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<document id="B0DF6D312D6727E301D12146A667A26F" ID-CLB-Dataset="34103" ID-DOI="http://dx.doi.org/10.3897/zookeys.346.6114" ID-GBIF-Dataset="892cdf91-8f73-426f-9143-b21e3d4df75d" ID-PMC="PMC3821061" ID-Pensoft-Pub="1313-2970-346-1" ID-PubMed="24223483" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-346-1" ModsDocOrigin="ZooKeys 346" ModsDocTitle="Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" checkinTime="1451246756994" checkinUser="pensoft" docAuthor="Perger, Robert &amp; Grossi, Paschoal Coelho" docDate="2013" docId="421B2C21647FB09372C611317C6E7A7E" docLanguage="en" docName="ZooKeys 346: 1-16" docOrigin="ZooKeys 346" docSource="http://dx.doi.org/10.3897/zookeys.346.6114" docTitle="Oryctophileurus armicollis Prell 1911" docType="treatment" docVersion="5" lastPageNumber="8" masterDocId="562FFFC4FFE7E0630407FF9EFF8CFFC5" masterDocTitle="Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="8" updateTime="1732733777238" updateUser="ExternalLinkService">
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<mods:title id="3D9C290D9EF98339D2E800E3E1F86B6A">Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)</mods:title>
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<mods:namePart id="2782F85640C400B1B16453C7BF138B33">Perger, Robert</mods:namePart>
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<mods:namePart id="027BBE7B2C9CD59108E83B495978C21B">Grossi, Paschoal Coelho</mods:namePart>
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<mods:classification id="85D0C189BF128248C08BA00445CFDB60">journal article</mods:classification>
<mods:identifier id="B0BF2F2CFC22D2A94E877681096E9F27" type="DOI">http://dx.doi.org/10.3897/zookeys.346.6114</mods:identifier>
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<treatment id="421B2C21647FB09372C611317C6E7A7E" ID-GBIF-Taxon="152049040" LSID="urn:lsid:plazi:treatment:421B2C21647FB09372C611317C6E7A7E" httpUri="http://treatment.plazi.org/id/421B2C21647FB09372C611317C6E7A7E" lastPageNumber="8" pageId="7" pageNumber="8" scope_subFamily="Dynastinae" scope_superFamily="Scarabaeoidea" scope_tribe="Phileurini">
<subSubSection id="60B9D409C8B9DD59FF58EB5AB06C68E9" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="B5BEE8FE0B408AAAE229B012E040F6AA" pageId="7" pageNumber="8">
<taxonomicName id="326CF223E05AB64AA072914126EF2848" ID-CoL="74ZBS" LSID="http://species-id.net/wiki/Oryctophileurus_armicollis" authority="Prell, 1911" authorityName="Prell" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus armicollis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="armicollis">Oryctophileurus armicollis Prell, 1911</taxonomicName>
Figs 5
<normalizedToken id="510BD90CE665D84223344E5C862C996E" originalValue="AB">A-B</normalizedToken>
; 6; 7A, C
</paragraph>
</subSubSection>
<subSubSection id="AE11D7E7CBA868E84E495D0D58DF1EC3" pageId="7" pageNumber="8" type="type material examined">
<paragraph id="961BB47ACC429875D510616CD88502FA" pageId="7" pageNumber="8">Type material examined.</paragraph>
<paragraph id="8F4A14665F1284D79FF1269CC05794DD" pageId="7" pageNumber="8">Peru: holotype, male, body length 18.5 mm (ZMHB).</paragraph>
</subSubSection>
<subSubSection id="3DC0C9B224FBBACB9DEF0A17B6C623BB" pageId="7" pageNumber="8" type="additional material examined">
<paragraph id="710D73ACF380F98F9C93C17F9F1459E7" pageId="7" pageNumber="8">Additional material examined.</paragraph>
<paragraph id="633282C25DC21E4AA530084CAA2A6485" pageId="7" pageNumber="8">Bolivia: La Paz dep., Nor Yungas prov., Caranavi, I-2003, 2 males with body length 18 mm and 20 mm and 1 female with body length 18 mm (EPGC); Route Coroico-Caranavi, XII-2008, 1 male (EPGC); La Paz dep., Calisaia, V-1925, G.L. Harrington col., 1 male (MZSP); Beni dep., Cosincho, VIII-1925, G.L. Harrington col., 1 female (MZSP).</paragraph>
</subSubSection>
<subSubSection id="015B3F46AE6EEB7D665356D682DAE5E4" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="13492FF2779BD9911E669FA15AA07F78" pageId="7" pageNumber="8">Diagnosis.</paragraph>
