373 lines
45 KiB
XML
373 lines
45 KiB
XML
<document id="2965197C09A2A29472C38E9D3B69E4F7" ID-DOI="10.5281/zenodo.197354" ID-GBIF-Dataset="a050f982-3c72-45f5-b15f-1672f34763df" ID-ISSN="1175-5326" ID-Zenodo-Dep="197354" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460266747342" checkinUser="plazi" docAuthor="Mcdonald, Andrew T., Barrett, Paul M. & Chapman, Sandra D." docDate="2010" docId="03A587FAFFBBE115FF76FC7AFB29AB70" docLanguage="en" docName="zt02569p043.pdf" docOrigin="Zootaxa 2569" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Iguanodon" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FF9CFF82FFBAE116FFE1FFDFFF98A902" masterDocTitle="A new basal iguanodont (Dinosauria: Ornithischia) from the Wealden (Lower Cretaceous) of England" masterLastPageNumber="43" masterPageNumber="1" pageNumber="2" updateTime="1698245971878" updateUser="plazi">
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<mods:title id="E6683E552024AC58C521C33E72B2D1F5">A new basal iguanodont (Dinosauria: Ornithischia) from the Wealden (Lower Cretaceous) of England</mods:title>
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<mods:name id="2D7F64C920AC8DB5EB350E8FD478D7FA" type="personal">
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<mods:roleTerm id="E16172918ECEE7D4C95B8BD435E5F9E2">Author</mods:roleTerm>
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<mods:namePart id="86EDCFC61E8B8099994F16AA337DAD5B">Mcdonald, Andrew T.</mods:namePart>
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<mods:namePart id="D00F22A2117F148D9439BFE2E8D75DD0">Barrett, Paul M.</mods:namePart>
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<mods:namePart id="43AD776C005FCF4BEE6E4762D40744CC">Chapman, Sandra D.</mods:namePart>
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<mods:title id="0D0FEECC13D5DF3EEDF396F1E58BED75">Zootaxa</mods:title>
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<mods:part id="34E97AC7F3E18ACFCCE622C55C8D7FE9">
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<mods:date id="C2382A2D9FD5C07217AD2DEDEB7782DB">2010</mods:date>
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<mods:number id="8380811B2B017900B87E984B3AE717A4">2569</mods:number>
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<mods:identifier id="46900AF80F2C8EC69D37A66B8C4761EC" type="DOI">10.5281/zenodo.197354</mods:identifier>
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<mods:identifier id="889FD007F77BDB2FFAC0DE539B0C46EA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9374B2E77B34E3A89451565F3601586E" type="Zenodo-Dep">197354</mods:identifier>
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<treatment id="03A587FAFFBBE115FF76FC7AFB29AB70" ID-DOI="http://doi.org/10.5281/zenodo.6208586" ID-GBIF-Taxon="119394381" ID-Zenodo-Dep="6208586" LSID="urn:lsid:plazi:treatment:03A587FAFFBBE115FF76FC7AFB29AB70" httpUri="http://treatment.plazi.org/id/03A587FAFFBBE115FF76FC7AFB29AB70" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="C3166567FFBBE117FF76FC7AFD97AABD" box="[151,527,933,959]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BB336ECFFBBE117FF76FC7AFD97AABD" blockId="1.[151,527,933,959]" box="[151,527,933,959]" pageId="1" pageNumber="2">
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<heading id="D0FB8180FFBBE117FF76FC7AFD97AABD" bold="true" box="[151,527,933,959]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="B978EAFEFFBBE117FF76FC7AFD97AABD" bold="true" box="[151,527,933,959]" pageId="1" pageNumber="2">
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Note on
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<taxonomicName id="4C0C4D6FFFBBE117FF1EFC7AFE1CAABD" box="[255,388,933,959]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis id="B978EAFEFFBBE117FF1EFC7AFE1CAABD" bold="true" box="[255,388,933,959]" italics="true" pageId="1" pageNumber="2">Iguanodon</emphasis>
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</taxonomicName>
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Taxonomy
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3166567FFBBE115FF76FC32FB29AB70" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
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<paragraph id="8BB336ECFFBBE117FF76FC32FCCAAC7E" blockId="1.[151,1437,1005,2002]" pageId="1" pageNumber="2">
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The taxonomy of
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<taxonomicName id="4C0C4D6FFFBBE117FE86FC32FE7EAD04" box="[359,486,1005,1030]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis id="B978EAFEFFBBE117FE86FC32FE7EAD04" box="[359,486,1005,1030]" italics="true" pageId="1" pageNumber="2">Iguanodon</emphasis>
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</taxonomicName>
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has been problematic ever since
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<bibRefCitation id="EF9D4B1DFFBBE117FC8BFC32FB83AD05" author="Mantell" box="[874,1051,1005,1031]" pageId="1" pageNumber="2" refString="Mantell, G. (1825) Notice on the Iguanodon, a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex. Philosophical Transactions of the Royal Society of London, 115, 179 - 186." type="journal article" year="1825">Mantell (1825)</bibRefCitation>
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erected the genus without a
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<typeStatus id="54B7884EFFBBE117FA88FC32FA04AD05" box="[1385,1436,1005,1031]" pageId="1" pageNumber="2">type</typeStatus>
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||
species.
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<bibRefCitation id="EF9D4B1DFFBBE117FF1EFBCDFE16AD2E" author="Holl" box="[255,398,1042,1068]" pageId="1" pageNumber="2" refString="Holl, F. (1829) Handbuch der Petrefactenkunde, Part I. Quedlingberg, Leipzig, 232 pp." type="book" year="1829">Holl (1829)</bibRefCitation>
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provided the species name
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<taxonomicName id="4C0C4D6FFFBBE117FD03FBCDFCF5AD29" box="[738,877,1042,1068]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="anglicum">
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<emphasis id="B978EAFEFFBBE117FD03FBCDFD74AD29" box="[738,748,1042,1067]" italics="true" pageId="1" pageNumber="2">I</emphasis>
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.
