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<document id="2EA79DFB6F490EF56029B58A3F26DDFD" ID-DOI="10.5281/zenodo.5460634" ID-ISSN="2345-7600" ID-Zenodo-Dep="5460634" ID-ZooBank="1E7214C5-24E8-4860-BF13-203FD93A2C19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630239335276" checkinUser="valdenar" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2018" docId="03A587FA782AA037FCBBB83B379D536B" docLanguage="en" docName="66rbz763-781.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Macrochlamys brunnea Mollendorff 1902" docType="treatment" docVersion="3" lastPageNumber="775" masterDocId="FF9CFF827821A03BFFC7BF1034285476" masterDocTitle="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)" masterLastPageNumber="781" masterPageNumber="763" pageNumber="774" updateTime="1699195389588" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="73145F9A936D805E5AABD01761822FC3">The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)</mods:title>
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<mods:namePart id="D9B561789D92D8139BA3271AC322C144">Pholyotha, Arthit</mods:namePart>
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<taxonomicName id="4C0C4D6F782AA030FCBBB83B31745335" ID-CoL="3X42W" authority="Mollendorff, 1902" authorityName="Mollendorff" authorityYear="1902" box="[892,1372,1835,1859]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782AA030FCBBB83B31745335" bold="true" box="[892,1372,1835,1859]" pageId="11" pageNumber="774">
<emphasis id="B978EAFE782AA030FCBBB83B30AD5335" bold="true" box="[892,1157,1835,1859]" italics="true" pageId="11" pageNumber="774">Macrochlamys brunnea</emphasis>
Möllendorff, 1902
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(
<figureCitation id="13372A69782AA030FB86B85B30BE5315" box="[1089,1174,1867,1891]" captionStart="Fig" captionStartId="13.[113,148,1229,1251]" captionTargetBox="[113,1451,189,1212]" captionTargetId="figure-10@13.[113,1451,189,1212]" captionTargetPageId="13" captionText="Fig. 8. Shells of Macrochlamys brunnea Möllendorff, 1902 (A), and M. kelantanensis Möllendorff, 1902 (BD). A, Macrochlamys brunnea holotype (SMF 227090/1); B, M. kelantanensis holotype (SMF 227098); C, shell from Kedah, Malaysia (CUMZ 7130); D, shell from Chiang Rai, Thailand (CUMZ 7120)." figureDoi="http://doi.org/10.5281/zenodo.5460654" httpUri="https://zenodo.org/record/5460654/files/figure.png" pageId="11" pageNumber="774">Fig. 8A</figureCitation>
)
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<paragraph id="8BB336EC782AA030FCEAB89A318453CA" blockId="11.[813,1452,1930,1981]" pageId="11" pageNumber="774">
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<treatmentCitation id="0AAD10FD782AA030FCEAB89A313F53D6" author="Mollendorff O von" box="[813,1303,1930,1952]" page="155" pageId="11" pageNumber="774" year="1902">
<taxonomicName id="4C0C4D6F782AA030FCEAB89A313F53D6" ID-CoL="3X42W" authority="Mollendorff, 1902 b: 155" authorityName="Mollendorff" authorityPageNumber="155" authorityYear="1902" box="[813,1303,1930,1952]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782AA030FCEAB89A303F53E9" box="[813,1047,1930,1951]" italics="true" pageId="11" pageNumber="774">Macrochlamys brunnea</emphasis>
<bibRefCitation id="EF9D4B1D782AA030FBE7B89A313F53D6" author="Mollendorff O von" box="[1056,1303,1930,1952]" pageId="11" pageNumber="774" pagination="153 - 160" refId="ref13613" refString="Mollendorff O von (1902 b) Binnenmollusken aus Hinterindien. 2. Neue arten und unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 34: 153 - 160." type="journal article" year="1902">Möllendorff, 1902b: 155</bibRefCitation>
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(
<typeStatus id="54B7884E782AA030FAE0B89A317A53E9" box="[1319,1362,1930,1951]" pageId="11" pageNumber="774">type</typeStatus>
locality:
