132 lines
17 KiB
XML
132 lines
17 KiB
XML
<document id="4422D1CB72B6D486296F69FEAFB7ACA9" ID-CLB-Dataset="27314" ID-DOI="10.5852/ejt.2019.515" ID-GBIF-Dataset="4d4cc07e-3729-4179-90e1-cf469370776a" ID-ISSN="2118-9773" ID-Zenodo-Dep="2638175" ID-ZooBank="urn:lsid:zoobank.org:pub:F7AB8B53-FEB1-4473-8B22-DFEC9CE98FDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555091115131" checkinUser="plazi" docAuthor="Josens, Guy & Deligne, Jean" docDate="2019" docId="03AD879FFF9BFFF5FF51FD9AFD70B091" docLanguage="en" docName="ejt-515_josens_deligne.pdf.imf" docOrigin="European Journal of Taxonomy 515" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cubitermes Josens & Deligne 2019" docType="treatment" docVersion="12" lastPageNumber="55" masterDocId="FF94FFE7FFAEFFC3FFECFFC2FFF6B201" masterDocTitle="Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology" masterLastPageNumber="72" masterPageNumber="1" pageNumber="54" updateTime="1698720336213" updateUser="ExternalLinkService">
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<mods:title id="FCE94536EF427625564E75B0AB6A3620">Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology</mods:title>
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<mods:namePart id="1605654FA69261E7F65323A675370D40">Josens, Guy</mods:namePart>
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<mods:namePart id="FF0DD8E7D9E2490DBBA4258708D4B447">Deligne, Jean</mods:namePart>
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<mods:title id="3AD122FD45759686733A9D8DF7E05FC6">European Journal of Taxonomy</mods:title>
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<mods:part id="E43D6F93A3CDD520E6CB34191B910E0D">
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<mods:date id="21D2020AE66453952F371D50514DBC61">2019</mods:date>
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<mods:number id="DEDE4768AEC694206FD45FC926F429F9">2019-04-11</mods:number>
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<mods:number id="CC9896C6F7D261CF181D77F3B4ED3753">515</mods:number>
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<mods:identifier id="35B1132F0703988465D8256C68983D9F" type="DOI">10.5852/ejt.2019.515</mods:identifier>
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<treatment id="03AD879FFF9BFFF5FF51FD9AFD70B091" ID-GBIF-Taxon="156199110" LSID="urn:lsid:plazi:treatment:03AD879FFF9BFFF5FF51FD9AFD70B091" httpUri="http://treatment.plazi.org/id/03AD879FFF9BFFF5FF51FD9AFD70B091" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
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<subSubSection id="C31E6502FF9BFFF6FF51FD9AFD7FB073" box="[189,649,600,627]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="8BBB3689FF9BFFF6FF51FD9AFD7FB073" blockId="53.[189,649,600,627]" box="[189,649,600,627]" pageId="53" pageNumber="54">
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<heading id="D0F381E5FF9BFFF6FF51FD9AFD7FB073" bold="true" box="[189,649,600,627]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
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<emphasis id="B970EA9BFF9BFFF6FF51FD9AFD7FB073" bold="true" box="[189,649,600,627]" pageId="53" pageNumber="54">
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9. The
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<taxonomicName id="4C044D0AFF9BFFF6FEFEFD9AFE7DB072" authorityName="Josens & Deligne" authorityYear="2019" box="[274,395,600,627]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="speciesGroup" speciesGroup="sulcifrons valve pattern">
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<emphasis id="B970EA9BFF9BFFF6FEFEFD9AFE7DB072" bold="true" box="[274,395,600,627]" italics="true" pageId="53" pageNumber="54">sulcifrons</emphasis>
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</taxonomicName>
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valve pattern group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E6502FF9BFFF5FF51FD45FAC8B026" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="description">
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<paragraph id="8BBB3689FF9BFFF6FF51FD45FEB6B175" blockId="53.[189,1399,647,884]" pageId="53" pageNumber="54">
