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128 lines
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<document id="5AC94FEBBB61803F48D1799D6C9299E3" ID-CLB-Dataset="32988" ID-DOI="http://dx.doi.org/10.3897/zookeys.675.13252" ID-GBIF-Dataset="724d6de0-f19a-457d-9339-84c836e19c38" ID-PMC="PMC5523889" ID-Pensoft-Pub="1313-2970-675-129" ID-PubMed="28769682" ID-ZooBank="E5C8F163D61547B98418CEE8D71A7DAB" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-675-129" ModsDocOrigin="ZooKeys 675" ModsDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" checkinTime="1495538522124" checkinUser="pensoft" docAuthor="Budha, Prem B., Naggs, Fred & Backeljau, Thierry" docDate="2017" docId="420EEDBFE04C8DE744FB1A39F2B69C7B" docLanguage="en" docName="ZooKeys 675: 129-156" docOrigin="ZooKeys 675" docSource="http://dx.doi.org/10.3897/zookeys.675.13252" docTitle="Glessula tamakoshi Budha & Backeljau, sp. n." docType="treatment" docUuid="1366B4C7-D3B6-4FE5-AFD0-55C7CDD7A400" docUuidSource="ZooBank" docVersion="9" lastPageNumber="133" masterDocId="FF8BE96F4A658939FFA88167FF8E6E18" masterDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" masterLastPageNumber="156" masterPageNumber="129" pageNumber="133" updateTime="1732877711202" updateUser="ExternalLinkService">
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<mods:title id="66170F8E191B05DC5B869E99B74BC09C">Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species</mods:title>
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<mods:roleTerm id="B8460BE93A7B0DB94606C78663AD70C8">Author</mods:roleTerm>
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<mods:namePart id="D2BEE036BEF99176B49502978FD7E77D">Budha, Prem B.</mods:namePart>
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<mods:namePart id="CFF38DA616EDDFB9C81EDCE7F9FCA034">Naggs, Fred</mods:namePart>
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<mods:roleTerm id="AA3089A62D7D454B1FB77BEEBCE50F1B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F3E23C44B235ED13FF9455501A75D28">Backeljau, Thierry</mods:namePart>
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<mods:typeOfResource id="B2C20EC0E29201050F012729B57AA0EC">text</mods:typeOfResource>
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<mods:title id="F92A65171EB72362379CF9E4E5E28405">ZooKeys</mods:title>
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<mods:part id="ED5E05E928461AB5C928FFEA51E05EB0">
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<mods:date id="ECA8349CA13873D97BC9003894F22496">2017</mods:date>
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<mods:detail id="55B1A196407D296BAE4BD63A9D19458D" type="volume">
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<mods:number id="24CBAC855946152262405AB764099166">675</mods:number>
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<mods:start id="969A60207843C22700C5A4C82A1833C1">129</mods:start>
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<mods:url id="785448A019400A1E4B19C5B1297E475F">http://dx.doi.org/10.3897/zookeys.675.13252</mods:url>
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<mods:classification id="CA215750FD431F83880892C78B2CC1ED">journal article</mods:classification>
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<mods:identifier id="DCFEBAE2A0C23A7704C774DF7876A0C9" type="DOI">http://dx.doi.org/10.3897/zookeys.675.13252</mods:identifier>
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<mods:identifier id="AE75BCC81113A0B304556285E184620E" type="Pensoft-Pub">1313-2970-675-129</mods:identifier>
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<treatment id="420EEDBFE04C8DE744FB1A39F2B69C7B" ID-GBIF-Taxon="130844520" LSID="urn:lsid:zoobank.org:act:1366B4C7-D3B6-4FE5-AFD0-55C7CDD7A400" httpUri="http://treatment.plazi.org/id/420EEDBFE04C8DE744FB1A39F2B69C7B" lastPageNumber="133" pageId="4" pageNumber="133" scope_class="Gastropoda" scope_order="Stylommatophora" scope_subFamily="Glessulinae">
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<subSubSection id="29452F3A18B3FF0E0624AF3DBDB941BF" pageId="4" pageNumber="133" type="nomenclature">
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<paragraph id="536A4F730014FABB694E3FA62D5F26BC" pageId="4" pageNumber="133">
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<taxonomicName id="AAC9AC8CCAB978FC836B7C496150BC4C" LSID="http://zoobank.org/1366B4C7-D3B6-4FE5-AFD0-55C7CDD7A400" authority="Budha & Backeljau" family="Subulinidae" genus="Glessula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Glessula tamakoshi" order="Gastropoda" pageId="4" pageNumber="133" rank="species" species="tamakoshi">Glessula tamakoshi Budha & Backeljau</taxonomicName>
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<taxonomicNameLabel id="AA23A0D830F459B09BEBEB72173EFCE3" pageId="4" pageNumber="133">sp. n.</taxonomicNameLabel>
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Figs 2C, 5, 13A
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</paragraph>
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</subSubSection>
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<subSubSection id="B7958B085850B540BA5A5F5A95F414EC" pageId="4" pageNumber="133" type="materials_examined">
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<paragraph id="FA47713652055EDE3570E5211C056A77" pageId="4" pageNumber="133">Material examined.</paragraph>
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<paragraph id="4AC579ED945936A1B5BDE9DF21B30B21" pageId="4" pageNumber="133">
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Holotype: CDZMTU057P/1, Suridobhan, left bank of Tamakoshi River, Dolakha District, Central Nepal, North face, rocky hill slope, mixed
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<taxonomicName id="BE83F75891515D2C306A68057F048A7B" class="Magnoliopsida" family="Theaceae" genus="Schima" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Schima wallichi" order="Ericales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="wallichi">Schima wallichi</taxonomicName>
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forest with dominant
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<taxonomicName id="DBBDFB24E9911C8B0560C7937F074415" class="Magnoliopsida" family="Lauraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Laurales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
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trees,1023 m,
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<geoCoordinate id="5A5E057AB67497925F62AFEAED705BC3" direction="north" orientation="latitude" precision="1" value="27.754753">27.754754N</geoCoordinate>
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,
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<geoCoordinate id="F6F907150AA48FD3A6E38172DDF65D98" direction="east" orientation="longitude" precision="1" value="86.21676">86.216755E</geoCoordinate>
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, 03.II.2009, leg. P.B. Budha. Paratypes: CDZMTU058/13 shells and CDZMTU058P/2 specimens (dissected) from the type locality (same data as holotype).
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</paragraph>
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</subSubSection>
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<subSubSection id="0253EDD596803E92DFB806775D2DB47B" pageId="4" pageNumber="133" type="distribution">
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<paragraph id="0D41D9BF3EB7A2640154DF3610DB6967" pageId="4" pageNumber="133">Distribution.</paragraph>
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<paragraph id="8B3C1851C703F1E651BD6B38637C95B8" pageId="4" pageNumber="133">Only known from the type locality.</paragraph>
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</subSubSection>
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<subSubSection id="D524B99B0ED4C47C2321E74ED6B6EDC9" pageId="4" pageNumber="133" type="etymology">
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<paragraph id="230416799215334DE1247988AC80E30E" pageId="4" pageNumber="133">Etymology.</paragraph>
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<paragraph id="88903D4626DFC058CA5437B3F000B01C" pageId="4" pageNumber="133">The species name refers to the type locality Tamakoshi River valley.</paragraph>
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</subSubSection>
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<subSubSection id="49E23A7ED3B373B146742250A35EBB59" pageId="4" pageNumber="133" type="shell">
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<paragraph id="217FFE36E739FFE2927BF4DF5BAC414E" pageId="4" pageNumber="133">Shell.</paragraph>
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<paragraph id="50A2DF81BA7FFE6BBB67EC95D3C5779F" pageId="4" pageNumber="133">
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Measurements (n = 4): SH 17.7-19.5 mm, SW 9.2-9.6 mm, HA 8.6-8.8 mm, WA 4.9-5.0 mm, Wh 7.0-8.0; holotype: SH 19.5 mm, SW 9.6 mm, HA 8.8 mm, WA 4.9 mm, Wh 7.8. The largest shell measured 19.5 mm, approx. 2
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<normalizedToken id="AB167927C2D3E0F4AE6B0CA3C2C09813" originalValue="×">x</normalizedToken>
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higher than wide, solid, ovate-conic, light yellowish. Surface glossy, with widely spaced incised radial striations, stronger towards the suture and faint at the middle to lower part of the whorls. The first whorl smooth, blunt with fine and dense striations near the suture. Sides convex, suture fairly impressed. Aperture nearly ovate, 1.7
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<normalizedToken id="C17EFC450FBED02A81A92ECC479960FB" originalValue="×">x</normalizedToken>
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higher than wide, peristome simple and thick, columellar margin abruptly truncate, columella slightly curved.
