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<document id="1112ACF4BA8293C54660F05F041391A1" ID-CLB-Dataset="30115" ID-DOI="10.11646/zootaxa.4418.3.6" ID-GBIF-Dataset="9ebe3063-1237-493c-90b6-804d0ceb1ad9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1244892" ID-ZooBank="7A2FDE23-93A2-4C62-8072-FA79A8204D1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525974261415" checkinUser="plazi" docAuthor="Koeda, Keita, Fujii, Takuma &amp; Motomura, Hiroyuki" docDate="2018" docId="122D8798FFFE4965FF2861354C5FFD0F" docLanguage="en" docName="zootaxa.4418.3.5.pdf" docOrigin="Zootaxa 4418 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteroconger fugax Koeda &amp; Fujii &amp; Motomura 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="294" masterDocId="EE14FFE0FFFF4962FFBF62564D2EFFC2" masterDocTitle="A new garden eel, Heteroconger fugax (Congridae: Heterocongrinae), from the northwestern Pacific Ocean" masterLastPageNumber="295" masterPageNumber="287" pageNumber="288" updateTime="1698543612679" updateUser="ExternalLinkService">
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<mods:title id="9C00B8B4F575D7789740432D928152F2">A new garden eel, Heteroconger fugax (Congridae: Heterocongrinae), from the northwestern Pacific Ocean</mods:title>
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<mods:namePart id="D5615053380F04B8C67F75D822E6379B">Koeda, Keita</mods:namePart>
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<mods:namePart id="BD26BF10320D6C59449659FAF19AD504">Fujii, Takuma</mods:namePart>
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<mods:namePart id="3836722B92B080C3632E5BB3D5AD0F5A">Motomura, Hiroyuki</mods:namePart>
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<mods:date id="6FFAFD65E3D1F31C767C4ABCD043D4BE">2018</mods:date>
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<emphasis id="A8F0EA9CFFFE4963FF2861354CC1FCBC" bold="true" box="[151,495,867,894]" pageId="1" pageNumber="288">
<taxonomicName id="5D844D0DFFFE4963FF2861354CA4FCBF" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[151,394,867,893]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="1" pageNumber="288" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFE4963FF2861354CA4FCBF" bold="true" box="[151,394,867,893]" italics="true" pageId="1" pageNumber="288">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFE4963FE2E61324CC1FCBC" box="[401,495,868,894]" pageId="1" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="9A3B368EFFFE4963FF2861D14C51FC25" blockId="1.[151,735,867,999]" pageId="1" pageNumber="288">
New standard Japanese name: Nigemizu-chin'anago New English name: Shy Garden Eel
<figureCitation id="02BF2A0BFFFE4963FF2861984DD0FC25" box="[151,254,974,999]" captionStart="FIGURE 1" captionStartId="2.[151,250,1029,1051]" captionTargetBox="[152,1435,193,1008]" captionTargetId="figure@2.[152,1435,193,1008]" captionTargetPageId="2" captionText="FIGURE 1. Fresh holotype of Heteroconger fugax sp. nov. from Amami-oshima island, Japan. KAUMI. 95673, 729.0 mm TL.Red and green arrows indiCate vertiCal positions of tip of appressed peCtoral fin and origin of dorsal fin, respeCtively." httpUri="https://zenodo.org/record/1244894/files/figure.png" pageId="1" pageNumber="288">Figures 1</figureCitation>
¯4;
<tableCitation id="D7060335FFFE4963FE9361984C51FC25" box="[300,383,974,999]" captionStart="TABLE 1" captionStartId="5.[151,239,296,319]" captionText="TABLE 1. Counts and measurements of type specimens of Heteroconger fugax sp. nov. and H. tomberua. Data for paratypes of H. tomberua extracted from Castle &amp; Randall (1999)." pageId="1" pageNumber="288">Table 1</tableCitation>
</paragraph>
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<paragraph id="9A3B368EFFFE4963FF2866414CDBFB89" blockId="1.[151,1436,1046,1099]" pageId="1" pageNumber="288">
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<taxonomicName id="5D844D0DFFFE4963FF2866414CB2FBEE" box="[151,412,1047,1068]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="1" pageNumber="288" phylum="Chordata" rank="species" species="lentignosus">
<emphasis id="A8F0EA9CFFFE4963FF2866414CB2FBEE" box="[151,412,1047,1068]" italics="true" pageId="1" pageNumber="288">Heteroconger lentignosus</emphasis>
</taxonomicName>
(not
<bibRefCitation id="FE154B7FFFFE4963FE6E66404FE8FBEE" box="[465,710,1046,1068]" pageId="1" pageNumber="288" refId="ref3371" refString="Bohlke, J. E. &amp; Randall, J. E. (1981) Four new garden eels (Congridae, HeteroCongrinae) from the PaCifiC and Indian oCeans. Bulletin of Marine Science, 31, 366 - 382." type="journal article">Böhlke &amp; Randall 1981</bibRefCitation>
):
<treatmentCitation id="1B25109FFFFE4963FD6866404E85FBEE" author="" box="[727,939,1046,1068]" page="48" pageId="1" pageNumber="288">
<bibRefCitation id="FE154B7FFFFE4963FD6866404EA2FBEE" box="[727,908,1046,1068]" pageId="1" pageNumber="288" refId="ref3636" refString="Senou, H., Kobayashi, Y. &amp; Kobayashi, N. (2007) Coastal fishes of the Miyako Group, the Ryukyu Islands, Japan. Bulletin of Kanagawa Prefectural Museum (Natural Science), 36, 47 - 74." type="journal article">
Senou
<emphasis id="A8F0EA9CFFFE4963FCA266414E67FBEE" box="[797,841,1047,1068]" italics="true" pageId="1" pageNumber="288">et al</emphasis>
. 2007
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:48
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, fig. 2 (Irabu-jima island, Miyako Islands,
<collectingCountry id="E293761EFFFE4963FAE3664148B6FBEE" box="[1372,1432,1047,1068]" name="Japan" pageId="1" pageNumber="288">Japan</collectingCountry>
, underwater photograph only).
</treatmentCitationGroup>
</paragraph>
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<paragraph id="9A3B368EFFFE4963FF28662148A2FB77" blockId="1.[151,1437,1143,1997]" pageId="1" pageNumber="288">
<materialsCitation id="2AEC3CD3FFFE4963FF28662148A2FB77" ID-GBIF-Occurrence="1844406408" collectingDate="2016-11-11" collectionCode="KAUM" collectorName="T. Fujii &amp; K. Koeda." country="Japan" county="Amami Islands" elevation="33" location="Shirahama" municipality="Amami-oshima island" pageId="1" pageNumber="288" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<emphasis id="A8F0EA9CFFFE4963FF2866214C25FB52" bold="true" box="[151,267,1143,1168]" pageId="1" pageNumber="288">
<typeStatus id="453F882CFFFE4963FF2866214C28FB52" box="[151,262,1143,1168]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="FC95AE4BFFFE4963FEA8662E4C5EFB52" box="[279,368,1144,1168]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="288">KAUM</collectionCode>
I. 95673, 729.0 mm TL, off
<location id="9F5B6055FFFE4963FD6A66214E7EFB52" LSID="urn:lsid:plazi:treatment:122D8798FFFE4965FF2861354C5FFD0F:9F5B6055FFFE4963FD6A66214E7EFB52" box="[725,848,1143,1168]" country="Japan" county="Amami Islands" municipality="Amami-oshima island" name="Shirahama" pageId="1" pageNumber="288" stateProvince="Kagoshima">Shirahama</location>
,
<location id="9F5B6055FFFE4963FCDF66214EEBFB52" LSID="urn:lsid:plazi:treatment:122D8798FFFE4965FF2861354C5FFD0F:9F5B6055FFFE4963FCDF66214EEBFB52" box="[864,965,1143,1168]" country="Japan" county="Amami Islands" municipality="Amami-oshima island" name="Setouchi" pageId="1" pageNumber="288" stateProvince="Kagoshima">Setouchi</location>
,
<collectingMunicipality id="7A5FACF4FFFE4963FC6A662149F8FB52" box="[981,1238,1143,1168]" pageId="1" pageNumber="288">Amami-oshima island</collectingMunicipality>
,
<collectingCounty id="735A4E02FFFE4963FB59662148B6FB52" box="[1254,1432,1143,1168]" pageId="1" pageNumber="288">Amami Islands</collectingCounty>
,
<collectingRegion id="5840F86CFFFE4963FF2866CA4C34FB77" box="[151,282,1180,1205]" country="Japan" name="Kagoshima" pageId="1" pageNumber="288">Kagoshima</collectingRegion>
,
<collectingCountry id="E293761EFFFE4963FE9B66CB4C4AFB76" box="[292,356,1181,1204]" name="Japan" pageId="1" pageNumber="288">Japan</collectingCountry>
(28°12ʹN, 129°16ʹE),
<elevation id="11A9D1BDFFFE4963FDDF66CA4F80FB76" box="[608,686,1180,1205]" metricMagnitude="1" metricUnit="m" metricValue="3.25" pageId="1" pageNumber="288" unit="m" value="32.5">
<quantity id="5D7C9B6BFFFE4963FDDF66CA4F80FB76" box="[608,686,1180,1205]" metricMagnitude="1" metricUnit="m" metricValue="3.25" pageId="1" pageNumber="288" unit="m" value="32.5">32.5 m</quantity>
</elevation>
depth,
<collectingDate id="FE7EE9A6FFFE4963FCBE66CA4EF5FB77" box="[769,987,1180,1205]" pageId="1" pageNumber="288" value="2016-11-11">
<date id="EE3A104EFFFE4963FCBE66CA4EF5FB77" box="[769,987,1180,1205]" pageId="1" pageNumber="288" value="2016-11-11">11 November 2016</date>
</collectingDate>
, coll. by hands,
<collectorName id="37715358FFFE4963FB2F66CB49CCFB77" box="[1168,1250,1180,1205]" pageId="1" pageNumber="288">T. Fujii</collectorName>
and
<collectorName id="37715358FFFE4963FAA666CB48A2FB77" box="[1305,1420,1180,1205]" pageId="1" pageNumber="288">K. Koeda.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D29E6505FFFE4963FF7866E9487CFA4F" pageId="1" pageNumber="288" type="diagnosis">
<paragraph id="9A3B368EFFFE4963FF7866E9487CFA4F" blockId="1.[151,1437,1143,1997]" pageId="1" pageNumber="288">
<emphasis id="A8F0EA9CFFFE4963FF7866E94C6FFB1A" bold="true" box="[199,321,1215,1240]" pageId="1" pageNumber="288">Diagnosis.</emphasis>
A species of
<taxonomicName id="5D844D0DFFFE4963FE6266974F59FB1A" authorityName="Bleeker" authorityYear="1868" box="[477,631,1217,1240]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="1" pageNumber="288" phylum="Chordata" rank="genus">
<emphasis id="A8F0EA9CFFFE4963FE6266974F59FB1A" box="[477,631,1217,1240]" italics="true" pageId="1" pageNumber="288">Heteroconger</emphasis>
</taxonomicName>
with the following combination of characters: body extremely slender, depth 1.2% TL and 1.0% TL at gill opening and anus, respectively; pre-dorsal-fin and pre-anal-fin vertebrae 8 and 67, respectively, total vertebrae 197; origin of dorsal fin slightly posterior to tip of appressed pectoral fin; numerus small conical cirri on chin; 23 rows of dark spots above and below lateral line; 67 rows of dark spots on interorbital space; large distinct white rectangular blotch on opercle; number and density of spots on head greater than those on body; lips spotted; ground color of body uniformly cream, without microscopic dark speckles.
</paragraph>
</subSubSection>
<subSubSection id="D29E6505FFFE4961FF7867C14ECFF851" lastPageId="3" lastPageNumber="290" pageId="1" pageNumber="288" type="description">
<paragraph id="9A3B368EFFFE4963FF7867C14E50F9DE" blockId="1.[151,1437,1143,1997]" pageId="1" pageNumber="288">
<emphasis id="A8F0EA9CFFFE4963FF7867C14C77FA72" bold="true" box="[199,345,1431,1456]" pageId="1" pageNumber="288">Description.</emphasis>
Pre-dorsal-fin vertebrae 8; Pre-anal-fin vertebrae 67; total vertebrae 197; lateral-line pores before pectoral fin 7; lateral-line pores before anus 65; pectoral-fin rays 9; dorsal-fin rays before anus 182; total dorsal-fin rays 590; anal-fin rays 425; caudal-fin rays 9 + 8. Head pores (left side): supraorbital series 1 + 4; infraorbital series 3 + 3; preoperculomandibular series 5 + 3; supratemporal series 3.
</paragraph>
<paragraph id="9A3B368EFFFE4963FF78647E49E3F96E" blockId="1.[151,1437,1143,1997]" pageId="1" pageNumber="288">As percent of TL: HL 3.4%; Snout-anus length 30.1%; pre-dorsal-fin length 4.0%, trunk length 26.4%; tail length 67.4%; depth at gill opening 1.2%; depth at anus 1.0%. As percent of snout-anus length: HL 11.5%; predorsal-fin length 13.7%. As percent of HL: snout length 16.8%; eye diameter 19.2%; mouth width 24.5%; gill opening 9.1%; interbranchial distance 21.4%; depth at gill opening 34.2%; depth at anus 27.9%.</paragraph>
<paragraph id="9A3B368EFFFE4963FF7864E14DE6F80F" blockId="1.[151,1437,1143,1997]" pageId="1" pageNumber="288">
Body extremely elongate, body depth similar throughout; greatest depth at gill opening. Body slightly compressed laterally along head, progressively more so to tail tip; tail much longer than body, anus at about 30% total length. Head slightly compressed ventrally; snout short, rounded. Mouth with fleshy lips; distinct fold extending to end of lips; upper lip confluent medially, enclosing anterior nostrils; posterior nostril located just in front of eye; no flap around posterior nostril; mouth strongly oblique, posterior margin of upper lip reaching below anterior rim of pupil; numerous conical cirri on chin (
<figureCitation id="02BF2A0BFFFE4963FD4C653A4E17F846" box="[755,825,1900,1925]" captionStart="FIGURE 2" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[293,1323,1157,1835]" captionTargetId="figure@2.[264,1323,1131,1857]" captionTargetPageId="2" captionText="FIGURE 2. Numerous small ConiCal Cirri on the Chin of the holotype of Heteroconger fugax sp. nov. KAUMI.95673, 729.0 mm TL." httpUri="https://zenodo.org/record/1244896/files/figure.png" pageId="1" pageNumber="288">Fig. 2</figureCitation>
). Eye large, oval, weakly intruding into dorsal profile. Throat with developed folds extending to branchial aperture. Gill opening vertically slit, located before pectoral fins.
</paragraph>
<caption id="CEFB6606FFFD4960FF286653484DFBF8" httpUri="https://zenodo.org/record/1244894/files/figure.png" pageId="2" pageNumber="289" startId="2.[151,250,1029,1051]" targetBox="[152,1435,193,1008]" targetPageId="2">
<paragraph id="9A3B368EFFFD4960FF286653484DFBF8" blockId="2.[151,1435,1029,1082]" pageId="2" pageNumber="289">
<emphasis id="A8F0EA9CFFFD4960FF2866534C20FBD8" bold="true" box="[151,270,1029,1051]" pageId="2" pageNumber="289">FIGURE 1</emphasis>
. Fresh holotype of
<taxonomicName id="5D844D0DFFFD4960FE6666504F8AFBD8" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[473,676,1029,1051]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFD4960FE6666504F8AFBD8" box="[473,676,1029,1051]" italics="true" pageId="2" pageNumber="289">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFD4960FD1366534FD0FBD8" box="[684,766,1029,1050]" pageId="2" pageNumber="289" rank="species">
<emphasis id="A8F0EA9CFFFD4960FD1366534FD0FBD8" bold="true" box="[684,766,1029,1050]" pageId="2" pageNumber="289">sp. nov.</emphasis>
</taxonomicNameLabel>
from Amami-oshima island, Japan. KAUMI. 95673, 729.0 mm TL. Red and green arrows indiCate vertiCal positions of tip of appressed peCtoral fin and origin of dorsal fin, respeCtively.
</paragraph>
</caption>
<caption id="CEFB6606FFFD4960FF2865014DC5F849" httpUri="https://zenodo.org/record/1244896/files/figure.png" pageId="2" pageNumber="289" startId="2.[151,250,1879,1901]" targetBox="[293,1323,1157,1835]" targetPageId="2">
<paragraph id="9A3B368EFFFD4960FF2865014DC5F849" blockId="2.[151,1436,1879,1931]" pageId="2" pageNumber="289">
<emphasis id="A8F0EA9CFFFD4960FF2865014C3DF8AE" bold="true" box="[151,275,1879,1901]" pageId="2" pageNumber="289">FIGURE 2.</emphasis>
Numerous small ConiCal Cirri on the Chin of the holotype of
<taxonomicName id="5D844D0DFFFD4960FCC2650E4966F8AE" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[893,1096,1879,1901]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="289" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFD4960FCC2650E4966F8AE" box="[893,1096,1879,1901]" italics="true" pageId="2" pageNumber="289">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFD4960FBF0650149B1F8AE" box="[1103,1183,1879,1900]" pageId="2" pageNumber="289" rank="species">
<emphasis id="A8F0EA9CFFFD4960FBF0650149B1F8AE" bold="true" box="[1103,1183,1879,1900]" pageId="2" pageNumber="289">sp. nov.</emphasis>
</taxonomicNameLabel>
KAUMI. 95673, 729.0 mm TL.
</paragraph>
</caption>
<paragraph id="9A3B368EFFFC4961FF7862C14938FF17" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="290">Pectoral fins a small rounded flap; origin of dorsal-fin posterior to tip of appressed pectoral fin; dorsal and anal fins very low; tip of tail hardened, inflexible; caudal-fin rays not readily visible.</paragraph>
<paragraph id="9A3B368EFFFC4961FF7862894E67FE82" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="290">
Teeth small, numerous, in broad multi-serial patches anteriorly on jaws and vomer (
<figureCitation id="02BF2A0BFFFC4961FBCC62894999FF3A" box="[1139,1207,223,248]" captionStart="FIGURE 3" captionStartId="3.[151,250,1570,1592]" captionTargetBox="[277,1266,391,1515]" captionTargetId="figure@3.[260,1288,373,1536]" captionTargetPageId="3" captionText="FIGURE 3. Dentition of upper and lower jaws of the holotype of Heteroconger fugax sp. nov. KAUMI. 95673, 729.0 mm TL." httpUri="https://zenodo.org/record/1244898/files/figure.png" pageId="3" pageNumber="290">Fig. 3</figureCitation>
); patches narrowing posteriorly to a point on vomer, or to 1 or 2 rows on jaws; posterior portion of maxillary patch flared laterally; anterior intermaxillary-vomerine teeth larger than other teeth.
</paragraph>
<caption id="CEFB6606FFFC4961FF2864744D90F995" httpUri="https://zenodo.org/record/1244898/files/figure.png" pageId="3" pageNumber="290" startId="3.[151,250,1570,1592]" targetBox="[277,1266,391,1515]" targetPageId="3">
<paragraph id="9A3B368EFFFC4961FF2864744D90F995" blockId="3.[151,1436,1570,1623]" pageId="3" pageNumber="290">
<emphasis id="A8F0EA9CFFFC4961FF2864744C20F9F5" bold="true" box="[151,270,1570,1592]" pageId="3" pageNumber="290">FIGURE 3</emphasis>
. Dentition of upper and lower jaws of the holotype of
<taxonomicName id="5D844D0DFFFC4961FCF86475493DF9FA" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[839,1043,1571,1592]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFC4961FCF86475493DF9FA" box="[839,1043,1571,1592]" italics="true" pageId="3" pageNumber="290">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFC4961FBA464754943F9FA" box="[1051,1133,1571,1592]" pageId="3" pageNumber="290" rank="species">
<emphasis id="A8F0EA9CFFFC4961FBA464754943F9FA" bold="true" box="[1051,1133,1571,1592]" pageId="3" pageNumber="290">sp. nov.</emphasis>
</taxonomicNameLabel>
KAUMI. 95673, 729.0 mm TL.
</paragraph>
</caption>
<paragraph id="9A3B368EFFFC4961FF7864294E78F889" blockId="3.[151,1436,1662,2010]" pageId="3" pageNumber="290">
<emphasis id="A8F0EA9CFFFC4961FF7864294CDFF954" box="[199,497,1662,1686]" italics="true" pageId="3" pageNumber="290">Live and fresh coloration</emphasis>
(
<figureCitation id="02BF2A0BFFFC4961FDBB64284F78F955" box="[516,598,1662,1687]" captionStart="FIGURE 1" captionStartId="2.[151,250,1029,1051]" captionTargetBox="[152,1435,193,1008]" captionTargetId="figure@2.[152,1435,193,1008]" captionTargetPageId="2" captionText="FIGURE 1. Fresh holotype of Heteroconger fugax sp. nov. from Amami-oshima island, Japan. KAUMI. 95673, 729.0 mm TL.Red and green arrows indiCate vertiCal positions of tip of appressed peCtoral fin and origin of dorsal fin, respeCtively." httpUri="https://zenodo.org/record/1244894/files/figure.png" pageId="3" pageNumber="290">Figs. 1</figureCitation>
,
<figureCitation id="02BF2A0BFFFC4961FDD664294F54F955" box="[617,634,1663,1687]" captionStart="FIGURE 4" captionStartId="4.[151,250,1087,1109]" captionTargetBox="[151,1436,193,1066]" captionTargetId="figure@4.[151,1436,193,1066]" captionTargetPageId="4" captionText="FIGURE 4. Underwater photographs of a Colony (A) and single individuals (BD) of Heteroconger fugax sp. nov. A: off Amami-oshima island, Japan; B: off Irabu-jima island, Miyako Islands, Japan, 1011 m depth (KPM-NR 67322); C: off Mindoro Island, Philippines, 10 m depth (KPM-NR 88296); D: off Mabul Island, Malaysia, 12 m depth (KPM-NR 23927)" httpUri="https://zenodo.org/record/1244900/files/figure.png" pageId="3" pageNumber="290">4</figureCitation>
). Body and head with small, round, discrete dark spots overlying a uniformly cream ground color, without microscopic dark speckles; dark spots in 2 or 3 irregular rows above and below lateral line; spots more numerous and crowded over entire head, smaller ventrally; ca. 20 spots surrounding eye; 67 rows of spots across interorbital area; a large distinct white rectangular blotch on opercle; three dark spots just anterior to white blotch confluent with each other to form an inverted fiddle-shaped blotch; fins semitransparent. Tip of numerous small conical cirri on chin dusky.
</paragraph>
<paragraph id="9A3B368EFFFC4961FF7865004ECFF851" blockId="3.[151,1436,1662,2010]" pageId="3" pageNumber="290">
<emphasis id="A8F0EA9CFFFC4961FF7865004CF6F8AC" box="[199,472,1878,1903]" italics="true" pageId="3" pageNumber="290">Coloration of preserved</emphasis>
. Coloration of body and head almost unchanged from live and fresh; dark spots on body and white blotch on opercle clearly distinguishable from background.
</paragraph>
</subSubSection>
<subSubSection id="D29E6505FFFC4961FF7865C84DCBF818" pageId="3" pageNumber="290" type="etymology">
<paragraph id="9A3B368EFFFC4961FF7865C84DCBF818" blockId="3.[151,1436,1662,2010]" pageId="3" pageNumber="290">
<emphasis id="A8F0EA9CFFFC4961FF7865C84C60F875" bold="true" box="[199,334,1950,1975]" pageId="3" pageNumber="290">Etymology.</emphasis>
The specific name of the new species,
<taxonomicName id="5D844D0DFFFC4961FCB265C94E65F875" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[781,843,1951,1975]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="290" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFC4961FCB265C94E65F875" box="[781,843,1951,1975]" italics="true" pageId="3" pageNumber="290">fugax</emphasis>
</taxonomicName>
, means “shy” in Greek, in reference to their bashful nature.
</paragraph>
</subSubSection>
<caption id="CEFB6606FFFB4966FF2866694840FB50" httpUri="https://zenodo.org/record/1244900/files/figure.png" pageId="4" pageNumber="291" startId="4.[151,250,1087,1109]" targetBox="[151,1436,193,1066]" targetPageId="4">
<paragraph id="9A3B368EFFFB4966FF2866694840FB50" blockId="4.[151,1436,1087,1170]" pageId="4" pageNumber="291">
<emphasis id="A8F0EA9CFFFB4966FF2866694C3BFB96" bold="true" box="[151,277,1087,1109]" pageId="4" pageNumber="291">FIGURE 4.</emphasis>
Underwater photographs of a Colony (A) and single individuals (BD) of
<taxonomicName id="5D844D0DFFFB4966FB96661649D8FB97" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[1065,1270,1088,1109]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFB4966FB96661649D8FB97" box="[1065,1270,1088,1109]" italics="true" pageId="4" pageNumber="291">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFB4966FB406616487DFB97" box="[1279,1363,1088,1109]" pageId="4" pageNumber="291" rank="species">
<emphasis id="A8F0EA9CFFFB4966FB406616487DFB97" bold="true" box="[1279,1363,1088,1109]" pageId="4" pageNumber="291">sp. nov.</emphasis>
</taxonomicNameLabel>
A: off Amami-oshima island, Japan; B: off Irabu-jima island, Miyako Islands, Japan, 1011 m depth (KPM-NR 67322); C: off Mindoro Island, Philippines, 10 m depth (KPM-NR 88296); D: off Mabul Island, Malaysia, 12 m depth (KPM-NR 23927)
</paragraph>
</caption>
<subSubSection id="D29E6505FFFB4966FF7866EC4F18F92D" pageId="4" pageNumber="291" type="distribution">
<paragraph id="9A3B368EFFFB4966FF7866EC4C1AFA69" blockId="4.[151,1436,1210,2027]" pageId="4" pageNumber="291">
<emphasis id="A8F0EA9CFFFB4966FF7866EC4C4EFB11" bold="true" box="[199,352,1210,1235]" pageId="4" pageNumber="291">Distribution.</emphasis>
<taxonomicName id="5D844D0DFFFB4966FED366ED4F7EFB10" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[364,592,1210,1234]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFB4966FED366ED4F7EFB10" box="[364,592,1210,1234]" italics="true" pageId="4" pageNumber="291">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFB4966FDE366EC4F95FB11" box="[604,699,1210,1235]" pageId="4" pageNumber="291" rank="species">
<emphasis id="A8F0EA9CFFFB4966FDE366EC4F95FB11" bold="true" box="[604,699,1210,1235]" pageId="4" pageNumber="291">sp. nov.</emphasis>
</taxonomicNameLabel>
is currently known from the northwestern Pacific Ocean. The
<typeStatus id="453F882CFFFB4966FF2866884DD7FB35" box="[151,249,1246,1271]" pageId="4" pageNumber="291" type="holotype">holotype</typeStatus>
was collected off Shirahama on the south coast of Amami-oshima island, Ryukyu Archipelago (
<collectingCountry id="E293761EFFFB4966FAF4668948BEFB35" box="[1355,1424,1247,1271]" name="Japan" pageId="4" pageNumber="291">Japan</collectingCountry>
), facing the Oshima Strait; no other specimens appear to have been collected. Underwater photographs of colonies of the species were taken at the
<typeStatus id="453F882CFFFB4966FE6267714F23FAFD" box="[477,525,1319,1343]" pageId="4" pageNumber="291">type</typeStatus>
locality in depths of 28.0
<quantity id="5D7C9B6BFFFB4966FC8467704EA6FAFC" box="[827,904,1318,1343]" metricMagnitude="1" metricUnit="m" metricValue="3.25" pageId="4" pageNumber="291" unit="m" value="32.5">32.5 m</quantity>
(
<figureCitation id="02BF2A0BFFFB4966FC2867704EF5FAFD" box="[919,987,1318,1343]" captionStart="FIGURE 4" captionStartId="4.[151,250,1087,1109]" captionTargetBox="[151,1436,193,1066]" captionTargetId="figure@4.[151,1436,193,1066]" captionTargetPageId="4" captionText="FIGURE 4. Underwater photographs of a Colony (A) and single individuals (BD) of Heteroconger fugax sp. nov. A: off Amami-oshima island, Japan; B: off Irabu-jima island, Miyako Islands, Japan, 1011 m depth (KPM-NR 67322); C: off Mindoro Island, Philippines, 10 m depth (KPM-NR 88296); D: off Mabul Island, Malaysia, 12 m depth (KPM-NR 23927)" httpUri="https://zenodo.org/record/1244900/files/figure.png" pageId="4" pageNumber="291">Fig. 4</figureCitation>
), in addition to underwater photographs of single individuals taken off Irabu-jima island, Miyako Islands (
<collectingCountry id="E293761EFFFB4966FC2D671D4EFDFAA1" box="[914,979,1355,1379]" name="Japan" pageId="4" pageNumber="291">Japan</collectingCountry>
)(KPM-NR 67322, 10
<quantity id="5D7C9B6BFFFB4966FB5F671D4834FAA0" box="[1248,1306,1355,1379]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="291" unit="m" value="11.0">11 m</quantity>
depth), off northern Mindoro Island (
<collectingCountry id="E293761EFFFB4966FE0367384F12FA45" box="[444,572,1390,1415]" name="Philippines" pageId="4" pageNumber="291">Philippines</collectingCountry>
)(KPM-NR 88296, 10 m) and Mabul Island, northeastern Borneo (KPM-NR 23927, 12 m).
</paragraph>
<paragraph id="9A3B368EFFFB4966FF7867E049E2F9D4" blockId="4.[151,1436,1210,2027]" pageId="4" pageNumber="291">
<emphasis id="A8F0EA9CFFFB4966FF7867E04F0EFA0D" bold="true" box="[199,544,1462,1487]" pageId="4" pageNumber="291">Habitat and ecological notes.</emphasis>
<taxonomicName id="5D844D0DFFFB4966FD9867E14E29FA0C" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[551,775,1462,1486]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFB4966FD9867E14E29FA0C" box="[551,775,1462,1486]" italics="true" pageId="4" pageNumber="291">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFB4966FCB167E04E47FA0D" box="[782,873,1462,1487]" pageId="4" pageNumber="291" rank="species">
<emphasis id="A8F0EA9CFFFB4966FCB167E04E47FA0D" bold="true" box="[782,873,1462,1487]" pageId="4" pageNumber="291">sp. nov.</emphasis>
</taxonomicNameLabel>
forms colonies in muddy bottoms at depths of ca.
<quantity id="5D7C9B6BFFFB4966FF28678D4DFCFA30" box="[151,210,1499,1523]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="291" unit="m" value="30.0">30 m</quantity>
off Amami-oshima island, being absent from shallower or deeper areas around the
<typeStatus id="453F882CFFFB4966FB2B678D49EAFA31" box="[1172,1220,1499,1523]" pageId="4" pageNumber="291">type</typeStatus>
locality. However, photographic records from other localities were taken over sandy bottoms at depths of
<quantity id="5D7C9B6BFFFB4966FBDD67A949E6F9D4" box="[1122,1224,1535,1559]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="4" pageNumber="291" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">
10
<specimenCount id="8C82FD07FFFB4966FB3267A949E6F9D4" box="[1165,1224,1535,1559]" pageId="4" pageNumber="291" type="generic">12 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="9A3B368EFFFB4966FF7864744F4CF941" blockId="4.[151,1436,1210,2027]" pageId="4" pageNumber="291">
Although sometimes exiting vertically more than
<quantity id="5D7C9B6BFFFB4966FC9664754E59F9F8" box="[809,887,1571,1595]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="291" unit="cm" value="30.0">30 cm</quantity>
from their burrows, individuals immediately disappeared into the burrow when approached, their head holding steadily at ca. several centimeters depth (still visible from directly above the burrows).
</paragraph>
<paragraph id="9A3B368EFFFB4966FF7864D94F18F92D" blockId="4.[151,1436,1210,2027]" pageId="4" pageNumber="291">
As a result of poor water visibility at the
<typeStatus id="453F882CFFFB4966FD2964D94FE8F965" box="[662,710,1679,1703]" pageId="4" pageNumber="291">type</typeStatus>
locality (&lt;
<quantity id="5D7C9B6BFFFB4966FCF664D94EA8F965" box="[841,902,1679,1703]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="291" unit="m" value="10.0">10 m</quantity>
), an accurate count of individuals in the colony was difficult. However, about 40 individuals were estimated for each colony, from a combination of underwater photographs and number of burrows.
</paragraph>
</subSubSection>
<subSubSection id="D29E6505FFFB4967FF7864AC4849FF3A" lastPageId="5" lastPageNumber="292" pageId="4" pageNumber="291" type="discussion">
<paragraph id="9A3B368EFFFB4967FF7864AC4849FF3A" blockId="4.[151,1436,1210,2027]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="292" pageId="4" pageNumber="291">
<emphasis id="A8F0EA9CFFFB4966FF7864AC4C15F8D1" bold="true" box="[199,315,1786,1811]" pageId="4" pageNumber="291">Remarks.</emphasis>
<taxonomicName id="5D844D0DFFFB4966FEFD64AD4F0DF8D0" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[322,547,1786,1810]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFB4966FEFD64AD4F0DF8D0" box="[322,547,1786,1810]" italics="true" pageId="4" pageNumber="291">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFB4966FD9564AC4FABF8D1" box="[554,645,1786,1811]" pageId="4" pageNumber="291" rank="species">
<emphasis id="A8F0EA9CFFFB4966FD9564AC4FABF8D1" bold="true" box="[554,645,1786,1811]" pageId="4" pageNumber="291">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles
<taxonomicName id="5D844D0DFFFB4966FCBB64AD49B3F8D1" authority="Castle &amp; Randall 1999" authorityName="Castle &amp; Randall" authorityYear="1999" box="[772,1181,1786,1811]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFB4966FCBB64AD4EBCF8D0" box="[772,914,1786,1810]" italics="true" pageId="4" pageNumber="291">H. tomberua</emphasis>
<bibRefCitation id="FE154B7FFFFB4966FC2664AC49B3F8D1" box="[921,1181,1786,1811]" pageId="4" pageNumber="291" refId="ref3414" refString="Castle, P. H. J. &amp; Randall, J. E. (1999) Revision of Indo-PaCifiC Garden Eels (Congridae: HeteroCongrinae), with desCriptions of five new speCies. Indo-Pacific Fishes, 30, 1 - 52." type="journal article">Castle &amp; Randall 1999</bibRefCitation>
</taxonomicName>
, both species having a very slender body with numerous small dark spots on the body and head, about 200 vertebrae, and similar counts of head pores (
<bibRefCitation id="FE154B7FFFFB4966FE9765144F12F899" box="[296,572,1858,1883]" pageId="4" pageNumber="291" refId="ref3414" refString="Castle, P. H. J. &amp; Randall, J. E. (1999) Revision of Indo-PaCifiC Garden Eels (Congridae: HeteroCongrinae), with desCriptions of five new speCies. Indo-Pacific Fishes, 30, 1 - 52." type="journal article">Castle &amp; Randall 1999</bibRefCitation>
). However, a comparison of
<taxonomicName id="5D844D0DFFFB4966FC2865154ED2F898" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[919,1020,1858,1882]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFB4966FC2865154ED2F898" box="[919,1020,1858,1882]" italics="true" pageId="4" pageNumber="291">H. fugax</emphasis>
</taxonomicName>
with the
<typeStatus id="453F882CFFFB4966FBCB651449F8F899" box="[1140,1238,1858,1883]" pageId="4" pageNumber="291" type="holotype">holotype</typeStatus>
and the original description of
<taxonomicName id="5D844D0DFFFB4966FE8565314CE7F8BC" authorityName="Castle &amp; Randall" authorityYear="1999" box="[314,457,1894,1918]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFB4966FE8565314CE7F8BC" box="[314,457,1894,1918]" italics="true" pageId="4" pageNumber="291">H. tomberua</emphasis>
</taxonomicName>
(USNM 349761:
<figureCitation id="02BF2A0BFFFB4966FD2A65304FF9F8BD" box="[661,727,1894,1919]" captionStart="FIGURE 5" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[152,1435,265,1952]" captionTargetId="figure@6.[152,1435,265,1952]" captionTargetPageId="6" captionText="FIGURE 5. Preserved holotype of Heteroconger tomberua. USNM 349761, 401 mm TL. Red and green arrows indiCate vertiCal positions of tip of appressed peCtoralfin and origin of dorsal fin, respeCtively." httpUri="https://zenodo.org/record/1244902/files/figure.png" pageId="4" pageNumber="291">Fig. 5</figureCitation>
) showed that the former can be distinguished by the dorsal fin originating posterior to the appressed pectoral-fin tip (vs. anterior in
<taxonomicName id="5D844D0DFFFB4966FC1965DD491BF860" authorityName="Castle &amp; Randall" authorityYear="1999" box="[934,1077,1930,1954]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="291" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFB4966FC1965DD491BF860" box="[934,1077,1930,1954]" italics="true" pageId="4" pageNumber="291">H. tomberua</emphasis>
</taxonomicName>
), numerous conical cirri on the chin (vs. absent) (
<figureCitation id="02BF2A0BFFFB4966FEDF65F84C8BF805" box="[352,421,1966,1991]" captionStart="FIGURE 2" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[293,1323,1157,1835]" captionTargetId="figure@2.[264,1323,1131,1857]" captionTargetPageId="2" captionText="FIGURE 2. Numerous small ConiCal Cirri on the Chin of the holotype of Heteroconger fugax sp. nov. KAUMI.95673, 729.0 mm TL." httpUri="https://zenodo.org/record/1244896/files/figure.png" pageId="4" pageNumber="291">Fig. 2</figureCitation>
), a large distinct white rectangular blotch on the opercle (vs. none), dense dark spots over the entire head, including lips (vs. absent on throat, snout and lower jaws), and lacking microscopic dark speckles on the body (vs. present) (
<figureCitation id="02BF2A0BFFFA4967FE0662C14CD4FF72" box="[441,506,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1029,1051]" captionTargetBox="[152,1435,193,1008]" captionTargetId="figure@2.[152,1435,193,1008]" captionTargetPageId="2" captionText="FIGURE 1. Fresh holotype of Heteroconger fugax sp. nov. from Amami-oshima island, Japan. KAUMI. 95673, 729.0 mm TL.Red and green arrows indiCate vertiCal positions of tip of appressed peCtoral fin and origin of dorsal fin, respeCtively." httpUri="https://zenodo.org/record/1244894/files/figure.png" pageId="5" pageNumber="292">Fig. 1</figureCitation>
). In addition,
<taxonomicName id="5D844D0DFFFA4967FD2762CF4FD6FF72" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[664,760,152,176]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFA4967FD2762CF4FD6FF72" box="[664,760,152,176]" italics="true" pageId="5" pageNumber="292">H. fugax</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5D844D0DFFFA4967FC3562CF4939FF72" authorityName="Castle &amp; Randall" authorityYear="1999" box="[906,1047,152,176]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFA4967FC3562CF4939FF72" box="[906,1047,152,176]" italics="true" pageId="5" pageNumber="292">H. tomberua</emphasis>
</taxonomicName>
in having slightly higher counts of preanal vertebrae (
<quantity id="5D7C9B6BFFFA4967FED762EA4C8DFF17" box="[360,419,188,213]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" pageId="5" pageNumber="292" unit="in" value="67.0">67 in</quantity>
<taxonomicName id="5D844D0DFFFA4967FE1562EB4F25FF16" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[426,523,188,212]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFA4967FE1562EB4F25FF16" box="[426,523,188,212]" italics="true" pageId="5" pageNumber="292">H. fugax</emphasis>
</taxonomicName>
vs.
<quantity id="5D7C9B6BFFFA4967FD8562EA4FB3FF17" box="[570,669,188,213]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" metricValueMax="1.6256" metricValueMin="1.4731999999999998" pageId="5" pageNumber="292" unit="in" value="61.0" valueMax="64.0" valueMin="58.0">5864 in</quantity>
<taxonomicName id="5D844D0DFFFA4967FD1B62EB4E1CFF16" authorityName="Castle &amp; Randall" authorityYear="1999" box="[676,818,188,212]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFA4967FD1B62EB4E1CFF16" box="[676,818,188,212]" italics="true" pageId="5" pageNumber="292">H. tomberua</emphasis>
</taxonomicName>
), lateral-line pores before the anus (65 vs. 5663), and shorter head length (11.5% in HL vs. 12.516.3%) and eye diameter (19.2% in HL vs. 22.827.6%) (
<tableCitation id="D7060335FFFA4967FABD62894878FF3A" box="[1282,1366,223,248]" captionStart="TABLE 1" captionStartId="5.[151,239,296,319]" captionText="TABLE 1. Counts and measurements of type specimens of Heteroconger fugax sp. nov. and H. tomberua. Data for paratypes of H. tomberua extracted from Castle &amp; Randall (1999)." pageId="5" pageNumber="292">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="CEFB6606FFFA4967FF28637E4E73FEA0" pageId="5" pageNumber="292">
<paragraph id="9A3B368EFFFA4967FF28637E4E73FEA0" blockId="5.[151,1436,295,354]" pageId="5" pageNumber="292">
<emphasis id="A8F0EA9CFFFA4967FF28637E4C22FEFD" bold="true" box="[151,268,296,319]" pageId="5" pageNumber="292">TABLE 1.</emphasis>
Counts and measurements of type specimens of
<taxonomicName id="5D844D0DFFFA4967FC8E637F4927FEFD" authority="Koeda &amp; Fujii &amp; Motomura, 2018" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[817,1033,295,320]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax" status="sp. nov.">
<emphasis id="A8F0EA9CFFFA4967FC8E637F4927FEFD" box="[817,1033,295,320]" italics="true" pageId="5" pageNumber="292">Heteroconger fugax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3C357E7FFFA4967FBAC637E4942FEFD" box="[1043,1132,296,319]" pageId="5" pageNumber="292" rank="species">
<emphasis id="A8F0EA9CFFFA4967FBAC637E4942FEFD" bold="true" box="[1043,1132,296,319]" pageId="5" pageNumber="292">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="5D844D0DFFFA4967FB19637F481EFEFD" authorityName="Castle &amp; Randall" authorityYear="1999" box="[1190,1328,295,320]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFA4967FB19637F481EFEFD" box="[1190,1328,295,320]" italics="true" pageId="5" pageNumber="292">H. tomberua</emphasis>
</taxonomicName>
. Data for paratypes of
<taxonomicName id="5D844D0DFFFA4967FE9F631D4C89FEA0" authorityName="Castle &amp; Randall" authorityYear="1999" box="[288,423,330,354]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFA4967FE9F631D4C89FEA0" box="[288,423,330,354]" italics="true" pageId="5" pageNumber="292">H. tomberua</emphasis>
</taxonomicName>
extracted from Castle &amp; Randall (1999).
</paragraph>
</caption>
<paragraph id="9A3B368EFFFA4967FD3B632A4834F9AB" pageId="5" pageNumber="292">
<table id="E884C42EFFFAB69DFF20632D481DF9AB" box="[159,1331,379,1641]" gridcols="4" gridrows="31" pageId="5" pageNumber="292">
<tr id="24B434CCFFFAB69DFF20632D481DFE53" box="[159,1331,379,401]" gridrow="0" pageId="5" pageNumber="292" rowspan-0="1" rowspan-3="1">
<th id="67655DB0FFFAB69DFD3B632D4E1DFE53" box="[644,819,379,401]" gridcol="1" gridrow="0" pageId="5" pageNumber="292">
<taxonomicName id="5D844D0DFFFA4967FD3B632A4FF5FE52" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[644,731,379,401]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFA4967FD3B632A4FF5FE52" box="[644,731,379,401]" italics="true" pageId="5" pageNumber="292">H. fugax</emphasis>
</taxonomicName>
</th>
<th id="67655DB0FFFAB69DFC10632D4964FE53" box="[943,1098,379,401]" gridcol="2" gridrow="0" pageId="5" pageNumber="292">
<taxonomicName id="5D844D0DFFFA4967FC10632A4901FE52" authorityName="Castle &amp; Randall" authorityYear="1999" box="[943,1071,379,401]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFFA4967FC10632A4901FE52" box="[943,1071,379,401]" italics="true" pageId="5" pageNumber="292">H. tomberua</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="24B434CCFFFAB69DFF2063F2481DFE78" box="[159,1331,420,442]" gridrow="1" pageId="5" pageNumber="292" rowspan-0="1">
<td id="67655DB0FFFAB69DFD3B63F24E1DFE78" box="[644,819,420,442]" gridcol="1" gridrow="1" pageId="5" pageNumber="292">Holotype</td>
<td id="67655DB0FFFAB69DFC1063F24964FE78" box="[943,1098,420,442]" gridcol="2" gridrow="1" pageId="5" pageNumber="292">Holotype</td>
<td id="67655DB0FFFAB69DFB0B63F2481DFE78" box="[1204,1331,420,442]" gridcol="3" gridrow="1" pageId="5" pageNumber="292">Paratypes</td>
</tr>
<tr id="24B434CCFFFAB69DFF20639B481DFE21" box="[159,1331,461,483]" gridrow="2" pageId="5" pageNumber="292" rowspan-0="1">
<td id="67655DB0FFFAB69DFD3B639B4E1DFE21" box="[644,819,461,483]" gridcol="1" gridrow="2" pageId="5" pageNumber="292">KAUMI. 95673</td>
<td id="67655DB0FFFAB69DFC10639B4964FE21" box="[943,1098,461,483]" gridcol="2" gridrow="2" pageId="5" pageNumber="292">USNM 349761</td>
<td id="67655DB0FFFAB69DFB0B639B481DFE21" box="[1204,1331,461,483]" gridcol="3" gridrow="2" pageId="5" pageNumber="292">
<emphasis id="A8F0EA9CFFFA4967FB0B639849EFFE21" box="[1204,1217,462,483]" italics="true" pageId="5" pageNumber="292">n</emphasis>
= 19
</td>
</tr>
<tr id="24B434CCFFFAB69DFF2063A1481DFDCF" box="[159,1331,503,525]" gridrow="3" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2063A14F3DFDCF" box="[159,531,503,525]" gridcol="0" gridrow="3" pageId="5" pageNumber="292">Total length (mm; TL)</th>
<td id="67655DB0FFFAB69DFD3B63A14E1DFDCF" box="[644,819,503,525]" gridcol="1" gridrow="3" pageId="5" pageNumber="292">729.0</td>
<td id="67655DB0FFFAB69DFC1063A14964FDCF" box="[943,1098,503,525]" gridcol="2" gridrow="3" pageId="5" pageNumber="292">401.0</td>
<td id="67655DB0FFFAB69DFB0B63A1481DFDCF" box="[1204,1331,503,525]" gridcol="3" gridrow="3" pageId="5" pageNumber="292">121.0432.0</td>
</tr>
<tr id="24B434CCFFFAB69DFF206076481DFDF4" box="[159,1331,544,566]" gridrow="4" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF206076481DFDF4" box="[159,1331,544,566]" colspan="4" colspanRight="3" gridcol="0" gridrow="4" pageId="5" pageNumber="292">Counts</th>
</tr>
<tr id="24B434CCFFFAB69DFF20601F481DFD9D" box="[159,1331,585,607]" gridrow="5" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20601F4F3DFD9D" box="[159,531,585,607]" gridcol="0" gridrow="5" pageId="5" pageNumber="292">Pre-dorsal-fin vertebrae</th>
<td id="67655DB0FFFAB69DFD3B601F4E1DFD9D" box="[644,819,585,607]" gridcol="1" gridrow="5" pageId="5" pageNumber="292">8</td>
<td id="67655DB0FFFAB69DFC10601F4964FD9D" box="[943,1098,585,607]" gridcol="2" gridrow="5" pageId="5" pageNumber="292">7</td>
<td id="67655DB0FFFAB69DFB0B601F481DFD9D" box="[1204,1331,585,607]" gridcol="3" gridrow="5" pageId="5" pageNumber="292">58</td>
</tr>
<tr id="24B434CCFFFAB69DFF206025481DFD4B" box="[159,1331,627,649]" gridrow="6" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2060254F3DFD4B" box="[159,531,627,649]" gridcol="0" gridrow="6" pageId="5" pageNumber="292">Pre-anal-fin vertebrae</th>
<td id="67655DB0FFFAB69DFD3B60254E1DFD4B" box="[644,819,627,649]" gridcol="1" gridrow="6" pageId="5" pageNumber="292">67</td>
<td id="67655DB0FFFAB69DFC1060254964FD4B" box="[943,1098,627,649]" gridcol="2" gridrow="6" pageId="5" pageNumber="292">62</td>
<td id="67655DB0FFFAB69DFB0B6025481DFD4B" box="[1204,1331,627,649]" gridcol="3" gridrow="6" pageId="5" pageNumber="292">5864</td>
</tr>
<tr id="24B434CCFFFAB69DFF2060CA481DFD70" box="[159,1331,668,690]" gridrow="7" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2060CA4F3DFD70" box="[159,531,668,690]" gridcol="0" gridrow="7" pageId="5" pageNumber="292">Total vertebrae</th>
<td id="67655DB0FFFAB69DFD3B60CA4E1DFD70" box="[644,819,668,690]" gridcol="1" gridrow="7" pageId="5" pageNumber="292">197</td>
<td id="67655DB0FFFAB69DFC1060CA4964FD70" box="[943,1098,668,690]" gridcol="2" gridrow="7" pageId="5" pageNumber="292">193</td>
<td id="67655DB0FFFAB69DFB0B60CA481DFD70" box="[1204,1331,668,690]" gridcol="3" gridrow="7" pageId="5" pageNumber="292">191204</td>
</tr>
<tr id="24B434CCFFFAB69DFF206093481DFD19" box="[159,1331,709,731]" gridrow="8" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2060934F3DFD19" box="[159,531,709,731]" gridcol="0" gridrow="8" pageId="5" pageNumber="292">Lateral-line pores before peCtoral fin</th>
<td id="67655DB0FFFAB69DFD3B60934E1DFD19" box="[644,819,709,731]" gridcol="1" gridrow="8" pageId="5" pageNumber="292">7</td>
<td id="67655DB0FFFAB69DFC1060934964FD19" box="[943,1098,709,731]" gridcol="2" gridrow="8" pageId="5" pageNumber="292">6</td>
<td id="67655DB0FFFAB69DFB0B6093481DFD19" box="[1204,1331,709,731]" gridcol="3" gridrow="8" pageId="5" pageNumber="292">57</td>
</tr>
<tr id="24B434CCFFFAB69DFF2060B9481DFCC7" box="[159,1331,751,773]" gridrow="9" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2060B94F3DFCC7" box="[159,531,751,773]" gridcol="0" gridrow="9" pageId="5" pageNumber="292">Lateral-line pores before anus</th>
<td id="67655DB0FFFAB69DFD3B60B94E1DFCC7" box="[644,819,751,773]" gridcol="1" gridrow="9" pageId="5" pageNumber="292">65</td>
<td id="67655DB0FFFAB69DFC1060B94964FCC7" box="[943,1098,751,773]" gridcol="2" gridrow="9" pageId="5" pageNumber="292">56</td>
<td id="67655DB0FFFAB69DFB0B60B9481DFCC7" box="[1204,1331,751,773]" gridcol="3" gridrow="9" pageId="5" pageNumber="292">5963</td>
</tr>
<tr id="24B434CCFFFAB69DFF20614E481DFCEC" box="[159,1331,792,814]" gridrow="10" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20614E4F3DFCEC" box="[159,531,792,814]" gridcol="0" gridrow="10" pageId="5" pageNumber="292">PeCtoral-fin rays</th>
<td id="67655DB0FFFAB69DFD3B614E4E1DFCEC" box="[644,819,792,814]" gridcol="1" gridrow="10" pageId="5" pageNumber="292">9</td>
<td id="67655DB0FFFAB69DFC10614E4964FCEC" box="[943,1098,792,814]" gridcol="2" gridrow="10" pageId="5" pageNumber="292">9</td>
<td id="67655DB0FFFAB69DFB0B614E481DFCEC" box="[1204,1331,792,814]" gridcol="3" gridrow="10" pageId="5" pageNumber="292">910</td>
</tr>
<tr id="24B434CCFFFAB69DFF206117481DFC95" box="[159,1331,833,855]" gridrow="11" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2061174F3DFC95" box="[159,531,833,855]" gridcol="0" gridrow="11" pageId="5" pageNumber="292">Dorsal-fin rays before anus</th>
<td id="67655DB0FFFAB69DFD3B61174E1DFC95" box="[644,819,833,855]" gridcol="1" gridrow="11" pageId="5" pageNumber="292">182</td>
<td id="67655DB0FFFAB69DFC1061174964FC95" box="[943,1098,833,855]" gridcol="2" gridrow="11" pageId="5" pageNumber="292">215</td>
<td id="67655DB0FFFAB69DFB0B6117481DFC95" box="[1204,1331,833,855]" gridcol="3" gridrow="11" pageId="5" pageNumber="292">149-220</td>
</tr>
<tr id="24B434CCFFFAB69DFF20613D481DFC43" box="[159,1331,875,897]" gridrow="12" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20613D4F3DFC43" box="[159,531,875,897]" gridcol="0" gridrow="12" pageId="5" pageNumber="292">Total dorsal-fin rays</th>
<td id="67655DB0FFFAB69DFD3B613D4E1DFC43" box="[644,819,875,897]" gridcol="1" gridrow="12" pageId="5" pageNumber="292">590</td>
<td id="67655DB0FFFAB69DFC10613D4964FC43" box="[943,1098,875,897]" gridcol="2" gridrow="12" pageId="5" pageNumber="292">718</td>
<td id="67655DB0FFFAB69DFB0B613D481DFC43" box="[1204,1331,875,897]" gridcol="3" gridrow="12" pageId="5" pageNumber="292">540731</td>
</tr>
<tr id="24B434CCFFFAB69DFF2061C2481DFC68" box="[159,1331,916,938]" gridrow="13" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2061C24F3DFC68" box="[159,531,916,938]" gridcol="0" gridrow="13" pageId="5" pageNumber="292">Anal-fin rays</th>
<td id="67655DB0FFFAB69DFD3B61C24E1DFC68" box="[644,819,916,938]" gridcol="1" gridrow="13" pageId="5" pageNumber="292">425</td>
<td id="67655DB0FFFAB69DFC1061C24964FC68" box="[943,1098,916,938]" gridcol="2" gridrow="13" pageId="5" pageNumber="292">487</td>
<td id="67655DB0FFFAB69DFB0B61C2481DFC68" box="[1204,1331,916,938]" gridcol="3" gridrow="13" pageId="5" pageNumber="292">402529</td>
</tr>
<tr id="24B434CCFFFAB69DFF2061EB481DFC11" box="[159,1331,957,979]" gridrow="14" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2061EB4F3DFC11" box="[159,531,957,979]" gridcol="0" gridrow="14" pageId="5" pageNumber="292">Caudal-fin rays</th>
<td id="67655DB0FFFAB69DFD3B61EB4E1DFC11" box="[644,819,957,979]" gridcol="1" gridrow="14" pageId="5" pageNumber="292">9+8</td>
<td id="67655DB0FFFAB69DFC1061EB4964FC11" box="[943,1098,957,979]" gridcol="2" gridrow="14" pageId="5" pageNumber="292">N/A</td>
<td id="67655DB0FFFAB69DFB0B61EB481DFC11" box="[1204,1331,957,979]" gridcol="3" gridrow="14" pageId="5" pageNumber="292">N/A</td>
</tr>
<tr id="24B434CCFFFAB69DFF2061B1481DFC3F" box="[159,1331,999,1021]" gridrow="15" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2061B1481DFC3F" box="[159,1331,999,1021]" colspan="4" colspanRight="3" gridcol="0" gridrow="15" pageId="5" pageNumber="292">As% of TL</th>
</tr>
<tr id="24B434CCFFFAB69DFF206646481DFBE4" box="[159,1331,1040,1062]" gridrow="16" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2066464F3DFBE4" box="[159,531,1040,1062]" gridcol="0" gridrow="16" pageId="5" pageNumber="292">Head length (HL)</th>
<td id="67655DB0FFFAB69DFD3B66464E1DFBE4" box="[644,819,1040,1062]" gridcol="1" gridrow="16" pageId="5" pageNumber="292">3.4</td>
<td id="67655DB0FFFAB69DFC1066464964FBE4" box="[943,1098,1040,1062]" gridcol="2" gridrow="16" pageId="5" pageNumber="292">4.1</td>
<td id="67655DB0FFFAB69DFB0B6646481DFBE4" box="[1204,1331,1040,1062]" gridcol="3" gridrow="16" pageId="5" pageNumber="292">N/A</td>
</tr>
<tr id="24B434CCFFFAB69DFF20666F481DFB8D" box="[159,1331,1081,1103]" gridrow="17" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20666F4F3DFB8D" box="[159,531,1081,1103]" gridcol="0" gridrow="17" pageId="5" pageNumber="292">Snout-anus length (SAL)</th>
<td id="67655DB0FFFAB69DFD3B666F4E1DFB8D" box="[644,819,1081,1103]" gridcol="1" gridrow="17" pageId="5" pageNumber="292">30.1</td>
<td id="67655DB0FFFAB69DFC10666F4964FB8D" box="[943,1098,1081,1103]" gridcol="2" gridrow="17" pageId="5" pageNumber="292">29.1</td>
<td id="67655DB0FFFAB69DFB0B666F481DFB8D" box="[1204,1331,1081,1103]" gridcol="3" gridrow="17" pageId="5" pageNumber="292">28.636.6</td>
</tr>
<tr id="24B434CCFFFAB69DFF206635481DFBBB" box="[159,1331,1123,1145]" gridrow="18" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2066354F3DFBBB" box="[159,531,1123,1145]" gridcol="0" gridrow="18" pageId="5" pageNumber="292">Depth at gill opening</th>
<td id="67655DB0FFFAB69DFD3B66354E1DFBBB" box="[644,819,1123,1145]" gridcol="1" gridrow="18" pageId="5" pageNumber="292">1.2</td>
<td id="67655DB0FFFAB69DFC1066354964FBBB" box="[943,1098,1123,1145]" gridcol="2" gridrow="18" pageId="5" pageNumber="292">1.4</td>
<td id="67655DB0FFFAB69DFB0B6635481DFBBB" box="[1204,1331,1123,1145]" gridcol="3" gridrow="18" pageId="5" pageNumber="292">N/A</td>
</tr>
<tr id="24B434CCFFFAB69DFF2066DA481DFB60" box="[159,1331,1164,1186]" gridrow="19" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2066DA4F3DFB60" box="[159,531,1164,1186]" gridcol="0" gridrow="19" pageId="5" pageNumber="292">Depth at anus</th>
<td id="67655DB0FFFAB69DFD3B66DA4E1DFB60" box="[644,819,1164,1186]" gridcol="1" gridrow="19" pageId="5" pageNumber="292">1.0</td>
<td id="67655DB0FFFAB69DFC1066DA4964FB60" box="[943,1098,1164,1186]" gridcol="2" gridrow="19" pageId="5" pageNumber="292">1.0</td>
<td id="67655DB0FFFAB69DFB0B66DA481DFB60" box="[1204,1331,1164,1186]" gridcol="3" gridrow="19" pageId="5" pageNumber="292">N/A</td>
</tr>
<tr id="24B434CCFFFAB69DFF2066E3481DFB09" box="[159,1331,1205,1227]" gridrow="20" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2066E3481DFB09" box="[159,1331,1205,1227]" colspan="4" colspanRight="3" gridcol="0" gridrow="20" pageId="5" pageNumber="292">As% of SAL</th>
</tr>
<tr id="24B434CCFFFAB69DFF206689481DFB37" box="[159,1331,1247,1269]" gridrow="21" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2066894F3DFB37" box="[159,531,1247,1269]" gridcol="0" gridrow="21" pageId="5" pageNumber="292">HL</th>
<td id="67655DB0FFFAB69DFD3B66894E1DFB37" box="[644,819,1247,1269]" gridcol="1" gridrow="21" pageId="5" pageNumber="292">11.5</td>
<td id="67655DB0FFFAB69DFC1066894964FB37" box="[943,1098,1247,1269]" gridcol="2" gridrow="21" pageId="5" pageNumber="292">14.2</td>
<td id="67655DB0FFFAB69DFB0B6689481DFB37" box="[1204,1331,1247,1269]" gridcol="3" gridrow="21" pageId="5" pageNumber="292">12.516.3</td>
</tr>
<tr id="24B434CCFFFAB69DFF20675E481DFADC" box="[159,1331,1288,1310]" gridrow="22" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20675E4F3DFADC" box="[159,531,1288,1310]" gridcol="0" gridrow="22" pageId="5" pageNumber="292">Pre-dorsal-fin length</th>
<td id="67655DB0FFFAB69DFD3B675E4E1DFADC" box="[644,819,1288,1310]" gridcol="1" gridrow="22" pageId="5" pageNumber="292">13.7</td>
<td id="67655DB0FFFAB69DFC10675E4964FADC" box="[943,1098,1288,1310]" gridcol="2" gridrow="22" pageId="5" pageNumber="292">14.6</td>
<td id="67655DB0FFFAB69DFB0B675E481DFADC" box="[1204,1331,1288,1310]" gridcol="3" gridrow="22" pageId="5" pageNumber="292">13.317.3</td>
</tr>
<tr id="24B434CCFFFAB69DFF206767481DFA85" box="[159,1331,1329,1351]" gridrow="23" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF206767481DFA85" box="[159,1331,1329,1351]" colspan="4" colspanRight="3" gridcol="0" gridrow="23" pageId="5" pageNumber="292">As% of HL</th>
</tr>
<tr id="24B434CCFFFAB69DFF20670D481DFAB3" box="[159,1331,1371,1393]" gridrow="24" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20670D4F3DFAB3" box="[159,531,1371,1393]" gridcol="0" gridrow="24" pageId="5" pageNumber="292">Snout length</th>
<td id="67655DB0FFFAB69DFD3B670D4E1DFAB3" box="[644,819,1371,1393]" gridcol="1" gridrow="24" pageId="5" pageNumber="292">16.8</td>
<td id="67655DB0FFFAB69DFC10670D4964FAB3" box="[943,1098,1371,1393]" gridcol="2" gridrow="24" pageId="5" pageNumber="292">16.5</td>
<td id="67655DB0FFFAB69DFB0B670D481DFAB3" box="[1204,1331,1371,1393]" gridcol="3" gridrow="24" pageId="5" pageNumber="292">12.518.2</td>
</tr>
<tr id="24B434CCFFFAB69DFF2067D2481DFA58" box="[159,1331,1412,1434]" gridrow="25" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2067D24F3DFA58" box="[159,531,1412,1434]" gridcol="0" gridrow="25" pageId="5" pageNumber="292">Eye diameter</th>
<td id="67655DB0FFFAB69DFD3B67D24E1DFA58" box="[644,819,1412,1434]" gridcol="1" gridrow="25" pageId="5" pageNumber="292">19.2</td>
<td id="67655DB0FFFAB69DFC1067D24964FA58" box="[943,1098,1412,1434]" gridcol="2" gridrow="25" pageId="5" pageNumber="292">22.8</td>
<td id="67655DB0FFFAB69DFB0B67D2481DFA58" box="[1204,1331,1412,1434]" gridcol="3" gridrow="25" pageId="5" pageNumber="292">23.327.6</td>
</tr>
<tr id="24B434CCFFFAB69DFF2067FB481DFA01" box="[159,1331,1453,1475]" gridrow="26" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2067FB4F3DFA01" box="[159,531,1453,1475]" gridcol="0" gridrow="26" pageId="5" pageNumber="292">Mouth width</th>
<td id="67655DB0FFFAB69DFD3B67FB4E1DFA01" box="[644,819,1453,1475]" gridcol="1" gridrow="26" pageId="5" pageNumber="292">24.5</td>
<td id="67655DB0FFFAB69DFC1067FB4964FA01" box="[943,1098,1453,1475]" gridcol="2" gridrow="26" pageId="5" pageNumber="292">24.0</td>
<td id="67655DB0FFFAB69DFB0B67FB481DFA01" box="[1204,1331,1453,1475]" gridcol="3" gridrow="26" pageId="5" pageNumber="292">18.225.0</td>
</tr>
<tr id="24B434CCFFFAB69DFF206781481DFA2F" box="[159,1331,1495,1517]" gridrow="27" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2067814F3DFA2F" box="[159,531,1495,1517]" gridcol="0" gridrow="27" pageId="5" pageNumber="292">Gill opening</th>
<td id="67655DB0FFFAB69DFD3B67814E1DFA2F" box="[644,819,1495,1517]" gridcol="1" gridrow="27" pageId="5" pageNumber="292">9.1</td>
<td id="67655DB0FFFAB69DFC1067814964FA2F" box="[943,1098,1495,1517]" gridcol="2" gridrow="27" pageId="5" pageNumber="292">N/A</td>
<td id="67655DB0FFFAB69DFB0B6781481DFA2F" box="[1204,1331,1495,1517]" gridcol="3" gridrow="27" pageId="5" pageNumber="292">4.49.1</td>
</tr>
<tr id="24B434CCFFFAB69DFF206456481DF9D4" box="[159,1331,1536,1558]" gridrow="28" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2064564F3DF9D4" box="[159,531,1536,1558]" gridcol="0" gridrow="28" pageId="5" pageNumber="292">BranChial interspaCe</th>
<td id="67655DB0FFFAB69DFD3B64564E1DF9D4" box="[644,819,1536,1558]" gridcol="1" gridrow="28" pageId="5" pageNumber="292">21.4</td>
<td id="67655DB0FFFAB69DFC1064564964F9D4" box="[943,1098,1536,1558]" gridcol="2" gridrow="28" pageId="5" pageNumber="292">18.9</td>
<td id="67655DB0FFFAB69DFB0B6456481DF9D4" box="[1204,1331,1536,1558]" gridcol="3" gridrow="28" pageId="5" pageNumber="292">13.821.2</td>
</tr>
<tr id="24B434CCFFFAB69DFF20647F481DF9FD" box="[159,1331,1577,1599]" gridrow="29" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF20647F4F3DF9FD" box="[159,531,1577,1599]" gridcol="0" gridrow="29" pageId="5" pageNumber="292">Depth at gill opening</th>
<td id="67655DB0FFFAB69DFD3B647F4E1DF9FD" box="[644,819,1577,1599]" gridcol="1" gridrow="29" pageId="5" pageNumber="292">34.2</td>
<td id="67655DB0FFFAB69DFC10647F4964F9FD" box="[943,1098,1577,1599]" gridcol="2" gridrow="29" pageId="5" pageNumber="292">33.1</td>
<td id="67655DB0FFFAB69DFB0B647F481DF9FD" box="[1204,1331,1577,1599]" gridcol="3" gridrow="29" pageId="5" pageNumber="292">25.835.3</td>
</tr>
<tr id="24B434CCFFFAB69DFF206405481DF9AB" box="[159,1331,1619,1641]" gridrow="30" pageId="5" pageNumber="292">
<th id="67655DB0FFFAB69DFF2064054F3DF9AB" box="[159,531,1619,1641]" gridcol="0" gridrow="30" pageId="5" pageNumber="292">Depth at anus</th>
<td id="67655DB0FFFAB69DFD3B64054E1DF9AB" box="[644,819,1619,1641]" gridcol="1" gridrow="30" pageId="5" pageNumber="292">27.9</td>
<td id="67655DB0FFFAB69DFC1064054964F9AB" box="[943,1098,1619,1641]" gridcol="2" gridrow="30" pageId="5" pageNumber="292">23.0</td>
<td id="67655DB0FFFAB69DFB0B6405481DF9AB" box="[1204,1331,1619,1641]" gridcol="3" gridrow="30" pageId="5" pageNumber="292">18.426.5</td>
</tr>
</table>
</paragraph>
<subSubSection id="D29E6505FFFA4965FF7864F94818FEDE" lastPageId="7" lastPageNumber="294" pageId="5" pageNumber="292" type="reference_group">
<paragraph id="9A3B368EFFFA4967FF7864F949D7F8B8" blockId="5.[151,1437,1709,2023]" pageId="5" pageNumber="292">
<taxonomicName id="5D844D0DFFFA4967FF7864F94FFDF904" ID-CoL="3L8RJ" authority="Bohlke &amp; Randall 1981" authorityName="Bohlke &amp; Randall" authorityYear="1981" box="[199,723,1709,1734]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="cobra">
<emphasis id="A8F0EA9CFFFA4967FF7864F94C81F904" box="[199,431,1710,1734]" italics="true" pageId="5" pageNumber="292">Heteroconger cobra</emphasis>
<bibRefCitation id="FE154B7FFFFA4967FE0664FB4FFDF904" box="[441,723,1709,1734]" pageId="5" pageNumber="292" refId="ref3371" refString="Bohlke, J. E. &amp; Randall, J. E. (1981) Four new garden eels (Congridae, HeteroCongrinae) from the PaCifiC and Indian oCeans. Bulletin of Marine Science, 31, 366 - 382." type="journal article">Böhlke &amp; Randall 1981</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="5D844D0DFFFA4967FD5C64F949A9F904" ID-CoL="6LT28" authority="Allen &amp; Erdmann 2009" authorityName="Allen &amp; Erdmann" authorityYear="2009" box="[739,1159,1709,1734]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="mercyae">
<emphasis id="A8F0EA9CFFFA4967FD5C64F94E46F904" box="[739,872,1711,1734]" italics="true" pageId="5" pageNumber="292">H. mercyae</emphasis>
<bibRefCitation id="FE154B7FFFFA4967FCCC64FB49A9F904" box="[883,1159,1709,1734]" pageId="5" pageNumber="292" refId="ref3142" refString="Allen, G. R. &amp; Erdmann, M. V. (2009) Heteroconger mercyae, a new speCies of garden eel (Congridae: HeteroCongridae) from West Papua, Indonesia. Aqua, International Journal of Ichthyology, 15, 135 - 142." type="journal article">Allen &amp; Erdmann 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="5D844D0DFFFA4967FB7964F94C03F928" ID-CoL="3L8S3" authority="Castle &amp; Randall 1999" authorityName="Castle &amp; Randall" authorityYear="1999" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tricia">
<emphasis id="A8F0EA9CFFFA4967FB7964F94804F904" box="[1222,1322,1711,1734]" italics="true" pageId="5" pageNumber="292">H. tricia</emphasis>
<bibRefCitation id="FE154B7FFFFA4967FA8A64FB4C03F928" pageId="5" pageNumber="292" refId="ref3414" refString="Castle, P. H. J. &amp; Randall, J. E. (1999) Revision of Indo-PaCifiC Garden Eels (Congridae: HeteroCongrinae), with desCriptions of five new speCies. Indo-Pacific Fishes, 30, 1 - 52." type="journal article">Castle &amp; Randall 1999</bibRefCitation>
</taxonomicName>
all have&gt; 196 vertebrae (as does
<taxonomicName id="5D844D0DFFFA4967FD1664854E27F928" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[681,777,1746,1770]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFFA4967FD1664854E27F928" box="[681,777,1746,1770]" italics="true" pageId="5" pageNumber="292">H. fugax</emphasis>
</taxonomicName>
). However, the coloration of these species, e.g. large black saddle on the head enclosing a U-shaped white marking, and pair of widely spaced black saddles with white central bar on the anterior body in
<taxonomicName id="5D844D0DFFFA4967FE6C654D4F16F8F0" ID-CoL="3L8RJ" authorityName="Bohlke &amp; Randall" authorityYear="1981" box="[467,568,1818,1842]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="cobra">
<emphasis id="A8F0EA9CFFFA4967FE6C654D4F16F8F0" box="[467,568,1818,1842]" italics="true" pageId="5" pageNumber="292">H. cobra</emphasis>
</taxonomicName>
; zebra-like pattern of black and white bars in
<taxonomicName id="5D844D0DFFFA4967FBF1654D49FEF8F0" ID-CoL="6LT28" authorityName="Allen &amp; Erdmann" authorityYear="2009" box="[1102,1232,1819,1842]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="mercyae">
<emphasis id="A8F0EA9CFFFA4967FBF1654D49FEF8F0" box="[1102,1232,1819,1842]" italics="true" pageId="5" pageNumber="292">H. mercyae</emphasis>
</taxonomicName>
; and two rows of large round spots on the body in
<taxonomicName id="5D844D0DFFFA4967FD8965694FB5F894" ID-CoL="3L8S3" authorityName="Castle &amp; Randall" authorityYear="1999" box="[566,667,1855,1878]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="tricia">
<emphasis id="A8F0EA9CFFFA4967FD8965694FB5F894" box="[566,667,1855,1878]" italics="true" pageId="5" pageNumber="292">H. tricia</emphasis>
</taxonomicName>
; are very different from that of
<emphasis id="A8F0EA9CFFFA4967FB90656949B0F894" box="[1071,1182,1854,1878]" italics="true" pageId="5" pageNumber="292">
<taxonomicName id="5D844D0DFFFA4967FB90656949B4F894" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[1071,1178,1854,1878]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="fugax">H. fugax</taxonomicName>
.
</emphasis>
The remaining eight
<taxonomicName id="5D844D0DFFFA4967FF2865354C1CF8B8" ID-CoL="4WCR" authorityName="Bleeker" authorityYear="1868" box="[151,306,1891,1914]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="genus">
<emphasis id="A8F0EA9CFFFA4967FF2865354C1CF8B8" box="[151,306,1891,1914]" italics="true" pageId="5" pageNumber="292">Heteroconger</emphasis>
</taxonomicName>
species have &lt;175 vertebrae (
<bibRefCitation id="FE154B7FFFFA4967FD3465344EA1F8B8" box="[651,911,1890,1915]" pageId="5" pageNumber="292" refId="ref3414" refString="Castle, P. H. J. &amp; Randall, J. E. (1999) Revision of Indo-PaCifiC Garden Eels (Congridae: HeteroCongrinae), with desCriptions of five new speCies. Indo-Pacific Fishes, 30, 1 - 52." type="journal article">Castle &amp; Randall 1999</bibRefCitation>
;
<bibRefCitation id="FE154B7FFFFA4967FC2465344988F8B8" box="[923,1190,1890,1915]" pageId="5" pageNumber="292" refId="ref3142" refString="Allen, G. R. &amp; Erdmann, M. V. (2009) Heteroconger mercyae, a new speCies of garden eel (Congridae: HeteroCongridae) from West Papua, Indonesia. Aqua, International Journal of Ichthyology, 15, 135 - 142." type="journal article">Allen &amp; Erdmann 2009</bibRefCitation>
,
<bibRefCitation id="FE154B7FFFFA4967FB0F653449C3F8B8" box="[1200,1261,1890,1914]" pageId="5" pageNumber="292" refId="ref3191" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol 1. TropiCal Reef ResearCh, Perth, xiii + 424 pp." type="book">2012</bibRefCitation>
).
</paragraph>
<paragraph id="9A3B368EFFFA4964FF7865D14F5EFF17" blockId="5.[151,1437,1709,2023]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292">
<taxonomicName id="5D844D0DFFFA4967FF7865D14FACF85C" ID-CoL="3L8RH" authority="(Herre 1923)" authorityName="Herre" authorityYear="1923" baseAuthorityName="Herre" baseAuthorityYear="1923" box="[199,642,1926,1950]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="species" species="chapmani">
<emphasis id="A8F0EA9CFFFA4967FF7865D14CF2F85C" box="[199,476,1926,1950]" italics="true" pageId="5" pageNumber="292">Heteroconger chapmani</emphasis>
(Herre 1923)
</taxonomicName>
was regarded as “unidentifiable” in the revision of the subfamily
<taxonomicName id="5D844D0DFFFA4967FF2865FC4C7CF801" authorityName="Günther" authorityYear="1870" box="[151,338,1962,1987]" class="Actinopterygii" family="Congridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="292" phylum="Chordata" rank="subFamily" subFamily="Heterocongrinae">Heterocongrinae</taxonomicName>
because of an inadequately description (of 2 poorly preserved Philippine specimens), compounded by a lack of
<typeStatus id="453F882CFFFA4967FE7265994CD3F825" box="[461,509,1999,2023]" pageId="5" pageNumber="292">type</typeStatus>
specimens, color description or vertebral counts (
<bibRefCitation id="FE154B7FFFFA4967FBF7659B4877F824" box="[1096,1369,1997,2022]" pageId="5" pageNumber="292" refId="ref3414" refString="Castle, P. H. J. &amp; Randall, J. E. (1999) Revision of Indo-PaCifiC Garden Eels (Congridae: HeteroCongrinae), with desCriptions of five new speCies. Indo-Pacific Fishes, 30, 1 - 52." type="journal article">Castle &amp; Randall 1999</bibRefCitation>
). The species was omitted by
<bibRefCitation id="FE154B7FFFF94964FE1562C14FE5FF72" box="[426,715,151,176]" pageId="6" pageNumber="293" refId="ref3142" refString="Allen, G. R. &amp; Erdmann, M. V. (2009) Heteroconger mercyae, a new speCies of garden eel (Congridae: HeteroCongridae) from West Papua, Indonesia. Aqua, International Journal of Ichthyology, 15, 135 - 142." type="journal article">Allen &amp; Erdmann (2009)</bibRefCitation>
in their comparison of a newly undescribed species with valid congeners, and we followed their opinion.
</paragraph>
<caption id="CEFB6606FFF94964FF2865E34ED3F828" httpUri="https://zenodo.org/record/1244902/files/figure.png" pageId="6" pageNumber="293" startId="6.[151,250,1973,1995]" targetBox="[152,1435,265,1952]" targetPageId="6">
<paragraph id="9A3B368EFFF94964FF2865E34ED3F828" blockId="6.[151,1435,1973,2026]" pageId="6" pageNumber="293">
<emphasis id="A8F0EA9CFFF94964FF2865E34C3EF808" bold="true" box="[151,272,1973,1995]" pageId="6" pageNumber="293">FIGURE 5</emphasis>
. Preserved holotype of
<taxonomicName id="5D844D0DFFF94964FDB265E04E2AF808" ID-CoL="3L8S2" authorityName="Castle &amp; Randall" authorityYear="1999" box="[525,772,1973,1995]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="293" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFF94964FDB265E04E2AF808" box="[525,772,1973,1995]" italics="true" pageId="6" pageNumber="293">Heteroconger tomberua</emphasis>
</taxonomicName>
. USNM 349761, 401 mm TL. Red and green arrows indiCate vertiCal positions of tip of appressed peCtoral fin and origin of dorsal fin, respeCtively.
</paragraph>
</caption>
<paragraph id="9A3B368EFFF84965FF7862CE4818FEDE" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="294">
<bibRefCitation id="FE154B7FFFF84965FF7862CE4C88FF72" box="[199,422,152,176]" pageId="7" pageNumber="294" refId="ref3636" refString="Senou, H., Kobayashi, Y. &amp; Kobayashi, N. (2007) Coastal fishes of the Miyako Group, the Ryukyu Islands, Japan. Bulletin of Kanagawa Prefectural Museum (Natural Science), 36, 47 - 74." type="journal article">
Senou
<emphasis id="A8F0EA9CFFF84965FEA862CF4C65FF72" box="[279,331,152,176]" italics="true" pageId="7" pageNumber="294">et al</emphasis>
. (2007)
</bibRefCitation>
reported
<taxonomicName id="5D844D0DFFF84965FDA862CF4ECCFF72" ID-CoL="3L8RR" authority="Bohlke &amp; Randall 1981" authorityName="Bohlke &amp; Randall" authorityYear="1981" box="[535,994,151,176]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="lentiginosus">
<emphasis id="A8F0EA9CFFF84965FDA862CF4FEAFF72" box="[535,708,152,176]" italics="true" pageId="7" pageNumber="294">H. lentiginosus</emphasis>
<bibRefCitation id="FE154B7FFFF84965FD7162C14ECCFF72" box="[718,994,151,176]" pageId="7" pageNumber="294" refId="ref3371" refString="Bohlke, J. E. &amp; Randall, J. E. (1981) Four new garden eels (Congridae, HeteroCongrinae) from the PaCifiC and Indian oCeans. Bulletin of Marine Science, 31, 366 - 382." type="journal article">Böhlke &amp; Randall 1981</bibRefCitation>
</taxonomicName>
from Irabu-jima island in the Miyako Islands,
<collectingCountry id="E293761EFFF84965FF4662EB4C12FF16" box="[249,316,189,212]" name="Japan" pageId="7" pageNumber="294">Japan</collectingCountry>
, based on an underwater photograph (KPM-NR 6 7322). The photograph shows clearly a large distinct white rectangular blotch on the opercle (vs. absent in
<taxonomicName id="5D844D0DFFF84965FCE462B74928FF3A" ID-CoL="3L8RR" authorityName="Bohlke &amp; Randall" authorityYear="1981" box="[859,1030,224,248]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="lentiginosus">
<emphasis id="A8F0EA9CFFF84965FCE462B74928FF3A" box="[859,1030,224,248]" italics="true" pageId="7" pageNumber="294">H. lentiginosus</emphasis>
</taxonomicName>
), circular numerous dark spots (vs. freckles) and head uniformly covered by spots (vs. uniformly dark brown), and is identical with
<emphasis id="A8F0EA9CFFF84965FB7263534818FEDE" box="[1229,1334,260,284]" italics="true" pageId="7" pageNumber="294">
<taxonomicName id="5D844D0DFFF84965FB726353481FFEDE" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[1229,1329,260,284]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="fugax">H. fugax</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="D29E6505FFF84965FF7863714C5FFD0F" pageId="7" pageNumber="294" type="description">
<paragraph id="9A3B368EFFF84965FF7863714C5FFD0F" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="294">
<bibRefCitation id="FE154B7FFFF84965FF7863714C94FE82" box="[199,442,295,320]" pageId="7" pageNumber="294" refId="ref3228" refString="Biswas, S., Mishra, S. S., Satpathy, K. K., Das, N. P. I., Selvanayagam, M. &amp; Nayak, L. (2012) A new reCord of a garden eel, Heteroconger tomberua (ACtinopterygii: Anguilliformes: Congridae), from the Indian OCean. Acta Ichthyologica et Piscatoria, 42, 65 - 68. https: // doi. org / 10.3750 / AIP 2011.42.1.08" type="journal article">
Biswas
<emphasis id="A8F0EA9CFFF84965FE9B637F4C74FE82" box="[292,346,296,320]" italics="true" pageId="7" pageNumber="294">et al</emphasis>
. (2012)
</bibRefCitation>
reported a species resembling
<taxonomicName id="5D844D0DFFF84965FC8F637F4EB8FE82" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[816,918,296,320]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFF84965FC8F637F4EB8FE82" box="[816,918,296,320]" italics="true" pageId="7" pageNumber="294">H. fugax</emphasis>
</taxonomicName>
from Kalakkam, east coast of
<collectingCountry id="E293761EFFF84965FAB063714865FE82" box="[1295,1355,295,320]" name="India" pageId="7" pageNumber="294">India</collectingCountry>
, as
<taxonomicName id="5D844D0DFFF84965FA3E637F4C2CFEA6" authorityName="Castle &amp; Randall" authorityYear="1999" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="tomberua">
<emphasis id="A8F0EA9CFFF84965FA3E637F4C2CFEA6" italics="true" pageId="7" pageNumber="294">H. tomberua</emphasis>
</taxonomicName>
, based on a specimen (ZSI F-10575/2) collected from the traveling water screen of the seawater cooling system of the nearby nuclear power station. The specimen was similar to
<taxonomicName id="5D844D0DFFF84965FC5963274967FE4A" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[998,1097,368,392]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFF84965FC5963274967FE4A" box="[998,1097,368,392]" italics="true" pageId="7" pageNumber="294">H. fugax</emphasis>
</taxonomicName>
, having an extremely slender body (depth 1.16% of TL), 191 total vertebrae, dark spots on the head and body, a large distinct white blotch on the opercle, spots on the head more numerous and denser than on the body, lips spotted, and body ground color uniformly cream, lacking microscopic dark speckles. However,
<taxonomicName id="5D844D0DFFF84965FCCB638B4EF9FE36" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[884,983,476,500]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFF84965FCCB638B4EF9FE36" box="[884,983,476,500]" italics="true" pageId="7" pageNumber="294">H. fugax</emphasis>
</taxonomicName>
differed in having the dorsal-fin origin posterior to tip of the appressed pectoral fin (vs. above middle of pectoral fin), 23 rows of dark spots above and below the lateral line (vs. 34 rows of dark spots restricted to anterior half of body, faint or lacking posteriorly), 67 row of spots across interorbital area (vs. 1113 rows of spots), and numerous crowded dark spots over entire head (vs. ventral half of head lacking dark spots) all from
<bibRefCitation id="FE154B7FFFF84965FCFD603A491EFD46" box="[834,1072,620,645]" pageId="7" pageNumber="294" refId="ref3228" refString="Biswas, S., Mishra, S. S., Satpathy, K. K., Das, N. P. I., Selvanayagam, M. &amp; Nayak, L. (2012) A new reCord of a garden eel, Heteroconger tomberua (ACtinopterygii: Anguilliformes: Congridae), from the Indian OCean. Acta Ichthyologica et Piscatoria, 42, 65 - 68. https: // doi. org / 10.3750 / AIP 2011.42.1.08" type="journal article">
Biswas
<emphasis id="A8F0EA9CFFF84965FC23603B4EF6FD46" box="[924,984,620,644]" italics="true" pageId="7" pageNumber="294">et al.</emphasis>
(2012)
</bibRefCitation>
. Clearly,
<bibRefCitation id="FE154B7FFFF84965FB20603A48B2FD46" box="[1183,1436,620,645]" pageId="7" pageNumber="294" refId="ref3228" refString="Biswas, S., Mishra, S. S., Satpathy, K. K., Das, N. P. I., Selvanayagam, M. &amp; Nayak, L. (2012) A new reCord of a garden eel, Heteroconger tomberua (ACtinopterygii: Anguilliformes: Congridae), from the Indian OCean. Acta Ichthyologica et Piscatoria, 42, 65 - 68. https: // doi. org / 10.3750 / AIP 2011.42.1.08" type="journal article">
Biswas
<emphasis id="A8F0EA9CFFF84965FB45603B4800FD46" box="[1274,1326,620,644]" italics="true" pageId="7" pageNumber="294">et al</emphasis>
.s (2012)
</bibRefCitation>
specimen is distinct from
<taxonomicName id="5D844D0DFFF84965FE7E60C74F0DFD6A" authorityName="Koeda &amp; Fujii &amp; Motomura" authorityYear="2018" box="[449,547,656,680]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="species" species="fugax">
<emphasis id="A8F0EA9CFFF84965FE7E60C74F0DFD6A" box="[449,547,656,680]" italics="true" pageId="7" pageNumber="294">H. fugax</emphasis>
</taxonomicName>
, as well as from all other known species of
<taxonomicName id="5D844D0DFFF84965FB9A60C74990FD6A" authorityName="Bleeker" authorityYear="1868" box="[1061,1214,657,680]" class="Actinopterygii" family="Congridae" genus="Heteroconger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="294" phylum="Chordata" rank="genus">
<emphasis id="A8F0EA9CFFF84965FB9A60C74990FD6A" box="[1061,1214,657,680]" italics="true" pageId="7" pageNumber="294">Heteroconger</emphasis>
</taxonomicName>
, and remains to be formally identified.
</paragraph>
</subSubSection>
</treatment>
</document>