310 lines
38 KiB
XML
310 lines
38 KiB
XML
<document id="38201659CD6A3DFBD06AC1B12C467865" ID-DOI="10.5852/ejt.2023.916.2381" ID-ISSN="2118-9773" ID-Zenodo-Dep="10453460" ID-ZooBank="0DC587F6-9DAA-4F6E-BA2A-AD528990BA24" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704271610563" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2023" docId="03ADDB11FF86FFA60A58F93FFA2DFC67" docLanguage="en" docName="EJT.2023.916.1-85.pdf" docOrigin="European Journal of Taxonomy 916 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2381/10463" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Aciandrena) nitidilabris Perez 1895" docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FF94A369FF88FFB6085AFFA3FE1EFFB2" masterDocTitle="Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="15" updateTime="1704272290945" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BA62634649BB4741E2F9F2FD36E21C62">Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="F23D02D9B02ED37A0A5CB5F2893E379C">Wood, Thomas James</mods:namePart>
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<mods:title id="67F9BBF8966FA258DC31D30BF19B3A9B">European Journal of Taxonomy</mods:title>
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<mods:part id="E7E12C0F163867C0E5B4F6D19C5D6B0E">
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<mods:date id="84240635D29DF84FE10B2ADF1535D888">2023</mods:date>
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<mods:number id="FC667E7E8000DF4315E997876D029BA9">2023-12-21</mods:number>
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<mods:identifier id="01F6F2275A4CFBC361409757BA72B752" type="DOI">10.5852/ejt.2023.916.2381</mods:identifier>
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<subSubSection id="C31E398CFF86FFB80A58F93FFA2CF905" box="[514,1074,1692,1719]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BBB6A07FF86FFB80A58F93FFA2CF905" blockId="14.[514,1074,1692,1754]" box="[514,1074,1692,1719]" pageId="14" pageNumber="15">
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<heading id="D0F3DD6BFF86FFB80A58F93FFA2CF905" box="[514,1074,1692,1719]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
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<taxonomicName id="4C041184FF86FFB80A58F93FFA2CF905" ID-CoL="66N5W" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[514,1074,1692,1719]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nitidilabris" subGenus="Aciandrena">
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<emphasis id="B970B615FF86FFB80A58F93FFDBEF905" bold="true" box="[514,928,1692,1719]" pageId="14" pageNumber="15">
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<emphasis id="B970B615FF86FFB80A58F93FFC72F905" bold="true" box="[514,620,1692,1719]" italics="true" pageId="14" pageNumber="15">Andrena</emphasis>
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(
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<emphasis id="B970B615FF86FFB80A27F93FFD15F905" bold="true" box="[637,779,1692,1719]" italics="true" pageId="14" pageNumber="15">Aciandrena</emphasis>
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)
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<emphasis id="B970B615FF86FFB80B46F93FFDBEF905" bold="true" box="[796,928,1692,1719]" italics="true" pageId="14" pageNumber="15">nitidilabris</emphasis>
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</emphasis>
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<bibRefCitation id="EF9517F6FF86FFB80BFDF93EFA2CF905" author="Perez J." box="[935,1074,1692,1719]" pageId="14" pageNumber="15" refId="ref40126" refString="Perez J. 1895. Especes nouvelles de Melliferes de Barbarie. (Diagnoses preliminaires). Gounouilhou, Bordeaux." type="book" year="1895">Pérez, 1895</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31E398CFF86FFB80AABF963FD5CF968" box="[753,834,1728,1754]" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BBB6A07FF86FFB80AABF963FD5CF968" blockId="14.[514,1074,1692,1754]" box="[753,834,1728,1754]" pageId="14" pageNumber="15">
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||
<figureCitation id="133F7682FF86FFB80AABF963FD5CF968" box="[753,834,1728,1754]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31E398CFF86FFB808E7F8A2FB01F88D" pageId="14" pageNumber="15" type="reference_group">
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||
<paragraph id="8BBB6A07FF86FFB808E7F8A2FA0DF8A9" blockId="14.[189,1311,1793,1857]" box="[189,1043,1793,1820]" pageId="14" pageNumber="15">
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||
<treatmentCitationGroup id="AB144D29FF86FFB808E7F8A2FA0DF8A9" box="[189,1043,1793,1820]" pageId="14" pageNumber="15">
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<treatmentCitation id="0AA54C16FF86FFB808E7F8A2FD11F8A9" author="Perez J." box="[189,783,1793,1820]" page="46" pageId="14" pageNumber="15" year="1895">
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<taxonomicName id="4C041184FF86FFB808E7F8A2FD11F8A9" ID-CoL="66N5W" authority="Perez, 1895: 46" authorityName="Perez" authorityPageNumber="46" authorityYear="1895" box="[189,783,1793,1820]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nitidilabris" subGenus="Aciandrena">
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||
<emphasis id="B970B615FF86FFB808E7F8A2FF3FF8A9" box="[189,289,1793,1819]" italics="true" pageId="14" pageNumber="15">Andrena</emphasis>
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||
(
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||
<emphasis id="B970B615FF86FFB80968F8A2FFA4F8A9" box="[306,442,1793,1819]" italics="true" pageId="14" pageNumber="15">Aciandrena</emphasis>
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||
)
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||
<emphasis id="B970B615FF86FFB80991F8A2FC53F8A9" box="[459,589,1793,1819]" italics="true" pageId="14" pageNumber="15">nitidilabris</emphasis>
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||
<bibRefCitation id="EF9517F6FF86FFB80A0FF8A2FD11F8A9" author="Perez J." box="[597,783,1793,1820]" pageId="14" pageNumber="15" refId="ref40126" refString="Perez J. 1895. Especes nouvelles de Melliferes de Barbarie. (Diagnoses preliminaires). Gounouilhou, Bordeaux." type="book" year="1895">Pérez, 1895: 46</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
,
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||
<specimenCount id="9D02A18EFF86FFB80B41F8A1FD2FF8AE" box="[795,817,1794,1820]" count="1" pageId="14" pageNumber="15" type="female">♀</specimenCount>
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||
(
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||
<collectingCountry id="F3132A97FF86FFB80B1AF8A2FD83F8A9" box="[832,925,1793,1819]" name="Tunisia" pageId="14" pageNumber="15">Tunisia</collectingCountry>
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: MNHN).
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BBB6A07FF86FFB808E7F887FB01F88D" blockId="14.[189,1311,1793,1857]" box="[189,1311,1828,1857]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="AB144D29FF86FFB808E7F887FB01F88D" box="[189,1311,1828,1857]" pageId="14" pageNumber="15">
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<treatmentCitation id="0AA54C16FF86FFB808E7F887FD91F88D" author="Warncke K." box="[189,911,1828,1855]" page="45" pageId="14" pageNumber="15" year="1974">
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||
<taxonomicName id="4C041184FF86FFB808E7F887FD91F88D" authority="Warncke, 1974: 45" authorityName="Warncke" authorityPageNumber="45" authorityYear="1974" box="[189,911,1828,1855]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="montarca" subGenus="Graecandrena" subSpecies="parva">
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||
<emphasis id="B970B615FF86FFB808E7F887FF3FF88C" box="[189,289,1828,1854]" italics="true" pageId="14" pageNumber="15">Andrena</emphasis>
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||
(
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||
<emphasis id="B970B615FF86FFB80968F887FFC2F88C" box="[306,476,1828,1854]" italics="true" pageId="14" pageNumber="15">Graecandrena</emphasis>
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||
)
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||
<emphasis id="B970B615FF86FFB809B7F886FCB6F88C" box="[493,680,1829,1854]" italics="true" pageId="14" pageNumber="15">montarca parva</emphasis>
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||
<bibRefCitation id="EF9517F6FF86FFB80AF5F887FD91F88D" author="Warncke K." box="[687,911,1828,1855]" pageId="14" pageNumber="15" pagination="3 - 54" refId="ref41080" refString="Warncke K. 1974. Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem zoologischen Museum in Berlin 50: 3 - 54." type="journal article" year="1974">Warncke, 1974: 45</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
,
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||
<specimenCount id="9D02A18EFF86FFB80BC1F886FDB3F88D" box="[923,941,1829,1855]" count="1" pageId="14" pageNumber="15" type="female">♀</specimenCount>
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||
<specimenCount id="9D02A18EFF86FFB80BF7F886FDD9F88D" box="[941,967,1829,1855]" count="1" pageId="14" pageNumber="15" type="male">♂</specimenCount>
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||
(
|
||
<collectingCountry id="F3132A97FF86FFB80B8DF887FA2AF88C" box="[983,1076,1828,1854]" name="Tunisia" pageId="14" pageNumber="15">Tunisia</collectingCountry>
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||
: OÖLM).