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</paragraph>
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</subSubSection>
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<subSubSection id="92C8FEBAC6C1957320C2E6773F33999E" pageId="4" pageNumber="133" type="radula">
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<paragraph id="4D7E07ABF7B21788F7FF75D7661E8F91" pageId="4" pageNumber="133">Radula</paragraph>
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<paragraph id="957914A063476E9A8987D431D33D2191" pageId="4" pageNumber="133">(Fig. 13A). Teeth pointed, lateral cusps not distinct. Central tooth smaller and with a narrower base than the lateral teeth.</paragraph>
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</subSubSection>
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<subSubSection id="E1ECB356D00D63082099CB769CC3E70F" pageId="4" pageNumber="133" type="genitalia">
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<paragraph id="87254CD5B2FB0FDCE6E9FBB1915E63FF" pageId="4" pageNumber="133">Genitalia</paragraph>
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<paragraph id="5170712B8DCE152424FD1D15D22C04A6" pageId="4" pageNumber="133">
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(n = 3) (Fig. 5). Vas deferens with a constant diameter. Flagellum wide comb-like with numerous notches ending in a short hook at the free end. Vagina very short nearly 1/6th length of the penis. Gametolytic sac is oval, connected to the gametolytic duct by a short neck. The penial retractor muscle close to the flagellum. The mass of the hermaphroditic duct highly developed in all dissected samples. Interior of penis with two bulky masses of 'brain-like
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<normalizedToken id="F27C79FEB9A65D039F00EB035A7A25F0" originalValue="folds’">folds'</normalizedToken>
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(Fig. 5B).
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</paragraph>
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<caption id="D938C809D6E45975A05B1F6E22C0E90E" pageId="4" pageNumber="133">
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<paragraph id="6971A748D3A06B8D106F47F77822B117" pageId="4" pageNumber="133">
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Figure 5. Genitalia of
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<taxonomicName id="9146904E0E4ED51CE33AAE928FD8BA37" lsidName="G. tamakoshi" pageId="4" pageNumber="133" rank="species" species="tamakoshi">G. tamakoshi</taxonomicName>
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sp. n., holotype: CDZMTU057P, A General view B Interior of penis of the same, Suridobhan, left bank of Tamakoshi River, Dolakha District, Central Nepal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="9FD30FD254F001F2CA72361453A1C085" pageId="4" pageNumber="133" type="remarks">
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<paragraph id="A76A9014E7DF6A58C290C98FE0A3A464" pageId="4" pageNumber="133">Remarks.</paragraph>
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<paragraph id="9EC7F3E8C8672DFF37A3DC7AE819598B" pageId="4" pageNumber="133">
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Conchologically,
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<taxonomicName id="665E8B5EFD00208D82605833ECF93907" lsidName="G. tamakoshi" pageId="4" pageNumber="133" rank="species" species="tamakoshi">G. tamakoshi</taxonomicName>
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sp. n. is similar to South Indian
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<taxonomicName id="149D61FFFA0852DB5D223090A9C49DB3" lsidName="G. indica" pageId="4" pageNumber="133" rank="species" species="indica">G. indica</taxonomicName>
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Gude, 1914 (genitalia unknown) and Sri Lankan
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<taxonomicName id="635350EBE5C828F22B438B7326158852" lsidName="G. serena" pageId="4" pageNumber="133" rank="species" species="serena">G. serena</taxonomicName>
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(Benson, 1860). Yet,
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<taxonomicName id="04992E8D3217D2ACCDE5842C185BE6D2" lsidName="G. indica" pageId="4" pageNumber="133" rank="species" species="indica">G. indica</taxonomicName>
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has a much wider shell and relatively more whorls (>9 whorls), while the penis of
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<taxonomicName id="14BF50DC41A6ECCABA13733A7CE337A3" lsidName="G. serena" pageId="4" pageNumber="133" rank="species" species="serena">G. serena</taxonomicName>
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contains one longitudinal fold and two folds proximal to this longitudinal fold.
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<bibRefCitation id="9EE369E7AAAA649FE02D5737C3CBA910" author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="14" pageNumber="143" pagination="437 - 564" title="Treatise on recent terrestrial pulmonate molluscs Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae." volume="2" year="1999">Schileyko (1999</bibRefCitation>
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, p. 542, fig. 711) referred the longitudinal fold to as pilaster and the two proximal folds to as the spiral stimulators. In
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<taxonomicName id="63C7EC7A91053EABCEC0E442B87BA192" lsidName="G. tamakoshi" pageId="4" pageNumber="133" rank="species" species="tamakoshi">G. tamakoshi</taxonomicName>
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sp. n. the interior of the penis contains a bulky masses of brain-like folds (Fig. 5B).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |