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<mods:title id="3D55A6F2BF0CE79270784BD0D2CEAACC">New insights into the identities of the elasmobranch fauna of Sri Lanka</mods:title>
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<taxonomicName id="7984E2A7DA263E400AC1FAB8FEB1FAD2" authorityName="Muller &amp; Henle" authorityYear="1837" box="[151,319,1337,1363]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA263E400AC1FAB8FEB1FAD2" bold="true" box="[151,319,1337,1363]" italics="true" pageId="28" pageNumber="229">Chiloscyllium</emphasis>
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sp. 1
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<paragraph id="BE3B9924DA263E400AC1FADDFED1FAF5" blockId="28.[151,383,1337,1397]" box="[151,351,1372,1397]" pageId="28" pageNumber="229">
(
<figureCitation id="26BF85A1DA263E400AC9FADDFE8DFAF4" box="[159,259,1372,1397]" captionStart="FIGURE 5" captionStartId="9.[151,250,1427,1449]" captionTargetBox="[166,1430,205,1398]" captionTargetId="figure@9.[151,1436,193,1406]" captionTargetPageId="9" captionText="FIGURE 5. Neighbor-Joining trees of p-distances based on 1,044 bp of aligned NADH2 sequence data for individual selachian genera or groups of genera. Taxon names are followed by country, host specimen number (in parentheses), and GenBank number. Outgroup taxa used in each analysis were omitted from the individual trees. Scale bar values indicate substitutions per site. Specimens deposited in the Blue Resources Trust Ichthyology Collection are indicted by stars (see text for details). A. Echinorhinidae. B. Hexanchidae. C. Lamniformes. D. Chiloscyllium (Hemiscylliidae). E. Centrophorus (Centrophoridae). F. Somniosidae/Oxynotidae (Somniosus omitted)." httpUri="https://zenodo.org/record/2637246/files/figure.png" pageId="28" pageNumber="229">Figs. 5D</figureCitation>
,
<figureCitation id="26BF85A1DA263E400B58FADDFEA6FAF5" box="[270,296,1372,1396]" captionStart="FIGURE 17" captionStartId="29.[151,250,1806,1828]" captionTargetBox="[151,1436,472,1775]" captionTargetId="figure@29.[151,1436,472,1775]" captionTargetPageId="29" captionText="FIGURE 17. Echinorhinus sp. 1 (Echinorhinidae) (AD), Hexanchus griseus (Hexanchidae) (EH), and Chiloscyllium sp. 1 (Hemiscylliidae) (IK). A. Lateral view of female (SL-108). B. Ventral view of head of female (SL-108). C. Teeth of upper jaw of female (SL-108). D. Teeth of lower jaw of female (SL-108). E. Lateral view of immature male (SL-106). F. Ventral view of head of female (SL-91). G. Teeth of upper jaw of female (SL-91). H. Teeth of lower jaw of female (SL-91). I. Lateral view of mature male. J. Ventral view of head of mature male (SL-59). K. Dorsal view of head and trunk of mature male (SL-59)." httpUri="https://zenodo.org/record/2637270/files/figure.png" pageId="28" pageNumber="229">17</figureCitation>
IK)
</paragraph>
<paragraph id="BE3B9924DA263E410AC1FA1FFB9FFE18" blockId="28.[151,1437,1438,2030]" lastBlockId="29.[151,1436,151,410]" lastPageId="29" lastPageNumber="230" pageId="28" pageNumber="229">
Six specimens of an interesting species of
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<emphasis id="8CF04536DA263E400821FA1FFC9DFA36" box="[631,787,1438,1463]" italics="true" pageId="28" pageNumber="229">Chiloscyllium</emphasis>
Müller &amp; Henle
</taxonomicName>
were examined from the Munai (SL-31) and Kottadi (SL-32, SL-59, SL-60) markets at Point Pedro, and Vankalai (SL-65, SL-70) in the Northern Province. Immature specimens of this species exhibited brown bands and saddles that persisted, but were less conspicuous, in adults. Morphologically, among its congeners, these specimens most closely resemble
<taxonomicName id="7984E2A7DA263E400EDCF982FA1BF99A" authorityName="Gubanov" authorityYear="1980" box="[1162,1429,1538,1563]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="arabicum">
<emphasis id="8CF04536DA263E400EDCF982FA1BF99A" box="[1162,1429,1538,1563]" italics="true" pageId="28" pageNumber="229">Chiloscyllium arabicum</emphasis>
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, particularly in fin shape and position. Our molecular analysis included six of the eight described and both undescribed members of the genus (i.e.,
<taxonomicName id="7984E2A7DA263E40080BF9C4FD77F9DF" authorityName="Muller &amp; Henle" authorityYear="1837" box="[605,761,1605,1630]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA263E40080BF9C4FD77F9DF" box="[605,761,1605,1630]" italics="true" pageId="28" pageNumber="229">Chiloscyllium</emphasis>
</taxonomicName>
cf.
<emphasis id="8CF04536DA263E400972F9C7FC14F9DC" box="[804,922,1606,1629]" italics="true" pageId="28" pageNumber="229">punctatum</emphasis>
1 of
<bibRefCitation id="DA15E4D5DA263E400983F9C4FB4FF9DF" author="Naylor" box="[981,1217,1605,1630]" etAl="et al." firstAuthor="Naylor" pageId="28" pageNumber="229" pagination="1 - 262" refId="ref19577" refString="Naylor, G. J. P., Caira, J. N., Jensen, K., Rosana, K. A. M., White, W. T. &amp; Last, P. R. (2012 a) A DNA sequence-based approach to the identification of shark and ray species and its implications for global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History, 367, 1 - 262." type="journal article" year="2012" yearSuffix="a">
Naylor
<emphasis id="8CF04536DA263E400E7CF9C7FBEDF9DF" box="[1066,1123,1605,1630]" italics="true" pageId="28" pageNumber="229">et al.</emphasis>
[2012a]
</bibRefCitation>
from
<collectingCountry id="C693D9B4DA263E400F50F9C4FAE2F9DF" box="[1286,1388,1605,1630]" name="Australia" pageId="28" pageNumber="229">Australia</collectingCountry>
and
<taxonomicName id="7984E2A7DA263E400AC1F9E7FEBDF9FE" authorityName="Muller &amp; Henle" authorityYear="1837" box="[151,307,1638,1663]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA263E400AC1F9E7FEBDF9FE" box="[151,307,1638,1663]" italics="true" pageId="28" pageNumber="229">Chiloscyllium</emphasis>
</taxonomicName>
cf.
<emphasis id="8CF04536DA263E400B36F9E6FE58F9FF" box="[352,470,1639,1662]" italics="true" pageId="28" pageNumber="229">punctatum</emphasis>
2 from
<collectingCountry id="C693D9B4DA263E400862F9E7FD08F9FE" box="[564,646,1638,1663]" name="Taiwan" pageId="28" pageNumber="229">Taiwan</collectingCountry>
of Straub
<emphasis id="8CF04536DA263E4008A9F9E6FCB7F9FE" box="[767,825,1638,1663]" italics="true" pageId="28" pageNumber="229">et al.</emphasis>
[2013]). Because of the resemblance to
<taxonomicName id="7984E2A7DA263E400F51F9E6FA1BF9FE" authorityName="Gubanov" authorityYear="1980" box="[1287,1429,1638,1663]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="arabicum">
<emphasis id="8CF04536DA263E400F51F9E6FA1BF9FE" box="[1287,1429,1638,1663]" italics="true" pageId="28" pageNumber="229">C. arabicum</emphasis>
</taxonomicName>
, we included newly generated NADH2 data for a reference specimen (MM-903;
<accessionNumber id="A1D704C7DA263E400E7DF906FB20F921" box="[1067,1198,1671,1696]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK335259" pageId="28" pageNumber="229">MK335259</accessionNumber>
) of the latter species from the Gulf of
<collectingCountry id="C693D9B4DA263E400B09F92BFE29F943" box="[351,423,1706,1730]" name="Oman" pageId="28" pageNumber="229">Oman</collectingCountry>
. In the tree resulting from the Neighbor-Joining analysis, the specimens from
<collectingCountry id="C693D9B4DA263E400F7AF928FA12F943" box="[1324,1436,1705,1730]" name="Sri Lanka" pageId="28" pageNumber="229">Sri Lanka</collectingCountry>
formed a distinct cluster, differing from one another by 14 bp. They grouped most closely with our reference specimen of
<taxonomicName id="7984E2A7DA263E400B70F96DFDF5F884" authority="Bleeker" authorityName="Bleeker" authorityYear="1852" box="[294,635,1772,1797]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="hasselti">
<emphasis id="8CF04536DA263E400B70F96DFD93F884" box="[294,541,1772,1797]" italics="true" pageId="28" pageNumber="229">Chiloscyllium hasselti</emphasis>
Bleeker
</taxonomicName>
(KA-164;
<accessionNumber id="A1D704C7DA263E4008A0F96DFCE2F884" box="[758,876,1772,1797]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ519066" pageId="28" pageNumber="229">JQ519066</accessionNumber>
) from Indonesian Borneo. However, they differed from this specimen by 3743 bp. They were even more divergent from
<taxonomicName id="7984E2A7DA263E400985F88FFBECF8A7" authorityName="Gubanov" authorityYear="1980" box="[979,1122,1805,1830]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="arabicum">
<emphasis id="8CF04536DA263E400985F88FFBECF8A7" box="[979,1122,1805,1830]" italics="true" pageId="28" pageNumber="229">C. arabicum</emphasis>
</taxonomicName>
and
<taxonomicName id="7984E2A7DA263E400ECBF88CFEC1F8C6" authority="Muller &amp; Henle" authorityName="Muller &amp; Henle" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="griseum">
<emphasis id="8CF04536DA263E400ECBF88CFA12F8A4" box="[1181,1436,1805,1830]" italics="true" pageId="28" pageNumber="229">Chiloscyllium griseum</emphasis>
Müller &amp; Henle
</taxonomicName>
, differing from the reference specimen of the former species by 109124 bp and from that of the latter species by 100115 bp. With respect to the two known species not included in our molecular analysis, the specimens from
<collectingCountry id="C693D9B4DA263E400B0BF8F0FE5FF80B" box="[349,465,1905,1930]" name="Sri Lanka" pageId="28" pageNumber="229">Sri Lanka</collectingCountry>
conspicuously differ from both. They lack the blue-white spots characteristic of
<taxonomicName id="7984E2A7DA263E400AC1F813FD1BF82A" authority="Pellegrin" authorityName="Pellegrin" authorityYear="1914" box="[151,661,1938,1963]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="caeruleopunctatum">
<emphasis id="8CF04536DA263E400AC1F813FD93F82A" box="[151,541,1938,1963]" italics="true" pageId="28" pageNumber="229">Chiloscyllium caeruleopunctatum</emphasis>
Pellegrin
</taxonomicName>
(see
<bibRefCitation id="DA15E4D5DA263E4008BFF812FC21F82A" author="Compagno" box="[745,943,1939,1963]" firstAuthor="Compagno" pageId="28" pageNumber="229" refId="ref17157" refString="Compagno, L. J. V. (1984) FAO Species Catalogue. Fol. 4. Sharks of the World. An Annotated and Illustrated Catalogue of Shark Species Known to Date. Part 1. Hexanchiformes to Lanniformes. FAO Fisheries Synopsis. No. 125, FAO, Rome, Italy, 249 pp." type="book" year="1984">Compagno 1984</bibRefCitation>
) and the diagnostic dark fin webs of
<taxonomicName id="7984E2A7DA263E400AC1F835FE20F84C" authorityName="Dingerkus &amp; DeFino" authorityYear="1983" box="[151,430,1972,1997]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="burmensis">
<emphasis id="8CF04536DA263E400AC1F835FE20F84C" box="[151,430,1972,1997]" italics="true" pageId="28" pageNumber="229">Chiloscyllium burmensis</emphasis>
</taxonomicName>
(see Dinkerkus &amp;
<bibRefCitation id="DA15E4D5DA263E4008D4F835FC97F84D" author="Dingerkus, G. &amp; DeFino, T. C." box="[642,793,1972,1997]" pageId="28" pageNumber="229" pagination="1 - 93" refId="ref17802" refString="Dingerkus, G. &amp; DeFino, T. C. (1983) A revision of the orectolobiform shark family Hemiscyllidae (Chondrichthyes, Selachii). Bulletin of the American Museum of Natural History, 176, 1 - 93." type="journal article" year="1983">DeFino 1983</bibRefCitation>
). It could be argued that these specimens raise a question regarding the validity of
<taxonomicName id="7984E2A7DA263E400BE6F854FCB5F86F" authority="Dingerkus" authorityName="Dingerkus" box="[432,827,2005,2030]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="28" pageNumber="229" phylum="Chordata" rank="species" species="confusum">
<emphasis id="8CF04536DA263E400BE6F854FD30F86F" box="[432,702,2005,2030]" italics="true" pageId="28" pageNumber="229">Chiloscyllium confusum</emphasis>
Dingerkus
</taxonomicName>
&amp; DeFino—a species originally described from
<collectingCountry id="C693D9B4DA263E400F34F854FA12F86F" box="[1378,1436,2005,2030]" name="India" pageId="28" pageNumber="229">India</collectingCountry>
by
<bibRefCitation id="DA15E4D5DA273E410AEAFF16FD8FFF31" author="Dingerkus &amp; DeFino" box="[188,513,151,176]" firstAuthor="Dingerkus" pageId="29" pageNumber="230" pagination="1 - 93" refId="ref17802" refString="Dingerkus, G. &amp; DeFino, T. C. (1983) A revision of the orectolobiform shark family Hemiscyllidae (Chondrichthyes, Selachii). Bulletin of the American Museum of Natural History, 176, 1 - 93." type="journal article" year="1983">Dingerkus &amp; DeFino (1983)</bibRefCitation>
, but synonymized with
<taxonomicName id="7984E2A7DA273E410943FF19FC2DFF31" authorityName="Gubanov" authorityYear="1980" box="[789,931,151,176]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="species" species="arabicum">
<emphasis id="8CF04536DA273E410943FF19FC2DFF31" box="[789,931,151,176]" italics="true" pageId="29" pageNumber="230">C. arabicum</emphasis>
</taxonomicName>
(see
<bibRefCitation id="DA15E4D5DA273E4109B4FF16FBF0FF31" author="Randall" box="[994,1150,151,176]" firstAuthor="Randall" pageId="29" pageNumber="230" refId="ref19790" refString="Randall, J. E. (1995) Coastal Fishes of Oman. University of Hawai'i Press, Honolulu, Hawaii, 432 pp." type="book" year="1995">Randall 1995</bibRefCitation>
). However,
<bibRefCitation id="DA15E4D5DA273E410F5FFF16FE1FFF50" author="Dingerkus, G. &amp; DeFino, T. C." pageId="29" pageNumber="230" pagination="1 - 93" refId="ref17802" refString="Dingerkus, G. &amp; DeFino, T. C. (1983) A revision of the orectolobiform shark family Hemiscyllidae (Chondrichthyes, Selachii). Bulletin of the American Museum of Natural History, 176, 1 - 93." type="journal article" year="1983">Dingerkus &amp; DeFino (1983; pg. 11)</bibRefCitation>
differentiated
<taxonomicName id="7984E2A7DA273E410860FF3BFD4DFF50" box="[566,707,185,209]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="species" species="confusum">
<emphasis id="8CF04536DA273E410860FF3BFD4DFF50" box="[566,707,185,209]" italics="true" pageId="29" pageNumber="230">C. confusum</emphasis>
</taxonomicName>
from all of its congeners, including
<taxonomicName id="7984E2A7DA273E410E0BFF3BFB67FF53" authorityName="Gubanov" authorityYear="1980" box="[1117,1257,185,210]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="species" species="arabicum">
<emphasis id="8CF04536DA273E410E0BFF3BFB67FF53" box="[1117,1257,185,210]" italics="true" pageId="29" pageNumber="230">C. arabicum</emphasis>
</taxonomicName>
, on the basis of the fact that &quot;This is the only species of
<taxonomicName id="7984E2A7DA273E410808FF5BFD74FF72" authorityName="Muller &amp; Henle" authorityYear="1837" box="[606,762,218,243]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA273E410808FF5BFD74FF72" box="[606,762,218,243]" italics="true" pageId="29" pageNumber="230">Chiloscyllium</emphasis>
</taxonomicName>
in which the immature body color is uniform, and identical with that of the adult&quot;, yet all of our specimens (immatures and adults) exhibit a clear pattern of brown bands and saddles. Furthermore, unlike
<taxonomicName id="7984E2A7DA273E410BB2FE9FFDFCFEB7" box="[484,626,286,310]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="species" species="confusum">
<emphasis id="8CF04536DA273E410BB2FE9FFDFCFEB7" box="[484,626,286,310]" italics="true" pageId="29" pageNumber="230">C. confusum</emphasis>
</taxonomicName>
our Sri Lankan specimens exhibit a first dorsal fin base that is longer, rather than shorter, than the second dorsal fin base. It thus seems apparent that our specimens represent an undescribed species. We have provisionally referred to them here as
<taxonomicName id="7984E2A7DA273E4109C1FEDEFBBDFEF9" authorityName="Muller &amp; Henle" authorityYear="1837" box="[919,1075,351,376]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA273E4109C1FEDEFBBDFEF9" box="[919,1075,351,376]" italics="true" pageId="29" pageNumber="230">Chiloscyllium</emphasis>
</taxonomicName>
sp. 1. An immature male of this species was deposited in the BRT Ichthyology Collection (SL-32; BRT-I 0011).
</paragraph>
<caption id="EAFBC9ACDA273E410AC1F88FFAEEF81E" httpUri="https://zenodo.org/record/2637270/files/figure.png" pageId="29" pageNumber="230" startId="29.[151,250,1806,1828]" targetBox="[151,1436,472,1775]" targetPageId="29">
<paragraph id="BE3B9924DA273E410AC1F88FFAEEF81E" blockId="29.[151,1436,1806,1951]" pageId="29" pageNumber="230">
<emphasis id="8CF04536DA273E410AC1F88FFEAFF8A5" bold="true" box="[151,289,1806,1828]" pageId="29" pageNumber="230">FIGURE 17.</emphasis>
<taxonomicName id="7984E2A7DA273E410B7EF88FFE3EF8A5" authorityName="De Blainville" authorityYear="1816" box="[296,432,1806,1828]" class="Chondrichthyes" family="Echinorhinidae" genus="Echinorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squaliformes" pageId="29" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA273E410B7EF88FFE3EF8A5" box="[296,432,1806,1828]" italics="true" pageId="29" pageNumber="230">Echinorhinus</emphasis>
</taxonomicName>
sp. 1 (
<taxonomicName id="7984E2A7DA273E410BAEF88FFD18F8A5" authorityName="Gill" authorityYear="1862" box="[504,662,1806,1828]" class="Chondrichthyes" family="Echinorhinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squaliformes" pageId="29" pageNumber="230" phylum="Chordata" rank="family">Echinorhinidae</taxonomicName>
) (
<emphasis id="8CF04536DA273E4108FAF88FFD50F8A5" bold="true" box="[684,734,1806,1828]" pageId="29" pageNumber="230">AD</emphasis>
),
<taxonomicName id="7984E2A7DA273E4108A2F88FFBDCF8A5" baseAuthorityName="Bonnaterre" baseAuthorityYear="1788" box="[756,1106,1806,1829]" class="Chondrichthyes" family="Hexanchidae" genus="Hexanchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hexanchiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="species" species="griseus">
<emphasis id="8CF04536DA273E4108A2F88FFC3BF8A4" box="[756,949,1806,1829]" italics="true" pageId="29" pageNumber="230">Hexanchus griseus</emphasis>
(Hexanchidae)
</taxonomicName>
(
<emphasis id="8CF04536DA273E410E34F88EFB1AF8A5" bold="true" box="[1122,1172,1807,1828]" pageId="29" pageNumber="230">EH</emphasis>
), and
<taxonomicName id="7984E2A7DA273E410E80F88FFAECF8A5" authorityName="Muller &amp; Henle" authorityYear="1837" box="[1238,1378,1806,1828]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CF04536DA273E410E80F88FFAECF8A5" box="[1238,1378,1806,1828]" italics="true" pageId="29" pageNumber="230">Chiloscyllium</emphasis>
</taxonomicName>
sp. 1 (
<taxonomicName id="7984E2A7DA273E410ACBF8ACFEB4F8C2" authorityName="Gill" authorityYear="1862" box="[157,314,1837,1859]" class="Chondrichthyes" family="Hemiscylliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="29" pageNumber="230" phylum="Chordata" rank="family">Hemiscylliidae</taxonomicName>
) (
<emphasis id="8CF04536DA273E410B19F8AFFEF7F8C2" bold="true" box="[335,377,1838,1859]" pageId="29" pageNumber="230">IK</emphasis>
).
<emphasis id="8CF04536DA273E410BD8F8ACFE28F8C3" bold="true" box="[398,422,1837,1858]" pageId="29" pageNumber="230">A.</emphasis>
Lateral view of female (SL-108).
<emphasis id="8CF04536DA273E4108A9F8AFFC98F8C2" bold="true" box="[767,790,1838,1859]" pageId="29" pageNumber="230">B.</emphasis>
Ventral view of head of female (SL-108).
<emphasis id="8CF04536DA273E410E97F8ACFB57F8C2" bold="true" box="[1217,1241,1837,1859]" pageId="29" pageNumber="230">C.</emphasis>
Teeth of upper jaw of female (SL-108).
<emphasis id="8CF04536DA273E410B31F8CDFE0EF8E0" bold="true" box="[359,384,1868,1889]" pageId="29" pageNumber="230">D.</emphasis>
Teeth of lower jaw of female (SL-108).
<emphasis id="8CF04536DA273E41094BF8CDFCBAF8E0" bold="true" box="[797,820,1868,1889]" pageId="29" pageNumber="230">E.</emphasis>
Lateral view of immature male (SL-106).
<emphasis id="8CF04536DA273E410EB3F8CDFB78F8E0" bold="true" box="[1253,1270,1868,1889]" pageId="29" pageNumber="230">F.</emphasis>
Ventral view of head of female (SL-91).
<emphasis id="8CF04536DA273E410BC5F8EAFE29F800" bold="true" box="[403,423,1899,1921]" pageId="29" pageNumber="230">G.</emphasis>
Teeth of upper jaw of female (SL-91).
<emphasis id="8CF04536DA273E41096AF8EAFCD9F801" bold="true" box="[828,855,1899,1920]" pageId="29" pageNumber="230">H.</emphasis>
Teeth of lower jaw of female (SL-91).
<emphasis id="8CF04536DA273E410EBFF8EAFB77F801" bold="true" box="[1257,1273,1899,1920]" pageId="29" pageNumber="230">I.</emphasis>
Lateral view of mature male.
<emphasis id="8CF04536DA273E410B77F80BFEBAF81E" bold="true" box="[289,308,1930,1951]" pageId="29" pageNumber="230">J.</emphasis>
Ventral view of head of mature male (SL-59).
<emphasis id="8CF04536DA273E410959F80BFCA4F81E" bold="true" box="[783,810,1930,1951]" pageId="29" pageNumber="230">K.</emphasis>
Dorsal view of head and trunk of mature male (SL-59).
</paragraph>
</caption>
<paragraph id="BE3B9924DA243E420A91FF19FD2CFF79" blockId="30.[151,1436,151,248]" pageId="30" pageNumber="231">
It seems likely that this is the species that has been previously reported as
<taxonomicName id="7984E2A7DA243E420E51FF19FB0FFF31" box="[1031,1153,152,176]" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="30" pageNumber="231" phylum="Chordata" rank="species" species="griseum">
<emphasis id="8CF04536DA243E420E51FF19FB94FF2E" box="[1031,1050,152,175]" italics="true" pageId="30" pageNumber="231">C</emphasis>
.
<emphasis id="8CF04536DA243E420E71FF18FB0FFF31" box="[1063,1153,153,176]" italics="true" pageId="30" pageNumber="231">griseum</emphasis>
</taxonomicName>
from
<collectingCountry id="C693D9B4DA243E420E90FF16FABAFF31" box="[1222,1332,151,176]" name="Sri Lanka" pageId="30" pageNumber="231">Sri Lanka</collectingCountry>
(e.g.,
<bibRefCitation id="DA15E4D5DA243E420F2DFF19FE9DFF54" author="De Silva" firstAuthor="De Silva" pageId="30" pageNumber="231" pagination="56 - 66" refId="ref17520" refString="De Silva, R. I. (1984 - 1985) The sharks of Sri Lanka. A key to the different species and a preliminary checklist. Ceylon Journal of Science (Biological Sciences), 17 - 18, 56 - 66." type="book chapter" year="1984">
De Silva
<date id="CA3ABFE4DA243E420A8CFF3DFE9DFF54" box="[218,275,188,213]" pageId="30" pageNumber="231" value="1984" valueMax="1985">1984</date>
</bibRefCitation>
1985, 2006, 2015;
<bibRefCitation id="DA15E4D5DA243E420BACFF3CFD62FF54" author="De Bruin" box="[506,748,188,213]" etAl="et al." firstAuthor="De Bruin" pageId="30" pageNumber="231" refId="ref17417" refString="De Bruin, G. H. P., Russell, B. C. &amp; Bogusch, A. (1995) FAO Species Identification Field Guide for Fisheries Purposes. The Marine Fishery Resources of Sri Lanka. FAO, Rome, Italy, 400 pp." type="book" year="1995">
De Bruin
<emphasis id="8CF04536DA243E42083BFF3CFD26FF54" box="[621,680,188,213]" italics="true" pageId="30" pageNumber="231">et al.</emphasis>
1995
</bibRefCitation>
;
<bibRefCitation id="DA15E4D5DA243E4208ADFF3DFC41FF55" author="Moron" box="[763,975,188,213]" etAl="et al." firstAuthor="Moron" pageId="30" pageNumber="231" pagination="142 - 157" refId="ref19181" refString="Moron, J., Bertrand, B. &amp; Last, P. (1998) A check-list of sharks and rays of western Sri Lanka. Journal of the Marine Biological Association of India, 40, 142 - 157." type="journal article" year="1998">
Morón
<emphasis id="8CF04536DA243E420906FF3CFC04FF54" box="[848,906,188,213]" italics="true" pageId="30" pageNumber="231">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="DA15E4D5DA243E42098BFF3DFBE1FF55" author="Joseph" box="[989,1135,188,213]" firstAuthor="Joseph" pageId="30" pageNumber="231" pagination="339 - 366" refId="ref18550" refString="Joseph, L. (1999) Management of shark fisheries in Sri Lanka. In: Shotton, R. (Ed.), Case Studies of the Management of Elasmobranch Fisheries. FAO Fisheries Technical Paper. No. 3780, Rome, Italy, pp. 339 - 366." type="book chapter" year="1999">Joseph 1999</bibRefCitation>
). If so, the presence of
<taxonomicName id="7984E2A7DA243E420FD4FF3CFF7FFF79" class="Chondrichthyes" family="Hemiscylliidae" genus="Chiloscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="30" pageNumber="231" phylum="Chordata" rank="species" species="griseum">
<emphasis id="8CF04536DA243E420FD4FF3CFF7FFF79" italics="true" pageId="30" pageNumber="231">C. griseum</emphasis>
</taxonomicName>
in
<collectingCountry id="C693D9B4DA243E420B43FF5EFE0CFF79" box="[277,386,223,248]" name="Sri Lanka" pageId="30" pageNumber="231">Sri Lanka</collectingCountry>
remains to be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>