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The worker within this group has an enteric valve with all the six primary cushions ending downstream in yellow to brown sclerotised spatulae that stick out of the valve (
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<figureCitation id="133F2A0CFF9BFFF6FC77FD68FBF1B0C5" box="[923,1031,682,708]" captionStart="Fig" captionStartId="53.[189,232,1896,1922]" captionTargetBox="[235,1361,1137,1823]" captionTargetId="figure@53.[189,1398,1096,1864]" captionTargetPageId="53" captionText="Fig. 20. A. Enteric valve of the “sulcifrons valve pattern”, from a worker of Cubitermes sulcifrons Wasmann, 1911: note the spearhead-shape of the secondary cushions; B. Idem from a soldier of C. sulcifrons; C. Idem from an imago of C. umbratus Williams, 1954, spatulae are sketched on each PC; A, B and C at the same scale; D. Geographical distribution of species with the sulcifrons valve pattern." httpUri="https://zenodo.org/record/2638215/files/figure.png" pageId="53" pageNumber="54">Fig. 20A</figureCitation>
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). The spiny parts of the primary cushions are more or less fusiform: the largest widths are located in the first upstream third and their lateral margins converge gradually towards both ends (
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<figureCitation id="133F2A0CFF9BFFF6FCA9FD32FC50B10B" box="[837,934,752,778]" captionStart="Fig" captionStartId="30.[189,232,1719,1745]" captionTargetBox="[189,1398,317,1678]" captionTargetId="figure@30.[189,1398,297,1681]" captionTargetPageId="30" captionText="Fig. 11. Representative primary cushions of the various patterns encountered in the workers of the genus Cubitermes; WVPD = downstream bristly part; WVPM = middle spiny part; WVPS = spatula; WVPU = upstream spiny part. A. PC1 of the bilobatus valve pattern. B. PC1 of the oculatus valve pattern. C. any odd PC of the bilobatodes valve pattern. D. PC1 of the muneris valve pattern. E. PC6 and PC1 of the fungifaber valve pattern. E’. PC3 of the fungifaber valve pattern, twisted and partly seen in profile. F. PC1 of the finitimus valve pattern. F’. PC3 of the finitimus valve pattern, twisted and seen in profile. G. PC3 of the sankurensis valve pattern. H and H’. PC3 and PC4 of the oblectatus valve pattern. I. Any PC of the sulcifrons valve pattern. I’. Any PC of the sulcifrons valve pattern, twisted and seen in profile. All drawings to the same scale." httpUri="https://zenodo.org/record/2638195/files/figure.png" pageId="53" pageNumber="54">
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Fig.
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<date id="FFBA1049FF9BFFF6FC91FD32FC50B10B" box="[893,934,752,778]" pageId="53" pageNumber="54" value="2011-01">11I</date>
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</figureCitation>
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). All of the spatulae end symmetrically with two right or acute angles and all the six primary cushions are more or less protruding into the lumen of the valve. All of the six primary cushions are of the same size, forming hexaradial symmetry (
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<figureCitation id="133F2A0CFF9BFFF6FF2AFC98FEC5B175" box="[198,307,858,884]" captionStart="Fig" captionStartId="53.[189,232,1896,1922]" captionTargetBox="[235,1361,1137,1823]" captionTargetId="figure@53.[189,1398,1096,1864]" captionTargetPageId="53" captionText="Fig. 20. A. Enteric valve of the “sulcifrons valve pattern”, from a worker of Cubitermes sulcifrons Wasmann, 1911: note the spearhead-shape of the secondary cushions; B. Idem from a soldier of C. sulcifrons; C. Idem from an imago of C. umbratus Williams, 1954, spatulae are sketched on each PC; A, B and C at the same scale; D. Geographical distribution of species with the sulcifrons valve pattern." httpUri="https://zenodo.org/record/2638215/files/figure.png" pageId="53" pageNumber="54">Fig. 20A</figureCitation>
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).
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</paragraph>
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<paragraph id="8BBB3689FF9BFFF6FF51FC5EFA80B61E" blockId="53.[189,1399,923,1056]" pageId="53" pageNumber="54">A PC is made of (a) an upstream spiny part (16–36% of total length) with relatively strong spines, (b) a middle spiny part (26–43% of total length) with somewhat weaker spines and with 14–23 lateral supporting bristles on each side, (c) a bristly part (13–27% of total length) with 45–65 straight or curved and apparently soft bristles on a low crest, and (d) a symmetric sclerotised spatula with two right or acute</paragraph>
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<caption id="DF7B6601FF9BFFF6FF51F8AAFA84B5ED" httpUri="https://zenodo.org/record/2638215/files/figure.png" pageId="53" pageNumber="54" startId="53.[189,232,1896,1922]" targetBox="[235,1361,1137,1823]" targetPageId="53">
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<paragraph id="8BBB3689FF9BFFF6FF51F8AAFA84B5ED" blockId="53.[189,1399,1896,2028]" pageId="53" pageNumber="54">
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<emphasis id="B970EA9BFF9BFFF6FF51F8AAFECBB583" bold="true" box="[189,317,1896,1922]" pageId="53" pageNumber="54">Fig. 20. A</emphasis>
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. Enteric valve of the “
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<taxonomicName id="4C044D0AFF9BFFF6FDB6F8AAFD3BB583" authorityName="Wasmann" authorityYear="1911" box="[602,717,1896,1922]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sulcifrons">
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<emphasis id="B970EA9BFF9BFFF6FDB6F8AAFD3BB583" box="[602,717,1896,1922]" italics="true" pageId="53" pageNumber="54">sulcifrons</emphasis>
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</taxonomicName>
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valve pattern”, from a worker of
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<taxonomicName id="4C044D0AFF9BFFF6FB9DF8AAFE75B5A4" authority="Wasmann, 1911" authorityName="Wasmann" authorityYear="1911" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sulcifrons">
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<emphasis id="B970EA9BFF9BFFF6FB9DF8AAFA81B583" box="[1137,1399,1896,1922]" italics="true" pageId="53" pageNumber="54">Cubitermes sulcifrons</emphasis>
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Wasmann, 1911
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</taxonomicName>
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: note the spearhead-shape of the secondary cushions;
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<emphasis id="B970EA9BFF9BFFF6FBC1F849FBB7B5A4" bold="true" box="[1069,1089,1931,1957]" pageId="53" pageNumber="54">B</emphasis>
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. Idem from a soldier of
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<taxonomicName id="4C044D0AFF9BFFF6FF51F86DFEA7B5C9" authorityName="Wasmann" authorityYear="1911" box="[189,337,1966,1993]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sulcifrons">
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<emphasis id="B970EA9BFF9BFFF6FF51F86DFEA7B5C9" box="[189,337,1966,1993]" italics="true" pageId="53" pageNumber="54">C. sulcifrons</emphasis>
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</taxonomicName>
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;
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<emphasis id="B970EA9BFF9BFFF6FE8CF86CFE83B5C9" bold="true" box="[352,373,1966,1992]" pageId="53" pageNumber="54">C</emphasis>
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. Idem from an imago of
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<taxonomicName id="4C044D0AFF9BFFF6FD79F86DFC17B5C8" authority="Williams, 1954" authorityName="Williams" authorityYear="1954" box="[661,993,1966,1993]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="umbratus">
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<emphasis id="B970EA9BFF9BFFF6FD79F86DFCD0B5C9" box="[661,806,1966,1993]" italics="true" pageId="53" pageNumber="54">C. umbratus</emphasis>
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Williams, 1954
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</taxonomicName>
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, spatulae are sketched on each PC; A, B and C at the same scale;
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<emphasis id="B970EA9BFF9BFFF6FDF2F810FDC5B5ED" bold="true" box="[542,563,2002,2028]" pageId="53" pageNumber="54">D</emphasis>
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. Geographical distribution of species with the
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<taxonomicName id="4C044D0AFF9BFFF6FBB4F810FB3DB5ED" authorityName="Wasmann" authorityYear="1911" box="[1112,1227,2002,2028]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sulcifrons">
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<emphasis id="B970EA9BFF9BFFF6FBB4F810FB3DB5ED" box="[1112,1227,2002,2028]" italics="true" pageId="53" pageNumber="54">sulcifrons</emphasis>
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</taxonomicName>
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valve pattern.
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</paragraph>
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</caption>
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<paragraph id="8BBB3689FF98FFF5FF51FEC9FDDBB348" blockId="54.[189,1399,267,329]" pageId="54" pageNumber="55">
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angles (13–27% of total length) bearing some short spines and, on each side, some 10–15 very short tooth-like setae (
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<figureCitation id="133F2A0CFF98FFF5FE6DFEEDFDFAB348" box="[385,524,303,329]" captionStart="Fig" captionStartId="30.[189,232,1719,1745]" captionTargetBox="[189,1398,317,1678]" captionTargetId="figure@30.[189,1398,297,1681]" captionTargetPageId="30" captionText="Fig. 11. Representative primary cushions of the various patterns encountered in the workers of the genus Cubitermes; WVPD = downstream bristly part; WVPM = middle spiny part; WVPS = spatula; WVPU = upstream spiny part. A. PC1 of the bilobatus valve pattern. B. PC1 of the oculatus valve pattern. C. any odd PC of the bilobatodes valve pattern. D. PC1 of the muneris valve pattern. E. PC6 and PC1 of the fungifaber valve pattern. E’. PC3 of the fungifaber valve pattern, twisted and partly seen in profile. F. PC1 of the finitimus valve pattern. F’. PC3 of the finitimus valve pattern, twisted and seen in profile. G. PC3 of the sankurensis valve pattern. H and H’. PC3 and PC4 of the oblectatus valve pattern. I. Any PC of the sulcifrons valve pattern. I’. Any PC of the sulcifrons valve pattern, twisted and seen in profile. All drawings to the same scale." httpUri="https://zenodo.org/record/2638195/files/figure.png" pageId="54" pageNumber="55">
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Fig.
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<date id="FFBA1049FF98FFF5FE54FEEDFE16B348" box="[440,480,303,329]" pageId="54" pageNumber="55" value="2011-01">11I</date>
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, 11
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</figureCitation>
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I’).
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</paragraph>
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<paragraph id="8BBB3689FF98FFF5FF51FEABFAC0B3C8" blockId="54.[189,1399,361,458]" pageId="54" pageNumber="55">
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The secondary cushions are either narrow in their upstream end, widen downstream and sketch spearheads (
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<figureCitation id="133F2A0CFF98FFF5FF2AFE4EFEC2B3A7" box="[198,308,396,422]" captionStart="Fig" captionStartId="53.[189,232,1896,1922]" captionTargetBox="[235,1361,1137,1823]" captionTargetId="figure@53.[189,1398,1096,1864]" captionTargetPageId="53" captionText="Fig. 20. A. Enteric valve of the “sulcifrons valve pattern”, from a worker of Cubitermes sulcifrons Wasmann, 1911: note the spearhead-shape of the secondary cushions; B. Idem from a soldier of C. sulcifrons; C. Idem from an imago of C. umbratus Williams, 1954, spatulae are sketched on each PC; A, B and C at the same scale; D. Geographical distribution of species with the sulcifrons valve pattern." httpUri="https://zenodo.org/record/2638215/files/figure.png" pageId="54" pageNumber="55">Fig. 20A</figureCitation>
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), or are about twice as wide as the primary cushions, with their largest widths located in the upstream fourth or third, narrowing noticeably downstream with a homogeneous spine scattering.
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</paragraph>
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<paragraph id="8BBB3689FF98FFF5FF51FE2BFAC8B026" blockId="54.[189,1398,489,551]" pageId="54" pageNumber="55">This valve pattern is therefore characterised by six spatulae in the workers’ and soldiers’ valves and by the workers’ SCs which are either rather narrow or with a spearhead shape; most species are large.</paragraph>
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</subSubSection>
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<subSubSection id="C31E6502FF98FFF5FF51FD85FD70B091" pageId="54" pageNumber="55" type="discussion">
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<paragraph id="8BBB3689FF98FFF5FF51FD85FE5CB060" blockId="54.[189,426,583,609]" box="[189,426,583,609]" pageId="54" pageNumber="55">
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<heading id="D0F381E5FF98FFF5FF51FD85FE5CB060" bold="true" box="[189,426,583,609]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
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<emphasis id="B970EA9BFF98FFF5FF51FD85FE5CB060" bold="true" box="[189,426,583,609]" pageId="54" pageNumber="55">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB3689FF98FFF5FF51FDB4FD70B091" blockId="54.[189,646,630,656]" box="[189,646,630,656]" pageId="54" pageNumber="55">Eight species have such enteric valves:</paragraph>
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</subSubSection>
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</treatment>
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</document> |