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<emphasis id="B978EAFEFFBBE117FD1CFBCDFCF5AD29" box="[765,877,1042,1067]" italics="true" pageId="1" pageNumber="2">anglicum</emphasis>
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</taxonomicName>
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(emended to
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<taxonomicName id="4C0C4D6FFFBBE117FBF4FBCDFAAFAD2E" authority="Norman 1986" authorityName="Norman" authorityYear="1986" box="[1045,1335,1042,1068]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="anglicus">
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<emphasis id="B978EAFEFFBBE117FBF4FBCDFBE4AD29" box="[1045,1148,1042,1067]" italics="true" pageId="1" pageNumber="2">anglicus</emphasis>
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:
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<bibRefCitation id="EF9D4B1DFFBBE117FB6CFBCDFAAFAD2E" author="Norman" box="[1165,1335,1042,1068]" pageId="1" pageNumber="2" refString="Norman, D. B. (1986) On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 56, 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
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</taxonomicName>
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) shortly thereafter based upon the teeth described by Mantell, including the
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<typeStatus id="54B7884EFFBBE117FC3EFBE7FBC8AD50" box="[991,1104,1080,1106]" pageId="1" pageNumber="2" type="lectotype">lectotype</typeStatus>
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NHMUK R2392 (
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<bibRefCitation id="EF9D4B1DFFBBE117FAD6FBE7FF40AD75" author="Norman" pageId="1" pageNumber="2" refString="Norman, D. B. (1986) On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 56, 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
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). However, these teeth possess no features that would definitively demarcate them from the teeth of other, better known iguanodontian species (
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<bibRefCitation id="EF9D4B1DFFBBE117FD7EFB5DFCD1AD9E" author="Norman" box="[671,841,1154,1180]" pageId="1" pageNumber="2" refString="Norman, D. B. (1986) On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 56, 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
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;
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<bibRefCitation id="EF9D4B1DFFBBE117FCB7FB5DFB10AD9E" author="Charig" box="[854,1160,1154,1180]" pageId="1" pageNumber="2" refString="Charig, A. J. & Chapman, S. D. (1998) Iguanodon Mantell, 1825 (Reptilia, Ornithischia): proposed designation of Iguanodon bernissartensis Boulenger in Beneden, 1881 as the type species, and proposed designation of a lectotype. Bulletin of Zoological Nomenclature, 55, 99 - 104." type="journal article" year="1998">Charig & Chapman 1998</bibRefCitation>
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). Thus,
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<taxonomicName id="4C0C4D6FFFBBE117FB06FB5CFAFDAD9E" box="[1255,1381,1154,1180]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="anglicus">
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||
<emphasis id="B978EAFEFFBBE117FB06FB5CFB69AD9E" box="[1255,1265,1155,1180]" italics="true" pageId="1" pageNumber="2">I</emphasis>
|
||
.
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||
<emphasis id="B978EAFEFFBBE117FAE0FB5CFAFDAD9E" box="[1281,1381,1155,1180]" italics="true" pageId="1" pageNumber="2">anglicus</emphasis>
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</taxonomicName>
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||
was declared a
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||
<emphasis id="B978EAFEFFBBE117FEF8FB77FE51ADC3" box="[281,457,1192,1217]" italics="true" pageId="1" pageNumber="2">nomen dubium</emphasis>
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and
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<taxonomicName id="4C0C4D6FFFBBE117FDE5FB77FD48ADC3" box="[516,720,1192,1218]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="B978EAFEFFBBE117FDE5FB77FD96ADC3" box="[516,526,1192,1217]" italics="true" pageId="1" pageNumber="2">I</emphasis>
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||
.
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||
<emphasis id="B978EAFEFFBBE117FDFCFB77FD48ADC3" box="[541,720,1192,1217]" italics="true" pageId="1" pageNumber="2">bernissartensis</emphasis>
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</taxonomicName>
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, by virtue of its complete
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<typeStatus id="54B7884EFFBBE117FBECFB77FBE2ADC0" box="[1037,1146,1192,1218]" pageId="1" pageNumber="2" type="lectotype">lectotype</typeStatus>
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skeleton from
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<collectingCountry id="F31B767CFFBBE117FACFFB77FA0FADC0" box="[1326,1431,1192,1218]" name="Belgium" pageId="1" pageNumber="2">Belgium</collectingCountry>
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, was made the
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<typeStatus id="54B7884EFFBBE117FEAFFB12FE1DADE5" box="[334,389,1229,1255]" pageId="1" pageNumber="2">type</typeStatus>
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species of
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<taxonomicName id="4C0C4D6FFFBBE117FDF7FB12FB9BADE5" authority="Charig & Chapman 1998" authorityName="Charig & Chapman" authorityYear="1998" box="[534,1027,1229,1255]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis id="B978EAFEFFBBE117FDF7FB12FD07ADE4" box="[534,671,1229,1254]" italics="true" pageId="1" pageNumber="2">Iguanodon</emphasis>
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(
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<bibRefCitation id="EF9D4B1DFFBBE117FD50FB12FC60ADE5" author="Charig" box="[689,1016,1229,1255]" pageId="1" pageNumber="2" refString="Charig, A. J. & Chapman, S. D. (1998) Iguanodon Mantell, 1825 (Reptilia, Ornithischia): proposed designation of Iguanodon bernissartensis Boulenger in Beneden, 1881 as the type species, and proposed designation of a lectotype. Bulletin of Zoological Nomenclature, 55, 99 - 104." type="journal article" year="1998">Charig & Chapman 1998</bibRefCitation>
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)
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</taxonomicName>
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. Furthermore, some species and specimens formerly regarded as pertaining to
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<taxonomicName id="4C0C4D6FFFBBE117FD29FB2CFCD3AC0E" box="[712,843,1267,1292]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis id="B978EAFEFFBBE117FD29FB2CFCD3AC0E" box="[712,843,1267,1292]" italics="true" pageId="1" pageNumber="2">Iguanodon</emphasis>
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||
</taxonomicName>
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have been recently removed (
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<bibRefCitation id="EF9D4B1DFFBBE117FB5AFB2DFF4EAC30" author="Norman" pageId="1" pageNumber="2" refString="Norman, D. B. & Barrett, P. M. (2002) Ornithischian dinosaurs from the Lower Cretaceous (Berriasian) of England. Special Papers in Palaeontology, 68, 161 - 189." type="journal article" year="2002">Norman & Barrett 2002</bibRefCitation>
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||
;
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||
<bibRefCitation id="EF9D4B1DFFBBE117FF03FAC7FEC5AC30" author="Paul" box="[226,349,1304,1330]" pageId="1" pageNumber="2" refString="Paul, G. S. (2007) Turning the old into the new: a separate genus for the gracile iguanodont from the Wealden of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington, pp. 69 - 77." type="book chapter" year="2007">Paul 2007</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FE88FAC7FE30AC30" author="Paul" box="[361,424,1304,1330]" pageId="1" pageNumber="2" refString="Paul, G. S. (2008) A revised taxonomy of the iguanodont dinosaur genera and species. Cretaceous Research, 29, 192 - 216." type="journal article" year="2008">2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FE54FAC7FDD5AC30" author="Galton" box="[437,589,1304,1330]" pageId="1" pageNumber="2" refString="Galton, P. M. (2009) Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, " Camptosaurus ", " Iguanodon " - and referred specimens from Romania and elsewhere. Revue de Paleobiologie, Geneve, 28, 211 - 273." type="journal article" year="2009">Galton 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FDB8FAC7FC9AAC30" author="Norman" box="[601,770,1304,1330]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman 2010</bibRefCitation>
|
||
); the name
|
||
<taxonomicName id="4C0C4D6FFFBBE117FC6AFAC7FB5DAC30" authority="Paul, 2007" authorityName="Paul" authorityYear="2007" box="[907,1221,1304,1330]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="B978EAFEFFBBE117FC6AFAC7FBA6AC33" box="[907,1086,1304,1329]" italics="true" pageId="1" pageNumber="2">Mantellisaurus</emphasis>
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FBA7FAC7FB5DAC30" author="Paul" box="[1094,1221,1304,1330]" pageId="1" pageNumber="2" refString="Paul, G. S. (2007) Turning the old into the new: a separate genus for the gracile iguanodont from the Wealden of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington, pp. 69 - 77." type="book chapter" year="2007">Paul, 2007</bibRefCitation>
|
||
</taxonomicName>
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||
is used herein for “
|
||
<emphasis id="B978EAFEFFBBE117FF45FAE2FF36AC54" box="[164,174,1341,1366]" italics="true" pageId="1" pageNumber="2">I</emphasis>
|
||
.”
|
||
<taxonomicName id="4C0C4D6FFFBBE117FF2BFAE2FD30AC55" authority="Owenodon Galton, 2009" authorityName="Owenodon Galton" authorityYear="2009" box="[202,680,1341,1367]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="B978EAFEFFBBE117FF2BFAE2FEEFAC54" box="[202,375,1341,1366]" italics="true" pageId="1" pageNumber="2">atherfieldensis</emphasis>
|
||
,
|
||
<emphasis id="B978EAFEFFBBE117FE66FAE2FD9EAC54" box="[391,518,1341,1366]" italics="true" pageId="1" pageNumber="2">Owenodon</emphasis>
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FDEFFAE2FD30AC55" author="Galton" box="[526,680,1341,1367]" pageId="1" pageNumber="2" refString="Galton, P. M. (2009) Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, " Camptosaurus ", " Iguanodon " - and referred specimens from Romania and elsewhere. Revue de Paleobiologie, Geneve, 28, 211 - 273." type="journal article" year="2009">Galton, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
for “
|
||
<emphasis id="B978EAFEFFBBE117FD09FAE2FD6AAC54" box="[744,754,1341,1366]" italics="true" pageId="1" pageNumber="2">I</emphasis>
|
||
.”
|
||
<emphasis id="B978EAFEFFBBE117FCEFFAE2FCC1AC54" box="[782,857,1341,1366]" italics="true" pageId="1" pageNumber="2">hoggii</emphasis>
|
||
, and
|
||
<emphasis id="B978EAFEFFBBE117FC7AFAE2FB9CAC54" box="[923,1028,1341,1366]" italics="true" pageId="1" pageNumber="2">Barilium</emphasis>
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FBEDFAE2FB2EAC55" author="Norman" box="[1036,1206,1341,1367]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman, 2010</bibRefCitation>
|
||
and
|
||
<emphasis id="B978EAFEFFBBE117FB10FAE2FA04AC54" box="[1265,1436,1341,1366]" italics="true" pageId="1" pageNumber="2">Hypselospinus</emphasis>
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FF76FABDFED9AC7E" author="Norman" box="[151,321,1378,1404]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman, 2010</bibRefCitation>
|
||
respectively for “
|
||
<emphasis id="B978EAFEFFBBE117FDF2FABCFD85AC7E" box="[531,541,1379,1404]" italics="true" pageId="1" pageNumber="2">I</emphasis>
|
||
.”
|
||
<emphasis id="B978EAFEFFBBE117FDD9FABCFD02AC7E" box="[568,666,1379,1404]" italics="true" pageId="1" pageNumber="2">dawsoni</emphasis>
|
||
and “
|
||
<emphasis id="B978EAFEFFBBE117FD01FABCFD72AC7E" box="[736,746,1379,1404]" italics="true" pageId="1" pageNumber="2">I</emphasis>
|
||
.”
|
||
<emphasis id="B978EAFEFFBBE117FCE4FABCFCD3AC7E" box="[773,843,1379,1404]" italics="true" pageId="1" pageNumber="2">fittoni</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFBBE114FF27FA57FE4EA820" blockId="1.[151,1437,1005,2002]" lastBlockId="2.[151,1438,152,1634]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FF27FA57FEE7ACA0" author="Norman" box="[198,383,1416,1442]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman (2010)</bibRefCitation>
|
||
discussed NHMUK 28660 and illustrated it in medial view (
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FB8DFA57FA16ACA0" author="Norman" box="[1132,1422,1416,1442]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman 2010: fig. 10C</bibRefCitation>
|
||
), suggesting that it might be referable to
|
||
<emphasis id="B978EAFEFFBBE117FD9DFA72FD70ACC4" box="[636,744,1453,1478]" italics="true" pageId="1" pageNumber="2">Barilium</emphasis>
|
||
, which occurs in the middle Valanginian Wadhurst Clay Formation with
|
||
<emphasis id="B978EAFEFFBBE117FEB5FA0CFE67ACEE" box="[340,511,1491,1516]" italics="true" pageId="1" pageNumber="2">Hypselospinus</emphasis>
|
||
. However, there are several reasons for not referring NHMUK 28660 to either of the Wadhurst Clay taxa. The unit in which NHMUK 28660 was found, the Grinstead Clay Member of the Tunbridge Wells Sand Formation, is stratigraphically above and slightly younger than the Wadhurst Clay Formation (
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FEC6F99DFDF7AF5E" author="Allen" box="[295,623,1602,1628]" pageId="1" pageNumber="2" refString="Allen, P. & Wimbledon, W. A. (1991) Correlation of NW European Purbeck - Wealden" type="book" year="1991">Allen & Wimbledon 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FD9DF99DFC84AF5E" author="Radley" box="[636,796,1602,1628]" pageId="1" pageNumber="2" refString="Radley, J. (2004) Demystifying the Wealden of the Weald (Lower Cretaceous, south-east England). Open University Geological Society Journal, 25, 6 - 16." type="journal article" year="2004">Radley 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FCCBF99DFC40AF5E" author="Rawson" box="[810,984,1602,1628]" pageId="1" pageNumber="2" refString="Rawson, P. F. (2006) Cretaceous: sea levels peak as the North Atlantic opens. In: Brenchley, P. J. & Rawson, P. F. (Eds.), The Geology of England and Wales, 2 nd Edition. The Geological Society of London, Bath, pp. 365 - 394." type="book chapter" year="2006">Rawson 2006</bibRefCitation>
|
||
). More importantly, NHMUK 28660 differs from the two basal iguanodont dentaries from the Wadhurst Clay: NHMUK R1834 and NHMUK R1831. NHMUK R1834 is a partial associated skeleton that includes a partial left dentary, numerous vertebral centra, a fragmented right scapula, a well preserved right ilium, and several pedal elements; the specimen is herein considered referable to
|
||
<emphasis id="B978EAFEFFBBE117FE18F907FDF9AFF3" box="[505,609,1752,1777]" italics="true" pageId="1" pageNumber="2">Barilium</emphasis>
|
||
due to the similar morphologies (e.g., smoothly convex dorsal margin) shared by its ilium and NHMUK R802, the
|
||
<typeStatus id="54B7884EFFBBE117FD44F922FC95AE15" box="[677,781,1789,1815]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
|
||
ilium of
|
||
<emphasis id="B978EAFEFFBBE117FC9CF922FC7EAE14" box="[893,998,1789,1814]" italics="true" pageId="1" pageNumber="2">Barilium</emphasis>
|
||
(
|
||
<emphasis id="B978EAFEFFBBE117FC18F922FBDEAE14" box="[1017,1094,1789,1814]" italics="true" pageId="1" pageNumber="2">contra</emphasis>
|
||
<bibRefCitation id="EF9D4B1DFFBBE117FBAFF922FB6FAE15" author="Norman" box="[1102,1271,1789,1815]" pageId="1" pageNumber="2" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman 2010</bibRefCitation>
|
||
, who referred NHMUK R1834 to
|
||
<emphasis id="B978EAFEFFBBE117FE65F8FCFDA8AE3E" box="[388,560,1827,1852]" italics="true" pageId="1" pageNumber="2">Hypselospinus</emphasis>
|
||
). The ventrally inflected rostral ramus of the dentary of NHMUK R1834 differs from the straight rostral ramus of NHMUK 28660 (
|
||
<figureCitation id="13372A69FFBBE117FCA5F897FC12AE60" box="[836,906,1864,1890]" captionStart="FIGURE 1" captionStartId="3.[151,259,1944,1968]" captionTargetBox="[434,1147,1447,1895]" captionTargetId="figure@3.[434,1148,1447,1895]" captionTargetPageId="3" captionText="FIGURE 1. Dentaries of basal iguanodonts from the Wadhurst Clay Formation. Left dentary of NHMUK R 1834, referred herein to Barilium dawsoni, in lateral (A) view; right dentary NHMUK R 1831 in lateral (B) view. Scale bars equal 10 cm." httpUri="https://zenodo.org/record/197355/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
|
||
A, 2A), suggesting that they do not represent the same taxon. Dentaries of basal iguanodonts known from multiple specimens from the same locality and horizon (e.g.,
|
||
<taxonomicName id="4C0C4D6FFFBBE117FEA5F84CFDD4AEAE" box="[324,588,1939,1964]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="B978EAFEFFBBE117FEA5F84CFDD4AEAE" box="[324,588,1939,1964]" italics="true" pageId="1" pageNumber="2">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
[USNM 4281, 5818, 5961; YPM 1886, 1888] from Quarry
|
||
<quantity id="4CF49B09FFBBE117FAD1F84DFAF6AEAE" box="[1328,1390,1938,1964]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="1" pageNumber="2" unit="in" value="13.0">13 in</quantity>
|
||
the Morrison Formation,
|
||
<taxonomicName id="4C0C4D6FFFBBE117FE78F867FD4EAED3" box="[409,726,1976,2001]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="B978EAFEFFBBE117FE78F867FD4EAED3" box="[409,726,1976,2001]" italics="true" pageId="1" pageNumber="2">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
[IRSNB 1536, 1561, 1639] from Bernissart, and
|
||
<emphasis id="B978EAFEFFBBE114FAC8F867FEB7A9B3" italics="true" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Eolambia caroljonesa</emphasis>
|
||
[CEUM 34357, 34447, 35482, 35714, 52139, 52995] from
|
||
<emphasis id="B978EAFEFFB8E114FBDFFF47FB23A9B3" box="[1086,1211,152,177]" italics="true" pageId="2" pageNumber="3">Eolambia</emphasis>
|
||
#2 Quarry in the Mussentuchit Member, Cedar Mountain Formation) indicate that there is no intraspecific variation in the morphology of the rostral ramus, i.e. straight versus ventrally inflected, and that this character is probably taxonomically informative.
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB8E114FF24FEF2FE51AAE5" blockId="2.[151,1438,152,1634]" pageId="2" pageNumber="3">
|
||
NHMUK R1831, a right dentary, is part of an associated skeleton with elements catalogued under numbers NHMUK R
|
||
<date id="FFB2102CFFB8E114FE78FE8DFD9CA86E" box="[409,516,338,364]" pageId="2" pageNumber="3" value="1832" valueMax="1833">1832–33</date>
|
||
and NHMUK R
|
||
<date id="FFB2102CFFB8E114FD32FE8DFCD9A86E" box="[723,833,338,364]" pageId="2" pageNumber="3" value="1835" valueMax="1836">1835–36</date>
|
||
. The taxonomic identification of this skeleton is undecided, with
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FE8DFEA7FD9AA890" author="Paul" box="[364,514,376,402]" pageId="2" pageNumber="3" refString="Paul, G. S. (2008) A revised taxonomy of the iguanodont dinosaur genera and species. Cretaceous Research, 29, 192 - 216." type="journal article" year="2008">Paul (2008)</bibRefCitation>
|
||
regarding it as a potential new taxon and
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FBC3FEA7FB7EA890" author="Norman" box="[1058,1254,376,402]" pageId="2" pageNumber="3" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman (2010)</bibRefCitation>
|
||
referring it to
|
||
<emphasis id="B978EAFEFFB8E114FF76FE42FEDBA8B4" box="[151,323,413,438]" italics="true" pageId="2" pageNumber="3">Hypselospinus</emphasis>
|
||
; however, the specimen includes only a shard of an ilium and a very poorly preserved femur, and thus possesses no overlapping features that would support referral to
|
||
<emphasis id="B978EAFEFFB8E114FBD7FE1DFB73A8D9" box="[1078,1259,450,475]" italics="true" pageId="2" pageNumber="3">Hypselospinus</emphasis>
|
||
(based on the
|
||
<typeStatus id="54B7884EFFB8E114FF76FE37FE98AB00" box="[151,256,488,514]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
NHMUK R1635 and referred material comprising the
|
||
<typeStatus id="54B7884EFFB8E114FC7DFE37FB9DAB00" box="[924,1029,488,514]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
of the subjective junior synonym “
|
||
<taxonomicName id="4C0C4D6FFFB8E114FF45FDD2FE75AB24" box="[164,493,525,550]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="hollingtoniensis">
|
||
<emphasis id="B978EAFEFFB8E114FF45FDD2FE75AB24" box="[164,493,525,550]" italics="true" pageId="2" pageNumber="3">Iguanodon hollingtoniensis</emphasis>
|
||
</taxonomicName>
|
||
”;
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FDEAFDD2FD2EAB25" author="Norman" box="[523,694,525,551]" pageId="2" pageNumber="3" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman 2010</bibRefCitation>
|
||
). NHMUK R1831 is similar to NHMUK
|
||
<quantity id="4CF49B09FFB8E114FB4AFDD2FA8DAB25" box="[1195,1301,525,551]" metricMagnitude="2" metricUnit="m" metricValue="7.279639999999999" pageId="2" pageNumber="3" unit="in" value="28660.0">28660 in</quantity>
|
||
its straight rostral ramus (
|
||
<figureCitation id="13372A69FFB8E114FEA1FDEDFE1AAB4E" box="[320,386,562,588]" captionStart="FIGURE 1" captionStartId="3.[151,259,1944,1968]" captionTargetBox="[434,1147,1447,1895]" captionTargetId="figure@3.[434,1148,1447,1895]" captionTargetPageId="3" captionText="FIGURE 1. Dentaries of basal iguanodonts from the Wadhurst Clay Formation. Left dentary of NHMUK R 1834, referred herein to Barilium dawsoni, in lateral (A) view; right dentary NHMUK R 1831 in lateral (B) view. Scale bars equal 10 cm." httpUri="https://zenodo.org/record/197355/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
B), but differs in having a longer diastema (though not as long as suggested by
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FAFFFDEDFA0FAB4E" author="Paul" box="[1310,1431,562,588]" pageId="2" pageNumber="3" refString="Paul, G. S. (2008) A revised taxonomy of the iguanodont dinosaur genera and species. Cretaceous Research, 29, 192 - 216." type="journal article" year="2008">Paul 2008</bibRefCitation>
|
||
; ATM pers. obs.;
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FE83FD87FD93AB70" author="Norman" box="[354,523,600,626]" pageId="2" pageNumber="3" refString="Norman, D. B. (2010) A taxonomy of iguanodontians (Dinosauria: Ornithopoda) from the lower Wealden Group (Cretaceous: Valanginian) of southern England. Zootaxa, 2489, 47 - 66." type="journal article" year="2010">Norman 2010</bibRefCitation>
|
||
) and teeth with more prominent accessory ridges arising from the marginal denticles directly mesial to the primary ridge. The coronoid process of NHMUK R1831 is broken both at its base and apex, thus rendering its inclination and expansion, or lack thereof, difficult to evaluate. Given that there are two named taxa (
|
||
<emphasis id="B978EAFEFFB8E114FE39FD17FDDAABE3" box="[472,578,712,737]" italics="true" pageId="2" pageNumber="3">Barilium</emphasis>
|
||
and
|
||
<emphasis id="B978EAFEFFB8E114FD9CFD17FCB3ABE3" box="[637,811,712,737]" italics="true" pageId="2" pageNumber="3">Hypselospinus</emphasis>
|
||
) and two different dentary morphologies (NHMUK R1834, referred herein to
|
||
<emphasis id="B978EAFEFFB8E114FE29FD32FDA8AA04" box="[456,560,749,774]" italics="true" pageId="2" pageNumber="3">Barilium</emphasis>
|
||
, and NHMUK R1831) from the Wadhurst Clay, it would be convenient to refer NHMUK R1831 and associated postcranial specimens to
|
||
<emphasis id="B978EAFEFFB8E114FC2CFCCDFB11AA29" box="[973,1161,786,811]" italics="true" pageId="2" pageNumber="3">Hypselospinus</emphasis>
|
||
; however, this is not supportable in the absence of shared morphologies and overlapping material. This situation is akin to that of the derived titanosaurs
|
||
<taxonomicName id="4C0C4D6FFFB8E114FE4CFC82FDFDAA74" box="[429,613,861,886]" class="Reptilia" family="Nemegtosauridae" genus="Nemegtosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B978EAFEFFB8E114FE4CFC82FDFDAA74" box="[429,613,861,886]" italics="true" pageId="2" pageNumber="3">Nemegtosaurus</emphasis>
|
||
</taxonomicName>
|
||
(known from a skull) and
|
||
<taxonomicName id="4C0C4D6FFFB8E114FC47FC82FB16AA74" box="[934,1166,861,886]" class="Reptilia" family="Camarasauridae" genus="Opisthocoelicaudia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B978EAFEFFB8E114FC47FC82FB16AA74" box="[934,1166,861,886]" italics="true" pageId="2" pageNumber="3">Opisthocoelicaudia</emphasis>
|
||
</taxonomicName>
|
||
(known from a partial postcranium) from the Nemegt Formation of
|
||
<collectingCountry id="F31B767CFFB8E114FD48FC5DFC83AA9E" box="[681,795,898,924]" name="Mongolia" pageId="2" pageNumber="3">Mongolia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FCCBFC5DFC5EAA9E" author="Wilson" box="[810,966,898,924]" pageId="2" pageNumber="3" refString="Wilson, J. A. (2005) Redescription of the Mongolian sauropod Nemegtosaurus mongoliensis Nowinski (Dinosauria: Saurischia) and comments on Late Cretaceous sauropod diversity. Journal of Systematic Palaeontology, 3, 283 - 318." type="journal article" year="2005">Wilson 2005</bibRefCitation>
|
||
), i.e. the possibilities that the specimens represent the same taxon and that they represent different diagnostic taxa must be considered equally likely given the known material.
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB8E114FF24FC2DFD9FAF60" blockId="2.[151,1438,152,1634]" pageId="2" pageNumber="3">
|
||
Both NHMUK R1834 and NHMUK R1831 lack the autapomorphic row of foramina observed in NHMUK 28660 (see below). These differences indicate that NHMUK 28660 should not be referred to the same taxon as either of the Wadhurst Clay dentaries. Because the distinctive dentary NHMUK 28660 was found in the same unit (Grinstead Clay Member of the Tunbridge Wells Sand Formation) and vicinity as the non-diagnostic teeth of
|
||
<taxonomicName id="4C0C4D6FFFB8E114FE4EFB57FD01ADA3" box="[431,665,1160,1185]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="anglicus">
|
||
<emphasis id="B978EAFEFFB8E114FE4EFB57FD01ADA3" box="[431,665,1160,1185]" italics="true" pageId="2" pageNumber="3">Iguanodon anglicus</emphasis>
|
||
</taxonomicName>
|
||
, it would be tempting to refer NHMUK 28660 to
|
||
<taxonomicName id="4C0C4D6FFFB8E114FB0CFB57FAF1ADA3" box="[1261,1385,1160,1186]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="anglicus">
|
||
<emphasis id="B978EAFEFFB8E114FB0CFB57FB6FADA3" box="[1261,1271,1160,1185]" italics="true" pageId="2" pageNumber="3">I</emphasis>
|
||
.
|
||
<emphasis id="B978EAFEFFB8E114FAE7FB57FAF1ADA3" box="[1286,1385,1160,1185]" italics="true" pageId="2" pageNumber="3">anglicus</emphasis>
|
||
</taxonomicName>
|
||
and thus render that taxon diagnostic. However, there are two reasons for not doing so. Firstly, the isolated dentary teeth of
|
||
<taxonomicName id="4C0C4D6FFFB8E114FF17FB0CFEEAADEE" box="[246,370,1234,1260]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="anglicus">
|
||
<emphasis id="B978EAFEFFB8E114FF17FB0CFE98ADEE" box="[246,256,1235,1260]" italics="true" pageId="2" pageNumber="3">I</emphasis>
|
||
.
|
||
<emphasis id="B978EAFEFFB8E114FEEFFB0CFEEAADEE" box="[270,370,1235,1260]" italics="true" pageId="2" pageNumber="3">anglicus</emphasis>
|
||
</taxonomicName>
|
||
do not share any synapomorphies with the
|
||
<emphasis id="B978EAFEFFB8E114FC8DFB0CFC2CADEE" box="[876,948,1235,1260]" italics="true" pageId="2" pageNumber="3">in situ</emphasis>
|
||
tooth of NHMUK 28660 to the exclusion of other iguanodontian specimens, such as NHMUK R1831,
|
||
<emphasis id="B978EAFEFFB8E114FCBEFB27FBA8AC13" box="[863,1072,1272,1297]" italics="true" pageId="2" pageNumber="3">Owenodon hoggii</emphasis>
|
||
(NHMUK R2998),
|
||
<taxonomicName id="4C0C4D6FFFB8E114FAFCFB27FED1AC34" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="B978EAFEFFB8E114FAFCFB27FED1AC34" italics="true" pageId="2" pageNumber="3">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
(MIWG 1997.55), and
|
||
<taxonomicName id="4C0C4D6FFFB8E114FD82FAC2FC52AC34" box="[611,970,1309,1334]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="B978EAFEFFB8E114FD82FAC2FC52AC34" box="[611,970,1309,1334]" italics="true" pageId="2" pageNumber="3">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK R5764). Secondly, although NHMUK 28660 and the teeth of
|
||
<taxonomicName id="4C0C4D6FFFB8E114FDC2FA9CFD38AC5E" box="[547,672,1346,1372]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="anglicus">
|
||
<emphasis id="B978EAFEFFB8E114FDC2FA9CFDB5AC5E" box="[547,557,1347,1372]" italics="true" pageId="2" pageNumber="3">I</emphasis>
|
||
.
|
||
<emphasis id="B978EAFEFFB8E114FDDDFA9CFD38AC5E" box="[572,672,1347,1372]" italics="true" pageId="2" pageNumber="3">anglicus</emphasis>
|
||
</taxonomicName>
|
||
both came from quarries near Cuckfield, it is evident that more than one lithostratigraphic horizon was exposed in those quarries (
|
||
<bibRefCitation id="EF9D4B1DFFB8E114FC25FAB7FB67AC80" author="Gallois" box="[964,1279,1384,1410]" pageId="2" pageNumber="3" refString="Gallois, R. W. & Worssam, B. C. (Eds) (1993) Geology of the country around Horsham: Memoir for 1: 50,000 geological sheet 302 (England and Wales). Her Majesty's Stationary Office, London, 130 pp." type="book" year="1993">Gallois & Worssam 1993</bibRefCitation>
|
||
). Among the numerous isolated and disarticulated iguanodontian bones found near Cuckfield, some are embedded in a tan, well sorted, fine-grained sandstone matrix, whilst others are in dark brown, poorly sorted, coarser sandstone. The disarticulated nature of the bones and presence of more than one bone-bearing layer make it impossible to refer disparate elements to the same taxon with any certainty. Considering these factors, it is best to continue to regard “
|
||
<taxonomicName id="4C0C4D6FFFB8E114FEFBF9FCFD94AF3E" box="[282,524,1571,1596]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="anglicus">
|
||
<emphasis id="B978EAFEFFB8E114FEFBF9FCFD94AF3E" box="[282,524,1571,1596]" italics="true" pageId="2" pageNumber="3">Iguanodon anglicus</emphasis>
|
||
</taxonomicName>
|
||
” as a
|
||
<emphasis id="B978EAFEFFB8E114FDBBF9FCFC96AF3E" box="[602,782,1571,1596]" italics="true" pageId="2" pageNumber="3">nomen dubium</emphasis>
|
||
and to create the novel genus and species
|
||
<emphasis id="B978EAFEFFB8E114FAF6F9FCFE88AF63" italics="true" pageId="2" pageNumber="3">Kukufeldia tilgatensis</emphasis>
|
||
for NHMUK 28660.
