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<document id="C21ABAACCAEAD7A57A789613E53BE2B7" ID-CLB-Dataset="20703" ID-DOI="10.15553/c2018v731a4" ID-GBIF-Dataset="023dbe5b-0635-4a3b-b1dc-d8a7532c75cc" ID-ISSN="2235-3658" ID-Zenodo-Dep="5722136" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637688271325" checkinUser="carolina" docAuthor="Applequist, Wendy L." docDate="2018" docId="03CAA9273A10FFB8E74EF990FA262FF7" docLanguage="en" docName="Candollea.73.1.27-48.pdf" docOrigin="Candollea 73 (1)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Homalium moniliforme H. Perrier" docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFF3D15F3A18FFB2E476FFA7FFE42D0E" masterDocTitle="A revision of Homalium sect. Odontolobus (Salicaceae) endemic to Madagascar" masterLastPageNumber="48" masterPageNumber="27" pageNumber="35" updateTime="1699469775107" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="154B4989719FDA0E4A83C46E93580E60">A revision of Homalium sect. Odontolobus (Salicaceae) endemic to Madagascar</mods:title>
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<mods:namePart id="DA5D608C2D604104560800C91E3951A1">Applequist, Wendy L.</mods:namePart>
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<mods:title id="BA00E2CB1FDAA50FD4939AA5E2D359D0">Candollea</mods:title>
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<mods:date id="D62A846094226E596E8FEF484C3A0F8A">2018</mods:date>
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<treatment id="03CAA9273A10FFB8E74EF990FA262FF7" ID-DOI="http://doi.org/10.5281/zenodo.6309037" ID-GBIF-Taxon="193333140" ID-Zenodo-Dep="6309037" LSID="urn:lsid:plazi:treatment:03CAA9273A10FFB8E74EF990FA262FF7" httpUri="http://treatment.plazi.org/id/03CAA9273A10FFB8E74EF990FA262FF7" lastPageId="10" lastPageNumber="37" pageId="8" pageNumber="35">
<subSubSection id="C3794BBA3A10FFBAE74EF990FB812B61" pageId="8" pageNumber="35" type="nomenclature">
<paragraph id="8BDC18313A10FFBAE74EF990FB812B61" blockId="8.[824,1474,1590,1647]" pageId="8" pageNumber="35">
6.
<taxonomicName id="4C6363B23A10FFBAE728F991FB862B61" ID-CoL="3MCNY" authority="H. Perrier" authorityName="H. Perrier" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="moniliforme">
<emphasis id="B917C4233A10FFBAE728F991FB842B40" bold="true" box="[862,1120,1590,1614]" italics="true" pageId="8" pageNumber="35">Homalium moniliforme</emphasis>
<bibRefCitation id="EFF265C03A10FFBAE01FF990FB862B61" author="PERRIER DE LA BATHIE, H." firstAuthor="H" pageId="8" pageNumber="35" pagination="261 - 301" refId="ref21624" refString="PERRIER DE LA BATHIE, H. (1940). Revision des Flacourtiacees de Madagascar et des Comores. Mem. Mus. Natl. Hist. Nat. 13: 261 - 301." type="journal article" year="1940">
H. Perrier in
<bibCitation id="5E49105D3A10FFBAE177F990FB862B61" journal="Mem. Mus. Natl. Hist. Nat." page="291" pageId="8" pageNumber="35" score="166.9911666303064" volume="13" year="1940">Mém. Mus. Natl. Hist. Nat. 13: 291. 1940</bibCitation>
</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3794BBA3A10FFBAE728F925FA9F2A37" pageId="8" pageNumber="35" type="materials_examined">
<paragraph id="8BDC18313A10FFBAE728F925FA9F2A37" blockId="8.[862,1474,1666,1850]" pageId="8" pageNumber="35">
<materialsCitation id="3B0B126C3A10FFBAE728F925FA5A2BD7" ID-GBIF-Occurrence="3464757388" collectionCode="P" collectorName="Decary" country="Madagascar" location="Vondrozo" pageId="8" pageNumber="35" specimenCode="P04679983" specimenCount="1" stateProvince="Prov. Fianarantsoa" typeStatus="lectotype">
<emphasis id="B917C4233A10FFBAE728F925FC0C2B94" bold="true" box="[862,1000,1666,1690]" pageId="8" pageNumber="35">
<typeStatus id="54D8A6933A10FFBAE728F925FC0C2B94" box="[862,1000,1666,1690]" pageId="8" pageNumber="35" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated by
<bibRefCitation id="EFF265C03A10FFBAE0C5F925FA572B94" author="SLEUMER, H." box="[1203,1459,1666,1690]" pageId="8" pageNumber="35" pagination="239 - 328" refId="ref21737" refString="SLEUMER, H. (1973). Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes). Bull. Jard. Bot. Natl. Belg. 43: 239 - 328." type="journal article" year="1973">SLEUMER, 1973: 308</bibRefCitation>
):
<emphasis id="B917C4233A10FFBAE728F905FB0C2BB7" bold="true" box="[862,1256,1697,1722]" pageId="8" pageNumber="35">
<collectingCountry id="F37458A13A10FFBAE728F905FC192BB4" box="[862,1021,1698,1722]" name="Madagascar" pageId="8" pageNumber="35" value="Madagascar">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49A7D6D33A10FFBAE07CF905FB072BB7" box="[1034,1251,1697,1722]" pageId="8" pageNumber="35">Prov. Fianarantsoa</collectingRegion>
:
</emphasis>
<location id="8EBC4EEA3A10FFBAE098F905FABD2BB4" LSID="urn:lsid:plazi:treatment:03CAA9273A10FFB8E74EF990FA262FF7:8EBC4EEA3A10FFBAE098F905FABD2BB4" box="[1262,1369,1698,1722]" country="Madagascar" name="Vondrozo" pageId="8" pageNumber="35" stateProvince="Prov. Fianarantsoa">Vondrozo</location>
,
<location id="8EBC4EEA3A10FFBAE113F905FBEE2BD4" LSID="urn:lsid:plazi:treatment:03CAA9273A10FFB8E74EF990FA262FF7:8EBC4EEA3A10FFBAE113F905FBEE2BD4" country="Madagascar" name="province de Farafangana" pageId="8" pageNumber="35" stateProvince="Prov. Fianarantsoa">province de Farafangana</location>
, s.d., fl.,
<emphasis id="B917C4233A10FFBAE018F965FB132BD7" box="[1134,1271,1729,1753]" italics="true" pageId="8" pageNumber="35">
<collectorName id="26967DE73A10FFBAE018F965FB5C2BD7" box="[1134,1208,1730,1753]" pageId="8" pageNumber="35">Decary</collectorName>
5469
</emphasis>
(
<collectionCode id="ED7280F43A10FFBAE17EF965FAFD2BD4" box="[1288,1305,1730,1754]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="8" pageNumber="35" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBC5B04A3A10FFBAE15EF966FA572BD7" box="[1320,1459,1729,1753]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="8" pageNumber="35" type="Herbarium">P04679983</specimenCode>
]!
</materialsCitation>
;
<materialsCitation id="3B0B126C3A10FFBAE728F945FC382A14" ID-GBIF-Occurrence="3464757475" collectionCode="K, L" country="Madagascar" pageId="8" pageNumber="35" specimenCode="K000231481, L0010965" specimenCount="1" stateProvince="Prov. Fianarantsoa">
isolecto-:
<collectionCode id="ED7280F43A10FFBAE7B8F945FC052BF7" box="[974,993,1762,1785]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="8" pageNumber="35" type="Herbarium">K</collectionCode>
[
<specimenCode id="DBC5B04A3A10FFBAE786F945FB632BF4" box="[1008,1159,1762,1786]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="8" pageNumber="35" type="Herbarium">K000231481</specimenCode>
] image seen,
<collectionCode id="ED7280F43A10FFBAE157F945FAD72BF7" box="[1313,1331,1762,1785]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="8" pageNumber="35" type="Herbarium">L</collectionCode>
[
<specimenCode id="DBC5B04A3A10FFBAE133F945FA5F2BF7" box="[1349,1467,1761,1785]" collectionCode="L" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="8" pageNumber="35" type="Herbarium">L0010965</specimenCode>
] image seen)
</materialsCitation>
.
<materialsCitation id="3B0B126C3A10FFBAE791F8A6FA932A37" ID-GBIF-Occurrence="3464757402" collectingDate="25. VIII" collectionCode="P" collectorName="Louvel" country="Madagascar" location="Tampina" pageId="8" pageNumber="35" specimenCode="P04679987" specimenCount="1" stateProvince="Prov. Toamasina" typeStatus="syntype">
<emphasis id="B917C4233A10FFBAE791F8A6FA262A17" bold="true" box="[999,1474,1793,1818]" pageId="8" pageNumber="35">
<typeStatus id="54D8A6933A10FFBAE791F8A6FBB42A17" box="[999,1104,1793,1817]" pageId="8" pageNumber="35" type="syntype">Syntypus</typeStatus>
:
<collectingCountry id="F37458A13A10FFBAE02CF8A5FB132A14" box="[1114,1271,1794,1818]" name="Madagascar" pageId="8" pageNumber="35" value="Madagascar">MADAGASCAR</collectingCountry>
.
<collectingRegion id="49A7D6D33A10FFBAE175F8A5FAD82A14" box="[1283,1340,1794,1818]" pageId="8" pageNumber="35">Prov.</collectingRegion>
Toamasina:
</emphasis>
<location id="8EBC4EEA3A10FFBAE728F885FC5B2A34" LSID="urn:lsid:plazi:treatment:03CAA9273A10FFB8E74EF990FA262FF7:8EBC4EEA3A10FFBAE728F885FC5B2A34" box="[862,959,1826,1850]" country="Madagascar" municipality="Louvel" name="Tampina" pageId="8" pageNumber="35" stateProvince="Prov. Toamasina">Tampina</location>
,
<date id="FFDD3EF13A10FFBAE7BFF886FBF82A37" box="[969,1052,1825,1849]" pageId="8" pageNumber="35">
<collectingDate id="EF99C7193A10FFBAE7BFF886FBF82A37" box="[969,1052,1825,1849]" pageId="8" pageNumber="35">25.VIII</collectingDate>
</date>
, fr.,
<emphasis id="B917C4233A10FFBAE03CF885FB542A34" box="[1098,1200,1826,1850]" italics="true" pageId="8" pageNumber="35">
<collectorName id="26967DE73A10FFBAE03CF885FB742A34" box="[1098,1168,1826,1850]" pageId="8" pageNumber="35">Louvel</collectorName>
53
</emphasis>
(
<collectionCode id="ED7280F43A10FFBAE0C9F885FB342A34" box="[1215,1232,1826,1850]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="8" pageNumber="35" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBC5B04A3A10FFBAE0ABF886FA812A37" box="[1245,1381,1825,1849]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="8" pageNumber="35" type="Herbarium">P04679987</specimenCode>
]!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3794BBA3A10FFB8E728F8C5FD992E01" lastPageId="10" lastPageNumber="37" pageId="8" pageNumber="35" type="description">
<paragraph id="8BDC18313A10FFB8E728F8C5FD992E01" blockId="8.[824,1474,1890,1946]" lastBlockId="10.[136,787,279,783]" lastPageId="10" lastPageNumber="37" pageId="8" pageNumber="35">
<emphasis id="B917C4233A10FFBAE728F8C5FC692A77" box="[862,909,1890,1913]" italics="true" pageId="8" pageNumber="35">Tree</emphasis>
to
<quantity id="4C9BB5D43A10FFBAE7C3F8C5FC0B2A77" box="[949,1007,1890,1914]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="35" unit="m" value="13.0">13 m</quantity>
tall,
<quantity id="4C9BB5D43A10FFBAE05CF8C5FB942A77" box="[1066,1136,1890,1914]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="35" unit="cm" value="30.0">30 cm</quantity>
dbh; young twigs moderately pubescent with minute hairs or glabrous (glabrate).
<emphasis id="B917C4233A10FFBAE102F825FA262A97" box="[1396,1474,1922,1945]" italics="true" pageId="8" pageNumber="35">Leaves:</emphasis>
petiole 1-7(-10) mm long, glabrous (minutely pubescent); blade narrowly elliptical to elliptical (oblanceolate) or obovate to elliptical, (1.8-)2.7-6.8 × 0.7-2.6(-3) cm; base cuneate to attenuate or convex; margins serrulate to crenate-serrate, shallowly wavy or subentire, sometimes slightly revolute; apex rounded-acute to rounded (slightly acuminate, rounded-cuspidate, aberrantly emarginate).
<emphasis id="B917C4233A12FFB8E5C0FE70FDD22CE1" box="[438,566,471,495]" italics="true" pageId="10" pageNumber="37">Inflorescences</emphasis>
spicate, (2-)4-7(-10) cm long, minutely pubescent or glabrous; flowers in severalflowered glomerules, sessile; bracts transversely oblong, often irregular; bracteoles reduced to minute teeth, rapidly caducous.
<emphasis id="B917C4233A12FFB8E4FEFDF0FF3D2F61" box="[136,217,599,623]" italics="true" pageId="10" pageNumber="37">Flowers</emphasis>
6-7(-8)-merous, greenish (corolla pale yellow); sepals deltoid to ovate,
<quantity id="4C9BB5D43A12FFB8E534FDD0FE212F80" box="[322,453,631,655]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="37" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1-0.3 mm</quantity>
long; petals transversely (very broadly) ovate (very broadly oblong-ovate), 0.5-0.7(-0.8) mm long, glabrous, apex rounded (rounded-obtuse); filaments 0.2-0.4(-0.6) mm long; upper surface of ovary densely shortpubescent; styles (2-)3, 0.2-0.3(-0.5) mm long.
</paragraph>
</subSubSection>
<caption id="DF1C48B93A11FFBBE407F8DDFDB32AA9" ID-DOI="http://doi.org/10.5281/zenodo.5722140" ID-Zenodo-Dep="5722140" httpUri="https://zenodo.org/record/5722140/files/figure.png" pageId="9" pageNumber="36" startId="9.[113,140,1914,1933]" targetBox="[227,1343,416,1809]" targetPageId="9">
<paragraph id="8BDC18313A11FFBBE407F8DDFDB32AA9" blockId="9.[113,744,1914,1959]" pageId="9" pageNumber="36">
<emphasis id="B917C4233A11FFBBE407F8DDFF4B2A83" bold="true" box="[113,175,1914,1933]" pageId="9" pageNumber="36">Fig. 2. -</emphasis>
<taxonomicName id="4C6363B23A11FFBBE4C4F8DDFDDC2A83" authority="Appleq. A. Flowering" box="[178,568,1914,1933]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="masoalense">
<emphasis id="B917C4233A11FFBBE4C4F8DDFE992A83" box="[178,381,1914,1933]" italics="true" pageId="9" pageNumber="36">Homalium masoalense</emphasis>
Appleq.
<emphasis id="B917C4233A11FFBBE5BCF8DDFE392A83" bold="true" box="[458,477,1914,1933]" pageId="9" pageNumber="36">A.</emphasis>
Flowering
</taxonomicName>
branch;
<emphasis id="B917C4233A11FFBBE6F3F8DDFD732A83" bold="true" box="[645,663,1914,1933]" pageId="9" pageNumber="36">B.</emphasis>
Flowers. [
<emphasis id="B917C4233A11FFBBE400F833FEEB2AA9" box="[118,271,1940,1959]" italics="true" pageId="9" pageNumber="36">Schatz et al.3367,</emphasis>
TAN] [Drawings: R.L. Andriamiarisoa]
</paragraph>
</caption>
<subSubSection id="C3794BBA3A12FFB8E4D8FC9FFDC52BA1" pageId="10" pageNumber="37" type="discussion">
<paragraph id="8BDC18313A12FFB8E4D8FC9FFDC52BA1" blockId="10.[136,787,823,1935]" pageId="10" pageNumber="37">
<emphasis id="B917C4233A12FFB8E4D8FC9FFE112E41" box="[174,501,823,847]" italics="true" pageId="10" pageNumber="37">
Notes.
<taxonomicName id="4C6363B23A12FFB8E570FC90FE112E41" box="[262,501,823,847]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="species" species="moniliforme">Homalium moniliforme</taxonomicName>
</emphasis>
displays geographic variation in its vegetative morphology and ecological preferences. Collections from the southern provinces of Toliara and Fianarantsoa occur consistently at low altitudes on sand and have small, short-petioled, often narrowly elliptical to elliptical leaves and minutely pubescent twigs. This
<taxonomicName id="4C6363B23A12FFB8E6F4FC70FCF52EE1" box="[642,785,983,1007]" form="extends" pageId="10" pageNumber="37" rank="form">form extends</taxonomicName>
to coastal forests in Toamasina province. Inland collections from moderate-elevation humid forests, which occur on more diverse substrates, range from Antsiranana to Toamasina provinces and rarely Fianarantsoa province. These have somewhat larger leaves that are longer-petioled and usually mostly obovate, less often elliptical; the twigs are glabrous or glabrate, rarely papillate at the extreme apices. The two geographically distinct forms are usually readily distinguishable. It therefore seems appropriate to recognize them as subspecies. The
<typeStatus id="54D8A6933A12FFB8E4FEFAB0FF082821" box="[136,236,1303,1327]" pageId="10" pageNumber="37" type="lectotype">lectotype</typeStatus>
is from Fianarantsoa province at a low-moderate elevation, in a region where no other material of the species has been collected. Its morphology is somewhat intermediate between the two forms, with mostly narrow, oblanceolate leaves, and the leaves dry relatively pale, which is much more common in the small-leaved subspecies (though that is not a fully reliable character in any related group observed to date). However, it is overall more similar to material from the moderate-elevation populations, for which it would represent a substantial range extension. These populations are therefore herein recognized as
<taxonomicName id="4C6363B23A12FFB8E51DF9F0FDCB2B61" box="[363,559,1623,1647]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="subSpecies" species="moniliforme" subSpecies="moniliforme">
subsp.
<emphasis id="B917C4233A12FFB8E5C5F9F0FDCB2B61" box="[435,559,1623,1647]" italics="true" pageId="10" pageNumber="37">moniliforme</emphasis>
</taxonomicName>
, and the coastal subspecies is described below as
<taxonomicName id="4C6363B23A12FFB8E5C1F9D0FD482B81" authority="Appleq." authorityName="Appleq." box="[439,684,1655,1679]" pageId="10" pageNumber="37" rank="subSpecies" subSpecies="littoralis">
subsp.
<emphasis id="B917C4233A12FFB8E58AF9D0FDB42B81" box="[508,592,1655,1679]" italics="true" pageId="10" pageNumber="37">littoralis</emphasis>
Appleq.
</taxonomicName>
, to which the other
<typeStatus id="54D8A6933A12FFB8E486F930FEA52BA1" box="[240,321,1687,1711]" pageId="10" pageNumber="37" type="syntype">syntype</typeStatus>
<emphasis id="B917C4233A12FFB8E53FF930FE262BA1" box="[329,450,1687,1711]" italics="true" pageId="10" pageNumber="37">(Louvel 53)</emphasis>
belongs.
</paragraph>
</subSubSection>
<subSubSection id="C3794BBA3A12FFB8E4D8F910FA5E2C41" pageId="10" pageNumber="37" type="reference_group">
<paragraph id="8BDC18313A12FFB8E4D8F910FA5E2C41" blockId="10.[136,787,823,1935]" lastBlockId="10.[824,1475,279,335]" pageId="10" pageNumber="37">
The original publication of
<taxonomicName id="4C6363B23A12FFB8E5A6F910FD5B2BC1" ID-CoL="3MCNY" authority="H. Perrier" box="[464,703,1719,1743]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="species" species="moniliforme">
<emphasis id="B917C4233A12FFB8E5A6F910FD5B2BC1" box="[464,703,1719,1743]" italics="true" pageId="10" pageNumber="37">Homalium moniliforme</emphasis>
</taxonomicName>
did not explicitly cite specimens, but did enumerate the vernacular names provided on the labels of both
<emphasis id="B917C4233A12FFB8E660F95FFD7C2A01" box="[534,664,1783,1807]" italics="true" pageId="10" pageNumber="37">Decary 5469</emphasis>
and
<emphasis id="B917C4233A12FFB8E6BAF950FF472A21" italics="true" pageId="10" pageNumber="37">Louvel 53</emphasis>
, which were the only two collections cited in the Flora treatment six years later (
<bibRefCitation id="EFF265C03A12FFB8E5D8F890FCE22A41" author="PERRIER DE LA BATHIE, H." box="[430,774,1841,1871]" pageId="10" pageNumber="37" pagination="140" refId="ref21660" refString="PERRIER DE LA BATHIE, H. (1946). Flacourtiacees. In: HUMBERT, H. (ed.), Fl. Madagascar Comores 140." type="book chapter" year="1946">PERRIER DE LA BÂTHIE, 1946</bibRefCitation>
).
<bibRefCitation id="EFF265C03A12FFB8E4FEF8F0FEDF2A61" author="SLEUMER, H." box="[136,315,1879,1903]" pageId="10" pageNumber="37" pagination="239 - 328" refId="ref21737" refString="SLEUMER, H. (1973). Revision du genre Homalium Jacq. (Flacourtiacees) en Afrique (y compris Madagascar et les Mascareignes). Bull. Jard. Bot. Natl. Belg. 43: 239 - 328." type="journal article" year="1973">SLEUMER (1973)</bibRefCitation>
therefore termed them
<typeStatus id="54D8A6933A12FFB8E64FF8F0FD722A61" box="[569,662,1879,1903]" pageId="10" pageNumber="37" type="syntype">syntypes</typeStatus>
. That interpretation could be questioned, since they were only indirectly cited, but they are certainly original material, and he appropriately designated the P duplicate of
<emphasis id="B917C4233A12FFB8E0D8FE9FFAD52C41" box="[1198,1329,311,335]" italics="true" pageId="10" pageNumber="37">Decary 5469</emphasis>
as
<typeStatus id="54D8A6933A12FFB8E122FE90FA522C41" box="[1364,1462,311,335]" pageId="10" pageNumber="37" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3794BBA3A12FFB8E74EFE33FA262FF7" pageId="10" pageNumber="37" type="key">
<paragraph id="8BDC18313A12FFB8E74EFE33FC042CDE" blockId="10.[824,1320,404,464]" pageId="10" pageNumber="37">
<heading id="D094AF5D3A12FFB8E74EFE33FC042CDE" bold="true" fontSize="11" level="1" pageId="10" pageNumber="37" reason="1">
<emphasis id="B917C4233A12FFB8E74EFE33FC042CDE" bold="true" pageId="10" pageNumber="37">
Key to the subspecies of
<taxonomicName id="4C6363B23A12FFB8E0EAFE33FC042CDE" ID-CoL="3MCNY" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="species" species="moniliforme">Homalium moniliforme</taxonomicName>
</emphasis>
</heading>
</paragraph>
<key id="FFF24F003A12FFB8E74EFE45FA262FF7" pageId="10" pageNumber="37">
<keyStep id="309700943A12FFB8E74EFE45FA262FF7" pageId="10" pageNumber="37">
<paragraph id="8BDC18313A12FFB8E74EFE45FA262F77" blockId="10.[824,1474,482,762]" pageId="10" pageNumber="37">
<keyLead id="3092BB043A12FFB8E74EFE45FA262F77" pageId="10" pageNumber="37">
1. Young twigs glabrous (glabrate, minutely papillate at the apex); petiole 2-7(-10) mm long; leaf blade obovate to elliptical, narrowly elliptical, or oblanceolate, 3-6.8 × (1.1-)1.3- 2.6(-3) cm; mid- to low-elevation humid forests ................. ...................................................... 6a.
<taxonomicName id="4C6363B23A12FFB8E08FFDC5FA262F77" ID-CoL="5HYXQ" box="[1273,1474,609,634]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="subSpecies" species="moniliforme" subSpecies="moniliforme">
subsp.
<emphasis id="B917C4233A12FFB8E136FDC6FA262F77" bold="true" box="[1344,1474,609,633]" italics="true" pageId="10" pageNumber="37">moniliforme</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDC18313A12FFB8E74EFD25FA262FF7" blockId="10.[824,1474,482,762]" pageId="10" pageNumber="37">
<keyLead id="3092BB043A12FFB8E74EFD25FA262FF7" pageId="10" pageNumber="37">
1a. Young twigs moderately minutely pubescent; petiole
<quantity id="4C9BB5D43A12FFB8E728FD05FC532FB7" box="[862,951,674,698]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="37" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mm</quantity>
long; leaf blade narrowly elliptical to elliptical (oblanceolate), (1.8-)2.7-4.7(-5.6) × 0.7-1.6(-1.9) cm; littoral or low-elevation coastal forests .... 6b.
<taxonomicName id="4C6363B23A12FFB8E15EFD45FA262FF7" ID-CoL="5HYXP" box="[1320,1474,737,762]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="subSpecies" species="moniliforme" subSpecies="littorale">
subsp.
<emphasis id="B917C4233A12FFB8E119FD46FA262FF7" bold="true" box="[1391,1474,737,761]" italics="true" pageId="10" pageNumber="37">littorale</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>