treatments-xml/data/42/63/87/426387C8B461FFCCFE37FB859AE9FB0B.xml

<document id="0EA6885D0E922C1CE97E2FDFA1F9CCD4" ID-CLB-Dataset="26998" ID-DOI="10.5852/ejt.2019.517" ID-GBIF-Dataset="d05987e2-6008-4deb-9773-1d988013beba" ID-Zenodo-Dep="2647891" ID-ZooBank="CAC6F8AF-ED37-4989-9672-68316920750B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555772398596" checkinUser="plazi" docAuthor="Fernandes, Maurício Romulo &amp; Pimenta, Alexandre Dias" docDate="2019" docId="426387C8B461FFCCFE37FB859AE9FB0B" docLanguage="en" docName="ejt-517_fernandes_pimenta.pdf" docOrigin="European Journal of Taxonomy 517" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nanaphora verbernei  1989" docType="treatment" docVersion="12" lastPageNumber="33" masterDocId="BE5AFFB0B47CFFECFFDEFFCA9B24FFA1" masterDocTitle="Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil" masterLastPageNumber="60" masterPageNumber="1" pageNumber="30" updateTime="1698723356892" updateUser="ExternalLinkService">
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<mods:title id="A2B94561802FCCC26372E45866128268">Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil</mods:title>
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<mods:namePart id="67708B3CEC88180CD6C743705C78196E">Fernandes, Maurício Romulo</mods:namePart>
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<mods:namePart id="D8DB905CCD2A17E679C4EDE2CF06D0AF">Pimenta, Alexandre Dias</mods:namePart>
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<mods:title id="A4DAD243C737327DF31EB2F4BDC36E22">European Journal of Taxonomy</mods:title>
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<mods:date id="C5626DE273FD3C020E4B1FCEF8797BD9">2019</mods:date>
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<treatment id="426387C8B461FFCCFE37FB859AE9FB0B" ID-DOI="http://doi.org/10.5281/zenodo.5586647" ID-GBIF-Taxon="156828161" ID-Zenodo-Dep="5586647" LSID="urn:lsid:plazi:treatment:426387C8B461FFCCFE37FB859AE9FB0B" httpUri="http://treatment.plazi.org/id/426387C8B461FFCCFE37FB859AE9FB0B" lastPageId="32" lastPageNumber="33" pageId="29" pageNumber="30">
<subSubSection id="82D06555B461FFF1FE37FB859F6EFBCB" box="[489,1098,1103,1130]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="CA7536DEB461FFF1FE37FB859F6EFBCB" blockId="29.[489,1098,1103,1165]" box="[489,1098,1103,1130]" pageId="29" pageNumber="30">
<heading id="913D81B2B461FFF1FE37FB859F6EFBCB" box="[489,1098,1103,1130]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="0DCA4D5DB461FFF1FE37FB859F6EFBCB" ID-CoL="45LPW" authority="(Moolenbeek &amp; Faber, 1989)" authorityName="" authorityYear="1989" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[489,1098,1103,1130]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB461FFF1FE37FB8599C9FBCB" bold="true" box="[489,749,1103,1130]" italics="true" pageId="29" pageNumber="30">Nanaphora verbernei</emphasis>
(Moolenbeek &amp; Faber, 1989)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="82D06555B461FFF1FD61FBB99850FB2C" box="[703,884,1139,1165]" pageId="29" pageNumber="30" type="description">
<paragraph id="CA7536DEB461FFF1FD61FBB99850FB2C" blockId="29.[489,1098,1103,1165]" box="[703,884,1139,1165]" pageId="29" pageNumber="30">
<figureCitation id="52F12A5BB461FFF1FD61FBB9983BFB2C" box="[703,799,1139,1165]" captionStart="Fig" captionStartId="7.[189,232,1707,1733]" captionTargetBox="[189,1398,742,1666]" captionTargetId="figure@7.[189,1399,742,1666]" captionTargetPageId="7" captionText="Fig. 2. Specimens studied. A. Metaxia excelsa Faber &amp; Moolenbeek, 1991 (MZSP 122353). B. Metaxia rugulosa (C.B. Adams, 1850) (BMSM 56034). C. Cosmotriphora melanura (C.B. Adams, 1850) (MNRJ 33980). D. Iniforis pseudothomae Rolán &amp; Fernández-Garcés, 1993 (MZSP 100957). E. Latitriphora albida (A. Adams, 1854) (BMSM 55442). F. Monophorus olivaceus (Dall, 1889) (MNRJ 34615). G. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989) (MZSP 85022). H. Nototriphora decorata (C.B. Adams, 1850) (MNRJ 28941). I. Sagenotriphora osclausum (Rolán &amp; Fernández-Garcés, 1995) (MNRJ 35061). J. Similiphora intermedia (C.B. Adams, 1850) (MNRJ 29763). K. Strobiligera gaesona (Dall, 1927) (MZSP 53685). L. “Inella” harryleei Rolán &amp; Fernández-Garcés, 2008 (FLMNH 450495). Scale bars: 1 mm." httpUri="https://zenodo.org/record/2647895/files/figure.png" pageId="29" pageNumber="30">Figs 2G</figureCitation>
, 
<figureCitation id="52F12A5BB461FFF1FCF5FBB99850FB2C" box="[811,884,1139,1165]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="30.[189,232,1915,1941]" captionStartId-1="31.[189,232,1953,1979]" captionTargetBox-0="[189,1398,1015,1868]" captionTargetBox-1="[189,1398,442,1913]" captionTargetId-0="figure@30.[189,1399,1015,1868]" captionTargetId-1="figure@31.[189,1399,442,1913]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="Fig. 19. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). A–B, F. MNRJ 28956. C. MZSP 85022. D–E. MNRJ 33139. A–D. External morphology. E–F. Operculum. Scale bars: A–D = 500 µm; E–F = 100 µm." captionText-1="Fig. 20. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989), radula. A–D. MNRJ 28956. E–F. MZSP 85022. Scale bars: A, E–F = 5 µm; B = 10 µm; C–D = 2 µm." httpUri-0="https://zenodo.org/record/2647933/files/figure.png" httpUri-1="https://zenodo.org/record/2647935/files/figure.png" pageId="29" pageNumber="30">19–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="82D06555B461FFF1FF63FB7E9F6EFA0F" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="CA7536DEB461FFF1FF63FB7E9A8EFB6F" blockId="29.[189,426,1204,1230]" box="[189,426,1204,1230]" pageId="29" pageNumber="30">
<heading id="913D81B2B461FFF1FF63FB7E9A8EFB6F" bold="true" box="[189,426,1204,1230]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="F8BEEACCB461FFF1FF63FB7E9A8EFB6F" bold="true" box="[189,426,1204,1230]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB461FFF1FF63FB299F6EFA0F" blockId="29.[189,1399,1250,1454]" pageId="29" pageNumber="30">
<materialsCitation id="7AA23C83B461FFF1FF63FB299A22FA80" ID-GBIF-Occurrence="2236664090" collectionCode="MNRJ" country="Brazil" county="State" latitude="-22.7" location="Campos Basin" longLatPrecision="1258" longitude="-40.666668" pageId="29" pageNumber="30" specimenCode="MNRJ 18756" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="B2DD764EB461FFF1FF63FB299A03FB5C" box="[189,295,1251,1277]" name="Brazil" pageId="29" pageNumber="30">BRAZIL</collectingCountry>
– 
<emphasis id="F8BEEACCB461FFF1FE96FB299968FB5C" bold="true" box="[328,588,1251,1277]" pageId="29" pageNumber="30">
<collectingRegion id="080EF83CB461FFF1FE96FB299926FB5C" box="[328,514,1251,1277]" country="Brazil" name="Rio de Janeiro" pageId="29" pageNumber="30">Rio de Janeiro</collectingRegion>
<collectingCounty id="23144E52B461FFF1FDD2FB299968FB5C" box="[524,588,1251,1277]" pageId="29" pageNumber="30">State</collectingCounty>
</emphasis>
• [4, 1 d] specs; 
<location id="CF156005B461FFF1FCC0FB2998E9FB5C" LSID="urn:lsid:plazi:treatment:426387C8B461FFCCFE37FB859AE9FB0B:CF156005B461FFF1FCC0FB2998E9FB5C" box="[798,973,1251,1277]" country="Brazil" county="State" latitude="-22.7" longLatPrecision="1258" longitude="-40.666668" name="Campos Basin" pageId="29" pageNumber="30" stateProvince="Rio de Janeiro">Campos Basin</location>
, 
<geoCoordinate id="AFFE5019B461FFF1FC05FB299F63FB5C" box="[987,1095,1251,1277]" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-22.7">22°42´S</geoCoordinate>
, 
<geoCoordinate id="AFFE5019B461FFF1FB88FB299FE9FB5C" box="[1110,1229,1251,1277]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="-40.666668">40°40´W</geoCoordinate>
; 2006; 
<specimenCode id="9A6C9EA5B461FFF1FAF6FB299A22FA80" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 18756</specimenCode>
</materialsCitation>
• 
<materialsCitation id="7AA23C83B461FFF1FEC2FACC98A8FA81" ID-GBIF-Occurrence="2236664078" box="[284,908,1285,1313]" collectionCode="MNRJ" country="Brazil" county="State" latitude="-22.7" location="Campos Basin" longLatPrecision="1258" longitude="-40.666668" pageId="29" pageNumber="30" specimenCode="MNRJ 33139" specimenCount="1" stateProvince="Rio de Janeiro">
[1, d] spec.; same data as for preceding; 
<specimenCode id="9A6C9EA5B461FFF1FD35FACD98A8FA81" box="[747,908,1286,1313]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 33139</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="7AA23C83B461FFF1FC74FACC98A4FAE5" ID-GBIF-Occurrence="2236664072" collectionCode="MZSP" collectorName="Itanhaem" country="Brazil" county="State" elevation="6" location="Ilha da Queimada Pequena" pageId="29" pageNumber="30" specimenCode="MZSP 85022" specimenCount="1" stateProvince="Sao Paulo">
<emphasis id="F8BEEACCB461FFF1FC74FACC9F4FFA81" bold="true" box="[938,1131,1286,1312]" pageId="29" pageNumber="30">
<collectingRegion id="080EF83CB461FFF1FC74FACC9F02FA81" box="[938,1062,1286,1312]" country="Brazil" name="Sao Paulo" pageId="29" pageNumber="30">São Paulo</collectingRegion>
<collectingCounty id="23144E52B461FFF1FBF5FACC9F4FFA81" box="[1067,1131,1286,1312]" pageId="29" pageNumber="30">State</collectingCounty>
</emphasis>
• [2, 1 d] specs; 
<location id="CF156005B461FFF1FAF9FACC9A87FAE5" LSID="urn:lsid:plazi:treatment:426387C8B461FFCCFE37FB859AE9FB0B:CF156005B461FFF1FAF9FACC9A87FAE5" country="Brazil" county="State" name="Ilha da Queimada Pequena" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Ilha da Queimada Pequena</location>
, 
<collectorName id="673F5308B461FFF1FE6EFAE39905FAE2" box="[432,545,1321,1347]" pageId="29" pageNumber="30">Itanhaém</collectorName>
; 
<elevation id="41E7D1EDB461FFF1FDF3FAE399A3FAE5" box="[557,647,1321,1348]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="29" pageNumber="30" unit="m" value="6.0" valueMax="12.0" valueMin="0.0">
<quantity id="0D329B3BB461FFF1FDF3FAE399A3FAE5" box="[557,647,1321,1348]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="29" pageNumber="30" unit="m" value="6.0" valueMax="12.0" valueMin="0.0">0–12 m</quantity>
</elevation>
depth; 
<specimenCode id="9A6C9EA5B461FFF1FD3EFAE398A4FAE5" box="[736,896,1321,1348]" collectionCode="MZSP" pageId="29" pageNumber="30">MZSP 85022</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="7AA23C83B461FFF1FC7DFAE39E52FAC7" ID-GBIF-Occurrence="2236664083" collectingDate="2016-12-17" collectionCode="MNRJ" collectorName="M. R. Fernandes" country="Brazil" county="Santa Catarina State" latitude="-27.140556" location="Bombinhas" longLatPrecision="20" longitude="-48.47861" municipality="Praia da Sepultura" pageId="29" pageNumber="30" specimenCode="MNRJ 28936" specimenCount="1" stateProvince="Sao Paulo">
<collectingCounty id="23144E52B461FFF1FC7DFAE39F8FFAE2" box="[931,1195,1321,1347]" pageId="29" pageNumber="30">
<emphasis id="F8BEEACCB461FFF1FC7DFAE39F8FFAE2" bold="true" box="[931,1195,1321,1347]" pageId="29" pageNumber="30">Santa Catarina State</emphasis>
</collectingCounty>
• [1] spec.; 
<collectingMunicipality id="2A11ACA4B461FFF1FAE4FAE39A76FAC7" pageId="29" pageNumber="30">Praia da Sepultura</collectingMunicipality>
, 
<location id="CF156005B461FFF1FE82FA869AC1FAC7" LSID="urn:lsid:plazi:treatment:426387C8B461FFCCFE37FB859AE9FB0B:CF156005B461FFF1FE82FA869AC1FAC7" box="[348,485,1356,1382]" country="Brazil" county="Santa Catarina State" latitude="-27.140556" longLatPrecision="20" longitude="-48.47861" municipality="Praia da Sepultura" name="Bombinhas" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Bombinhas</location>
; 
<geoCoordinate id="AFFE5019B461FFF1FE2EFA8199A6FAC6" box="[496,642,1355,1383]" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="-27.140556">27°08´26″ S</geoCoordinate>
, 
<geoCoordinate id="AFFE5019B461FFF1FD50FA81980FFAC6" box="[654,811,1355,1383]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-48.47861">48°28´43″ W</geoCoordinate>
; 
<collectingDate id="AE30E9F6B461FFF1FCE8FA8798F0FAC7" box="[822,980,1356,1383]" pageId="29" pageNumber="30" value="2016-12-17">
<date id="BE74101EB461FFF1FCE8FA8798F0FAC7" box="[822,980,1356,1383]" pageId="29" pageNumber="30" value="2016-12-17">17 Dec. 2016</date>
</collectingDate>
; 
<collectorName id="673F5308B461FFF1FC00FA879FBEFAC7" box="[990,1178,1356,1383]" pageId="29" pageNumber="30">M.R. Fernandes</collectorName>
leg.; 
<specimenCode id="9A6C9EA5B461FFF1FB07FA879E52FAC7" box="[1241,1398,1356,1383]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 28936</specimenCode>
</materialsCitation>
• 
<materialsCitation id="7AA23C83B461FFF1FF0FFAA59AB2FA0F" ID-GBIF-Occurrence="2236664044" collectingDate="2016-12-21" collectionCode="MNRJ" collectorName="M. R. Fernandes" country="Brazil" county="Santa Catarina State" latitude="-27.216389" location="Bombinhas" longLatPrecision="20" longitude="-48.511112" municipality="Praia da Tainha" pageId="29" pageNumber="30" specimenCode="MNRJ 28951" specimenCount="1" stateProvince="Sao Paulo">
[2] specs; 
<collectingMunicipality id="2A11ACA4B461FFF1FE90FABA9928FA2B" box="[334,524,1392,1418]" pageId="29" pageNumber="30">Praia da Tainha</collectingMunicipality>
, 
<location id="CF156005B461FFF1FDC7FABA9986FA2B" LSID="urn:lsid:plazi:treatment:426387C8B461FFCCFE37FB859AE9FB0B:CF156005B461FFF1FDC7FABA9986FA2B" box="[537,674,1392,1418]" country="Brazil" county="Santa Catarina State" latitude="-27.216389" longLatPrecision="20" longitude="-48.511112" municipality="Praia da Tainha" name="Bombinhas" pageId="29" pageNumber="30" stateProvince="Sao Paulo">Bombinhas</location>
; 
<geoCoordinate id="AFFE5019B461FFF1FD6EFAA49862FA2B" box="[688,838,1390,1418]" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="-27.216389">27°12´59″ S</geoCoordinate>
, 
<geoCoordinate id="AFFE5019B461FFF1FC8BFAA498D2FA2B" box="[853,1014,1390,1418]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-48.511112">48°30´40″ W</geoCoordinate>
; 
<collectingDate id="AE30E9F6B461FFF1FBDAFABA9F8DFA2B" box="[1028,1193,1392,1418]" pageId="29" pageNumber="30" value="2016-12-21">
<date id="BE74101EB461FFF1FBDAFABA9F8DFA2B" box="[1028,1193,1392,1418]" pageId="29" pageNumber="30" value="2016-12-21">21 Dec. 2016</date>
</collectingDate>
; 
<collectorName id="673F5308B461FFF1FB66FABA9E53FA2B" box="[1208,1399,1392,1418]" pageId="29" pageNumber="30">M.R. Fernandes</collectorName>
leg.; 
<specimenCode id="9A6C9EA5B461FFF1FF26FA599AB2FA0F" box="[248,406,1427,1454]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 28951</specimenCode>
</materialsCitation>
• 
<materialsCitation id="7AA23C83B461FFF1FE71FA599F62FA0F" ID-GBIF-Occurrence="2236664059" box="[431,1094,1426,1454]" collectingDate="2016-12-21" collectionCode="MNRJ" collectorName="M. R. Fernandes" country="Brazil" county="Santa Catarina State" latitude="-27.216389" location="Bombinhas" longLatPrecision="20" longitude="-48.511112" municipality="Praia da Tainha" pageId="29" pageNumber="30" specimenCode="MNRJ 28956" specimenCount="1" stateProvince="Sao Paulo">
[5, 2 d] specs; same data as for preceding; 
<specimenCode id="9A6C9EA5B461FFF1FC7AFA599F62FA0F" box="[932,1094,1427,1454]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 28956</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="82D06555B461FFF2FF63FA1E9875FC94" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="CA7536DEB461FFF1FF63FA1E990DFA4F" blockId="29.[189,553,1492,1518]" box="[189,553,1492,1518]" pageId="29" pageNumber="30">
<heading id="913D81B2B461FFF1FF63FA1E990DFA4F" bold="true" box="[189,553,1492,1518]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="F8BEEACCB461FFF1FF63FA1E990DFA4F" bold="true" box="[189,553,1492,1518]" pageId="29" pageNumber="30">Description of basic anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB461FFF1FF63F9C99F0BF9C2" blockId="29.[189,1398,1539,1636]" pageId="29" pageNumber="30">
EXTERNAL MORPHOLOGY. Body cream-yellowish, head-foot reddish, but the vivid pigmentation on headfoot vanishes after fixation; pedal slit covering about 68% of foot length; some black encapsulated fecal pellets present along intestine, with a vesicular shape and 
<quantity id="0D329B3BB461FFF1FCBAF98398C8F9C5" box="[868,1004,1609,1636]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" metricValueMax="10.0" metricValueMin="8.2" pageId="29" pageNumber="30" unit="mm" value="91.0" valueMax="100.0" valueMin="82.0">82–100 µm</quantity>
long.