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB8E114FF76F94DFE52AFAE" blockId="2.[151,458,1682,1708]" box="[151,458,1682,1708]" pageId="2" pageNumber="3">Institutional abbreviations</paragraph>
|
||
<paragraph id="8BB336ECFFB8E114FF76F902FC3FAE1E" blockId="2.[151,1108,1757,2007]" pageId="2" pageNumber="3">
|
||
AMNH American Museum of Natural History, New York, NY,
|
||
<collectingCountry id="F31B767CFFB8E114FC57F902FC69AFF5" box="[950,1009,1757,1783]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
|
||
ANSP Academy of Natural Sciences, Philadelphia, PA,
|
||
<collectingCountry id="F31B767CFFB8E114FC8DF8DDFC3FAE1E" box="[876,935,1794,1820]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB8E114FF76F8F7FC96AEB0" blockId="2.[151,1108,1757,2007]" pageId="2" pageNumber="3">
|
||
CEUM College of Eastern Utah Prehistoric Museum, Price, UT,
|
||
<collectingCountry id="F31B767CFFB8E114FC28F8F7FB9CAE40" box="[969,1028,1832,1858]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
|
||
CM Carnegie Museum of Natural History, Pittsburgh, PA,
|
||
<collectingCountry id="F31B767CFFB8E114FC49F892FC7BAE65" box="[936,995,1869,1895]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
|
||
DMNH Denver Museum of Nature and Science, Denver, CO,
|
||
<collectingCountry id="F31B767CFFB8E114FC46F8ADFC7AAE8E" box="[935,994,1906,1932]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
|
||
HERM Hull and East Riding Museum, Hull,
|
||
<collectingCountry id="F31B767CFFB8E114FD05F847FC96AEB0" box="[740,782,1944,1970]" name="United Kingdom" pageId="2" pageNumber="3">UK</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB8E115FF76F862FC16A9FE" blockId="2.[151,1108,1757,2007]" lastBlockId="3.[151,1408,152,626]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
IRSNB Institut royal des Sciences naturelles de
|
||
<collectingCountry id="F31B767CFFB8E114FCE5F862FCE9AED5" box="[772,881,1981,2007]" name="Belgium" pageId="2" pageNumber="3">Belgique</collectingCountry>
|
||
, Brussels,
|
||
<collectingCountry id="F31B767CFFB8E114FC0EF862FBCCAED5" box="[1007,1108,1981,2007]" name="Belgium" pageId="2" pageNumber="3">Belgium</collectingCountry>
|
||
IVPP Institute of Vertebrate Paleontology and Paleoanthropology, Beijing,
|
||
<collectingCountry id="F31B767CFFB9E115FBB6FF47FB05A9B0" box="[1111,1181,152,178]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
|
||
MIWG Museum of Isle of Wight Geology (Dinosaur Isle Museum), Sandown,
|
||
<collectingCountry id="F31B767CFFB9E115FB90FF62FB03A9D5" box="[1137,1179,189,215]" name="United Kingdom" pageId="3" pageNumber="4">UK</collectingCountry>
|
||
MNHN Muséum national d’Histoire naturelle, Paris,
|
||
<collectingCountry id="F31B767CFFB9E115FCDFFF3DFC16A9FE" box="[830,910,226,252]" name="France" pageId="3" pageNumber="4">France</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FED7FC73A86E" blockId="3.[151,1408,152,626]" pageId="3" pageNumber="4">
|
||
MSM Mesa Southwest Museum (now Arizona Museum of Natural History), Mesa, AZ,
|
||
<collectingCountry id="F31B767CFFB9E115FB0FFED7FAB0A820" box="[1262,1320,264,290]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
NHMUK Natural History Museum (formerly BMNH, British Museum of Natural History), London,
|
||
<collectingCountry id="F31B767CFFB9E115FAB7FEF2FA18A845" box="[1366,1408,301,327]" name="United Kingdom" pageId="3" pageNumber="4">UK</collectingCountry>
|
||
OXFUM Oxford University Museum of Natural History, Oxford,
|
||
<collectingCountry id="F31B767CFFB9E115FC20FE8DFC73A86E" box="[961,1003,338,364]" name="United Kingdom" pageId="3" pageNumber="4">UK</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FEA7FCBFA890" blockId="3.[151,1408,152,626]" box="[151,807,376,402]" pageId="3" pageNumber="4">
|
||
PIN Palaeontological Institute, Moscow,
|
||
<collectingCountry id="F31B767CFFB9E115FD38FEA7FCBFA890" box="[729,807,376,402]" name="Russia" pageId="3" pageNumber="4">Russia</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FE42FCFBA8B5" blockId="3.[151,1408,152,626]" box="[151,867,413,439]" pageId="3" pageNumber="4">
|
||
QM Queensland Museum, South Brisbane,
|
||
<collectingCountry id="F31B767CFFB9E115FD19FE42FCFBA8B5" box="[760,867,413,439]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FE1DFC7CAB25" blockId="3.[151,1408,152,626]" pageId="3" pageNumber="4">
|
||
SDSM South Dakota School of Mines and Technology, Rapid City, SD,
|
||
<collectingCountry id="F31B767CFFB9E115FBC6FE1DFBFAA8DE" box="[1063,1122,450,476]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
SMU Southern Methodist University Shuler Museum of Paleontology, Dallas, TX,
|
||
<collectingCountry id="F31B767CFFB9E115FB59FE37FB6BAB00" box="[1208,1267,488,514]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
UMNH Utah Museum of Natural History, Salt Lake City, UT,
|
||
<collectingCountry id="F31B767CFFB9E115FC48FDD2FC7CAB25" box="[937,996,525,551]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FDEDFC60AB4E" blockId="3.[151,1408,152,626]" box="[151,1016,562,588]" pageId="3" pageNumber="4">
|
||
USNM National Museum of Natural History, Washington, DC,
|
||
<collectingCountry id="F31B767CFFB9E115FC5CFDEDFC60AB4E" box="[957,1016,562,588]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8BB336ECFFB9E115FF76FD87FB29AB70" blockId="3.[151,1408,152,626]" box="[151,1201,600,626]" pageId="3" pageNumber="4">
|
||
YPM Yale University Peabody Museum of Natural History, New Haven, CT,
|
||
<collectingCountry id="F31B767CFFB9E115FB97FD87FB29AB70" box="[1142,1201,600,626]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |