treatments-xml/data/42/37/87/423787F5920DFF8DFEBCFC076C90128E.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4765398" ID-GBIF-Dataset="cbcdb060-6372-4004-bcbb-327f8c67893d" ID-GBIF-Taxon="182138342" ID-ISSN="1854-0392" ID-Zenodo-Dep="4765398" ID-ZooBank="49D10BC2-1FA5-4926-87C8-391EE43A1C89" checkinTime="1620921834960" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2014" docId="423787F5920DFF8DFEBCFC076C90128E" docLanguage="en" docName="Illiesia.10.04.32-42.pdf" docOrigin="Illiesia 10 (4)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Tyloperla agumbe Stark &amp; Sivec 2014, sp. n." docType="treatment" docVersion="3" lastPageId="2" lastPageNumber="34" masterDocId="BE0EFF8D920CFF8FFFE3FFF16F261508" masterDocTitle="Three New Species Of Tyloperla Sivec &amp; Stark (Plecoptera: Perlidae) From India" masterLastPageNumber="42" masterPageNumber="32" pageId="1" pageNumber="33" updateTime="1643524802742" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Three New Species Of Tyloperla Sivec &amp; Stark (Plecoptera: Perlidae) From India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
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<mods:title>Illiesia</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>10</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:start>32</mods:start>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4757611</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4765398" ID-GBIF-Taxon="182138342" ID-Zenodo-Dep="4765398" LSID="urn:lsid:plazi:treatment:423787F5920DFF8DFEBCFC076C90128E" httpUri="http://treatment.plazi.org/id/423787F5920DFF8DFEBCFC076C90128E" lastPageId="2" lastPageNumber="34" pageId="1" pageNumber="33">
<subSubSection box="[351,635,1014,1038]" pageId="1" pageNumber="33" type="nomenclature">
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<heading bold="true" box="[351,635,1014,1038]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="33" reason="2">
<emphasis bold="true" box="[351,635,1014,1038]" pageId="1" pageNumber="33">
<taxonomicName authority="Stark &amp; Sivec, 2014" authorityName="Stark &amp; Sivec" authorityYear="2014" box="[351,564,1014,1038]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="agumbe" status="sp. n.">
<emphasis bold="true" box="[351,564,1014,1038]" italics="true" pageId="1" pageNumber="33">Tyloperla agumbe</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[571,635,1014,1038]" pageId="1" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[429,558,1050,1074]" pageId="1" pageNumber="33" type="description">
<paragraph blockId="1.[429,558,1050,1074]" box="[429,558,1050,1074]" pageId="1" pageNumber="33">
(
<figureCitation box="[437,550,1050,1074]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[189,241,1475,1499]" captionStartId-1="3.[189,241,1795,1819]" captionTargetBox-0="[298,1342,250,1399]" captionTargetBox-1="[230,1402,271,1717]" captionTargetId-0="figure-210@2.[287,1345,223,1420]" captionTargetId-1="figure-45@3.[230,1402,271,1717]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." captionText-1="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi-0="http://doi.org/10.5281/zenodo.4757615" figureDoi-1="http://doi.org/10.5281/zenodo.4757619" httpUri-0="https://zenodo.org/record/4757615/files/figure.png" httpUri-1="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Figs. 1-12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="33" type="materials_examined">
<paragraph blockId="1.[189,798,1121,1928]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[189,427,1121,1145]" pageId="1" pageNumber="33">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111937306" collectingDate="2004-10-09" collectionCode="USNM" collectorName="G. Svenson" country="India" latitude="13.489767" location="Agumbe Ghats" longLatPrecision="1" longitude="75.075615" pageId="1" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus box="[435,547,1121,1145]" pageId="1" pageNumber="33">Holotype</typeStatus>
<specimenCount box="[555,575,1121,1145]" pageId="1" pageNumber="33" type="male">♂</specimenCount>
, 
<emphasis bold="true" box="[590,661,1121,1145]" pageId="1" pageNumber="33">
<collectingCountry box="[590,656,1121,1145]" name="India" pageId="1" pageNumber="33">India</collectingCountry>
,
</emphasis>
<collectingRegion box="[670,793,1121,1145]" country="India" name="Karnataka" pageId="1" pageNumber="33">Karnataka</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:423787F5920DFF8DFEBCFC076C90128E:CF416038920DFF8EFF5EFB746E491195" box="[189,367,1157,1181]" country="India" latitude="13.489767" longLatPrecision="1" longitude="75.075615" name="Agumbe Ghats" pageId="1" pageNumber="33" stateProvince="Karnataka">Agumbe Ghats</location>
, 
<geoCoordinate box="[377,525,1156,1181]" degrees="13" direction="north" minutes="29.386" orientation="latitude" pageId="1" pageNumber="33" precision="1" value="13.489767">
13 
<emphasis bold="true" box="[404,415,1156,1181]" pageId="1" pageNumber="33">°</emphasis>
29.386’N
</geoCoordinate>
, 
<geoCoordinate box="[534,677,1156,1181]" degrees="75" direction="east" minutes="04.537" orientation="longitude" pageId="1" pageNumber="33" precision="1" value="75.075615">
75 
<emphasis bold="true" box="[561,572,1156,1181]" pageId="1" pageNumber="33">°</emphasis>
04.537’E
</geoCoordinate>
, 
<date pageId="1" pageNumber="33" value="2004-10-09">
<collectingDate pageId="1" pageNumber="33" value="2004-10-09">9 October 2004</collectingDate>
</date>
, 
<collectorName box="[259,394,1193,1217]" pageId="1" pageNumber="33">G. Svenson</collectorName>
(
<collectionCode box="[411,498,1193,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="33" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111937303" collectingDate="2004-10-11" collectionCode="BYUC" collectorName="G. Svenson" country="India" location="Same" pageId="1" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus box="[519,638,1193,1217]" pageId="1" pageNumber="33">Paratypes</typeStatus>
: 
<emphasis bold="true" box="[653,718,1192,1216]" pageId="1" pageNumber="33">
<collectingCountry box="[653,718,1192,1216]" name="India" pageId="1" pageNumber="33">India</collectingCountry>
</emphasis>
, 
<location LSID="urn:lsid:plazi:treatment:423787F5920DFF8DFEBCFC076C90128E:CF416038920DFF8EFD3DFB586C3B11C9" box="[734,797,1193,1217]" country="India" name="Same" pageId="1" pageNumber="33">Same</location>
site as 
<typeStatus box="[285,390,1228,1252]" pageId="1" pageNumber="33">holotype</typeStatus>
specimen but 
<date box="[586,792,1228,1253]" pageId="1" pageNumber="33" value="2004-10-11">
<collectingDate box="[586,792,1228,1253]" pageId="1" pageNumber="33" value="2004-10-11">11 October 2004</collectingDate>
</date>
, 
<collectingMethod box="[189,429,1264,1288]" pageId="1" pageNumber="33">canopy light trap</collectingMethod>
, 
<collectorName box="[459,612,1264,1288]" pageId="1" pageNumber="33">G. Svenson</collectorName>
, 
<specimenCount box="[641,675,1264,1288]" pageId="1" pageNumber="33" type="male">1♂</specimenCount>
(
<collectionCode box="[707,787,1264,1288]" pageId="1" pageNumber="33">BYUC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111937305" collectingDate="2004-10-03" collectionCode="BYUC" collectorName="Svenson &amp; Cameron &amp; Miller" country="India" latitude="15.9307165" location="Amboli" longLatPrecision="1" longitude="73.963936" pageId="1" pageNumber="33" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Maharashtra" typeStatus="paratype">
<collectingRegion box="[189,341,1299,1323]" country="India" name="Maharashtra" pageId="1" pageNumber="33">Maharashtra</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:423787F5920DFF8DFEBCFC076C90128E:CF416038920DFF8EFE81FAE26E981023" box="[354,446,1299,1323]" country="India" latitude="15.9307165" longLatPrecision="1" longitude="73.963936" name="Amboli" pageId="1" pageNumber="33" stateProvince="Maharashtra">Amboli</location>
, 
<geoCoordinate box="[459,611,1298,1323]" degrees="15" direction="north" minutes="55.843" orientation="latitude" pageId="1" pageNumber="33" precision="1" value="15.9307165">
15 
<emphasis bold="true" box="[486,497,1298,1323]" pageId="1" pageNumber="33">°</emphasis>
55.843’N
</geoCoordinate>
, 
<geoCoordinate box="[625,771,1298,1324]" degrees="73" direction="east" minutes="57.836" orientation="longitude" pageId="1" pageNumber="33" precision="1" value="73.963936">
73 
<emphasis bold="true" box="[652,663,1298,1323]" pageId="1" pageNumber="33">°</emphasis>
57.836’E
</geoCoordinate>
, 
<date pageId="1" pageNumber="33" value="2004-10-03">
<collectingDate pageId="1" pageNumber="33" value="2004-10-03">3 October 2004</collectingDate>
</date>
, 
<collectorName box="[370,471,1335,1359]" pageId="1" pageNumber="33">Svenson</collectorName>
, 
<collectorName box="[488,599,1335,1359]" pageId="1" pageNumber="33">Cameron</collectorName>
, 
<collectorName box="[617,692,1335,1359]" pageId="1" pageNumber="33">Miller</collectorName>
, 
<specimenCount box="[710,744,1335,1359]" pageId="1" pageNumber="33" type="male">9♂</specimenCount>
, 
<specimenCount box="[764,797,1335,1359]" pageId="1" pageNumber="33" type="female">6♀</specimenCount>
(
<collectionCode box="[197,277,1371,1395]" pageId="1" pageNumber="33">BYUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="33" type="description">
<paragraph blockId="1.[189,798,1121,1928]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[189,365,1406,1430]" pageId="1" pageNumber="33">Adult habitus.</emphasis>
General color brown, patterned with pale brown. Dorsum of head covered with moderately dense patch of upright, long, brown hair; head mostly pale but with darker areas between ocelli; lappets and basal antennal segment pale (
<figureCitation captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 1</figureCitation>
). Pronotum with narrow median yellow band and wider lateral pale band; disc mostly brown or pale brown, reflexed margin bearing a dark brown band. Wings amber, veins darker except for pale C vein. Femora pale except narrow, dark brown band at distal tips, tibiae dark brown.
</paragraph>
<paragraph blockId="1.[189,798,1121,1928]" lastBlockId="1.[835,1444,197,1927]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[189,258,1797,1821]" pageId="1" pageNumber="33">Male.</emphasis>
Forewings ca. 
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long. Abdominal sterna 6-8 bearing mesal patches of short, thick sensilla basiconica; sternum 8 with ca. 
<quantity box="[633,688,1868,1893]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="33" unit="in" value="20.0">20 in</quantity>
scattered patch, sternum 7 bearing a low, posteromedian tubercle covered rather densely with sensilla basiconica which extend to anterior third of segment in a more scattered patch (
<figureCitation box="[1163,1232,268,292]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 3</figureCitation>
), sternum 6 with only a few (ca. 4-5) sensilla basiconica in a marginal patch, sterna 8-9 without sensilla basiconica. Hemiterga with a well-developed basal callus and a short anterior process which reaches far short of tergum 9; a small arc of sensilla basiconica occurs between hemiterga on tergum 10 (
<figureCitation box="[1247,1316,481,505]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 2</figureCitation>
); terga 8-9 each with a small mesal lobe covered with sensilla basiconica. Aedeagus without spiny lobes; tube and sac in lateral aspect curved gradually ventrad and bearing a ventromedian saddle of moderately long spines, and a subapical ring of short, thick, spines (
<figureCitation box="[843,910,694,718]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 4</figureCitation>
); subapical ring interrupted by a longitudinal median groove in which few spines occur, and from which a small membranous bag-like structure is everted (
<figureCitation box="[942,1013,800,824]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 5</figureCitation>
). A single seta occurs on each side between ventral saddle of spines and subapical ring (
<figureCitation box="[843,909,871,896]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 5</figureCitation>
).
</paragraph>
<paragraph blockId="1.[835,1444,197,1927]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[835,929,907,931]" pageId="1" pageNumber="33">Female.</emphasis>
Forewing length 
<quantity box="[1133,1251,907,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="1" pageNumber="33" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16-17 mm</quantity>
. Subgenital plate on sternum 8 triangular in outline, projecting over about 1/3 of sternum 9, and bearing a small U-shaped mesal notch (
<figureCitation box="[995,1064,1014,1038]" captionStart="Figs" captionStartId="2.[189,241,1475,1499]" captionTargetBox="[298,1342,250,1399]" captionTargetId="figure-210@2.[287,1345,223,1420]" captionTargetPageId="2" captionText="Figs. 1-6. Tyloperla agumbe structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="1" pageNumber="33">Fig. 6</figureCitation>
). U-shaped region surrounding notch hairless; plate darker than background pigment of segment. Width of plate at base ca. ½ of sternal width.
</paragraph>
<paragraph blockId="1.[835,1444,197,1927]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[835,887,1156,1180]" pageId="1" pageNumber="33">Egg.</emphasis>
Outline almost spherical (
<figureCitation box="[1193,1259,1156,1180]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 7</figureCitation>
). Length ca. 335 μm, equatorial width ca. 328 μm. Collar short and wide, width ca. 89 μm, rim not distinctly flanged, sides bearing ca. 11 vertical ridges in lateral aspect (
<figureCitation box="[843,927,1298,1323]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 10</figureCitation>
). Anchor a thin, umbrella-shaped structure with short pedicel, an irregular margin and apparently few surface globular bodies (
<figureCitation box="[1353,1428,1369,1394]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 8</figureCitation>
). Chorion covered throughout with mostly pentagonal and hexagonal follicle cell impressions (
<figureCitation box="[1313,1430,1441,1465]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Figs. 9-12</figureCitation>
); walls consist of six low, rounded tubercles set at FCI angles, forming cells ca. 27 μm across; most FCI floors contain a single central and six, fine submarginal aeropyles, but some contain 8-9 aeropyles and the central one may sometimes be displaced (
<figureCitation box="[965,1034,1654,1678]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 9</figureCitation>
). FCI’s on lid have tubercles more tightly grouped and slightly larger than those nearer the equatorial region (
<figureCitation box="[1090,1166,1725,1749]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 11</figureCitation>
). Subequatorial eclosion ring distinct, surface of ring smooth and ca. 12 μm wide (
<figureCitation box="[909,991,1796,1820]" captionStart="Figs" captionStartId="3.[189,241,1795,1819]" captionTargetBox="[230,1402,271,1717]" captionTargetId="figure-45@3.[230,1402,271,1717]" captionTargetPageId="3" captionText="Figs. 7-12. Tyloperla agumbe egg structures. 7. Entire egg, lateral aspect. 8. Collar end with anchor. 9. Chorionic detail, equatorial zone. 10. Collar end with most of anchor removed. 11.Chorionic detail, lid. 12. Equatorial zone, eclosion line and lid." figureDoi="http://doi.org/10.5281/zenodo.4757619" httpUri="https://zenodo.org/record/4757619/files/figure.png" pageId="1" pageNumber="33">Fig. 12</figureCitation>
). Micropyles not detected.
</paragraph>
<paragraph blockId="1.[835,1444,197,1927]" box="[835,1042,1831,1855]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[835,910,1831,1855]" pageId="1" pageNumber="33">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="33" type="etymology">
<paragraph blockId="1.[835,1444,197,1927]" pageId="1" pageNumber="33">
<emphasis bold="true" box="[835,973,1867,1891]" pageId="1" pageNumber="33">Etymology.</emphasis>
The species name, based on the 
<typeStatus box="[1391,1443,1868,1892]" pageId="1" pageNumber="33">type</typeStatus>
locality, is used as a noun in apposition.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757615" ID-Zenodo-Dep="4757615" httpUri="https://zenodo.org/record/4757615/files/figure.png" pageId="2" pageNumber="34" startId="2.[189,241,1475,1499]" targetBox="[298,1342,250,1399]" targetPageId="2">
<paragraph blockId="2.[189,1442,1475,1535]" pageId="2" pageNumber="34">
Figs. 1-6. 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2014" box="[301,492,1475,1499]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="agumbe">
<emphasis box="[301,492,1475,1499]" italics="true" pageId="2" pageNumber="34">Tyloperla agumbe</emphasis>
</taxonomicName>
structures. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Male abdominal sterna 7-8. 4. Aedeagus, lateral. 5. Aedeagus, ventral. 6. Female subgenital plate.
</paragraph>
</caption>
<subSubSection pageId="2" pageNumber="34" type="diagnosis">
<paragraph blockId="2.[189,797,1653,1926]" lastBlockId="2.[835,1443,1653,1926]" pageId="2" pageNumber="34">
<emphasis bold="true" box="[189,321,1653,1677]" pageId="2" pageNumber="34">Diagnosis.</emphasis>
The short hemitergal lobes and median sclerite with sensilla basiconica on tergum 10 are features unique to this species and 
<taxonomicName authority="Stark &amp; Sivec, 2014" authorityName="Stark &amp; Sivec" authorityYear="2014" box="[616,754,1724,1748]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="karnataka" status="sp. n.">
<emphasis box="[616,754,1724,1748]" italics="true" pageId="2" pageNumber="34">T. karnataka</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
among known 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2014" box="[413,517,1760,1784]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis box="[413,517,1760,1784]" italics="true" pageId="2" pageNumber="34">Tyloperla</emphasis>
</taxonomicName>
; males of the two are distinguished on the basis of the presence of a prominent lobe on sternum 
<quantity box="[548,598,1831,1855]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="34" unit="in" value="7.0">7 in</quantity>
<emphasis box="[612,740,1831,1855]" italics="true" pageId="2" pageNumber="34">
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2014" box="[612,735,1831,1855]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="agumbe">T. agumbe</taxonomicName>
.
</emphasis>
The female subgenital plate, reaching near midlength of sternum 9, is somewhat shorter than that of previously described species, and the egg is generally similar to those of 
<taxonomicName authority="Stark &amp; Sivec" authorityName="Stark &amp; Sivec" authorityYear="2005" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="courtneyi">
<emphasis box="[1199,1338,1689,1712]" italics="true" pageId="2" pageNumber="34">T. courtneyi</emphasis>
Stark &amp; Sivec
</taxonomicName>
, 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2014" box="[917,1061,1724,1748]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="karnataka">
<emphasis box="[917,1061,1724,1748]" italics="true" pageId="2" pageNumber="34">T. karnataka</emphasis>
</taxonomicName>
(see below), 
<taxonomicName authority="Stark &amp; Sivec" authorityName="Stark &amp; Sivec" authorityYear="2005" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="khang">
<emphasis box="[1233,1335,1724,1748]" italics="true" pageId="2" pageNumber="34">T. khang</emphasis>
Stark &amp; Sivec
</taxonomicName>
and 
<taxonomicName authority="Cao &amp; Bae" authorityName="Cao &amp; Bae" box="[966,1188,1760,1784]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="trui">
<emphasis box="[966,1043,1760,1783]" italics="true" pageId="2" pageNumber="34">T. trui</emphasis>
Cao &amp; Bae
</taxonomicName>
in having a distinct eclosion ring and prominent FCI’s over the entire egg (
<bibRefCitation author="Cao, T. K. T. &amp; Y. J. Bae" box="[894,1089,1831,1855]" pageId="2" pageNumber="34" pagination="329 - 334" refId="ref4124" refString="Cao, T. K. T. &amp; Y. J. Bae. 2007. Vietnamese stonefly species of the genus Tyloperla (Plecoptera: Perlidae). Journal of Asia-Pacific Entomology, 10: 329 - 334." type="journal article" year="2007">Cao &amp; Bae 2007</bibRefCitation>
; 
<bibRefCitation author="Stark, B. P. &amp; I. Sivec" box="[1101,1332,1831,1855]" pageId="2" pageNumber="34" pagination="1 - 7" refId="ref4395" refString="Stark, B. P. &amp; I. Sivec. 2005. New species of Tyloperla (Plecoptera: Perlidae) from Vietnam and Thailand. Illiesia, 1: 1 - 7." type="journal article" year="2005">Stark &amp; Sivec 2005</bibRefCitation>
). A more complete diagnosis is given below under 
<emphasis italics="true" pageId="2" pageNumber="34">
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2014" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="karnataka">T. karnataka</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>