<paragraph id="56747D501DA05D73BC8F5012DBD0005F" pageId="7" pageNumber="8">Color black, strongly shining. Head anteriorly flat, not concave; canthus subquadrate, in some specimens weakly projected forward; horn in males strongly recurved, on about the same level or slightly higher than posterior pronotal protuberance. Mentum with longitudinal furrow flat and wide, as well as posterior margin. Distance between inner teeth of dorsal pronotal protuberance in male wider that width between eyes (Fig. 6A); in females separated by a concavity with occelate punctures; pronotal concavity accounts for about 40-50 % of dorsal pronotal surface in males. Ocellate punctures above posterior-lateral pronotal margin continuous (Fig. 7C). Each elytron with 10 striae; punctures ocellate, sometimes coalescent and elongated, decreasing in size to sides; interstriae smooth, convex. Striae and interstitial punctures arranged in regular lines Mesotibia with 3 to 5 apical teeth; metatibia with 4 to 6 apical teeth. Pygidium in males distinctly more convex than in females, moderately punctate, denser at base and sides. Aedeagus symmetric (Fig. 5A); parameres with apex rounded to subtriangle shaped (Fig. 5B).</paragraph>
<caption id="E1E02EE801E052432CBD11654636857B" pageId="7" pageNumber="8">
<paragraph id="67A35F4E3283732003E907083B2F1827" pageId="7" pageNumber="8">
Figure 6. Dorsal and lateral habitus of
<taxonomicName id="2923C3FDC42D11FB018E873CFE399876" genus="Oryctophileurusarmicollis" lsidName="Oryctophileurusarmicollis" pageId="7" pageNumber="8" rank="genus">Oryctophileurusarmicollis</taxonomicName>
Prell, 1911, A and B holotype male, body length 18.5 mm C and D female, body length 18 mm, scale bar 5 mm.
</paragraph>
</caption>
<caption id="B30526073A9D788B1B518B122EE9000D" pageId="7" pageNumber="8">
<paragraph id="B5D4A89CA54AA04B07E352B941674A8C" pageId="7" pageNumber="8">
Figure 7. Pronotum of males with body length 20 mm, dorsal, higher areas light, inner teeth of transversal pronotal carina indicated by white arrow: A
<taxonomicName id="9056FA5F54F22E8F7E10FB213A529095" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus armicollis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="armicollis">Oryctophileurus armicollis</taxonomicName>
Prell B
<taxonomicName id="6099CE984F16E1B19D0501F473172256" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus guerrai" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guerrai">Oryctophileurus guerrai</taxonomicName>
; ocellate punctures at posteriolateral pronotal surface, character state does not differ between sexes C
<taxonomicName id="B0B73B30E473DC4F32CEF4DD6BCAAEB8" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus armicollis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="armicollis">Oryctophileurus armicollis</taxonomicName>
D
<taxonomicName id="1444D8890F6D7B98D602C92AEF60C41A" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus guerrai" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guerrai">Oryctophileurus guerrai</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4B7DC0247F66A5B607A398BD0F6C379E" pageId="7" pageNumber="8" type="geographical distribution">
<paragraph id="C11DC7227616BD49D239C153BC47400C" pageId="7" pageNumber="8">Geographical distribution.</paragraph>
<paragraph id="77F69190DBCEA6F09B19A321FEB1514F" pageId="7" pageNumber="8">
<taxonomicName id="BD2E5B902FAE8DEE1AE57BBE17E5057E" class="Insecta" family="Scarabaeidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus armicollis" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="armicollis">Oryctophileurus armicollis</taxonomicName>
is known from Peru (
<bibRefCitation id="3439954F38D754E0B23C1A6B0D1EA07A" author="Prell, H" journalOrPublisher="Entomologische Zeitschrift" pageId="14" pageNumber="15" pagination="105 - 107" title="Beitraege zur Kenntnis der Dynastinen." volume="25" year="1911">Prell 1911</bibRefCitation>
; location not specified) and the Andean (Nor Yungas, La Paz dep.) and Subandean (Cosincho, Beni dep.) areas of Bolivia (Fig. 1A). The ecosystem between 800 and 2000 m a.s.l. in this area is considered as South Yungas submontane, subhumid forest (
<bibRefCitation id="1189AED29D8BDF858FA56AA86395FB24" author="Josse, C" journalOrPublisher="NatureServe, Arlington, VA" pageId="13" pageNumber="14" title="Ecological systems of Latin America and the Caribbean: A working classification of terrestrial systems." year="2003">Josse et al. 2003</bibRefCitation>
) and receives an annual precipitation between 1500-6000 mm (
<bibRefCitation id="98E9B098C541A29E5BA3121D11EEE324" author="Ibisch, PL" editor="Ibisch, PL" journalOrPublisher="Editorial Fundacion Amigos de la Naturaleza (FAN), Santa Cruz, Bolivia" pageId="13" pageNumber="14" pagination="47 - 88" title="La diversidad biologica: ecorregiones y ecosistemas." year="2003 a">Ibisch et al. 2003a</bibRefCitation>
). The locality data suggests that this species is closely associated with the Bolivian Yungas forest and might also occur in the Peruvian Yungas forest.
</paragraph>
</subSubSection>
</treatment>
</document>