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<document id="0065DA255A8D9A10166913FA2F349FB1" ID-CLB-Dataset="26998" ID-DOI="10.5852/ejt.2019.517" ID-GBIF-Dataset="d05987e2-6008-4deb-9773-1d988013beba" ID-Zenodo-Dep="2647891" ID-ZooBank="CAC6F8AF-ED37-4989-9672-68316920750B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555772398596" checkinUser="plazi" docAuthor="Fernandes, Maurício Romulo &amp; Pimenta, Alexandre Dias" docDate="2019" docId="426387C8B477FFE2FE37FC77982FFB2A" docLanguage="en" docName="ejt-517_fernandes_pimenta.pdf" docOrigin="European Journal of Taxonomy 517" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cheirodonta dupliniana Fernandes &amp; Pimenta 2019, comb. nov." docType="treatment" docVersion="11" lastPageNumber="15" masterDocId="BE5AFFB0B47CFFECFFDEFFCA9B24FFA1" masterDocTitle="Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil" masterLastPageNumber="60" masterPageNumber="1" pageNumber="12" updateTime="1698723356892" updateUser="ExternalLinkService">
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<mods:title id="36618899AA1C92B5C94BF54102DAFFB6">Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil</mods:title>
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<mods:namePart id="5D204FE390F9FF25317F10BE08D38C92">Fernandes, Maurício Romulo</mods:namePart>
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<mods:namePart id="3C07D29CF4ABADB30303887119E16CF4">Pimenta, Alexandre Dias</mods:namePart>
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<treatment id="426387C8B477FFE2FE37FC77982FFB2A" ID-DOI="http://doi.org/10.5281/zenodo.5586627" ID-GBIF-Taxon="156828145" ID-Zenodo-Dep="5586627" LSID="urn:lsid:plazi:treatment:426387C8B477FFE2FE37FC77982FFB2A" httpUri="http://treatment.plazi.org/id/426387C8B477FFE2FE37FC77982FFB2A" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
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<paragraph id="CA7536DEB477FFE7FE37FC779F6FFC79" blockId="11.[489,1099,957,1020]" box="[489,1099,957,985]" pageId="11" pageNumber="12">
<heading id="913D81B2B477FFE7FE37FC779F6FFC79" box="[489,1099,957,985]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="0DCA4D5DB477FFE7FE37FC7798E5FC78" ID-CoL="7QMXS" authority="(Olsson, 1916)" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[489,961,957,985]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB477FFE7FE37FC779828FC79" bold="true" box="[489,780,957,984]" italics="true" pageId="11" pageNumber="12">Cheirodonta dupliniana</emphasis>
(Olsson, 1916)
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<taxonomicNameLabel id="E38D57B7B477FFE7FC17FC749F6FFC79" box="[969,1099,958,984]" pageId="11" pageNumber="12" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="82D06555B477FFE7FD39FC2B9868FC5D" box="[743,844,993,1020]" pageId="11" pageNumber="12" type="description">
<paragraph id="CA7536DEB477FFE7FD39FC2B9868FC5D" blockId="11.[489,1099,957,1020]" box="[743,844,993,1020]" pageId="11" pageNumber="12">
<figureCitation id="52F12A5BB477FFE7FD39FC2B9868FC5D" box="[743,844,993,1020]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[189,232,1954,1980]" captionStartId-1="13.[189,232,1948,1974]" captionTargetBox-0="[189,1398,599,1913]" captionTargetBox-1="[189,1398,756,1907]" captionTargetId-0="figure@12.[189,1398,599,1915]" captionTargetId-1="figure@13.[189,1399,754,1907]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 5. Cheirodonta dupliniana (Olsson, 1916) comb. nov. AE. USNM 1438722. AD. Jaw, outer side. E. Operculum. Scale bars: A = 50 µm; B, D = 10 µm; C = 5 µm; E = 100 µm." captionText-1="Fig. 6. Cheirodonta dupliniana (Olsson, 1916) comb. nov. AD. Radula (USNM 1438722). Scale bars: AB, D = 5 µm; C = 10 µm." httpUri-0="https://zenodo.org/record/2647901/files/figure.png" httpUri-1="https://zenodo.org/record/2647903/files/figure.png" pageId="11" pageNumber="12">Figs 56</figureCitation>
</paragraph>
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<subSubSection id="82D06555B477FFE7FF63FBE99E65FB9C" box="[189,1345,1058,1085]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="CA7536DEB477FFE7FF63FBE99E65FB9C" blockId="11.[189,1345,1058,1085]" box="[189,1345,1058,1085]" pageId="11" pageNumber="12">
<treatmentCitation id="4B6B10CFB477FFE7FF63FBE999DCFB9C" box="[189,760,1058,1085]" pageId="11" pageNumber="12">
<taxonomicName id="0DCA4D5DB477FFE7FF63FBE999DCFB9C" ID-CoL="7QMXT" author="Fernandes M. R. &amp; PimentaA. D." authority="Fernandes &amp; Pimenta, 2015" authorityName="Fernandes &amp; Pimenta" authorityYear="2015" box="[189,760,1058,1085]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" journalOrPublisher="Zootaxa" kingdom="Animalia" pageId="11" pageNumber="57" pagination="493 - 513" part="4012" phylum="Mollusca" rank="species" refId="ref20415" species="mizifio" title="Five new species and two records of Triphorinae (Caenogastropoda, Triphoridae) from Brazil" type="journal article" year="2015">
<emphasis id="F8BEEACCB477FFE7FF63FBE99A81FB9C" box="[189,421,1059,1085]" italics="true" pageId="11" pageNumber="12">Cheirodonta mizifio</emphasis>
<bibRefCitation id="AE5B4B2FB477FFE7FE73FBE899DCFB9C" author="Fernandes M. R. &amp; PimentaA. D." box="[429,760,1058,1085]" journalOrPublisher="Zootaxa" pageId="11" pageNumber="57" pagination="493 - 513" part="4012" refId="ref20415" title="Five new species and two records of Triphorinae (Caenogastropoda, Triphoridae) from Brazil" type="journal article" year="2015">Fernandes &amp; Pimenta, 2015</bibRefCitation>
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; see Fernandes &amp; Pimenta in prep. for synonymy.
</paragraph>
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<subSubSection id="82D06555B477FFE7FF63FBAE9AA3FB71" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="CA7536DEB477FFE7FF63FBAE9A8EFBDF" blockId="11.[189,426,1124,1150]" box="[189,426,1124,1150]" pageId="11" pageNumber="12">
<heading id="913D81B2B477FFE7FF63FBAE9A8EFBDF" bold="true" box="[189,426,1124,1150]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="F8BEEACCB477FFE7FF63FBAE9A8EFBDF" bold="true" box="[189,426,1124,1150]" pageId="11" pageNumber="12">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB477FFE7FF63FB599AA3FB71" blockId="11.[189,1398,1169,1232]" pageId="11" pageNumber="12">
<materialsCitation id="7AA23C83B477FFE7FF63FB599AA7FB71" ID-GBIF-Occurrence="2236664067" collectingDate="1980-09-04" collectionCode="USNM" collectorName="RV Bagby" country="United States of America" elevation="35" latitude="30.904999" location="Georgia" longLatPrecision="20" longitude="-80.60333" pageId="11" pageNumber="12" specimenCode="USNM 1438722" specimenCount="1" stateProvince="Georgia">
<collectingCountry id="B2DD764EB477FFE7FF63FB599BD2FB0C" box="[189,246,1171,1197]" name="United States of America" pageId="11" pageNumber="12">USA</collectingCountry>
• [1, d] spec.;
<collectingRegion id="080EF83CB477FFE7FE6EFB589936FB0D" box="[432,530,1170,1196]" name="Georgia" pageId="11" pageNumber="12">Georgia</collectingRegion>
;
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,
<geoCoordinate id="AFFE5019B477FFE7FD12FB5B9849FB0C" box="[716,877,1169,1197]" degrees="80" direction="west" minutes="36" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="12" value="-80.60333">80°36´12″ W</geoCoordinate>
;
<elevation id="41E7D1EDB477FFE7FCA3FB58989FFB0C" box="[893,955,1170,1197]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="m" value="35.0">
<quantity id="0D329B3BB477FFE7FCA3FB58989FFB0C" box="[893,955,1170,1197]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="m" value="35.0">35 m</quantity>
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depth;
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<date id="BE74101EB477FFE7FBC7FB599F8AFB0C" box="[1049,1198,1171,1197]" pageId="11" pageNumber="12" value="1980-09-04">4 Sep. 1980</date>
</collectingDate>
;
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RV
<emphasis id="F8BEEACCB477FFE7FB30FB599E1DFB0C" box="[1262,1337,1171,1197]" italics="true" pageId="11" pageNumber="12">Bagby</emphasis>
</collectorName>
leg.;
<specimenCode id="9A6C9EA5B477FFE7FF63FB7C9AA7FB71" box="[189,387,1206,1232]" pageId="11" pageNumber="12">USNM 1438722</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="82D06555B477FFE7FF63FB3D9E53F873" pageId="11" pageNumber="12" type="description">
<paragraph id="CA7536DEB477FFE7FF63FB3D990DFAB0" blockId="11.[189,553,1271,1297]" box="[189,553,1271,1297]" pageId="11" pageNumber="12">
<heading id="913D81B2B477FFE7FF63FB3D990DFAB0" bold="true" box="[189,553,1271,1297]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="F8BEEACCB477FFE7FF63FB3D990DFAB0" bold="true" box="[189,553,1271,1297]" pageId="11" pageNumber="12">Description of basic anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB477FFE7FF63FAEC9866FAC2" blockId="11.[189,1399,1318,1379]" pageId="11" pageNumber="12">OPERCULUM. Ovate-elliptical, thin, semi-transparent, membranous, poorly distinct whorls, nucleus subcentral, dislocated 22% from center toward margin.</paragraph>
<paragraph id="CA7536DEB477FFE7FF63FA419993F9F5" blockId="11.[189,1399,1418,1621]" pageId="11" pageNumber="12">
JAW. Wing-shaped; outer side with scales rectangular/squared, rectangular-bilobed, acute-lanceolate or puzzle-like; rectangular scales 9.0
<quantity id="0D329B3BB477FFE7FD4CFA6499DCFA69" box="[658,760,1454,1480]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="11" pageNumber="12" unit="mm" value="10.3">10.3 µm</quantity>
long,
<quantity id="0D329B3BB477FFE7FC93FA6798FEFA69" box="[845,986,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.1" metricValueMin="5.2" pageId="11" pageNumber="12" unit="mm" value="5.65" valueMax="6.1" valueMin="5.2">5.26.1 µm</quantity>
wide, ratio length/width 1.61.9, rectangular-bilobed scales
<quantity id="0D329B3BB477FFE7FDDDFA1B99BBFA4A" box="[515,671,1489,1515]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.03" metricValueMin="0.99" pageId="11" pageNumber="12" unit="mm" value="10.1" valueMax="10.3" valueMin="9.9">9.910.3 µm</quantity>
long,
<quantity id="0D329B3BB477FFE7FD2BFA1A98A6FA4A" box="[757,898,1488,1515]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.2" metricValueMin="4.6" pageId="11" pageNumber="12" unit="mm" value="4.9" valueMax="5.2" valueMin="4.6">4.65.2 µm</quantity>
wide, ratio length/width 2.02.2, acutelanceolate scales
<quantity id="0D329B3BB477FFE7FE53FA3E9910F9AF" box="[397,564,1524,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" metricValueMax="1.81" metricValueMin="1.67" pageId="11" pageNumber="12" unit="mm" value="17.4" valueMax="18.1" valueMin="16.7">16.718.1 µm</quantity>
long,
<quantity id="0D329B3BB477FFE7FD5CFA3E9828F9AF" box="[642,780,1524,1551]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.7" metricValueMin="5.6" pageId="11" pageNumber="12" unit="mm" value="6.15" valueMax="6.7" valueMin="5.6">5.66.7 µm</quantity>
wide, ratio length/width 2.63.2, puzzle-like scales
<quantity id="0D329B3BB477FFE7FF63F9DD9A40F993" box="[189,356,1559,1586]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.56" metricValueMin="1.29" pageId="11" pageNumber="12" unit="mm" value="14.25" valueMax="15.6" valueMin="12.9">12.915.6 µm</quantity>
long (largest lobe to largest lobe),
<quantity id="0D329B3BB477FFE7FCD5F9DD98B0F993" box="[779,916,1559,1586]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="9.9" metricValueMin="5.6" pageId="11" pageNumber="12" unit="mm" value="7.75" valueMax="9.9" valueMin="5.6">5.69.9 µm</quantity>
wide (perpendicular to the length at the center of scale), ratio length/width 1.42.8.
</paragraph>
<paragraph id="CA7536DEB477FFE7FF63F9B69E53F873" blockId="11.[189,1399,1660,2003]" pageId="11" pageNumber="12">
RADULA. Formula 8-1-1-1-8; central tooth usually with seven cusps, median one (cusp 4) is the smallest or even absent, cusps 1, 3, 5 and 7 medium-sized, cusps 2 and 6 considerably broader and robust (1.9 3.0× longer than cusp 4); lateral teeth with seven main cusps, all with similar length, and an additional small one (cusp 1); M1 with eight to nine cusps, with an increasing length from cusp 1 to cusp 7 (or cusp 8, when nine cusps are present), the latter usually being very elongated, up to 1.7× longer than median cusps, and with a filiform distal half (cusp 6 or
<date id="BE74101EB477FFE7FD0CF8E6983DF8E7" box="[722,793,1836,1862]" pageId="11" pageNumber="12">7 may</date>
also have a filiform elongation); remaining marginal teeth hand-like, with a long basal plate, usually with nine or ten cusps, also increasing in size from cusp 1 to cusps 7 (or 8) and 8 (or 9), the latter cusps usually having a filiform elongation in the distal half, up to 2.3× longer than median cusps; central tooth 4.95.0 µm wide, lateral teeth
<quantity id="0D329B3BB477FFE7FBA6F85F9FDBF811" box="[1144,1279,1941,1968]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="11" pageNumber="12" unit="mm" value="4.35" valueMax="4.6" valueMin="4.1">4.14.6 µm</quantity>
wide, M1
<quantity id="0D329B3BB477FFE7FF63F8739A67F872" box="[189,323,1977,2003]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="11" pageNumber="12" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.14.9 µm</quantity>
wide, M2 usually
<quantity id="0D329B3BB477FFE7FDC7F8729948F872" box="[537,620,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="11" pageNumber="12" unit="mm" value="5.4">5.4 µm</quantity>
wide, M3
<quantity id="0D329B3BB477FFE7FD36F872984AF872" box="[744,878,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="11" pageNumber="12" unit="mm" value="5.05" valueMax="5.3" valueMin="4.8">4.85.3 µm</quantity>
wide, outer marginal teeth
<quantity id="0D329B3BB477FFE7FB74F8729E14F872" box="[1194,1328,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="11" pageNumber="12" unit="mm" value="5.3" valueMax="5.4" valueMin="5.2">5.25.4 µm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="82D06555B470FFE2FF63FEC1982FFB2A" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="discussion">
<paragraph id="CA7536DEB470FFE0FF63FEC19A0BFE84" blockId="12.[189,303,267,293]" box="[189,303,267,293]" pageId="12" pageNumber="13">
<heading id="913D81B2B470FFE0FF63FEC19A0BFE84" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="F8BEEACCB470FFE0FF63FEC19A0BFE84" bold="true" box="[189,303,267,293]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB470FFE1FF63FEF39AACFECD" blockId="12.[189,1399,313,551]" lastBlockId="13.[189,1399,267,364]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
The operculum of
<taxonomicName id="0DCA4D5DB470FFE0FE4DFEF09917FEF2" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[403,563,313,340]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB470FFE0FE4DFEF09A83FEF5" box="[403,423,314,340]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="F8BEEACCB470FFE0FE6AFEF39917FEF2" box="[436,563,313,339]" italics="true" pageId="12" pageNumber="13">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B470FFE0FDE4FEF3999EFEF5" box="[570,698,313,340]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
agrees with that of the
<typeStatus id="1571887CB470FFE0FC19FEF098DEFEF5" box="[967,1018,314,340]" pageId="12" pageNumber="13">type</typeStatus>
species
<taxonomicName id="0DCA4D5DB470FFE0FB85FEF09FD2FEF2" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[1115,1270,313,340]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB470FFE0FB85FEF09F4BFEF5" box="[1115,1135,314,340]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="F8BEEACCB470FFE0FBA2FEF39FD2FEF2" box="[1148,1270,313,339]" italics="true" pageId="12" pageNumber="13">pallescens</emphasis>
</taxonomicName>
(described in
<bibRefCitation id="AE5B4B2FB470FFE0FF01FE979AAFFED6" author="Bouchet P." box="[223,395,348,375]" journalOrPublisher="Lavori della Societa Italiana di Malacologia" pageId="12" pageNumber="13" pagination="5 - 58" part="21" refId="ref20065" refString="Bouchet P. 1985. Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda). Lavori della Societa Italiana di Malacologia 21: 5 - 58." title="Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda)" type="journal article" year="1985">Bouchet 1985</bibRefCitation>
). Its radula has a superficial resemblance with the description of
<taxonomicName id="0DCA4D5DB470FFE0FB70FE979E6AFED6" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[1198,1358,349,375]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB470FFE0FB70FE979FE6FED6" box="[1198,1218,349,375]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="F8BEEACCB470FFE0FB0AFE979E6AFED6" box="[1236,1358,349,375]" italics="true" pageId="12" pageNumber="13">pallescens</emphasis>
</taxonomicName>
by
<bibRefCitation id="AE5B4B2FB470FFE0FF63FE4A990DFE3B" author="Bouchet P. &amp; Guillemot H." box="[189,553,384,410]" journalOrPublisher="Journal of Molluscan Studies" pageId="12" pageNumber="13" pagination="344 - 356" part="44" refId="ref20123" refString="Bouchet P. &amp; Guillemot H. 1978. The Triphora perversa - complex in Western Europe. Journal of Molluscan Studies 44: 344 - 356." title="The Triphora perversa - complex in Western Europe" type="journal article" year="1978">Bouchet &amp; Guillemot (1978)</bibRefCitation>
and
<bibRefCitation id="AE5B4B2FB470FFE0FDACFE4A9810FE3A" author="Bouchet P." box="[626,820,384,411]" journalOrPublisher="Lavori della Societa Italiana di Malacologia" pageId="12" pageNumber="13" pagination="5 - 58" part="21" refId="ref20065" refString="Bouchet P. 1985. Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda). Lavori della Societa Italiana di Malacologia 21: 5 - 58." title="Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda)" type="journal article" year="1985">Bouchet (1985)</bibRefCitation>
, despite significant differences:
<taxonomicName id="0DCA4D5DB470FFE0FB13FE4A9E52FE3B" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[1229,1398,384,410]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB470FFE0FB13FE4A9FC5FE3B" box="[1229,1249,384,410]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="F8BEEACCB470FFE0FB29FE4A9E52FE3B" box="[1271,1398,384,410]" italics="true" pageId="12" pageNumber="13">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B470FFE0FF63FE699A64FE1F" box="[189,320,419,446]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
has more marginal teeth (eight, instead of five or six), a completely different central tooth (seven cusps with varied sizes, instead of six to eight homogeneous cusps separated in two groups of three or four cusps by an internal diastema), lateral teeth with seven main cusps plus a smaller one (instead of six cusps in
<bibRefCitation id="AE5B4B2FB470FFE0FE0CFDC79952FD86" author="Bouchet P." box="[466,630,524,551]" journalOrPublisher="Lavori della Societa Italiana di Malacologia" pageId="12" pageNumber="13" pagination="5 - 58" part="21" refId="ref20065" refString="Bouchet P. 1985. Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda). Lavori della Societa Italiana di Malacologia 21: 5 - 58." title="Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda)" type="journal article" year="1985">Bouchet 1985</bibRefCitation>
but seven or eight cusps were similarly indicated in
<bibRefCitation id="AE5B4B2FB470FFE1FB28FDC79A5DFE84" author="Bouchet P. &amp; Guillemot H." journalOrPublisher="Journal of Molluscan Studies" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" pagination="344 - 356" part="44" refId="ref20123" refString="Bouchet P. &amp; Guillemot H. 1978. The Triphora perversa - complex in Western Europe. Journal of Molluscan Studies 44: 344 - 356." title="The Triphora perversa - complex in Western Europe" type="journal article" year="1978">Bouchet &amp; Guillemot 1978</bibRefCitation>
), and marginal teeth hand-like with a maximum of 10 long cusps distributed along their length (instead of broad spoon-like teeth with 12 to 20 cusps concentrated in the rounded end of teeth in
<taxonomicName id="0DCA4D5DB471FFE1FF05FE989A53FECD" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[219,375,338,364]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB471FFE1FF05FE989BCBFECD" box="[219,239,338,364]" italics="true" pageId="13" pageNumber="14">C</emphasis>
.
<emphasis id="F8BEEACCB471FFE1FF23FE989A53FECD" box="[253,375,338,364]" italics="true" pageId="13" pageNumber="14">pallescens</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="9EB56656B470FFE0FF63F8689F68F841" httpUri="https://zenodo.org/record/2647901/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1954,1980]" targetBox="[189,1398,599,1913]" targetPageId="12">
<paragraph id="CA7536DEB470FFE0FF63F8689F68F841" blockId="12.[189,1399,1954,2016]" pageId="12" pageNumber="13">
<emphasis id="F8BEEACCB470FFE0FF63F8689A2DF81D" bold="true" box="[189,265,1954,1980]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
<taxonomicName id="0DCA4D5DB470FFE0FECEF86899F8F81C" authority="(Olsson, 1916)" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[272,732,1954,1981]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB470FFE0FECEF868990CF81D" box="[272,552,1954,1980]" italics="true" pageId="12" pageNumber="13">Cheirodonta dupliniana</emphasis>
(Olsson, 1916)
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B470FFE0FD3CF8689847F81D" box="[738,867,1954,1980]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="F8BEEACCB470FFE0FCB7F8689884F81D" bold="true" box="[873,928,1954,1980]" pageId="12" pageNumber="13">AE</emphasis>
. USNM 1438722.
<emphasis id="F8BEEACCB470FFE0FBA3F8689F92F81D" bold="true" box="[1149,1206,1954,1980]" pageId="12" pageNumber="13">AD</emphasis>
. Jaw, outer side.
<emphasis id="F8BEEACCB470FFE0FF63F80F9BF5F87E" bold="true" box="[189,209,1989,2015]" pageId="12" pageNumber="13">E</emphasis>
. Operculum. Scale bars: A = 50 µm; B, D = 10 µm; C = 5 µm; E = 100 µm.
</paragraph>
</caption>
<paragraph id="CA7536DEB471FFE2FF63FE5999A9FE74" blockId="13.[189,1399,403,711]" lastBlockId="14.[189,1399,267,470]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
There are significant differences in the illustrated radulae of
<taxonomicName id="0DCA4D5DB471FFE1FC5AFE5E9F05FE0C" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[900,1057,403,430]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB471FFE1FC5AFE5E98BCFE0F" box="[900,920,404,430]" italics="true" pageId="13" pageNumber="14">C</emphasis>
.
<emphasis id="F8BEEACCB471FFE1FC79FE599F05FE0C" box="[935,1057,403,429]" italics="true" pageId="13" pageNumber="14">pallescens</emphasis>
</taxonomicName>
from French material (based on specimens from the eastern Atlantic and Mediterranean) described by
<bibRefCitation id="AE5B4B2FB471FFE1FBFFFE7C9E57FE71" author="Bouchet P. &amp; Guillemot H." box="[1057,1395,438,464]" journalOrPublisher="Journal of Molluscan Studies" pageId="13" pageNumber="14" pagination="344 - 356" part="44" refId="ref20123" refString="Bouchet P. &amp; Guillemot H. 1978. The Triphora perversa - complex in Western Europe. Journal of Molluscan Studies 44: 344 - 356." title="The Triphora perversa - complex in Western Europe" type="journal article" year="1978">Bouchet &amp; Guillemot (1978</bibRefCitation>
: fig. 19) and
<bibRefCitation id="AE5B4B2FB471FFE1FE9CFE139AD5FE55" author="Bouchet P." box="[322,497,473,500]" journalOrPublisher="Lavori della Societa Italiana di Malacologia" pageId="13" pageNumber="14" pagination="5 - 58" part="21" refId="ref20065" refString="Bouchet P. 1985. Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda). Lavori della Societa Italiana di Malacologia 21: 5 - 58." title="Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda)" type="journal article" year="1985">Bouchet (1985</bibRefCitation>
: figs 1011), especially regarding the central and lateral teeth. More differences emerge when considering the drawn radula of specimens from southern
<collectingCountry id="B2DD764EB471FFE1FBECFE379FB0FDB6" box="[1074,1172,509,535]" name="United Kingdom" pageId="13" pageNumber="14">England</collectingCountry>
by
<bibRefCitation id="AE5B4B2FB471FFE1FB16FE379E56FDB9" author="Fretter V." box="[1224,1394,509,536]" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="13" pageNumber="14" pagination="567 - 586" part="29" refId="ref20563" refString="Fretter V. 1951. Observations on the life history and functional morphology of Cerithiopsis tubercularis (Montagu) and Triphora perversa (L.). Journal of the Marine Biological Association of the United Kingdom 29: 567 - 586. https: // doi. org / 10.1017 / S 0025315400052784" title="Observations on the life history and functional morphology of Cerithiopsis tubercularis (Montagu) and Triphora perversa (L.)" type="journal article" year="1951">Fretter (1951)</bibRefCitation>
, bearing lateral teeth with nine cusps and marginal teeth with much longer cusps than those illustrated by Bouchet, in addition to a presumably different axis orientation of marginal teeth (downward instead of sideward). With the current limited amount of information, it is uncertain whether
<taxonomicName id="0DCA4D5DB471FFE1FB08FDAD9E52FD21" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[1238,1398,614,641]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB471FFE1FB08FDAD9FCEFD20" box="[1238,1258,615,641]" italics="true" pageId="13" pageNumber="14">C</emphasis>
.
<emphasis id="F8BEEACCB471FFE1FB22FDAC9E52FD21" box="[1276,1398,614,640]" italics="true" pageId="13" pageNumber="14">pallescens</emphasis>
</taxonomicName>
represents a single species with a variable radular morphology, or whether the Mediterranean population and perhaps also the population from
<collectingCountry id="B2DD764EB471FFE1FD5CFD6799C0FD66" box="[642,740,685,711]" name="United Kingdom" pageId="13" pageNumber="14">England</collectingCountry>
constitute one or two new species. The morphology of the central and marginal teeth of specimens from
<collectingCountry id="B2DD764EB472FFE2FCC4FEC6984EFE87" box="[794,874,268,294]" name="France" pageId="14" pageNumber="15">France</collectingCountry>
(especially from Mediterranean) is slightly similar to that of
<taxonomicName id="0DCA4D5DB472FFE2FE51FEE59F5EFEE8" authority="(Rolan &amp; Fernandez-Garces, 1995)" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1995" box="[399,1146,302,329]" class="Gastropoda" family="Triphoridae" genus="Sagenotriphora" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="osclausum">
<emphasis id="F8BEEACCB472FFE2FE51FEE599E9FEE8" box="[399,717,303,329]" italics="true" pageId="14" pageNumber="15">Sagenotriphora osclausum</emphasis>
(
<bibRefCitation id="AE5B4B2FB472FFE2FD00FEE49F56FEE8" author="Rolan E. &amp; Fernandez-Garces R." box="[734,1138,302,329]" journalOrPublisher="Apex" pageId="14" pageNumber="15" pagination="9 - 24" part="10" refId="ref21693" refString="Rolan E. &amp; Fernandez-Garces R. 1995. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 5. The genera Marshallora, Mesophora, Similiphora, Eutriphora, Latitriphora, Aclophora and other species without generic affiliation. Apex 10: 9 - 24." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 5. The genera Marshallora, Mesophora, Similiphora, Eutriphora, Latitriphora, Aclophora and other species without generic affiliation." type="journal article" year="1995">Rolán &amp; Fernández-Garcés, 1995</bibRefCitation>
)
</taxonomicName>
(see below), in spite of consistent differences mainly related to the lateral teeth. Considering only species from the Atlantic, both genera show similarities in shell (e.g., supranumerical cords, smooth subperipheral and basal cords, similar protoconch sculpture) and radula, as herein indicated. They may be proved to be phylogenetically close after a comprehensive phylogeny.
</paragraph>
<caption id="9EB56656B471FFE1FF63F856992FF87B" httpUri="https://zenodo.org/record/2647903/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1948,1974]" targetBox="[189,1398,756,1907]" targetPageId="13">
<paragraph id="CA7536DEB471FFE1FF63F856992FF87B" blockId="13.[189,1399,1948,2010]" pageId="13" pageNumber="14">
<emphasis id="F8BEEACCB471FFE1FF63F8569A2EF817" bold="true" box="[189,266,1948,1974]" pageId="13" pageNumber="14">Fig. 6.</emphasis>
<taxonomicName id="0DCA4D5DB471FFE1FECFF85699C4F816" authority="(Olsson, 1916)" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[273,736,1948,1975]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB471FFE1FECFF856990FF817" box="[273,555,1948,1974]" italics="true" pageId="13" pageNumber="14">Cheirodonta dupliniana</emphasis>
(Olsson, 1916)
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B471FFE1FD36F856984DF816" box="[744,873,1948,1975]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="F8BEEACCB471FFE1FCAEF856988DF817" bold="true" box="[880,937,1948,1974]" pageId="13" pageNumber="14">AD</emphasis>
. Radula (USNM 1438722). Scale bars: AB, D = 5 µm; C = 10 µm.
</paragraph>
</caption>
<paragraph id="CA7536DEB472FFE2FF63FE379A00FCF2" blockId="14.[189,1399,509,851]" pageId="14" pageNumber="15">
The radula of the southwestern Pacific species
<taxonomicName id="0DCA4D5DB472FFE2FCD0FE379FD7FDB9" authority="(A. Adams, 1854)" baseAuthorityName="A. Adams" baseAuthorityYear="1854" box="[782,1267,509,536]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="labiata">
<emphasis id="F8BEEACCB472FFE2FCD0FE379F21FDB6" box="[782,1029,509,535]" italics="true" pageId="14" pageNumber="15">Cheirodonta labiata</emphasis>
(A. Adams, 1854)
</taxonomicName>
, described and illustrated by
<bibRefCitation id="AE5B4B2FB472FFE2FE42FDEA9940FD9B" author="Marshall B. A." box="[412,612,544,570]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="14" pageNumber="15" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall (1983)</bibRefCitation>
, is much similar to that of
<taxonomicName id="0DCA4D5DB472FFE2FC68FDEA9F79FD9B" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[950,1117,544,570]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FC68FDEA98EEFD9B" box="[950,970,544,570]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FC05FDEA9F79FD9B" box="[987,1117,544,570]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FBB9FDEA9FCAFD9B" box="[1127,1262,544,570]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
They have the same number of marginal teeth (eight), a similar number of cusps and dimensions of the central tooth (
<taxonomicName id="0DCA4D5DB472FFE2FED0FDAD9A9BFD21" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[270,447,614,641]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FED0FDAD9A06FD20" box="[270,290,615,641]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FEE3FDAC9A9BFD21" box="[317,447,614,640]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FE16FDAC996BFD20" box="[456,591,614,641]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
usually with seven heterogeneous cusps, 5.0 µm wide;
<taxonomicName id="0DCA4D5DB472FFE2FB23FDAD9E53FD21" baseAuthorityName="A. Adams" baseAuthorityYear="1854" box="[1277,1399,614,641]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="labiata">
<emphasis id="F8BEEACCB472FFE2FB23FDAD9E35FD20" box="[1277,1297,615,641]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FAFCFDAC9E53FD21" box="[1314,1399,614,640]" italics="true" pageId="14" pageNumber="15">labiata</emphasis>
</taxonomicName>
with seven to nine heterogeneous cusps,
<quantity id="0D329B3BB472FFE2FD78FD4399DFFD05" box="[678,763,649,676]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="15" unit="mm" value="6.8">6.8 µm</quantity>
wide), idem for lateral teeth (both usually with eight cusps;
<quantity id="0D329B3BB472FFE2FECFFD679ABFFD66" box="[273,411,685,711]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="14" pageNumber="15" unit="mm" value="4.35" valueMax="4.6" valueMin="4.1">4.14.6 µm</quantity>
wide in
<taxonomicName id="0DCA4D5DB472FFE2FDD8FD679988FD66" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[518,684,685,711]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FDD8FD67993EFD66" box="[518,538,685,711]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FDF4FD679988FD66" box="[554,684,685,711]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FD6AFD679818FD66" box="[692,828,685,711]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
,
<quantity id="0D329B3BB472FFE2FC94FD6798BBFD66" box="[842,927,685,711]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="15" unit="mm" value="6.8">6.8 µm</quantity>
wide in
<taxonomicName id="0DCA4D5DB472FFE2FBD5FD679FA0FD66" baseAuthorityName="A. Adams" baseAuthorityYear="1854" box="[1035,1156,685,711]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="labiata">
<emphasis id="F8BEEACCB472FFE2FBD5FD679F3BFD66" box="[1035,1055,685,711]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FBF1FD679FA0FD66" box="[1071,1156,685,711]" italics="true" pageId="14" pageNumber="15">labiata</emphasis>
</taxonomicName>
) and marginal teeth (M1M3 with eight to ten cusps in
<taxonomicName id="0DCA4D5DB472FFE2FDB0FD1A9830FD4B" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[622,788,720,746]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FDB0FD1A99A6FD4B" box="[622,642,720,746]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FD4CFD1A9830FD4B" box="[658,788,720,746]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FCC3FD1A9881FD4B" box="[797,933,720,746]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
,
<quantity id="0D329B3BB472FFE2FC6DFD1A9F1AFD4B" box="[947,1086,720,747]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.4" metricValueMin="4.1" pageId="14" pageNumber="15" unit="mm" value="4.75" valueMax="5.4" valueMin="4.1">4.15.4 µm</quantity>
wide; M1M3 with eight to nine cusps in
<taxonomicName id="0DCA4D5DB472FFE2FE5EFD3E9ADCFCAC" baseAuthorityName="A. Adams" baseAuthorityYear="1854" box="[384,504,755,782]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="labiata">
<emphasis id="F8BEEACCB472FFE2FE5EFD3E9AB0FCAF" box="[384,404,756,782]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FE7DFD399ADCFCAC" box="[419,504,755,781]" italics="true" pageId="14" pageNumber="15">labiata</emphasis>
</taxonomicName>
, about 5.0 µm wide). In this sense, the radula of
<taxonomicName id="0DCA4D5DB472FFE2FB98FD3E9FCFFCAC" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[1094,1259,755,782]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FB98FD3E9F7EFCAF" box="[1094,1114,756,782]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FBB7FD399FCFFCAC" box="[1129,1259,755,781]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FB2DFD399E53FCAF" box="[1267,1399,755,782]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
is much more similar to
<taxonomicName id="0DCA4D5DB472FFE2FE3EFCDD997CFC91" baseAuthorityName="A. Adams" baseAuthorityYear="1854" box="[480,600,790,817]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="labiata">
<emphasis id="F8BEEACCB472FFE2FE3EFCDD9AD0FC90" box="[480,500,791,817]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FDDDFCDC997CFC91" box="[515,600,790,816]" italics="true" pageId="14" pageNumber="15">labiata</emphasis>
</taxonomicName>
than to
<taxonomicName id="0DCA4D5DB472FFE2FD64FCDD987DFC91" baseAuthorityName="Jeffreys" baseAuthorityYear="1867" box="[698,857,790,817]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="pallescens">
<emphasis id="F8BEEACCB472FFE2FD64FCDD99EAFC90" box="[698,718,791,817]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FD02FCDC987DFC91" box="[732,857,790,816]" italics="true" pageId="14" pageNumber="15">pallescens</emphasis>
</taxonomicName>
, despite the opposite can be affirmed to shell features.
</paragraph>
<paragraph id="CA7536DEB472FFE2FF63FCB1982FFB2A" blockId="14.[189,1399,891,1164]" pageId="14" pageNumber="15">
Radulae of Caribbean species of
<taxonomicName id="0DCA4D5DB472FFE2FD93FCB198A3FC37" authority="Laseron, 1958" authorityName="Laseron" authorityYear="1958" box="[589,903,891,918]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB472FFE2FD93FCB199F7FC34" box="[589,723,891,917]" italics="true" pageId="14" pageNumber="15">Nanaphora</emphasis>
Laseron, 1958
</taxonomicName>
illustrated in
<bibRefCitation id="AE5B4B2FB472FFE2FBEEFCB19A2AFC19" author="Rolan E. &amp; Fernandez-Garces R." journalOrPublisher="Apex" pageId="14" pageNumber="15" pagination="17 - 27" part="9" refId="ref21646" refString="Rolan E. &amp; Fernandez-Garces R. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species. Apex 9: 17 - 27." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species." type="journal article" year="1994">Rolán &amp; Fernández-Garcés (1994)</bibRefCitation>
, at that time under the name
<taxonomicName id="0DCA4D5DB472FFE2FDBEFC5499D7FC19" authorityName="Marshall" authorityYear="1983" box="[608,755,926,952]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB472FFE2FDBEFC5499D7FC19" box="[608,755,926,952]" italics="true" pageId="14" pageNumber="15">Cheirodonta</emphasis>
</taxonomicName>
, are also superficially similar to those of
<taxonomicName id="0DCA4D5DB472FFE2FB0BFC559E52FC19" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" baseAuthorityName="Olsson" baseAuthorityYear="1916" box="[1237,1398,926,953]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="dupliniana" status="comb. nov.">
<emphasis id="F8BEEACCB472FFE2FB0BFC559FCDFC18" box="[1237,1257,927,953]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="F8BEEACCB472FFE2FB29FC549E52FC19" box="[1271,1398,926,952]" italics="true" pageId="14" pageNumber="15">dupliniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38D57B7B472FFE2FF63FC0B9A67FC7D" box="[189,323,961,988]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
, particularly regarding the elongation of the basal plate of marginal teeth. Another species of
<taxonomicName id="0DCA4D5DB472FFE2FF01FC2F9A41FC5E" authorityName="Laseron" authorityYear="1958" box="[223,357,997,1023]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB472FFE2FF01FC2F9A41FC5E" box="[223,357,997,1023]" italics="true" pageId="14" pageNumber="15">Nanaphora</emphasis>
</taxonomicName>
from the Pacific was already indicated as similar to
<taxonomicName id="0DCA4D5DB472FFE2FC3AFC2F9F53FC5E" authorityName="Marshall" authorityYear="1983" box="[996,1143,997,1023]" class="Gastropoda" family="Triphoridae" genus="Cheirodonta" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB472FFE2FC3AFC2F9F53FC5E" box="[996,1143,997,1023]" italics="true" pageId="14" pageNumber="15">Cheirodonta</emphasis>
</taxonomicName>
in terms of marginal tooth morphology (
<bibRefCitation id="AE5B4B2FB472FFE2FE7AFBC29972FB83" author="Marshall B. A." box="[420,598,1032,1058]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="14" pageNumber="15" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall 1983</bibRefCitation>
). However, shell and operculum are consistently different between both genera (
<bibRefCitation id="AE5B4B2FB472FFE2FE88FBE19921FBE4" author="Marshall B. A." box="[342,517,1067,1093]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="14" pageNumber="15" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall 1983</bibRefCitation>
; Fernandes &amp; Pimenta 2015). Phylogenetic hypotheses will reveal whether these genera are analogous in radular morphology owing to feeding on similar sponge hosts, or whether they share a common and recent ancestral lineage.
</paragraph>
</subSubSection>
</treatment>
</document>