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<document id="1419B46BDB396C1595708C94B2A3F6B3" ID-CLB-Dataset="302422" ID-DOI="10.11646/phytotaxa.345.3.7" ID-GBIF-Dataset="d7e7b859-be05-4953-a885-0d439c726210" ID-ISSN="1179-3163" ID-Zenodo-Dep="13708421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725592911505" checkinUser="felipe" docAuthor="Henderson, Andrew, Iqbal, Moh., Rusydi, Muhammad &amp; Pitopang, Ramadhanil" docDate="2018" docId="423A87CDB300FFF838C4F9409625FB6D" docLanguage="en" docName="phytotaxa.345.3.7.pdf" docOrigin="Phytotaxa 345 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.345.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Calamus tadulakoensis Henderson, Iqbal, Rusydi &amp; Pitopang 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="300" masterDocId="BE03FFB5B300FFFA384CFFB09529FFEC" masterDocTitle="A new species of Calamus (Calaminae, Calamoideae, Arecaceae) from Sulawesi, Indonesia" masterLastPageNumber="300" masterPageNumber="298" pageNumber="298" updateTime="1725605308921" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="259E18B75F7215AFF894D67ECB540A65">A new species of Calamus (Calaminae, Calamoideae, Arecaceae) from Sulawesi, Indonesia</mods:title>
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<mods:namePart id="09AF734729E517EFB65A05450BECC4C9">Henderson, Andrew</mods:namePart>
<mods:affiliation id="7E5E5BC96458BDBEB0336676211CE1EE">The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
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<mods:namePart id="2A26F0D01BD8A60170A85546FE5DD193">Iqbal, Moh.</mods:namePart>
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<mods:namePart id="A38AA0B73B2B26308C9FCACBBAA3C1A9">Rusydi, Muhammad</mods:namePart>
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<mods:namePart id="3F4808875B6BF00C029C930A8C3B0517">Pitopang, Ramadhanil</mods:namePart>
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<subSubSection id="82896550B300FFFA38C4F94096D7F8E7" box="[136,1022,1776,1803]" pageId="0" pageNumber="298" type="nomenclature">
<paragraph id="CA2C36DBB300FFFA38C4F94096D7F8E7" blockId="0.[136,1022,1776,1803]" box="[136,1022,1776,1803]" pageId="0" pageNumber="298">
<heading id="916481B7B300FFFA38C4F94096D7F8E7" box="[136,1022,1776,1803]" fontSize="11" level="2" pageId="0" pageNumber="298" reason="3">
<taxonomicName id="0D934D58B300FFFA38C4F9409668F8E7" ID-CoL="8WS2B" authority="Henderson, Iqbal, Rusydi &amp; Pitopang" authorityName="Henderson, Iqbal, Rusydi &amp; Pitopang" authorityYear="2018" box="[136,833,1776,1803]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="0" pageNumber="298" phylum="Tracheophyta" rank="species" species="tadulakoensis" status="sp. nov.">
<emphasis id="F8E7EAC9B300FFFA38C4F94094BCF8E7" bold="true" box="[136,405,1776,1803]" italics="true" pageId="0" pageNumber="298">Calamus tadulakoensis</emphasis>
Henderson, Iqbal, Rusydi &amp; Pitopang
</taxonomicName>
<emphasis id="F8E7EAC9B300FFFA3B04F94296B4F8E7" box="[840,925,1778,1803]" italics="true" pageId="0" pageNumber="298">
<taxonomicNameLabel id="E3D457B2B300FFFA3B04F94296B4F8E7" box="[840,925,1778,1803]" pageId="0" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="52A82A5EB300FFFA3BE0F94296C4F8E7" box="[940,1005,1778,1803]" captionStart="FIGURE 1" captionStartId="1.[136,229,1917,1938]" captionTargetBox="[151,1436,188,1893]" captionTargetId="figure-17@1.[151,1436,188,1893]" captionTargetPageId="1" captionText="FIGURE 1. Calamus tadulakoensis. A. Cirrus with irregular spines. B. Grayish-white indumentose abaxial pinna surface. C. Leaf sheath with non-spiny knee. D. Fruits borne on short, stout pedicels." figureDoi="http://doi.org/10.5281/zenodo.13708423" httpUri="https://zenodo.org/record/13708423/files/figure.png" pageId="0" pageNumber="298">Fig. 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="82896550B300FFF838C4F8A7961EFC80" lastPageId="2" lastPageNumber="300" pageId="0" pageNumber="298" type="materials_examined">
<paragraph id="CA2C36DBB300FFFA38C4F8A79425F89D" blockId="0.[136,1452,1815,1905]" pageId="0" pageNumber="298">
Type:—
<collectingCountry id="B284764BB300FFFA389AF8A79474F8C0" box="[214,349,1815,1836]" name="Indonesia" pageId="0" pageNumber="298">INDONESIA</collectingCountry>
.
<collectingRegion id="0857F839B300FFFA3928F8A79723F8C0" box="[356,522,1815,1836]" country="Indonesia" name="Sulawesi Tengah" pageId="0" pageNumber="298">Sulawesi Tengah</collectingRegion>
: Banggai Regency, Lamala District, road from Labotan to Boloak, disturbed forest on steep slope, limestone soil, 00˚52.520
<collectionCode id="AC82AE1EB300FFFA39F0F88994E1F8A2" box="[444,456,1849,1870]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="0" pageNumber="298" type="Herbarium">S</collectionCode>
, 123˚14.423
<collectionCode id="AC82AE1EB300FFFA3A05F8899770F8A2" box="[585,601,1849,1870]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="0" pageNumber="298" type="Herbarium">E</collectionCode>
,
<quantity id="0D6B9B3EB300FFFA3A2FF8899788F8A2" box="[611,673,1849,1870]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="0" pageNumber="298" unit="m" value="146.0">146 m</quantity>
,
<date id="BE2D101BB300FFFA3AE7F889966BF8A2" box="[683,834,1849,1870]" pageId="0" pageNumber="298" value="2017-10-07">7 October 2017</date>
,
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<collectionCode id="AC82AE1EB300FFFA3B07F8899674F8A2" box="[843,861,1849,1870]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="0" pageNumber="298" type="Herbarium">A</collectionCode>
. Henderson &amp; Moh. Iqbal 4258
</emphasis>
(
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,
<typeStatus id="15288879B300FFFA3D6EF8899058F8A2" box="[1314,1393,1849,1870]" pageId="0" pageNumber="298" type="isotype">isotypes</typeStatus>
<collectionCode id="AC82AE1EB300FFFA3D3BF8899081F8A2" box="[1399,1448,1849,1870]" country="Indonesia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14168" lsid="urn:lsid:biocol.org:col:14168" name="Tadulako University" pageId="0" pageNumber="298" type="Herbarium">CEB</collectionCode>
,
<collectionCode id="AC82AE1EB300FFFA38F1F8EC95F9F89D" box="[189,208,1884,1905]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="0" pageNumber="298" type="Herbarium">K</collectionCode>
,
<collectionCode id="AC82AE1EB300FFFA3897F8EC942BF89D" box="[219,258,1884,1905]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="0" pageNumber="298" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph id="CA2C36DBB300FFFA38C4F81095D9F837" blockId="0.[136,1451,1952,2011]" pageId="0" pageNumber="298">
It differs from similar species (
<taxonomicName id="0D934D58B300FFFA398AF8109762F85B" authorityName="Beccari ex Heyne" authorityYear="1922" box="[454,587,1952,1975]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="0" pageNumber="298" phylum="Tracheophyta" rank="species" species="leiocaulis">
<emphasis id="F8E7EAC9B300FFFA398AF8109762F85B" box="[454,587,1952,1975]" italics="true" pageId="0" pageNumber="298">C. leiocaulis</emphasis>
</taxonomicName>
,
<taxonomicName id="0D934D58B300FFFA3A19F81097CEF85B" authorityName="Baker &amp; Dransfield" authorityYear="2004" box="[597,743,1952,1975]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="0" pageNumber="298" phylum="Tracheophyta" rank="species" species="suaveolens">
<emphasis id="F8E7EAC9B300FFFA3A19F81097CEF85B" box="[597,743,1952,1975]" italics="true" pageId="0" pageNumber="298">C. suaveolens</emphasis>
</taxonomicName>
) in its regularly arranged pinnae that are grayish-white indumentose abaxially.
</paragraph>
<caption id="9EEC6653B301FFFB38C4F8CD97FCF85A" ID-DOI="http://doi.org/10.5281/zenodo.13708423" ID-Zenodo-Dep="13708423" httpUri="https://zenodo.org/record/13708423/files/figure.png" pageId="1" pageNumber="299" startId="1.[136,229,1917,1938]" targetBox="[151,1436,188,1893]" targetPageId="1" targetType="figure">
<paragraph id="CA2C36DBB301FFFB38C4F8CD97FCF85A" blockId="1.[136,1451,1917,1974]" pageId="1" pageNumber="299">
<emphasis id="F8E7EAC9B301FFFB38C4F8CD95D7F87E" bold="true" box="[136,254,1917,1938]" pageId="1" pageNumber="299">FIGURE 1.</emphasis>
<taxonomicName id="0D934D58B301FFFB394FF8CD94CBF87E" authorityName="Henderson, Iqbal, Rusydi &amp; Pitopang" authorityYear="2018" box="[259,482,1917,1938]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="299" phylum="Tracheophyta" rank="species" species="tadulakoensis">
<emphasis id="F8E7EAC9B301FFFB394FF8CD94CBF87E" box="[259,482,1917,1938]" italics="true" pageId="1" pageNumber="299">Calamus tadulakoensis</emphasis>
</taxonomicName>
. A. Cirrus with irregular spines. B. Grayish-white indumentose abaxial pinna surface. C. Leaf sheath with non-spiny knee. D. Fruits borne on short, stout pedicels.
</paragraph>
</caption>
<paragraph id="CA2C36DBB302FFF838F1FF10961EFC80" blockId="2.[136,1452,160,1153]" pageId="2" pageNumber="300">
<emphasis id="F8E7EAC9B302FFF838F1FF109428FF5B" bold="true" box="[189,257,160,183]" pageId="2" pageNumber="300">Stems</emphasis>
solitary, climbing, 5.030.0 m long, 3.0
<quantity id="0D6B9B3EB302FFF83AFAFF1097D7FF5B" box="[694,766,160,183]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="300" unit="cm" value="4.2">4.2 cm</quantity>
diameter.
<emphasis id="F8E7EAC9B302FFF83B27FF1096D0FF5B" bold="true" box="[875,1017,160,183]" pageId="2" pageNumber="300">Leaf sheaths</emphasis>
tubular, closed opposite the petiole, with a knee below the petiole; leaf sheath spines stout, triangular, concave at the base proximally, horizontally spreading, dense, yellowish-brown; ocreas short, membranous, spiny, extending into an acute apex above the petiole; flagella absent; petioles 2.015.0 cm long; rachises 2.0 m long, the apices extended into an elongate cirrus without well-developed pinnae, adaxially flat, abaxially with irregularly arranged, closely spaced, solitary or clustered dark-tipped, recurved spines, terminating in a shallow groove adaxially;
<emphasis id="F8E7EAC9B302FFF839E9FEFD94DBFE88" bold="true" box="[421,498,333,356]" pageId="2" pageNumber="300">pinnae</emphasis>
15 per side of rachis, regularly or slightly irregularly arranged, elliptic, with spinules on veins adaxially, grayish-white indumentose abaxially, the lateral veins diverging then converging, both terminating apically or slightly subapically; proximalmost pinnae swept back across the sheath; middle pinnae 38.556.0 cm long,
<quantity id="0D6B9B3EB302FFF83D7FFE229085FE45" box="[1331,1452,402,425]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="2" pageNumber="300" unit="cm" value="5.9" valueMax="6.3" valueMin="5.5">5.56.3 cm</quantity>
wide.
<emphasis id="F8E7EAC9B302FFF83886FE059442FE20" bold="true" box="[202,363,437,460]" pageId="2" pageNumber="300">Inflorescences</emphasis>
diverging from sheath well below sheath apex, with two vertical ridges distal to point of divergence; inflorescences arching, slender and elongate, usually without recurved spines, terminating in a long, tubular structure in line with the main inflorescence axis and covered with overlapping bracts; partial inflorescences not stalked, with a pulvinus in axil of rachis and partial inflorescence; partial inflorescences with elongate rachis bracts, recurved rachises, and rachillae subtended by short, funnel-shaped, equally long rachillae bracts (
<emphasis id="F8E7EAC9B302FFF83B73FDF09649FDBB" box="[831,864,576,599]" italics="true" pageId="2" pageNumber="300">i.e.</emphasis>
, rachis bracts and rachillae bracts strongly dimorphic); rachis bracts tubular but not closely sheathing the rachis, abruptly narrowed proximally; rachillae sessile; staminate inflorescences not recorded; pistillate inflorescences branched to 2 orders,
<quantity id="0D6B9B3EB302FFF83BF5FD35912FFD70" box="[953,1030,645,668]" metricMagnitude="0" metricUnit="m" metricValue="2.45" pageId="2" pageNumber="300" unit="m" value="2.45">2.45 m</quantity>
long; pistillate rachillae 12.015.0 cm long, with dyads of 1 pistillate and 1 neuter flower, the dyads arranged in alternate rows along recurved rachillae but not opposite so that one side of rachillae without dyads, borne on short, stout pedicels;
<emphasis id="F8E7EAC9B302FFF83C03FD7A91A4FD0D" bold="true" box="[1103,1165,714,737]" pageId="2" pageNumber="300">fruits</emphasis>
globose to ellipsoid, 12.0 mm long, 8.0 mm diameter, whitish; fruiting perianths tubular; fruit scales channeled vertically; seeds 1 per fruit, basally attached, small, globose, with a ventral depression, reniform in longitudinal section, the dorsal surfaces pitted, covered with a tanniniferous, non-fibrous sarcotesta; raphe branches 2, bifurcating from the attachment of seed and running along ventral side; endosperm homogeneous; embryos at or near base of seed.
</paragraph>
</subSubSection>
<subSubSection id="82896550B302FFF838F1FCC89625FB6D" pageId="2" pageNumber="300" type="distribution">
<paragraph id="CA2C36DBB302FFF838F1FCC89005FC63" blockId="2.[136,1452,160,1153]" box="[189,1324,888,911]" pageId="2" pageNumber="300">
<emphasis id="F8E7EAC9B302FFF838F1FCC894FBFC63" bold="true" box="[189,466,888,911]" pageId="2" pageNumber="300">Distribution and habitat</emphasis>
:—Eastern
<collectingRegion id="0857F839B302FFF83A07FCC89782FC63" box="[587,683,888,911]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="300">Sulawesi</collectingRegion>
in lowland rainforest on limestone soils at
<quantity id="0D6B9B3EB302FFF83C34FCC89195FC63" box="[1144,1212,888,911]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="300" unit="m" value="146.0">146 m</quantity>
elevation.
</paragraph>
<paragraph id="CA2C36DBB302FFF838F1FC2A97AAFC38" blockId="2.[136,1452,160,1153]" pageId="2" pageNumber="300">
<emphasis id="F8E7EAC9B302FFF838F1FC2A9488FC5D" bold="true" box="[189,417,922,945]" pageId="2" pageNumber="300">Taxonomic notes:—</emphasis>
<taxonomicName id="0D934D58B302FFF839EDFC2A97B2FC5D" authorityName="Henderson, Iqbal, Rusydi &amp; Pitopang" authorityYear="2018" box="[417,667,922,945]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="300" phylum="Tracheophyta" rank="species" species="tadulakoensis">
<emphasis id="F8E7EAC9B302FFF839EDFC2A97B2FC5D" box="[417,667,922,945]" italics="true" pageId="2" pageNumber="300">Calamus tadulakoensis</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="0D934D58B302FFF83B39FC2A9128FC5D" authorityName="Blume" authorityYear="1847" baseAuthorityName="Blume" box="[885,1025,922,945]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="300" phylum="Tracheophyta" rank="species" species="adspersus">
<emphasis id="F8E7EAC9B302FFF83B39FC2A9128FC5D" box="[885,1025,922,945]" italics="true" pageId="2" pageNumber="300">C. adspersus</emphasis>
</taxonomicName>
species group. It is, as far as we know, confined to limestone soils in eastern
<collectingRegion id="0857F839B302FFF83A57FC0D9756FC38" box="[539,639,957,980]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="300">Sulawesi</collectingRegion>
.
</paragraph>
<paragraph id="CA2C36DBB302FFF838F1FC509172FBF5" blockId="2.[136,1452,160,1153]" pageId="2" pageNumber="300">
The specific epithet honors Tadulako University, our research base. The term tadulako is also a common term in the region of
<collectingRegion id="0857F839B302FFF838BCFBB29481FBF5" box="[240,424,1026,1049]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="300">Central Sulawesi</collectingRegion>
, meaning a person of good virtue, wisdom, courage, and morals.
</paragraph>
<paragraph id="CA2C36DBB302FFF838F1FB959625FB6D" blockId="2.[136,1452,160,1153]" pageId="2" pageNumber="300">
<emphasis id="F8E7EAC9B302FFF838F1FB959704FBD0" box="[189,557,1061,1084]" inLineHeading="true" pageId="2" pageNumber="300" reason="1">
<emphasis id="F8E7EAC9B302FFF838F1FB95970FFBD0" bold="true" box="[189,550,1061,1084]" pageId="2" pageNumber="300">Additional specimen examined</emphasis>
.
</emphasis>
<collectingCountry id="B284764BB302FFF83A75FB9597E7FBD0" box="[569,718,1061,1084]" name="Indonesia" pageId="2" pageNumber="300">INDONESIA</collectingCountry>
.
<collectingRegion id="0857F839B302FFF83A91FB9596B3FBD0" box="[733,922,1061,1084]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="300">Sulawesi Tengah</collectingRegion>
: Banggai Regency, Lamala District, road from Labotan to Boloak, disturbed forest on steep slope, limestone soil, 00˚52.520S, 123˚14.423E,
<quantity id="0D6B9B3EB302FFF83CCBFBF891E2FBB3" box="[1159,1227,1096,1119]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="300" unit="m" value="146.0">146 m</quantity>
elevation,
<date id="BE2D101BB302FFF83D0DFBF895E9FB6D" pageId="2" pageNumber="300" value="2017-10-06">6 October 2017</date>
,
<emphasis id="F8E7EAC9B302FFF83887FBDA971BFB6D" box="[203,562,1130,1153]" italics="true" pageId="2" pageNumber="300">A. Henderson &amp; Moh. Iqbal 4256</emphasis>
(BO, CEB, K, NY).
</paragraph>
</subSubSection>
</treatment>
</document>