<collectingRegion id="49C8F80E782AA030FC95B8B7378253CB" box="[850,938,1959,1981]" country="Thailand" name="Krung Thep Mahanakhon" pageId="11" pageNumber="774">Bangkok</collectingRegion>
, Muok Lek, Hinlap)
<treatmentCitation id="0AAD10FD782AA030FBB9B8B7318053CA" author="Hemmen J &amp; Hemmen C" box="[1150,1448,1959,1980]" page="44" pageId="11" pageNumber="774" year="2001">
<bibRefCitation id="EF9D4B1D782AA030FBB9B8B7318053CA" author="Hemmen J &amp; Hemmen C" box="[1150,1448,1959,1980]" pageId="11" pageNumber="774" pagination="35 - 70" refId="ref13112" refString="Hemmen J &amp; Hemmen C (2001) Aktualisierte liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie aus dem Haus der Natur-Cismar, 18: 35 - 70." type="journal article" year="2001">Hemmen &amp; Hemmen, 2001: 44</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BB336EC782AA030FCEAB8F531835C4B" blockId="11.[813,1452,2021,2109]" pageId="11" pageNumber="774">
<emphasis id="B978EAFE782AA030FCEAB8F53025538B" bold="true" box="[813,1037,2021,2045]" pageId="11" pageNumber="774">Material examined.</emphasis>
<materialsCitation id="3B643CB1782AA030FBD2B8F530405C6B" collectionCode="SMF" country="Thailand" location="Bangkok" pageId="11" pageNumber="774" specimenCode="SMF 227090" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
<typeStatus id="54B7884E782AA030FBD2B8F53050538B" box="[1045,1144,2021,2045]" pageId="11" pageNumber="774" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBAA9E97782AA030FB4FB8F53137538B" box="[1160,1311,2021,2045]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="774">SMF 227090</specimenCode>
/1),
<collectingRegion id="49C8F80E782AA030FA83B8F5318F538B" box="[1348,1447,2021,2045]" country="Thailand" name="Krung Thep Mahanakhon" pageId="11" pageNumber="774">Bangkok</collectingRegion>
, Siam [
<collectingRegion id="49C8F80E782AA030FCBDB71537CE5C6B" box="[890,998,2053,2077]" country="Thailand" name="Krung Thep Mahanakhon" pageId="11" pageNumber="774">Bangkok</collectingRegion>
,
<collectingCountry id="F31B767C782AA030FC32B71530495C6B" box="[1013,1121,2053,2077]" name="Thailand" pageId="11" pageNumber="774">Thailand</collectingCountry>
]
</materialsCitation>
.
<materialsCitation id="3B643CB1782AA030FBB1B715318F5C4B" collectionCode="SMF" country="Thailand" location="Bangkok" pageId="11" pageNumber="774" specimenCode="SMF 227091, SMF 227171" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<typeStatus id="54B7884E782AA030FBB1B71530C15C6B" box="[1142,1257,2053,2077]" pageId="11" pageNumber="774" type="paratype">Paratypes</typeStatus>
: one shell (
<specimenCode id="DBAA9E97782AA030FAB3B71537AC5C4B" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="774">SMF 227091</specimenCode>
/1),
<collectingRegion id="49C8F80E782AA030FC61B73530215C4B" box="[934,1033,2085,2109]" country="Thailand" name="Krung Thep Mahanakhon" pageId="11" pageNumber="774">Bangkok</collectingRegion>
,
<collectingCountry id="F31B767C782AA030FBD4B735305D5C4B" box="[1043,1141,2085,2109]" name="Thailand" pageId="11" pageNumber="774">Thailand</collectingCountry>
; four shells (
<specimenCode id="DBAA9E97782AA030FB3CB73531A25C4B" box="[1275,1418,2085,2109]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="774">SMF 227171</specimenCode>
/4)
</materialsCitation>
,
</paragraph>
<caption id="DF736664782DA037FF4FBFAD37CA54A5" box="[136,994,189,211]" pageId="12" pageNumber="775" startId="12.[136,191,189,211]">
<paragraph id="8BB336EC782DA037FF4FBFAD37CA54A5" blockId="12.[136,994,189,211]" box="[136,994,189,211]" pageId="12" pageNumber="775">
Table 1. Comparison of shell and genital system of
<taxonomicName id="4C0C4D6F782DA037FD4DBFAD374B54A5" box="[650,867,189,211]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B978EAFE782DA037FD4DBFAD373154A4" box="[650,793,189,210]" italics="true" pageId="12" pageNumber="775">Macrochlamys</emphasis>
species
</taxonomicName>
in Thailand.
</paragraph>
</caption>
<paragraph id="8BB336EC782DA037FF50BE0331B2512D" pageId="12" pageNumber="775">
<table id="F90CC44C782D5FC4FF50BE14319B51E3" box="[151,1459,260,1429]" gridcols="8" gridrows="13" pageId="12" pageNumber="775">
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<emphasis id="B978EAFE782DA037FF50BE033522555F" bold="true" box="[151,266,275,297]" pageId="12" pageNumber="775">Characters</emphasis>
</th>
<th id="76ED5DD2782D5FC4FE9BBE1435C95541" box="[348,481,260,311]" gridcol="1" gridrow="0" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FEA5BE1535F25541" bold="true" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FEA5BE1535F2556C" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[354,474,261,282]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE782DA037FEA5BE1535F2556C" bold="true" box="[354,474,261,282]" italics="true" pageId="12" pageNumber="775">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782DA037FEA5BE3235F25541" box="[354,474,289,311]" pageId="12" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<th id="76ED5DD2782D5FC4FDC7BE1436AA5541" box="[512,642,260,311]" gridcol="2" gridrow="0" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FDD7BE1536555541" bold="true" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FDD7BE15365A556C" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[528,626,260,282]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE782DA037FDD7BE15365A556C" bold="true" box="[528,626,260,282]" italics="true" pageId="12" pageNumber="775">M. coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782DA037FDC2BE3236555541" box="[517,637,289,311]" pageId="12" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
<th id="76ED5DD2782D5FC4FD65BE1437325541" box="[674,794,260,311]" gridcol="3" gridrow="0" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FD62BE1537325541" bold="true" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FD62BE15373F556C" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[677,791,261,282]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE782DA037FD62BE15373F556C" bold="true" box="[677,791,261,282]" italics="true" pageId="12" pageNumber="775">M. caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782DA037FD65BE3237325541" box="[674,794,289,311]" pageId="12" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<th id="76ED5DD2782D5FC4FCFEBE14379F5541" box="[825,951,260,311]" gridcol="4" gridrow="0" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FC82BE15379C5541" bold="true" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FC82BE153783556C" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[837,939,260,282]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="lemma" status="sp. nov.">
<emphasis id="B978EAFE782DA037FC82BE153783556C" bold="true" box="[837,939,260,282]" italics="true" pageId="12" pageNumber="775">M. lemma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782DA037FCFBBE32379C5541" box="[828,948,289,311]" pageId="12" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
<th id="76ED5DD2782D5FC4FC1EBE1430505541" box="[985,1144,260,311]" gridcol="5" gridrow="0" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FC1EBE15304D5541" bold="true" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FC1EBE153050556C" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[985,1144,260,282]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782DA037FC1EBE153050556C" bold="true" box="[985,1144,260,282]" italics="true" pageId="12" pageNumber="775">M. tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782DA037FC2ABE32304D5541" box="[1005,1125,289,311]" pageId="12" pageNumber="775" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<taxonomicName id="4C0C4D6F782DA037FB06BE1530D15541" authorityName="Mollendorff" authorityYear="1902" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782DA037FB06BE1530D15541" bold="true" italics="true" pageId="12" pageNumber="775">M. brunnea</emphasis>
</taxonomicName>
</th>
<th id="76ED5DD2782D5FC4FAE1BE14319B5541" box="[1318,1459,260,311]" gridcol="7" gridrow="0" pageId="12" pageNumber="775">
<taxonomicName id="4C0C4D6F782DA037FA9ABE1531865541" authorityName="Mollendorff" authorityYear="1902" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="kelantanensis">
<emphasis id="B978EAFE782DA037FA9ABE1531865541" bold="true" italics="true" pageId="12" pageNumber="775">M. kelantanensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353C34AE782D5FC4FF50BE48319B55B2" box="[151,1459,344,452]" gridrow="1" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BE48351655B2" box="[151,318,344,452]" gridcol="0" gridrow="1" pageId="12" pageNumber="775">Shell width (mm) ranges, means ± SD and no. of specimens</th>
<td id="76ED5DD2782D5FC4FE9BBE4835C955B2" box="[348,481,344,452]" gridcol="1" gridrow="1" pageId="12" pageNumber="775">11.714.0 12.8 ± 0.98 (n=6)</td>
<td id="76ED5DD2782D5FC4FDC7BE4836AA55B2" box="[512,642,344,452]" gridcol="2" gridrow="1" pageId="12" pageNumber="775">10.213.0 11.6 ± 0.99 (n=6)</td>
<td id="76ED5DD2782D5FC4FD65BE48373255B2" box="[674,794,344,452]" gridcol="3" gridrow="1" pageId="12" pageNumber="775">13.217.2 14.3 ± 1.09 (n=11)</td>
<td id="76ED5DD2782D5FC4FCFEBE48379F55B2" box="[825,951,344,452]" gridcol="4" gridrow="1" pageId="12" pageNumber="775">13.117.0 14.7 ± 1.54 (n=6)</td>
<td id="76ED5DD2782D5FC4FC1EBE48305055B2" box="[985,1144,344,452]" gridcol="5" gridrow="1" pageId="12" pageNumber="775">9.611.1 10.6 ± 0.58 (n=5)</td>
<td id="76ED5DD2782D5FC4FB5ABE48312855B2" box="[1181,1280,344,452]" gridcol="6" gridrow="1" pageId="12" pageNumber="775">9.0*</td>
<td id="76ED5DD2782D5FC4FAE1BE48319B55B2" box="[1318,1459,344,452]" gridcol="7" gridrow="1" pageId="12" pageNumber="775">16.724.0 20.0 ± 2.56 (n=9)</td>
</tr>
<tr id="353C34AE782D5FC4FF50BECD319B563C" box="[151,1459,477,586]" gridrow="2" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BECD3516563C" box="[151,318,477,586]" gridcol="0" gridrow="2" pageId="12" pageNumber="775">Shell height (mm) ranges, means ± SD and no. of specimens</th>
<td id="76ED5DD2782D5FC4FE9BBECD35C9563C" box="[348,481,477,586]" gridcol="1" gridrow="2" pageId="12" pageNumber="775">6.58.2 7.5 ± 0.66 (n=6)</td>
<td id="76ED5DD2782D5FC4FDC7BECD36AA563C" box="[512,642,477,586]" gridcol="2" gridrow="2" pageId="12" pageNumber="775">5.76.6 6.0 ± 0.35 (n=6)</td>
<td id="76ED5DD2782D5FC4FD65BECD3732563C" box="[674,794,477,586]" gridcol="3" gridrow="2" pageId="12" pageNumber="775">7.38.6 7.6 ± 0.51 (n=11)</td>
<td id="76ED5DD2782D5FC4FCFEBECD379F563C" box="[825,951,477,586]" gridcol="4" gridrow="2" pageId="12" pageNumber="775">6.57.6 7.1 ± 0.40 (n=6)</td>
<td id="76ED5DD2782D5FC4FC1EBECD3050563C" box="[985,1144,477,586]" gridcol="5" gridrow="2" pageId="12" pageNumber="775">4.95.5 5.2 ± 0.27 (n=5)</td>
<td id="76ED5DD2782D5FC4FB5ABECD3128563C" box="[1181,1280,477,586]" gridcol="6" gridrow="2" pageId="12" pageNumber="775">5.0*</td>
<td id="76ED5DD2782D5FC4FAE1BECD319B563C" box="[1318,1459,477,586]" gridcol="7" gridrow="2" pageId="12" pageNumber="775">9.013.0 11.4 ± 1.21 (n=9)</td>
</tr>
<tr id="353C34AE782D5FC4FF50BD73319B56E3" box="[151,1459,611,661]" gridrow="3" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BD73351656E3" box="[151,318,611,661]" gridcol="0" gridrow="3" pageId="12" pageNumber="775">Shape of suture</th>
<td id="76ED5DD2782D5FC4FE9BBD7335C956E3" box="[348,481,611,661]" gridcol="1" gridrow="3" pageId="12" pageNumber="775">Deep channel</td>
<td id="76ED5DD2782D5FC4FDC7BD7336AA56E3" box="[512,642,611,661]" gridcol="2" gridrow="3" pageId="12" pageNumber="775">Shallow channel</td>
<td id="76ED5DD2782D5FC4FD65BD73373256E3" box="[674,794,611,661]" gridcol="3" gridrow="3" pageId="12" pageNumber="775">Impressed</td>
<td id="76ED5DD2782D5FC4FCFEBD73379F56E3" box="[825,951,611,661]" gridcol="4" gridrow="3" pageId="12" pageNumber="775">Impressed</td>
<td id="76ED5DD2782D5FC4FC1EBD73305056E3" box="[985,1144,611,661]" gridcol="5" gridrow="3" pageId="12" pageNumber="775">Impressed</td>
<td id="76ED5DD2782D5FC4FB5ABD73312856E3" box="[1181,1280,611,661]" gridcol="6" gridrow="3" pageId="12" pageNumber="775">Impressed</td>
<td id="76ED5DD2782D5FC4FAE1BD73319B56E3" box="[1318,1459,611,661]" gridcol="7" gridrow="3" pageId="12" pageNumber="775">Impressed</td>
</tr>
<tr id="353C34AE782D5FC4FF50BDBE319B5696" box="[151,1459,686,736]" gridrow="4" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BDBE35165696" box="[151,318,686,736]" gridcol="0" gridrow="4" pageId="12" pageNumber="775">Umbilicus</th>
<td id="76ED5DD2782D5FC4FE9BBDBE35C95696" box="[348,481,686,736]" gridcol="1" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
<td id="76ED5DD2782D5FC4FDC7BDBE36AA5696" box="[512,642,686,736]" gridcol="2" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
<td id="76ED5DD2782D5FC4FD65BDBE37325696" box="[674,794,686,736]" gridcol="3" gridrow="4" pageId="12" pageNumber="775">Widely open</td>
<td id="76ED5DD2782D5FC4FCFEBDBE379F5696" box="[825,951,686,736]" gridcol="4" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
<td id="76ED5DD2782D5FC4FC1EBDBE30505696" box="[985,1144,686,736]" gridcol="5" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
<td id="76ED5DD2782D5FC4FB5ABDBE31285696" box="[1181,1280,686,736]" gridcol="6" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
<td id="76ED5DD2782D5FC4FAE1BDBE319B5696" box="[1318,1459,686,736]" gridcol="7" gridrow="4" pageId="12" pageNumber="775">Narrowly open</td>
</tr>
<tr id="353C34AE782D5FC4FF50BDEA319B5766" box="[151,1459,762,784]" gridrow="5" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BDEA35165766" box="[151,318,762,784]" gridcol="0" gridrow="5" pageId="12" pageNumber="775">Penial caecum</th>
<td id="76ED5DD2782D5FC4FE9BBDEA35C95766" box="[348,481,762,784]" gridcol="1" gridrow="5" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FDC7BDEA36AA5766" box="[512,642,762,784]" gridcol="2" gridrow="5" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FD65BDEA37325766" box="[674,794,762,784]" gridcol="3" gridrow="5" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FCFEBDEA379F5766" box="[825,951,762,784]" gridcol="4" gridrow="5" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FC1EBDEA30505766" box="[985,1144,762,784]" gridcol="5" gridrow="5" pageId="12" pageNumber="775">Absent</td>
<td id="76ED5DD2782D5FC4FB5ABDEA31285766" box="[1181,1280,762,784]" gridcol="6" gridrow="5" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BDEA319B5766" box="[1318,1459,762,784]" gridcol="7" gridrow="5" pageId="12" pageNumber="775">Absent</td>
</tr>
<tr id="353C34AE782D5FC4FF50BC39319B57B9" box="[151,1459,809,975]" gridrow="6" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BC39351657B9" box="[151,318,809,975]" gridcol="0" gridrow="6" pageId="12" pageNumber="775">Inner wall of penis sculpture (proximal / distal)</th>
<td id="76ED5DD2782D5FC4FE9BBC3935C957B9" box="[348,481,809,975]" gridcol="1" gridrow="6" pageId="12" pageNumber="775">Both with small wrinkles</td>
<td id="76ED5DD2782D5FC4FDC7BC3936AA57B9" box="[512,642,809,975]" gridcol="2" gridrow="6" pageId="12" pageNumber="775">Small wrinkled / small longitudinal and wrinkled folds</td>
<td id="76ED5DD2782D5FC4FD65BC39373257B9" box="[674,794,809,975]" gridcol="3" gridrow="6" pageId="12" pageNumber="775">Trapezoid / lateral folds</td>
<td id="76ED5DD2782D5FC4FCFEBC39379F57B9" box="[825,951,809,975]" gridcol="4" gridrow="6" pageId="12" pageNumber="775">Trapezoid / lateral folds</td>
<td id="76ED5DD2782D5FC4FC1EBC39305057B9" box="[985,1144,809,975]" gridcol="5" gridrow="6" pageId="12" pageNumber="775">Both with weak wrinkles</td>
<td id="76ED5DD2782D5FC4FB5ABC39312857B9" box="[1181,1280,809,975]" gridcol="6" gridrow="6" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BC39319B57B9" box="[1318,1459,809,975]" gridcol="7" gridrow="6" pageId="12" pageNumber="775">Both with weak wrinkles</td>
</tr>
<tr id="353C34AE782D5FC4FF50BCF8319B5788" box="[151,1459,1000,1022]" gridrow="7" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BCF835165788" box="[151,318,1000,1022]" gridcol="0" gridrow="7" pageId="12" pageNumber="775">Epiphallus</th>
<td id="76ED5DD2782D5FC4FE9BBCF835C95788" box="[348,481,1000,1022]" gridcol="1" gridrow="7" pageId="12" pageNumber="775">Short</td>
<td id="76ED5DD2782D5FC4FDC7BCF836AA5788" box="[512,642,1000,1022]" gridcol="2" gridrow="7" pageId="12" pageNumber="775">Short</td>
<td id="76ED5DD2782D5FC4FD65BCF837325788" box="[674,794,1000,1022]" gridcol="3" gridrow="7" pageId="12" pageNumber="775">Short</td>
<td id="76ED5DD2782D5FC4FCFEBCF8379F5788" box="[825,951,1000,1022]" gridcol="4" gridrow="7" pageId="12" pageNumber="775">Rather long</td>
<td id="76ED5DD2782D5FC4FC1EBCF830505788" box="[985,1144,1000,1022]" gridcol="5" gridrow="7" pageId="12" pageNumber="775">Very long</td>
<td id="76ED5DD2782D5FC4FB5ABCF831285788" box="[1181,1280,1000,1022]" gridcol="6" gridrow="7" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BCF8319B5788" box="[1318,1459,1000,1022]" gridcol="7" gridrow="7" pageId="12" pageNumber="775">Slightly long</td>
</tr>
<tr id="353C34AE782D5FC4FF50BB07319B503F" box="[151,1459,1047,1097]" gridrow="8" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BB073516503F" box="[151,318,1047,1097]" gridcol="0" gridrow="8" pageId="12" pageNumber="775">Epiphallic caecum</th>
<td id="76ED5DD2782D5FC4FE9BBB0735C9503F" box="[348,481,1047,1097]" gridcol="1" gridrow="8" pageId="12" pageNumber="775">One circle</td>
<td id="76ED5DD2782D5FC4FDC7BB0736AA503F" box="[512,642,1047,1097]" gridcol="2" gridrow="8" pageId="12" pageNumber="775">One circle</td>
<td id="76ED5DD2782D5FC4FD65BB073732503F" box="[674,794,1047,1097]" gridcol="3" gridrow="8" pageId="12" pageNumber="775">One circle</td>
<td id="76ED5DD2782D5FC4FCFEBB07379F503F" box="[825,951,1047,1097]" gridcol="4" gridrow="8" pageId="12" pageNumber="775">One circle</td>
<td id="76ED5DD2782D5FC4FC1EBB073050503F" box="[985,1144,1047,1097]" gridcol="5" gridrow="8" pageId="12" pageNumber="775">One circle</td>
<td id="76ED5DD2782D5FC4FB5ABB073128503F" box="[1181,1280,1047,1097]" gridcol="6" gridrow="8" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BB07319B503F" box="[1318,1459,1047,1097]" gridcol="7" gridrow="8" pageId="12" pageNumber="775">Two circles</td>
</tr>
<tr id="353C34AE782D5FC4FF50BB72319B500E" box="[151,1459,1122,1144]" gridrow="9" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BB723516500E" box="[151,318,1122,1144]" gridcol="0" gridrow="9" pageId="12" pageNumber="775">Tissue casing</th>
<td id="76ED5DD2782D5FC4FE9BBB7235C9500E" box="[348,481,1122,1144]" gridcol="1" gridrow="9" pageId="12" pageNumber="775">Absent</td>
<td id="76ED5DD2782D5FC4FDC7BB7236AA500E" box="[512,642,1122,1144]" gridcol="2" gridrow="9" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FD65BB723732500E" box="[674,794,1122,1144]" gridcol="3" gridrow="9" pageId="12" pageNumber="775">Absent</td>
<td id="76ED5DD2782D5FC4FCFEBB72379F500E" box="[825,951,1122,1144]" gridcol="4" gridrow="9" pageId="12" pageNumber="775">Present</td>
<td id="76ED5DD2782D5FC4FC1EBB723050500E" box="[985,1144,1122,1144]" gridcol="5" gridrow="9" pageId="12" pageNumber="775">Absent</td>
<td id="76ED5DD2782D5FC4FB5ABB723128500E" box="[1181,1280,1122,1144]" gridcol="6" gridrow="9" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BB72319B500E" box="[1318,1459,1122,1144]" gridcol="7" gridrow="9" pageId="12" pageNumber="775">Absent</td>
</tr>
<tr id="353C34AE782D5FC4FF50BB81319B50D1" box="[151,1459,1169,1191]" gridrow="10" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BB81351650D1" box="[151,318,1169,1191]" gridcol="0" gridrow="10" pageId="12" pageNumber="775">Vagina length</th>
<td id="76ED5DD2782D5FC4FE9BBB8135C950D1" box="[348,481,1169,1191]" gridcol="1" gridrow="10" pageId="12" pageNumber="775">Long</td>
<td id="76ED5DD2782D5FC4FDC7BB8136AA50D1" box="[512,642,1169,1191]" gridcol="2" gridrow="10" pageId="12" pageNumber="775">Slightly long</td>
<td id="76ED5DD2782D5FC4FD65BB81373250D1" box="[674,794,1169,1191]" gridcol="3" gridrow="10" pageId="12" pageNumber="775">Long</td>
<td id="76ED5DD2782D5FC4FCFEBB81379F50D1" box="[825,951,1169,1191]" gridcol="4" gridrow="10" pageId="12" pageNumber="775">Slightly long</td>
<td id="76ED5DD2782D5FC4FC1EBB81305050D1" box="[985,1144,1169,1191]" gridcol="5" gridrow="10" pageId="12" pageNumber="775">Short</td>
<td id="76ED5DD2782D5FC4FB5ABB81312850D1" box="[1181,1280,1169,1191]" gridcol="6" gridrow="10" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BB81319B50D1" box="[1318,1459,1169,1191]" gridcol="7" gridrow="10" pageId="12" pageNumber="775">Short</td>
</tr>
<tr id="353C34AE782D5FC4FF50BBD0319B5166" box="[151,1459,1216,1296]" gridrow="11" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BBD035165166" box="[151,318,1216,1296]" gridcol="0" gridrow="11" pageId="12" pageNumber="775">Position of thicken tissue on free oviduct</th>
<td id="76ED5DD2782D5FC4FE9BBBD035C95166" box="[348,481,1216,1296]" gridcol="1" gridrow="11" pageId="12" pageNumber="775">Base</td>
<td id="76ED5DD2782D5FC4FDC7BBD036AA5166" box="[512,642,1216,1296]" gridcol="2" gridrow="11" pageId="12" pageNumber="775">Base</td>
<td id="76ED5DD2782D5FC4FD65BBD037325166" box="[674,794,1216,1296]" gridcol="3" gridrow="11" pageId="12" pageNumber="775">One-fourth from base</td>
<td id="76ED5DD2782D5FC4FCFEBBD0379F5166" box="[825,951,1216,1296]" gridcol="4" gridrow="11" pageId="12" pageNumber="775">Near middle</td>
<td id="76ED5DD2782D5FC4FC1EBBD030505166" box="[985,1144,1216,1296]" gridcol="5" gridrow="11" pageId="12" pageNumber="775">Base</td>
<td id="76ED5DD2782D5FC4FB5ABBD031285166" box="[1181,1280,1216,1296]" gridcol="6" gridrow="11" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BBD0319B5166" box="[1318,1459,1216,1296]" gridcol="7" gridrow="11" pageId="12" pageNumber="775">Base</td>
</tr>
<tr id="353C34AE782D5FC4FF50BA38319B51E3" box="[151,1459,1320,1429]" gridrow="12" pageId="12" pageNumber="775">
<th id="76ED5DD2782D5FC4FF50BA38351651E3" box="[151,318,1320,1429]" gridcol="0" gridrow="12" pageId="12" pageNumber="775">Location of dart apparatus</th>
<td id="76ED5DD2782D5FC4FE9BBA3835C951E3" box="[348,481,1320,1429]" gridcol="1" gridrow="12" pageId="12" pageNumber="775">Proximal end of vagina</td>
<td id="76ED5DD2782D5FC4FDC7BA3836AA51E3" box="[512,642,1320,1429]" gridcol="2" gridrow="12" pageId="12" pageNumber="775">Proximal end of vagina</td>
<td id="76ED5DD2782D5FC4FD65BA38373251E3" box="[674,794,1320,1429]" gridcol="3" gridrow="12" pageId="12" pageNumber="775">Proximal end of vagina</td>
<td id="76ED5DD2782D5FC4FCFEBA38379F51E3" box="[825,951,1320,1429]" gridcol="4" gridrow="12" pageId="12" pageNumber="775">Near proximal end of vagina</td>
<td id="76ED5DD2782D5FC4FC1EBA38305051E3" box="[985,1144,1320,1429]" gridcol="5" gridrow="12" pageId="12" pageNumber="775">Proximal end of vagina</td>
<td id="76ED5DD2782D5FC4FB5ABA38312851E3" box="[1181,1280,1320,1429]" gridcol="6" gridrow="12" pageId="12" pageNumber="775"></td>
<td id="76ED5DD2782D5FC4FAE1BA38319B51E3" box="[1318,1459,1320,1429]" gridcol="7" gridrow="12" pageId="12" pageNumber="775">Proximal end of vagina</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB336EC782DA037FF4FBAA737B551BB" blockId="12.[136,925,1463,1485]" box="[136,925,1463,1485]" pageId="12" pageNumber="775">
*Shell measurement taken from the original description of
<bibRefCitation id="EF9D4B1D782DA037FD0ABAA737B151BB" author="Mollendorff O von" box="[717,921,1463,1485]" pageId="12" pageNumber="775" pagination="153 - 160" refId="ref13613" refString="Mollendorff O von (1902 b) Binnenmollusken aus Hinterindien. 2. Neue arten und unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 34: 153 - 160." type="journal article" year="1902">Möllendorff (1902b)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB336EC782DA037FF4FB91534DF524B" blockId="12.[136,775,1541,1597]" pageId="12" pageNumber="775">
Muok Lek,
<collectingCountry id="F31B767C782DA037FEC9B915356D526B" box="[270,325,1541,1565]" name="Thailand" pageId="12" pageNumber="775">Siam</collectingCountry>
[Muak Lek District,
<collectingRegion id="49C8F80E782DA037FDFEB915372B526B" box="[569,771,1541,1565]" country="Thailand" name="Sara Buri" pageId="12" pageNumber="775">Saraburi Province</collectingRegion>
,
<collectingCountry id="F31B767C782DA037FF4FB93534C5524B" box="[136,237,1573,1597]" name="Thailand" pageId="12" pageNumber="775">Thailand</collectingCountry>
].
</paragraph>
</subSubSection>
<subSubSection id="C3166567782DA037FF4FB975379D536B" pageId="12" pageNumber="775" type="discussion">
<paragraph id="8BB336EC782DA037FF4FB97535C953EB" blockId="12.[135,776,1637,1949]" pageId="12" pageNumber="775">
<emphasis id="B978EAFE782DA037FF4FB97534DE520B" bold="true" box="[136,246,1637,1661]" pageId="12" pageNumber="775">Remarks.</emphasis>
<taxonomicName id="4C0C4D6F782DA037FF3CB97535DD520B" authorityName="Mollendorff" authorityYear="1902" box="[251,501,1637,1661]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782DA037FF3CB97535DD520B" box="[251,501,1637,1661]" italics="true" pageId="12" pageNumber="775">Macrochlamys brunnea</emphasis>
</taxonomicName>
differs from its congeners in
<collectingCountry id="F31B767C782DA037FF6FB995352452EB" box="[168,268,1669,1693]" name="Thailand" pageId="12" pageNumber="775">Thailand</collectingCountry>
by its small size and globose shell (
<tableCitation id="C68E0357782DA037FD03B99534BD52CB" captionStart="Table 1" captionStartId="12.[136,191,189,211]" captionText="Table 1. Comparison of shell and genital system of Macrochlamys species in Thailand." pageId="12" pageNumber="775">Table 1</tableCitation>
). Other distinguishing characters of this species are the depressed globose, shiny dark brown shell with a smooth surface and rounded periphery. The shell has a low conical spire, which is rather convex, with a slightly raised apex, and shallow suture. Its aperture is ovate-lunate in shape, opens obliquely and has a simple peristome. The columellar margin is gradually reflected near the narrowly open and very small umbilicus (
<figureCitation id="13372A69782DA037FEBAB89535FC53EB" box="[381,468,1925,1949]" captionStart="Fig" captionStartId="13.[113,148,1229,1251]" captionTargetBox="[113,1451,189,1212]" captionTargetId="figure-10@13.[113,1451,189,1212]" captionTargetPageId="13" captionText="Fig. 8. Shells of Macrochlamys brunnea Möllendorff, 1902 (A), and M. kelantanensis Möllendorff, 1902 (BD). A, Macrochlamys brunnea holotype (SMF 227090/1); B, M. kelantanensis holotype (SMF 227098); C, shell from Kedah, Malaysia (CUMZ 7130); D, shell from Chiang Rai, Thailand (CUMZ 7120)." figureDoi="http://doi.org/10.5281/zenodo.5460654" httpUri="https://zenodo.org/record/5460654/files/figure.png" pageId="12" pageNumber="775">Fig. 8A</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC782DA037FF4FB8D5379D536B" blockId="12.[135,775,1989,2077]" lastBlockId="12.[834,1474,1541,1821]" pageId="12" pageNumber="775">
The generic placement of this species is tentative and requires verification since anatomical data is thus far unavailable. It is herein provisionally retained in the
<taxonomicName id="4C0C4D6F782DA037FD8FB71536CE5C6B" authorityName="Gray" authorityYear="1847" box="[584,742,2053,2077]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="genus">
<emphasis id="B978EAFE782DA037FD8FB71536CE5C6B" box="[584,742,2053,2077]" italics="true" pageId="12" pageNumber="775">Macrochlamys</emphasis>
</taxonomicName>
as originally proposed although this species can be assigned to either
<taxonomicName id="4C0C4D6F782DA037FC6EB93537C7524B" authorityName="Godwin-Austen" authorityYear="1907" box="[937,1007,1573,1597]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="genus">
<emphasis id="B978EAFE782DA037FC6EB93537C7524B" box="[937,1007,1573,1597]" italics="true" pageId="12" pageNumber="775">Sarika</emphasis>
</taxonomicName>
or
<taxonomicName id="4C0C4D6F782DA037FBDFB935309E524B" authorityName="Gray" authorityYear="1847" box="[1048,1206,1573,1597]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="genus">
<emphasis id="B978EAFE782DA037FBDFB935309E524B" box="[1048,1206,1573,1597]" italics="true" pageId="12" pageNumber="775">Macrochlamys</emphasis>
</taxonomicName>
if based solely on shell morphology. Unfortunately, attempts to obtain topotypic specimens from
<collectingRegion id="49C8F80E782DA037FC34B975307B520B" box="[1011,1107,1637,1661]" country="Thailand" name="Krung Thep Mahanakhon" pageId="12" pageNumber="775">Bangkok</collectingRegion>
and accessible limestone outcrops in the
<collectingRegion id="49C8F80E782DA037FC45B995301752EB" box="[898,1087,1669,1693]" country="Thailand" name="Sara Buri" pageId="12" pageNumber="775">Saraburi Province</collectingRegion>
were unsuccessful and no specimens identifiable as
<taxonomicName id="4C0C4D6F782DA037FC20B9B5304F52CB" authorityName="Mollendorff" authorityYear="1902" box="[999,1127,1701,1725]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="775" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782DA037FC20B9B5304F52CB" box="[999,1127,1701,1725]" italics="true" pageId="12" pageNumber="775">M. brunnea</emphasis>
</taxonomicName>
could be found. This could be due to habitat loss and population decline as many limestone outcrops in this region are being quarried. Its current status is unclear.
</paragraph>
</subSubSection>
</treatment>
</document>