</paragraph>
<paragraph id="CA7536DEB461FFF1FF63F941992EF94A" blockId="29.[189,1399,1674,1771]" pageId="29" pageNumber="30">OPERCULUM. Rounded, flat, moderately thin, semi-transparent, multispiral, 4.75 whorls, nucleus subcentral, dislocated 15% from center toward margin; diameter of operculum exceeds diameter of opercular pouch in 13–23%.</paragraph>
<paragraph id="CA7536DEB461FFF2FF63F8D99875FC94" blockId="29.[189,1399,1810,2013]" lastBlockId="30.[189,1399,267,822]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
RADULA. Formula 8-1-1-1-8; teeth row arranged in a ‘V-shape, with adjacent teeth aligned in a somewhat diagonal position to each other; central tooth with three main broad and triangular cusps, outer cusps 1.1–1.3× more elongated than median one, in addition to two marginal cusps very reduced in size; in one radula (
<figureCitation id="52F12A5BB461FFF1FEC9F8B69A49F837" box="[279,365,1916,1942]" captionStart="Fig" captionStartId="31.[189,232,1953,1979]" captionTargetBox="[189,1398,442,1913]" captionTargetId="figure@31.[189,1399,442,1913]" captionTargetPageId="31" captionText="Fig. 20. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989), radula. A–D. MNRJ 28956. E–F. MZSP 85022. Scale bars: A, E–F = 5 µm; B = 10 µm; C–D = 2 µm." httpUri="https://zenodo.org/record/2647935/files/figure.png" pageId="29" pageNumber="30">Fig. 20</figureCitation>
A–D), lateral teeth with three (left of central tooth) or four (right of central tooth) main broad and triangular cusps, in addition to two marginal cusps reduced in size, resulting in lateral teeth with five (left) or six (right) cusps; left lateral teeth with cusps 1 and 5 with length 39–63% of cusps 2 and 4, cusp 3 with length 70–88% of cusps 2 and 4; right lateral teeth with cusps 1 and 6 with length 29–47% of remaining cusps, which are similarly sized; in another radula (
<figureCitation id="52F12A5BB462FFF2FB55FEE49FC6FEE8" box="[1163,1250,302,329]" captionStart="Fig" captionStartId="31.[189,232,1953,1979]" captionTargetBox="[189,1398,442,1913]" captionTargetId="figure@31.[189,1399,442,1913]" captionTargetPageId="31" captionText="Fig. 20. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989), radula. A–D. MNRJ 28956. E–F. MZSP 85022. Scale bars: A, E–F = 5 µm; B = 10 µm; C–D = 2 µm." httpUri="https://zenodo.org/record/2647935/files/figure.png" pageId="30" pageNumber="31">Fig. 20</figureCitation>
E–F), lateral teeth always with four cusps; left M1 with three broad and triangular cusps, median one 1.3–1.5× more elongated than outer cusps; right M1 with four triangular cusps, cusp 1 much reduced, with length 49–60% of remaining cusps, which are similarly sized; M2–M6 with three main triangular, claw-like and equally-sized cusps in a posterior portion of radula, in addition to a fourth small cusp (cusp 1) that gradually strengthens towards the anterior portion of radula and may become equal in size to remaining cusps (at least in M3–M6), with right M5 being supposedly the first tooth to show a fully developed cusp 1; an additional fifth cusp may appear in M3 and M4 as an abnormal local asymmetry; M7 with four triangular, curved and similarly-sized cusps in a posterior region of radula, with cusp 1 gradually reducing in size and finally disappearing in the anterior region of radula, resulting in M7 with three cusps; M8 very narrow, with three finger-like cusps (median one 1.3–1.5× more elongated than outer cusps), fully formed only in the anterior region of radula; central tooth 
<quantity id="0D329B3BB462FFF2FBCFFD789FBEFD6D" box="[1041,1178,690,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="30" pageNumber="31" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 µm</quantity>
wide, lateral teeth 
<quantity id="0D329B3BB462FFF2FF63FD1F9A67FD4E" box="[189,323,725,751]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="30" pageNumber="31" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.8–4.1 µm</quantity>
wide, M1–M2 
<quantity id="0D329B3BB462FFF2FE2BFD1F995FFD4E" box="[501,635,725,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="30" pageNumber="31" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 µm</quantity>
wide, M3–M6 (with three main cusps) 
<quantity id="0D329B3BB462FFF2FB9AFD1F9FEEFD4E" box="[1092,1226,725,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="30" pageNumber="31" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 µm</quantity>
wide, M3–M6 (with four main cusps) 
<quantity id="0D329B3BB462FFF2FE13FD329970FCB2" box="[461,596,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="7.8999999999999995" metricValueMax="9.2" metricValueMin="6.6" pageId="30" pageNumber="31" unit="mm" value="7.8999999999999995" valueMax="9.2" valueMin="6.6">6.6–9.2 µm</quantity>
wide, M7 (with four cusps) 
<quantity id="0D329B3BB462FFF2FC7EFD329F03FCB2" box="[928,1063,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="30" pageNumber="31" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7–3.2 µm</quantity>
wide, M7 (with three cusps) 
<quantity id="0D329B3BB462FFF2FF63FCD19A60FC97" box="[189,324,795,822]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="30" pageNumber="31" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 µm</quantity>
wide, fully developed M8 
<quantity id="0D329B3BB462FFF2FD5CFCD1982DFC97" box="[642,777,795,822]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="30" pageNumber="31" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.9–3.3 µm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="82D06555B462FFCCFF63FC979AE9FB0B" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="discussion">
<paragraph id="CA7536DEB462FFF2FF63FC979A0BFCD6" blockId="30.[189,303,861,887]" box="[189,303,861,887]" pageId="30" pageNumber="31">
<heading id="913D81B2B462FFF2FF63FC979A0BFCD6" bold="true" box="[189,303,861,887]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="F8BEEACCB462FFF2FF63FC979A0BFCD6" bold="true" box="[189,303,861,887]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB462FFF3FF63FC419E53FECD" blockId="30.[189,1399,907,969]" lastBlockId="31.[189,1399,267,364]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
Figured specimens show a gradual loss of coloration of the reddish-pigmented soft parts in 
<taxonomicName id="0DCA4D5DB462FFF2FB3AFC469E52FC04" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1252,1398,907,934]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB462FFF2FB3AFC469FDCFC04" box="[1252,1272,908,933]" italics="true" pageId="30" pageNumber="31">N</emphasis>
. 
<emphasis id="F8BEEACCB462FFF2FAD8FC419E52FC04" box="[1286,1398,907,933]" italics="true" pageId="30" pageNumber="31">verbernei</emphasis>
</taxonomicName>
after many years conserved in ethanol, from fresh specimens (
<figureCitation id="52F12A5BB462FFF2FC7AFC6498DCFC68" box="[932,1016,942,969]" captionStart="Fig" captionStartId="30.[189,232,1915,1941]" captionTargetBox="[189,1398,1015,1868]" captionTargetId="figure@30.[189,1399,1015,1868]" captionTargetPageId="30" captionText="Fig. 19. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). A–B, F. MNRJ 28956. C. MZSP 85022. D–E. MNRJ 33139. A–D. External morphology. E–F. Operculum. Scale bars: A–D = 500 µm; E–F = 100 µm." httpUri="https://zenodo.org/record/2647933/files/figure.png" pageId="30" pageNumber="31">Fig. 19</figureCitation>
A–B) to old ones (
<figureCitation id="52F12A5BB462FFF2FB06FC649E09FC68" box="[1240,1325,942,969]" captionStart="Fig" captionStartId="30.[189,232,1915,1941]" captionTargetBox="[189,1398,1015,1868]" captionTargetId="figure@30.[189,1399,1015,1868]" captionTargetPageId="30" captionText="Fig. 19. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). A–B, F. MNRJ 28956. C. MZSP 85022. D–E. MNRJ 33139. A–D. External morphology. E–F. Operculum. Scale bars: A–D = 500 µm; E–F = 100 µm." httpUri="https://zenodo.org/record/2647933/files/figure.png" pageId="30" pageNumber="31">Fig. 19</figureCitation>
C–D). 
<bibRefCitation id="AE5B4B2FB463FFF3FF63FEC19977FE84" author="Rolan E. &amp; Fernandez-Garces R." box="[189,595,267,293]" journalOrPublisher="Apex" pageId="31" pageNumber="32" pagination="17 - 27" part="9" refId="ref21646" refString="Rolan E. &amp; Fernandez-Garces R. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species. Apex 9: 17 - 27." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species." type="journal article" year="1994">Rolán &amp; Fernández-Garcés (1994)</bibRefCitation>
described the external morphology of 
<taxonomicName id="0DCA4D5DB463FFF3FBC5FEC69F89FE84" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1051,1197,267,294]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB463FFF3FBC5FEC69F0BFE84" box="[1051,1071,268,293]" italics="true" pageId="31" pageNumber="32">N</emphasis>
. 
<emphasis id="F8BEEACCB463FFF3FBE3FEC19F89FE84" box="[1085,1197,267,293]" italics="true" pageId="31" pageNumber="32">verbernei</emphasis>
</taxonomicName>
and 
<taxonomicName id="0DCA4D5DB463FFF3FB39FEC69954FEE8" authority="(Rolan &amp; Fernandez-Garces, 1994)" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1994" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="decollata">
<emphasis id="F8BEEACCB463FFF3FB39FEC69FDFFE84" box="[1255,1275,268,293]" italics="true" pageId="31" pageNumber="32">N</emphasis>
. 
<emphasis id="F8BEEACCB463FFF3FAD7FEC19E52FE84" box="[1289,1398,267,293]" italics="true" pageId="31" pageNumber="32">decollata</emphasis>
(
<bibRefCitation id="AE5B4B2FB463FFF3FF1BFEE4994CFEE8" author="Rolan E. &amp; Fernandez-Garces R." box="[197,616,302,329]" journalOrPublisher="Apex" pageId="31" pageNumber="32" pagination="17 - 27" part="9" refId="ref21646" refString="Rolan E. &amp; Fernandez-Garces R. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species. Apex 9: 17 - 27." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species." type="journal article" year="1994">Rolán &amp; Fernández-Garcés, 1994</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="B2DD764EB463FFF3FD18FEE49820FEE9" box="[710,772,302,328]" name="Cuba" pageId="31" pageNumber="32">Cuba</collectingCountry>
as being translucent white with milk-white spots distributed on the dorsum. This could be a result of long-time storage of animals and consequent loss
</paragraph>
<caption id="9EB56656B462FFF2FF63F8B19A3BF87D" httpUri="https://zenodo.org/record/2647933/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1915,1941]" targetBox="[189,1398,1015,1868]" targetPageId="30">
<paragraph id="CA7536DEB462FFF2FF63F8B19A3BF87D" blockId="30.[189,1399,1915,2012]" pageId="30" pageNumber="31">
<emphasis id="F8BEEACCB462FFF2FF63F8B19A3EF834" bold="true" box="[189,282,1915,1941]" pageId="30" pageNumber="31">Fig. 19.</emphasis>
<taxonomicName id="0DCA4D5DB462FFF2FEFDF8B198ACF834" authority="(Moolenbeek &amp; Faber, 1989)" authorityName="" authorityYear="1989" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[291,904,1915,1941]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB462FFF2FEFDF8B19905F834" box="[291,545,1915,1941]" italics="true" pageId="30" pageNumber="31">Nanaphora verbernei</emphasis>
(Moolenbeek &amp; Faber, 1989)
</taxonomicName>
. 
<emphasis id="F8BEEACCB462FFF2FC4AF8B198C9F834" bold="true" box="[916,1005,1915,1941]" pageId="30" pageNumber="31">A–B, F</emphasis>
. MNRJ 28956. 
<emphasis id="F8BEEACCB462FFF2FB73F8B19FE6F834" bold="true" box="[1197,1218,1915,1941]" pageId="30" pageNumber="31">C</emphasis>
. MZSP 85022. 
<emphasis id="F8BEEACCB462FFF2FF63F8559BD0F818" bold="true" box="[189,244,1951,1977]" pageId="30" pageNumber="31">D–E</emphasis>
. MNRJ 33139. 
<emphasis id="F8BEEACCB462FFF2FE6AF8549AC9F819" bold="true" box="[436,493,1950,1976]" pageId="30" pageNumber="31">A–D</emphasis>
. External morphology. 
<emphasis id="F8BEEACCB462FFF2FCD8F855981EF818" bold="true" box="[774,826,1951,1977]" pageId="30" pageNumber="31">E–F</emphasis>
. Operculum. Scale bars: A–D = 500 µm; E–F = 100 µm.
</paragraph>
</caption>
<caption id="9EB56656B463FFF3FF63F86B98A2F87E" httpUri="https://zenodo.org/record/2647935/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1953,1979]" targetBox="[189,1398,442,1913]" targetPageId="31">
<paragraph id="CA7536DEB463FFF3FF63F86B98A2F87E" blockId="31.[189,1399,1953,2015]" pageId="31" pageNumber="32">
<emphasis id="F8BEEACCB463FFF3FF63F86B9A3EF81A" bold="true" box="[189,282,1953,1979]" pageId="31" pageNumber="32">Fig. 20.</emphasis>
<taxonomicName id="0DCA4D5DB463FFF3FEFDF86B98ACF81A" authority="(Moolenbeek &amp; Faber, 1989)" authorityName="" authorityYear="1989" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[291,904,1953,1979]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB463FFF3FEFDF86B9905F81A" box="[291,545,1953,1979]" italics="true" pageId="31" pageNumber="32">Nanaphora verbernei</emphasis>
(Moolenbeek &amp; Faber, 1989)
</taxonomicName>
, radula. 
<emphasis id="F8BEEACCB463FFF3FC30F86B9F03F81A" bold="true" box="[1006,1063,1953,1979]" pageId="31" pageNumber="32">A–D</emphasis>
. MNRJ 28956. 
<emphasis id="F8BEEACCB463FFF3FB39F86B9E3FF81A" bold="true" box="[1255,1307,1953,1979]" pageId="31" pageNumber="32">E–F</emphasis>
. MZSP 85022. Scale bars: A, E–F = 5 µm; B = 10 µm; C–D = 2 µm.
</paragraph>
</caption>
<paragraph id="CA7536DEB45CFFCCFF63FEC19ADAFEE8" blockId="32.[189,1399,267,329]" pageId="32" pageNumber="33">
of pigmentation, or even suggest that 
<taxonomicName id="0DCA4D5DB45CFFCCFD5FFEC69830FE84" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[641,788,267,294]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFD5FFEC699B1FE84" box="[641,661,268,293]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFD7AFEC19830FE84" box="[676,788,267,293]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFCBEFEC1988CFE84" box="[864,936,267,293]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
constitutes a different species than the Caribbean one (see below).
</paragraph>
<paragraph id="CA7536DEB45CFFCCFF63FEBA9E54FE55" blockId="32.[189,1399,368,500]" pageId="32" pageNumber="33">
<bibRefCitation id="AE5B4B2FB45CFFCCFF63FEBA9976FE2B" author="Rolan E. &amp; Fernandez-Garces R." box="[189,594,368,394]" journalOrPublisher="Apex" pageId="32" pageNumber="33" pagination="17 - 27" part="9" refId="ref21646" refString="Rolan E. &amp; Fernandez-Garces R. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species. Apex 9: 17 - 27." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species." type="journal article" year="1994">Rolán &amp; Fernández-Garcés (1994)</bibRefCitation>
briefly described, but did not illustrate, opercula of 
<taxonomicName id="0DCA4D5DB45CFFCCFB6DFEBB9E61FE2B" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1203,1349,368,395]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFB6DFEBB9FE3FE2B" box="[1203,1223,369,394]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFB0BFEBA9E61FE2B" box="[1237,1349,368,394]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
and 
<taxonomicName id="0DCA4D5DB45CFFCCFF63FE5E9A68FE0C" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1994" box="[189,332,403,430]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="decollata">
<emphasis id="F8BEEACCB45CFFCCFF63FE5E9BF5FE0C" box="[189,209,404,429]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFF01FE599A68FE0C" box="[223,332,403,429]" italics="true" pageId="32" pageNumber="33">decollata</emphasis>
</taxonomicName>
, which are similar to those herein illustrated (
<figureCitation id="52F12A5BB45CFFCCFCB4FE599898FE0C" box="[874,956,403,429]" captionStart="Fig" captionStartId="30.[189,232,1915,1941]" captionTargetBox="[189,1398,1015,1868]" captionTargetId="figure@30.[189,1399,1015,1868]" captionTargetPageId="30" captionText="Fig. 19. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). A–B, F. MNRJ 28956. C. MZSP 85022. D–E. MNRJ 33139. A–D. External morphology. E–F. Operculum. Scale bars: A–D = 500 µm; E–F = 100 µm." httpUri="https://zenodo.org/record/2647933/files/figure.png" pageId="32" pageNumber="33">Fig. 19</figureCitation>
E–F) by being multispiral and having a subcentral nucleus. The operculum of 
<taxonomicName id="0DCA4D5DB45CFFCCFD4FFE7D9F20FE70" authority="Laseron, 1958" authorityName="Laseron" authorityYear="1958" box="[657,1028,438,465]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="albogemmata">
<emphasis id="F8BEEACCB45CFFCCFD4FFE7D9981FE71" box="[657,677,439,464]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFD6AFE7C9870FE71" box="[692,852,438,464]" italics="true" pageId="32" pageNumber="33">albogemmata</emphasis>
Laseron, 1958
</taxonomicName>
, illustrated by 
<bibRefCitation id="AE5B4B2FB45CFFCCFB6FFE7C9E56FE70" author="Marshall B. A." box="[1201,1394,438,465]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="32" pageNumber="33" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall (1983)</bibRefCitation>
, possesses a remarkably prominent peg accessory, which was not observed in 
<taxonomicName id="0DCA4D5DB45CFFCCFB99FE109FFDFE55" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1095,1241,474,500]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFB99FE109F7FFE52" box="[1095,1115,474,499]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFBB7FE109FFDFE55" box="[1129,1241,474,500]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFAFFFE109E48FE55" box="[1313,1388,474,500]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="CA7536DEB45CFFCCFF63FDD1998EFD63" blockId="32.[189,1399,539,706]" pageId="32" pageNumber="33">
The radula of 
<taxonomicName id="0DCA4D5DB45CFFCCFEBBFDD69ADCFD94" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[357,504,539,566]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFEBBFDD69A5DFD94" box="[357,377,540,565]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFE56FDD19ADCFD94" box="[392,504,539,565]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFD9FFDD199ADFD94" box="[577,649,539,565]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
has a variable number of cusps in some teeth, in addition to an abnormal local asymmetry. The peculiar arrangement of teeth rows in a distinct ‘V-shape’ generates a pattern in which one tooth is exactly situated in the center of the six teeth adjacent to it (
<figureCitation id="52F12A5BB45CFFCCFB24FDAB9E4EFDDD" box="[1274,1386,609,636]" captionStart="Fig" captionStartId="31.[189,232,1953,1979]" captionTargetBox="[189,1398,442,1913]" captionTargetId="figure@31.[189,1399,442,1913]" captionTargetPageId="31" captionText="Fig. 20. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989), radula. A–D. MNRJ 28956. E–F. MZSP 85022. Scale bars: A, E–F = 5 µm; B = 10 µm; C–D = 2 µm." httpUri="https://zenodo.org/record/2647935/files/figure.png" pageId="32" pageNumber="33">Fig. 20D</figureCitation>
). Other genera of 
<taxonomicName id="0DCA4D5DB45CFFCCFE5FFD4F992EFD3E" authorityName="Gray" authorityYear="1847" box="[385,522,645,671]" class="Gastropoda" family="Triphoridae" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="family">Triphoridae</taxonomicName>
also seem to present a slight level of ‘V-shape’ arrangement of rows, but apparently never like the observed herein.
</paragraph>
<paragraph id="CA7536DEB45CFFCCFF63FD239AE9FB0B" blockId="32.[189,1399,745,1194]" pageId="32" pageNumber="33">
The most surprising feature of the radula of 
<taxonomicName id="0DCA4D5DB45CFFCCFCDAFD2098BAFCA2" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[772,926,745,772]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFCDAFD20983CFCA2" box="[772,792,746,771]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFCF0FD2398BAFCA2" box="[814,926,745,771]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFC29FD239F1BFCA2" box="[1015,1087,745,771]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
is its radically different tooth morphology when compared to 
<taxonomicName id="0DCA4D5DB45CFFCCFD7CFCC79818FC87" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[674,828,780,807]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFD7CFCC79992FC87" box="[674,694,781,806]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFD12FCC69818FC87" box="[716,828,780,806]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFC4BFCC698F7FC87" box="[917,979,780,806]" name="Cuba" pageId="32" pageNumber="33">Cuba</collectingCountry>
and other species (
<taxonomicName id="0DCA4D5DB45CFFCCFB09FCC79E4BFC87" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1994" box="[1239,1391,780,807]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="decollata">
<emphasis id="F8BEEACCB45CFFCCFB09FCC79FCFFC87" box="[1239,1259,781,806]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFADCFCC69E4BFC87" box="[1282,1391,780,806]" italics="true" pageId="32" pageNumber="33">decollata</emphasis>
</taxonomicName>
, 
<taxonomicName id="0DCA4D5DB45CFFCCFF63FCFB9A5BFCEB" authorityName="Laseron" authorityYear="1958" box="[189,383,816,842]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="albogemmata">
<emphasis id="F8BEEACCB45CFFCCFF63FCFB9BF5FCEB" box="[189,209,817,842]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFF01FCFA9A5BFCEB" box="[223,383,816,842]" italics="true" pageId="32" pageNumber="33">albogemmata</emphasis>
</taxonomicName>
and 
<taxonomicName id="0DCA4D5DB45CFFCCFE69FCFB9983FCEB" authorityName="Laseron" authorityYear="1958" box="[439,679,816,842]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="albogemmata">
<emphasis id="F8BEEACCB45CFFCCFE69FCFB9AEFFCEB" box="[439,459,817,842]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. aff. 
<emphasis id="F8BEEACCB45CFFCCFDD9FCFA9983FCEB" box="[519,679,816,842]" italics="true" pageId="32" pageNumber="33">albogemmata</emphasis>
</taxonomicName>
). Main differences comprise the central tooth of 
<taxonomicName id="0DCA4D5DB45CFFCCFB3AFCFB9E52FCEB" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1252,1398,816,842]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFB3AFCFB9FDCFCEB" box="[1252,1272,817,842]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFAD8FCFA9E52FCEB" box="[1286,1398,816,842]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFF22FC999A60FCCC" box="[252,324,851,877]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
bearing three main cusps (instead of two, four, six or nine cusps), lateral teeth with up to five or six cusps (instead of seven or more cusps in remaining species) and the main three claw-like cusps of marginal teeth (which completely differ from the several elongated cusps of the hand-shaped marginal teeth in other species). The radular formula of 
<taxonomicName id="0DCA4D5DB45CFFCCFD2DFC7798A1FC77" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1994" box="[755,901,956,983]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="decollata">
<emphasis id="F8BEEACCB45CFFCCFD2DFC779823FC77" box="[755,775,957,982]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFCC6FC7698A1FC77" box="[792,901,956,982]" italics="true" pageId="32" pageNumber="33">decollata</emphasis>
</taxonomicName>
is 7-1-1-1-7 (
<bibRefCitation id="AE5B4B2FB45CFFCCFBF1FC769BDFFC5B" author="Rolan E. &amp; Fernandez-Garces R." journalOrPublisher="Apex" pageId="32" pageNumber="33" pagination="17 - 27" part="9" refId="ref21646" refString="Rolan E. &amp; Fernandez-Garces R. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species. Apex 9: 17 - 27." title="The family Triphoridae (Mollusca, Gastropoda) in Cuba. 4. The genera Monophorus, Nototriphora, Cosmotriphora and Cheirodonta, with the description of three new species." type="journal article" year="1994">Rolán &amp; Fernández-Garcés 1994</bibRefCitation>
), similar to the eight marginal teeth observed in 
<taxonomicName id="0DCA4D5DB45CFFCCFC9EFC2B98F0FC5B" authorityName="" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[832,980,992,1018]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB45CFFCCFC9EFC2B9870FC5B" box="[832,852,993,1018]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFCBAFC2A98F0FC5B" box="[868,980,992,1018]" italics="true" pageId="32" pageNumber="33">verbernei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B2DD764EB45CFFCCFBC1FC2A9F4EFC5B" box="[1055,1130,992,1018]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
, whereas the radula of 
<taxonomicName id="0DCA4D5DB45CFFCCFF63FBCE9AA5FBBC" authorityName="Laseron" authorityYear="1958" box="[189,385,1027,1053]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="albogemmata">
<emphasis id="F8BEEACCB45CFFCCFF63FBCE9BF5FBBC" box="[189,209,1028,1053]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. 
<emphasis id="F8BEEACCB45CFFCCFF3FFBC99AA5FBBC" box="[225,385,1027,1053]" italics="true" pageId="32" pageNumber="33">albogemmata</emphasis>
</taxonomicName>
and 
<taxonomicName id="0DCA4D5DB45CFFCCFE63FBCE9996FBBC" authorityName="Laseron" authorityYear="1958" box="[445,690,1027,1053]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="albogemmata">
<emphasis id="F8BEEACCB45CFFCCFE63FBCE9AF5FBBC" box="[445,465,1028,1053]" italics="true" pageId="32" pageNumber="33">N</emphasis>
. aff. 
<emphasis id="F8BEEACCB45CFFCCFDCCFBC99996FBBC" box="[530,690,1027,1053]" italics="true" pageId="32" pageNumber="33">albogemmata</emphasis>
</taxonomicName>
is 1-1-1-1-1 (as argued by 
<bibRefCitation id="AE5B4B2FB45CFFCCFC24FBC99F8CFBBC" author="Marshall B. A." box="[1018,1192,1027,1053]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="32" pageNumber="33" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall 1983</bibRefCitation>
) or 2-0-1-0-2 (as argued by 
<bibRefCitation id="AE5B4B2FB45CFFCCFEEAFBEC9AE2FBE1" author="Nutzel A." box="[308,454,1062,1088]" journalOrPublisher="Berliner Geowissenschaftliche Abhandlungen" pageId="32" pageNumber="33" pagination="1 - 229" part="26" refId="ref21310" refString="Nutzel A. 1998. Ueber die Stammesgeschichte der Ptenoglossa (Gastropoda). Berliner Geowissenschaftliche Abhandlungen, ser. E (Palaeobiologie) 26: 1 - 229." title="Ueber die Stammesgeschichte der Ptenoglossa (Gastropoda)" type="journal article" year="1998">Nützel 1998</bibRefCitation>
). This extreme variation in the composition of the radula indicates that 
<taxonomicName id="0DCA4D5DB45CFFCCFB2FFBEC9E53FBE1" authorityName="Laseron" authorityYear="1958" box="[1265,1399,1062,1088]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB45CFFCCFB2FFBEC9E53FBE1" box="[1265,1399,1062,1088]" italics="true" pageId="32" pageNumber="33">Nanaphora</emphasis>
</taxonomicName>
shows an atypical evolutionary flexibility in order to colonize new sponge hosts or, alternatively, it is constituted by different monophyletic lineages that are masked by a convergence of inflated and reduced shells (
<bibRefCitation id="AE5B4B2FB45CFFCCFED1FB5A9AE4FB0B" author="Marshall B. A." box="[271,448,1168,1194]" journalOrPublisher="Records of the Australian Museum, suppl." pageId="32" pageNumber="33" pagination="1 - 119" part="2" refId="ref21274" refString="Marshall B. A. 1983. A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda). Records of the Australian Museum, suppl. 2: 1 - 119." title="A revision of the recent Triphoridae of Southern Australia (Mollusca: Gastropoda)" type="journal article" year="1983">Marshall 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>