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<emphasis id="B970B615FF86FFB80CF4F887FB01F88D" bold="true" box="[1198,1311,1828,1855]" pageId="14" pageNumber="15">Syn. nov.</emphasis>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31E398CFF86FFB808E7F8C5FD99F866" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="8BBB6A07FF86FFB808E7F8C5FFB4F832" blockId="14.[189,426,1894,1920]" box="[189,426,1894,1920]" pageId="14" pageNumber="15">
|
||
<heading id="D0F3DD6BFF86FFB808E7F8C5FFB4F832" bold="true" box="[189,426,1894,1920]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
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||
<emphasis id="B970B615FF86FFB808E7F8C5FFB4F832" bold="true" box="[189,426,1894,1920]" pageId="14" pageNumber="15">Material examined</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BBB6A07FF86FFB808E7F836FD99F866" blockId="14.[189,1398,1940,2004]" pageId="14" pageNumber="15">
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||
<materialsCitation id="3B6C605AFF86FFB808E7F836FAAFF81D" box="[189,1201,1940,1967]" collectionCode="MNHN" country="Tunisia" location="Kerouan" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Kairouan" typeStatus="lectotype">
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||
<collectingCountry id="F3132A97FF86FFB808E7F836FF2CF81D" box="[189,306,1941,1967]" name="Tunisia" pageId="14" pageNumber="15">TUNISIA</collectingCountry>
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||
•
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||
<specimenCount id="9D02A18EFF86FFB80916F836FF7CF81D" box="[332,354,1941,1967]" count="1" pageId="14" pageNumber="15" type="female">♀</specimenCount>
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||
,
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||
<typeStatus id="54BFD4A5FF86FFB8092BF837FFC3F81C" box="[369,477,1940,1966]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
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||
of
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||
<taxonomicName id="4C041184FF86FFB80A5FF836FCB6F81C" authorityName="Perez" authorityYear="1895" box="[517,680,1940,1967]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF86FFB80A5FF836FCB6F81C" box="[517,680,1940,1967]" italics="true" pageId="14" pageNumber="15">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133F7682FF86FFB80AE2F837FD5BF81D" box="[696,837,1940,1967]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="14" pageNumber="15">Fig. 11A–F</figureCitation>
|
||
);
|
||
<location id="8EDB3CDCFF86FFB80B00F836FDA1F81D" LSID="urn:lsid:plazi:treatment:03ADDB11FF86FFA60A58F93FFA2DFC67:8EDB3CDCFF86FFB80B00F836FDA1F81D" box="[858,959,1941,1967]" country="Tunisia" name="Kerouan" pageId="14" pageNumber="15" stateProvince="Kairouan">Kérouan</location>
|
||
[
|
||
<collectingRegion id="49C0A4E5FF86FFB80B97F837FA5AF81C" box="[973,1092,1940,1966]" country="Tunisia" name="Kairouan" pageId="14" pageNumber="15">Kairouan</collectingRegion>
|
||
];
|
||
<collectionCode id="ED15F2C2FF86FFB80C0DF836FAAFF81D" box="[1111,1201,1941,1967]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B6C605AFF86FFB80C91F836FD9DF866" collectionCode="OOLM" country="Tunisia" location="Gafsa" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Gafsa" typeStatus="holotype">
|
||
<specimenCount id="9D02A18EFF86FFB80C91F836FAFFF81D" box="[1227,1249,1941,1967]" count="1" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
,
|
||
<typeStatus id="54BFD4A5FF86FFB80CAAF837FB49F81C" box="[1264,1367,1940,1966]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C041184FF86FFB808E7F81BFF86F863" authorityName="Warncke" authorityYear="1974" box="[189,408,1976,2002]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="montarca" subSpecies="parva">
|
||
<emphasis id="B970B615FF86FFB808E7F81BFF86F863" box="[189,408,1976,2002]" italics="true" pageId="14" pageNumber="15">A. montarca parva</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<collectingRegion id="49C0A4E5FF86FFB809F2F81BFFEEF860" box="[424,496,1976,2002]" country="Tunisia" name="Gafsa" pageId="14" pageNumber="15">Gafsa</collectingRegion>
|
||
; 17 Mar. [year unknown];
|
||
<collectionCode id="ED15F2C2FF86FFB80B73F81BFD9DF866" box="[809,899,1976,2004]" pageId="14" pageNumber="15">OÖLM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E398CFF87FFA608E7FEAAFDC2FCDF" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="8BBB6A07FF87FFB908E7FEAAFF31FE91" blockId="15.[189,303,265,291]" box="[189,303,265,291]" pageId="15" pageNumber="16">
|
||
<heading id="D0F3DD6BFF87FFB908E7FEAAFF31FE91" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B970B615FF87FFB908E7FEAAFF31FE91" bold="true" box="[189,303,265,291]" pageId="15" pageNumber="16">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBB6A07FF87FFA608E7FE94FF62FED8" blockId="15.[188,1399,311,478]" lastBlockId="16.[188,1401,265,362]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
Inspection of the MNHN collection revealed that the
|
||
<typeStatus id="54BFD4A5FF87FFB90B0BFE94FDA3FEE3" box="[849,957,311,337]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C041184FF87FFB90BB6FE94FA8DFEE3" authorityName="Perez" authorityYear="1895" box="[1004,1171,311,337]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF87FFB90BB6FE94FA8DFEE3" box="[1004,1171,311,337]" italics="true" pageId="15" pageNumber="16">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133F7682FF87FFB90CF2FE94FB2AFEE3" box="[1192,1332,311,337]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="15" pageNumber="16">Fig. 11A–F</figureCitation>
|
||
) was correctly designated by
|
||
<bibRefCitation id="EF9517F6FF87FFB90980FEF9FC85FEC6" author="Warncke K." box="[474,667,346,372]" pageId="15" pageNumber="16" pagination="171 - 318" refId="ref41031" refString="Warncke K. 1967. Beitrag zur Klarung palaarktischer Andrena - Arten. Eos 43: 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
|
||
, but that the species was misinterpreted and later redescribed as
|
||
<taxonomicName id="4C041184FF87FFB908BFFEDDFFD1FE25" authorityName="Warncke" authorityYear="1974" box="[229,463,382,408]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="montarca" subSpecies="parva">
|
||
<emphasis id="B970B615FF87FFB908BFFEDDFFD1FE25" box="[229,463,382,408]" italics="true" pageId="15" pageNumber="16">A. montarca parva</emphasis>
|
||
</taxonomicName>
|
||
(type photographs at https://www.zobodat.at/belege.php). The
|
||
<typeStatus id="54BFD4A5FF87FFB90CB9FEDEFB51FE25" box="[1251,1359,381,407]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C041184FF87FFB908E7FE02FF41FE09" authorityName="Perez" authorityYear="1895" box="[189,351,417,443]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF87FFB908E7FE02FF41FE09" box="[189,351,417,443]" italics="true" pageId="15" pageNumber="16">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
has the clypeus weakly but distinctively domed and weakly shagreened, therefore weakly shining (
|
||
<figureCitation id="133F7682FF87FFB90978FE67FF92FE6C" box="[290,396,452,478]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="15" pageNumber="16">Fig. 11C</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C041184FF87FFB9099BFE67FC7DFE6C" authorityName="Perez" authorityYear="1895" box="[449,611,452,478]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF87FFB9099BFE67FC7DFE6C" box="[449,611,452,478]" italics="true" pageId="15" pageNumber="16">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
auctorum the clypeus is strongly flattened and shagreened over the majority of its area (cf.
|
||
<taxonomicName id="4C041184FF98FFA60996FEAAFCF5FE96" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[460,747,265,292]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="B970B615FF98FFA60996FEAAFC47FE91" box="[460,601,265,291]" italics="true" pageId="16" pageNumber="17">A. verticalis</emphasis>
|
||
<bibRefCitation id="EF9517F6FF98FFA60A3AFEAAFCF5FE96" author="Perez J." box="[608,747,265,292]" pageId="16" pageNumber="17" refId="ref40126" refString="Perez J. 1895. Especes nouvelles de Melliferes de Barbarie. (Diagnoses preliminaires). Gounouilhou, Bordeaux." type="book" year="1895">Pérez, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
) and only becomes shiny at its apical margin. The taxon referred to as
|
||
<taxonomicName id="4C041184FF98FFA60932FE8EFC13FEF4" authorityName="Perez" authorityYear="1895" box="[360,525,300,327]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF98FFA60932FE8EFC13FEF4" box="[360,525,300,327]" italics="true" pageId="16" pageNumber="17">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
by Warncke is actually undescribed, and is newly described below as
|
||
<taxonomicName id="4C041184FF98FFA60D05FE8EFF04FEDB" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pisantyi" status="sp. nov.">
|
||
<emphasis id="B970B615FF98FFA60D05FE8EFF04FEDB" italics="true" pageId="16" pageNumber="17">A. pisantyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2430B6EFF98FFA6097BFEF3FF62FED8" box="[289,380,336,362]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF7B3A8FFF87FFB908E7F825FF54F854" ID-DOI="http://doi.org/10.5281/zenodo.10453486" ID-Zenodo-Dep="10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1926,1952]" targetBox="[189,1399,533,1891]" targetPageId="15">
|
||
<paragraph id="8BBB6A07FF87FFB908E7F825FF54F854" blockId="15.[188,1399,1926,2022]" pageId="15" pageNumber="16">
|
||
<emphasis id="B970B615FF87FFB908E7F825FF02F812" bold="true" box="[189,284,1926,1952]" pageId="15" pageNumber="16">Fig. 11.</emphasis>
|
||
<taxonomicName id="4C041184FF87FFB9097DF825FD63F813" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[295,893,1926,1953]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nitidilabris" subGenus="Graecandrena">
|
||
<emphasis id="B970B615FF87FFB9097DF825FF95F812" box="[295,395,1926,1952]" italics="true" pageId="15" pageNumber="16">Andrena</emphasis>
|
||
(
|
||
<emphasis id="B970B615FF87FFB909FAF825FC54F812" box="[416,586,1926,1952]" italics="true" pageId="15" pageNumber="16">Graecandrena</emphasis>
|
||
)
|
||
<emphasis id="B970B615FF87FFB90A05F825FCFFF812" box="[607,737,1926,1952]" italics="true" pageId="15" pageNumber="16">nitidilabris</emphasis>
|
||
<bibRefCitation id="EF9517F6FF87FFB90AB6F825FD63F813" author="Perez J." box="[748,893,1926,1953]" pageId="15" pageNumber="16" refId="ref40126" refString="Perez J. 1895. Especes nouvelles de Melliferes de Barbarie. (Diagnoses preliminaires). Gounouilhou, Bordeaux." type="book" year="1895">Pérez, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype, ♀ (MNHN).
|
||
<emphasis id="B970B615FF87FFB90CF7F825FADCF812" bold="true" box="[1197,1218,1926,1952]" pageId="15" pageNumber="16">A</emphasis>
|
||
. Label details.
|
||
<emphasis id="B970B615FF87FFB908E7F80AFECFF871" bold="true" box="[189,209,1961,1987]" pageId="15" pageNumber="16">B</emphasis>
|
||
. Lateral habitus.
|
||
<emphasis id="B970B615FF87FFB909C6F80AFFAFF871" bold="true" box="[412,433,1961,1987]" pageId="15" pageNumber="16">C</emphasis>
|
||
. Face, frontal view.
|
||
<emphasis id="B970B615FF87FFB90AFBF80AFCA8F871" bold="true" box="[673,694,1961,1987]" pageId="15" pageNumber="16">D</emphasis>
|
||
. Scutum, dorsal view.
|
||
<emphasis id="B970B615FF87FFB90B9BF80AFDCBF871" bold="true" box="[961,981,1961,1987]" pageId="15" pageNumber="16">E</emphasis>
|
||
. Propodeum, dorsal view.
|
||
<emphasis id="B970B615FF87FFB90D57F80AFB01F871" bold="true" box="[1293,1311,1961,1987]" pageId="15" pageNumber="16">F</emphasis>
|
||
. Terga, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBB6A07FF98FFA608E7FE2AFDC2FCDF" blockId="16.[188,1399,393,877]" pageId="16" pageNumber="17">
|
||
<bibRefCitation id="EF9517F6FF98FFA608E7FE2AFF61FE11" author="Warncke K." box="[189,383,393,419]" pageId="16" pageNumber="17" pagination="3 - 54" refId="ref41080" refString="Warncke K. 1974. Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem zoologischen Museum in Berlin 50: 3 - 54." type="journal article" year="1974">Warncke (1974)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C041184FF98FFA60A53FE29FCF0FE11" authorityName="Warncke" authorityYear="1974" box="[521,750,394,420]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="montarca" subSpecies="parva">
|
||
<emphasis id="B970B615FF98FFA60A53FE29FCF0FE11" box="[521,750,394,420]" italics="true" pageId="16" pageNumber="17">A. montarca parva</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49C0A4E5FF98FFA60B1BFE2AFD9BFE11" box="[833,901,393,419]" country="Tunisia" name="Gafsa" pageId="16" pageNumber="17">Gafsa</collectingRegion>
|
||
in central
|
||
<collectingCountry id="F3132A97FF98FFA60C4BFE2AFA72FE11" box="[1041,1132,393,419]" name="Tunisia" pageId="16" pageNumber="17">Tunisia</collectingCountry>
|
||
. He described it as a subspecies of
|
||
<taxonomicName id="4C041184FF98FFA60933FE0EFD9BFE75" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[361,901,428,455]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="montarca" subGenus="Graecandrena">
|
||
<emphasis id="B970B615FF98FFA60933FE0EFF9DFE75" box="[361,387,429,455]" italics="true" pageId="16" pageNumber="17">A.</emphasis>
|
||
(
|
||
<emphasis id="B970B615FF98FFA609CDFE0FFC5FFE74" box="[407,577,428,454]" italics="true" pageId="16" pageNumber="17">Graecandrena</emphasis>
|
||
)
|
||
<emphasis id="B970B615FF98FFA60A0CFE0EFCDBFE74" box="[598,709,429,454]" italics="true" pageId="16" pageNumber="17">montarca</emphasis>
|
||
<bibRefCitation id="EF9517F6FF98FFA60A8AFE0FFD9BFE75" author="Warncke K." box="[720,901,428,455]" pageId="16" pageNumber="17" pagination="293 - 314" refId="ref41115" refString="Warncke K. 1975. Die Bienengattung Andrena F., in Iberien (Hym. Apidae). Teil A. Eos 49: 293 - 314." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
|
||
</taxonomicName>
|
||
(the difference in year priority is due to publication delays; see below for discussion and resolution of priority). This association was always morphologically tenuous –
|
||
<bibRefCitation id="EF9517F6FF98FFA60A5BFE50FD66FDBF" author="Gusenleitner F. & Schwarz M." box="[513,888,499,525]" pageId="16" pageNumber="17" pagination="1 - 1280" refId="ref39230" refString="Gusenleitner F. & Schwarz M. 2002. Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna Suppl. 10: 1 - 1280." type="journal article" year="2002">Gusenleitner & Schwarz (2002)</bibRefCitation>
|
||
noted the weak association of
|
||
<taxonomicName id="4C041184FF98FFA60CBCFE50FF1CFD82" authorityName="Warncke" authorityYear="1974" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="montarca" subSpecies="parva">
|
||
<emphasis id="B970B615FF98FFA60CBCFE50FF1CFD82" italics="true" pageId="16" pageNumber="17">A. montarca parva</emphasis>
|
||
</taxonomicName>
|
||
with the other taxa in this grouping and argued that it deserved species status in future revisions due to the weaker terga bands (
|
||
<figureCitation id="133F7682FF98FFA60A76FD9AFC8CFDE1" box="[556,658,569,595]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="16" pageNumber="17">Fig. 11F</figureCitation>
|
||
), the more granular shagreen of the scutum (
|
||
<figureCitation id="133F7682FF98FFA60CFAFD9AFB15FDE1" box="[1184,1291,569,595]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="16" pageNumber="17">Fig. 11D</figureCitation>
|
||
), and the smoother and shinier clypeus (
|
||
<figureCitation id="133F7682FF98FFA60A7AFDFFFC97FDC5" box="[544,649,604,631]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="16" pageNumber="17">Fig. 11C</figureCitation>
|
||
). All of these characters clearly place it as the junior synonym of
|
||
<taxonomicName id="4C041184FF98FFA608E6FD23FF40FD28" authorityName="Perez" authorityYear="1895" box="[188,350,640,666]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF98FFA608E6FD23FF40FD28" box="[188,350,640,666]" italics="true" pageId="16" pageNumber="17">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<collectingRegion id="49C0A4E5FF98FFA60937FD23FFC4FD28" box="[365,474,640,666]" country="Tunisia" name="Kairouan" pageId="16" pageNumber="17">Kairouan</collectingRegion>
|
||
and
|
||
<collectingRegion id="49C0A4E5FF98FFA60A4BFD23FC4BFD28" box="[529,597,640,666]" country="Tunisia" name="Gafsa" pageId="16" pageNumber="17">Gafsa</collectingRegion>
|
||
are approximately
|
||
<quantity id="4CFCC7E2FF98FFA60B6DFD23FD90FD28" box="[823,910,640,666]" metricMagnitude="5" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="17" unit="km" value="180.0">180 km</quantity>
|
||
apart, so this synonymy makes additional ecological sense.
|
||
<taxonomicName id="4C041184FF98FFA609D1FD00FC66FD0F" authorityName="Perez" authorityYear="1895" box="[395,632,675,701]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF98FFA609D1FD00FC66FD0F" box="[395,632,675,701]" italics="true" pageId="16" pageNumber="17">Andrena nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
is placed in the subgenus
|
||
<taxonomicName id="4C041184FF98FFA60BF1FD00FA2DFD0F" authorityName="sensu Warncke" authorityYear="1968" box="[939,1075,675,701]" pageId="16" pageNumber="17" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="B970B615FF98FFA60BF1FD00FA2DFD0F" box="[939,1075,675,701]" italics="true" pageId="16" pageNumber="17">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
based on its morphological features (e.g., the antefurcal nervulus of the forewing, the propodeal triangle with uniform granular shagreen and without basal rugae,
|
||
<figureCitation id="133F7682FF98FFA60A04FD4AFCD9FCB1" box="[606,711,745,771]" captionStart="Fig" captionStartId="15.[189,232,1926,1952]" captionTargetBox="[189,1399,533,1891]" captionTargetId="figure-123@15.[189,1399,533,1891]" captionTargetPageId="15" captionText="Fig. 11. Andrena (Graecandrena) nitidilabris Pérez, 1895, lectotype, ♀ (MNHN). A. Label details. B. Lateral habitus. C. Face, frontal view. D. Scutum, dorsal view. E. Propodeum, dorsal view. F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453486" httpUri="https://zenodo.org/record/10453486/files/figure.png" pageId="16" pageNumber="17">Fig. 11E</figureCitation>
|
||
), particularly the simple male genital capsule. Additional genetic data are required to conclusively demonstrate its phylogenetic placement, as for
|
||
<taxonomicName id="4C041184FF98FFA60C85FCAEFB6EFC94" authorityName="Warncke" authorityYear="1975" box="[1247,1392,781,807]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="montarca">
|
||
<emphasis id="B970B615FF98FFA60C85FCAEFB6EFC94" box="[1247,1392,781,807]" italics="true" pageId="16" pageNumber="17">A. montarca</emphasis>
|
||
</taxonomicName>
|
||
. From this point onwards,
|
||
<taxonomicName id="4C041184FF98FFA609AAFC93FC8DFCF8" authorityName="Perez" authorityYear="1895" box="[496,659,816,842]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF98FFA609AAFC93FC8DFCF8" box="[496,659,816,842]" italics="true" pageId="16" pageNumber="17">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
without qualification refers to the species in the correct sense;
|
||
<taxonomicName id="4C041184FF98FFA608E7FCF0FF7EFCDF" authorityName="Perez" authorityYear="1895" box="[189,352,851,877]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nitidilabris">
|
||
<emphasis id="B970B615FF98FFA608E7FCF0FF7EFCDF" box="[189,352,851,877]" italics="true" pageId="16" pageNumber="17">A. nitidilabris</emphasis>
|
||
</taxonomicName>
|
||
auctorum is used to refer to the taxon sensu Warncke.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E398CFF98FFA608E7FC2FFA2DFC67" box="[189,1075,908,982]" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="8BBB6A07FF98FFA608E7FC2FFF48FC14" blockId="16.[189,342,908,934]" box="[189,342,908,934]" pageId="16" pageNumber="17">
|
||
<heading id="D0F3DD6BFF98FFA608E7FC2FFF48FC14" bold="true" box="[189,342,908,934]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B970B615FF98FFA608E7FC2FFF48FC14" bold="true" box="[189,342,908,934]" pageId="16" pageNumber="17">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBB6A07FF98FFA608E7FC18FA2DFC67" blockId="16.[189,1075,955,982]" box="[189,1075,955,982]" pageId="16" pageNumber="17">
|
||
<collectingCountry id="F3132A97FF98FFA608E7FC18FF0BFC67" box="[189,277,955,981]" name="Algeria" pageId="16" pageNumber="17">Algeria</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3132A97FF98FFA60917FC18FFBAFC67" box="[333,420,955,981]" name="Tunisia" pageId="16" pageNumber="17">Tunisia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9517F6FF98FFA609E9FC18FD05FC67" author="Gusenleitner F. & Schwarz M." box="[435,795,955,981]" pageId="16" pageNumber="17" pagination="1 - 1280" refId="ref39230" refString="Gusenleitner F. & Schwarz M. 2002. Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna Suppl. 10: 1 - 1280." type="journal article" year="2002">Gusenleitner & Schwarz 2002</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C041184FF98FFA60B1CFC1FFA3CFC67" authorityName="Warncke" authorityYear="1974" box="[838,1058,956,982]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="montarca" subSpecies="parva">
|
||
<emphasis id="B970B615FF98FFA60B1CFC1FFA3CFC67" box="[838,1058,956,982]" italics="true" pageId="16" pageNumber="17">A. montarca parva</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |