
|
||
). In this context, we here propose the recognition of MLS 13 as the
|
||
<typeStatus id="D15E884DFF84145EFD455A22FC8DAA69" box="[681,779,548,572]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
based on the similarity of morphometric and meristic data with the original description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |