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<document id="E8C9FB87716329E801C48D3C988F0194" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A87FFFFB0CBA1554FB8A910A" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus obscurus Sharp" docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="29" updateTime="1698775642300" updateUser="ExternalLinkService">
<mods:mods id="4146B7470989A6131CEC6E513BC05BC9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4616FF4E223140BA3BE5D787C2966E4">
<mods:title id="AEAC8FB2DA0CA7B7A60CB81244ED155E">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="B602EC6AC1CC2EDEEF3D4EEB23541C2B" type="personal">
<mods:role id="5557025F4DB5590C928C27ADA0A8450A">
<mods:roleTerm id="27F4E62410FC6905BE35E7532CA38F60">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0676AA545CB0C163C392D52AF7D7C971">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="09376D9890BA9040475C21EFC50301E7">text</mods:typeOfResource>
<mods:relatedItem id="2CE785CB4852B812FA91232307326FF6" type="host">
<mods:titleInfo id="4785455BCE92F61311AC7EA49A19369B">
<mods:title id="E502A820730E30E8ED39D711011BB5B3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="742BCD58A9A4CC4F0BF565A860D4F1FA">
<mods:date id="E5BF84835BDED9116BF4128D562AFB1B">2019</mods:date>
<mods:detail id="80B75B3F75A4FA83F58C08AD65321D76" type="pubDate">
<mods:number id="382D8A708219E55067FB76ACF9711B37">2019-12-12</mods:number>
</mods:detail>
<mods:detail id="4076D94420D06D5085052A97D8409EE6" type="volume">
<mods:number id="FA3C0CEB36C832ABE5FFFA48756A978F">4708</mods:number>
</mods:detail>
<mods:detail id="8BCDC41401D5209BFA0876ACD13E3521" type="issue">
<mods:number id="A9DBAD7BA0641A6E2A633C5F66A6C9B3">1</mods:number>
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<mods:extent id="225FE11E32B565B007325051370DEABA" unit="page">
<mods:start id="B56CC3EF76139A9DAC0B5B693123CB6B">1</mods:start>
<mods:end id="A90B460F9D90263A4115444906BBEFDC">59</mods:end>
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<mods:classification id="4E897BE4374FB3C09AC29ABE05610687">journal article</mods:classification>
<mods:identifier id="60BB8AA1206D25297F408472AC56EC55" type="CLB-Dataset">24540</mods:identifier>
<mods:identifier id="BC23A10EE5061D1C06335EF18A249D56" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
<mods:identifier id="730A3DA46CF37E7ADC65170E62E41AB8" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
<mods:identifier id="A8EB534B8130B04DF2704A40BFEE1CBE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F1BFF6528916E6E1428DA28BD098BDAC" type="Zenodo-Dep">3587181</mods:identifier>
<mods:identifier id="2B1A1B70D9D22D3DE71681FAF32928FE" type="ZooBank">09F24F0B-5090-4089-A525-8E5150C91EF7</mods:identifier>
</mods:mods>
<treatment id="03A587E7A87FFFFB0CBA1554FB8A910A" ID-GBIF-Taxon="160798398" LSID="urn:lsid:plazi:treatment:03A587E7A87FFFFB0CBA1554FB8A910A" httpUri="http://treatment.plazi.org/id/03A587E7A87FFFFB0CBA1554FB8A910A" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<subSubSection id="C316657AA87FFFF80CBA1554FE07957F" box="[151,486,1808,1835]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BB336F1A87FFFF80CBA1554FE07957F" blockId="28.[151,486,1808,1871]" box="[151,486,1808,1835]" pageId="28" pageNumber="29">
<heading id="D0FB819DA87FFFF80CBA1554FE07957F" bold="true" box="[151,486,1808,1835]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<taxonomicName id="4C0C4D72A87FFFF80CBA1554FE07957F" ID-CoL="6MK96" authority="Sharp" authorityName="Sharp" box="[151,486,1808,1835]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B978EAE3A87FFFF80CBA1554FE07957F" bold="true" box="[151,486,1808,1835]" pageId="28" pageNumber="29">
<emphasis id="B978EAE3A87FFFF80CBA1554FE70957E" bold="true" box="[151,401,1808,1835]" italics="true" pageId="28" pageNumber="29">Hydrochus obscurus</emphasis>
Sharp
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87FFFF80CBA1570FEEC951A" box="[151,269,1844,1871]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB336F1A87FFFF80CBA1570FEEC951A" blockId="28.[151,486,1808,1871]" box="[151,269,1844,1871]" pageId="28" pageNumber="29">
<figureCitation id="13372A74A87FFFF80CBA1570FF04951A" box="[151,229,1844,1871]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="28" pageNumber="29">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A87FFFF80CDC1571FEEC951A" box="[241,269,1845,1871]" captionStart="FIGURE 18" captionStartId="36.[151,250,1885,1910]" captionTargetBox="[226,1371,295,1841]" captionTargetId="figure@36.[204,1384,273,1860]" captionTargetPageId="36" captionText="FIGURE 18. Hydrochus obscurus Sharp, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587219" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="28" pageNumber="29">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87FFFF80CBA153DFCF195E5" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8BB336F1A87FFFF80CBA153DFDEE95C4" blockId="28.[151,784,1912,1968]" box="[151,527,1912,1937]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB1C11DFA87FFFF80CBA153DFDEE95C4" box="[151,527,1912,1937]" pageId="28" pageNumber="29">
<taxonomicName id="4C0C4D72A87FFFF80CBA153DFDEA95C4" ID-CoL="6MK96" authority="Sharp 1882: 90" authorityName="Sharp" authorityPageNumber="90" authorityYear="1882" box="[151,523,1912,1937]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B978EAE3A87FFFF80CBA153DFE8695C4" box="[151,359,1913,1937]" italics="true" pageId="28" pageNumber="29">Hydrochus obscurus</emphasis>
<treatmentCitation id="0AAD10E0A87FFFF80D43153CFDEA95C4" author="Sharp, D." box="[366,523,1912,1937]" page="90" pageId="28" pageNumber="29" year="1882">
<bibRefCitation id="EF9D4B00A87FFFF80D43153CFDEA95C4" author="Sharp, D." box="[366,523,1912,1937]" pageId="28" pageNumber="29" refId="ref30814" refString="Sharp, D. (1882) Biologia Centrali-Americana (Coleoptera). Insecta Coleoptera. Vol. 1. Part 2. Published for the editors by R. H. Porter, London, 824 pp., 19 tabs." type="book" year="1882">Sharp 1882: 90</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB336F1A87FFFF80CBA15D3FCF195E5" blockId="28.[151,784,1912,1968]" box="[151,784,1943,1968]" pageId="28" pageNumber="29">
<taxonomicName id="4C0C4D72A87FFFF80CBA15D3FD6595E5" ID-CoL="3N6CV" authority="Makhan 1992" authorityName="Makhan" authorityYear="1992" box="[151,644,1943,1968]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vanbergehenegouweni">
<emphasis id="B978EAE3A87FFFF80CBA15D3FE0E95FA" box="[151,495,1943,1967]" italics="true" pageId="28" pageNumber="29">Hydrochus vanbergehenegouweni</emphasis>
<bibRefCitation id="EF9D4B00A87FFFF80DD815D3FD6595E5" author="Makhan, D." box="[501,644,1943,1968]" pageId="28" pageNumber="29" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF9D4B00A87FFFF80EA315D3FCF195E5" author="Oliva, A." box="[654,784,1943,1968]" pageId="28" pageNumber="29" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87FFFFA0CBA1599FA7A95F3" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB336F1A87FFFF80CBA1599FA7D95AD" blockId="28.[151,1436,2013,2040]" box="[151,1436,2013,2040]" pageId="28" pageNumber="29">
<emphasis id="B978EAE3A87FFFF80CBA1599FE2595AD" bold="true" box="[151,452,2013,2040]" pageId="28" pageNumber="29">Material examined (65):</emphasis>
<materialsCitation id="3B643CACA87FFFF80DFD159AFBF095AD" ID-GBIF-Occurrence="2521263873" box="[464,1041,2013,2040]" collectionCode="SEMC" country="Venezuela" location="Aragua" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Aragua">
<emphasis id="B978EAE3A87FFFF80DFD159AFD2195AD" bold="true" box="[464,704,2013,2040]" pageId="28" pageNumber="29">
<collectingCountry id="F31B7661A87FFFF80DFD159AFDAC95AD" box="[464,589,2014,2040]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A87FFFF80E701599FD5D95AD" box="[605,700,2013,2040]" country="Venezuela" name="Aragua" pageId="28" pageNumber="29">Aragua</collectingRegion>
:
</emphasis>
VZ09-0104-01A (1
<collectionCode id="ED1DAE34A87FFFF80F95159AFBE895AD" box="[952,1033,2014,2040]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="28" pageNumber="29" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<emphasis id="B978EAE3A87FFFF8080C1599FB5395AD" bold="true" box="[1057,1202,2013,2040]" pageId="28" pageNumber="29">
<collectingRegion id="49C8F813A87FFFF8080C1599FB4C95AD" box="[1057,1197,2013,2040]" country="Venezuela" name="Anzoategui" pageId="28" pageNumber="29">Anzoategui</collectingRegion>
:
</emphasis>
VZ10-0203-03A (1
</paragraph>
<paragraph id="8BB336F1A87EFFF90CBA12DDFDA193DE" blockId="29.[151,1437,152,395]" pageId="29" pageNumber="30">
<materialsCitation id="3B643CACA87EFFF90CBA12DDFE179282" ID-GBIF-Occurrence="2521263813" collectingDate="1989-01-25" collectionCode="SEMC, NMNH" collectorName="Faitoute" location="Rio Horero" municipality="Puerto Ayacucho" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Amazonas">
<collectionCode id="ED1DAE34A87EFFF90CBA12DDFF0992E6" box="[151,232,153,179]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
).
<emphasis id="B978EAE3A87EFFF90CD112DCFE6392E6" bold="true" box="[252,386,152,179]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90CD112DCFE9C92E6" box="[252,381,152,179]" country="Venezuela" name="Amazonas" pageId="29" pageNumber="30">Amazonas</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BD7AC8BA87EFFF90DA612DDFDAB92E6" box="[395,586,153,179]" pageId="29" pageNumber="30">Puerto Ayacucho</collectingMunicipality>
(
<quantity id="4CF49B14A87EFFF90E7612DDFD4492E6" box="[603,677,153,179]" metricMagnitude="4" metricUnit="m" metricValue="6.6" pageId="29" pageNumber="30" unit="km" value="66.0">66 km</quantity>
N),
<location id="8ED3602AA87EFFF90EF712DDFCBF92E6" LSID="urn:lsid:plazi:treatment:03A587E7A87FFFFB0CBA1554FB8A910A:8ED3602AA87EFFF90EF712DDFCBF92E6" box="[730,862,153,179]" municipality="Puerto Ayacucho" name="Rio Horero" pageId="29" pageNumber="30" stateProvince="Amazonas">Rio Horero</location>
,
<date id="FFB21031A87EFFF90F4612DCFC0692E6" box="[875,999,152,179]" pageId="29" pageNumber="30" value="1989-01-25">
<collectingDate id="EFF6E9D9A87EFFF90F4612DCFC0692E6" box="[875,999,152,179]" pageId="29" pageNumber="30" value="1989-01-25">25Jan1989</collectingDate>
</date>
, netted, stream edge, PJSpangler, RA-
<collectorName id="26F95327A87EFFF90CBA12F9FF129282" box="[151,243,189,215]" pageId="29" pageNumber="30">Faitoute</collectorName>
&amp; CBBarr (2
<collectionCode id="ED1DAE34A87EFFF90DBE12F9FE0F9282" box="[403,494,189,215]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA87EFFF90E2C12F9FBE69282" ID-GBIF-Occurrence="2521263900" box="[513,1031,188,215]" collectingDate="1965-04-12" collectionCode="NMNH" collectorName="F. Fernandez" location="Ocamo" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Amazonas">
<location id="8ED3602AA87EFFF90E2C12F9FDB49282" LSID="urn:lsid:plazi:treatment:03A587E7A87FFFFB0CBA1554FB8A910A:8ED3602AA87EFFF90E2C12F9FDB49282" box="[513,597,189,215]" name="Ocamo" pageId="29" pageNumber="30" stateProvince="Amazonas">Ocamo</location>
,
<date id="FFB21031A87EFFF90E7212F8FD059282" box="[607,740,188,215]" pageId="29" pageNumber="30" value="1965-04-12">
<collectingDate id="EFF6E9D9A87EFFF90E7212F8FD059282" box="[607,740,188,215]" pageId="29" pageNumber="30" value="1965-04-12">12-IV-1965</collectingDate>
</date>
,
<collectorName id="26F95327A87EFFF90EC212F9FC9E9282" box="[751,895,189,215]" pageId="29" pageNumber="30">F. Fernandez</collectorName>
(1
<collectionCode id="ED1DAE34A87EFFF90F8912F9FC1E9282" box="[932,1023,189,215]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF9083C12F8FA879282" ID-GBIF-Occurrence="2521263804" box="[1041,1382,188,215]" collectionCode="SEMC" location="Apure" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Apure">
<emphasis id="B978EAE3A87EFFF9083C12F8FB879282" bold="true" box="[1041,1126,188,215]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF9083C12F8FB809282" box="[1041,1121,188,215]" country="Venezuela" name="Apure" pageId="29" pageNumber="30">Apure</collectingRegion>
:
</emphasis>
AS-06-009 (1
<collectionCode id="ED1DAE34A87EFFF9092012F9FABC9282" box="[1293,1373,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
, AS- 06-019 (2
<collectionCode id="ED1DAE34A87EFFF90D2612A5FEBA92AE" box="[267,347,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
),
<materialsCitation id="3B643CACA87EFFF90D4312A5FD4692AE" ID-GBIF-Occurrence="2521263822" box="[366,679,225,251]" collectingDate="2003-10" collectionCode="SEMC" pageId="29" pageNumber="30" specimenCode="VZ09-0118" specimenCount="1" stateProvince="Apure">
<specimenCode id="DBAA9E8AA87EFFF90D4312A5FE1392AE" box="[366,498,225,251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">VZ09-0118</specimenCode>
-
<date id="FFB21031A87EFFF90DD412A5FDC892AE" box="[505,553,225,251]" pageId="29" pageNumber="30" value="2003-10">
<collectingDate id="EFF6E9D9A87EFFF90DD412A5FDC892AE" box="[505,553,225,251]" pageId="29" pageNumber="30" value="2003-10">03X</collectingDate>
</date>
(2
<collectionCode id="ED1DAE34A87EFFF90E6012A5FD7F92AE" box="[589,670,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF90E9D12A4FBB492AE" ID-GBIF-Occurrence="2521263824" box="[688,1109,224,251]" collectionCode="SEMC" location="Barinas" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Barinas">
<emphasis id="B978EAE3A87EFFF90E9D12A4FCF692AE" bold="true" box="[688,791,224,251]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90E9D12A4FCF392AE" box="[688,786,224,251]" country="Venezuela" name="Barinas" pageId="29" pageNumber="30">Barinas</collectingRegion>
:
</emphasis>
VZ12-0126-01B (2
<collectionCode id="ED1DAE34A87EFFF90FD112A5FBAC92AE" box="[1020,1101,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA87EFFF9084D12A4FA7992AE" ID-GBIF-Occurrence="2521263808" box="[1120,1432,224,251]" collectionCode="SEMC" pageId="29" pageNumber="30" specimenCode="VZ12-0125, 02" specimenCount="1" stateProvince="Barinas">
<specimenCode id="DBAA9E8AA87EFFF9084D12A4FB0392AE" box="[1120,1250,224,251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">VZ12-0125</specimenCode>
-
<specimenCode id="DBAA9E8AA87EFFF908C412A4FAE792AE" box="[1257,1286,224,251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">02</specimenCode>
A (1
<collectionCode id="ED1DAE34A87EFFF9091212A5FA7192AE" box="[1343,1424,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA87EFFF90CBA1340FE35934A" ID-GBIF-Occurrence="2521263868" box="[151,468,260,287]" collectionCode="SEMC" pageId="29" pageNumber="30" specimenCode="VZ12-0125, 01" specimenCount="1" stateProvince="Barinas">
<specimenCode id="DBAA9E8AA87EFFF90CBA1340FEFB934A" box="[151,282,260,287]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">VZ12-0125</specimenCode>
-
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A (1
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)
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.
<materialsCitation id="3B643CACA87EFFF90DCD1340FC66934A" ID-GBIF-Occurrence="2521263829" box="[480,903,260,287]" collectingDate="2002-10" collectionCode="SEMC" location="Bolivar" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Bolivar">
<emphasis id="B978EAE3A87EFFF90DCD1340FDA3934A" bold="true" box="[480,578,260,287]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90DCD1340FDDC934A" box="[480,573,260,287]" country="Venezuela" name="Bolivar" pageId="29" pageNumber="30">Bolivar</collectingRegion>
:
</emphasis>
VZ09-0111-
<date id="FFB21031A87EFFF90EF91341FCE4934A" box="[724,773,261,287]" pageId="29" pageNumber="30" value="2002-10">
<collectingDate id="EFF6E9D9A87EFFF90EF91341FCE4934A" box="[724,773,261,287]" pageId="29" pageNumber="30" value="2002-10">02X</collectingDate>
</date>
(1
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)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF90FBE1341FA78934A" ID-GBIF-Occurrence="2521263893" box="[915,1433,260,287]" collectionCode="SEMC" location="Delta Amacuro" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Delta Amacuro">
<emphasis id="B978EAE3A87EFFF90FBE1341FBB2934A" bold="true" box="[915,1107,260,287]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90FBE1341FBAF934A" box="[915,1102,260,287]" country="Venezuela" name="Delta Amacuro" pageId="29" pageNumber="30">Delta Amacuro</collectingRegion>
:
</emphasis>
VZ10-0203-01A (1
<collectionCode id="ED1DAE34A87EFFF9096D1341FA70934A" box="[1344,1425,261,287]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF90CBA136CFDA39316" ID-GBIF-Occurrence="2521263817" box="[151,578,296,323]" collectingDate="2003-10" collectionCode="SEMC" location="Guarico" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Guarico">
<emphasis id="B978EAE3A87EFFF90CBA136CFEE39316" bold="true" box="[151,258,296,323]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90CBA136CFF1C9316" box="[151,253,296,323]" country="Venezuela" name="Guarico" pageId="29" pageNumber="30">Guarico</collectingRegion>
:
</emphasis>
VZ09-0108-
<date id="FFB21031A87EFFF90DB9136DFE249316" box="[404,453,297,323]" pageId="29" pageNumber="30" value="2003-10">
<collectingDate id="EFF6E9D9A87EFFF90DB9136DFE249316" box="[404,453,297,323]" pageId="29" pageNumber="30" value="2003-10">03X</collectingDate>
</date>
(2
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)
</materialsCitation>
;
<materialsCitation id="3B643CACA87EFFF90E60136DFAC99316" ID-GBIF-Occurrence="2521263845" box="[589,1320,297,324]" collectingDate="1984-06-11" collectionCode="NMNH" collectorName="F. W. Eiland" location="Corozo" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Pando">
<location id="8ED3602AA87EFFF90E60136DFD7E9316" LSID="urn:lsid:plazi:treatment:03A587E7A87FFFFB0CBA1554FB8A910A:8ED3602AA87EFFF90E60136DFD7E9316" box="[589,671,297,323]" name="Corozo" pageId="29" pageNumber="30" stateProvince="Pando">Corozo</location>
<collectingRegion id="49C8F813A87EFFF90E88136DFD0F9316" box="[677,750,297,323]" country="Bolivia" name="Pando" pageId="29" pageNumber="30">Pando</collectingRegion>
, (8kmN),
<date id="FFB21031A87EFFF90F72136DFC149316" box="[863,1013,297,323]" pageId="29" pageNumber="30" value="1984-06-11">
<collectingDate id="EFF6E9D9A87EFFF90F72136DFC149316" box="[863,1013,297,323]" pageId="29" pageNumber="30" value="1984-06-11">11 June 1984</collectingDate>
</date>
,
<collectorName id="26F95327A87EFFF9082D136DFB699316" box="[1024,1160,297,323]" pageId="29" pageNumber="30">F. W. Eiland</collectorName>
II (1
<collectionCode id="ED1DAE34A87EFFF908E8136DFAC19316" box="[1221,1312,297,323]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA87EFFF9091E136DFC149332" ID-GBIF-Occurrence="2521263815" collectingDate="1986-03-05" collectionCode="NMNH" location="Hato Masaguaral" municipality="Calabozo" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Pando">
<collectingMunicipality id="6BD7AC8BA87EFFF9091E136DFA7C9316" box="[1331,1437,297,323]" pageId="29" pageNumber="30">Calabozo</collectingMunicipality>
(
<quantity id="4CF49B14A87EFFF90CB21309FF069332" box="[159,231,333,359]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="29" pageNumber="30" unit="km" value="44.0">44 km</quantity>
S),
<location id="8ED3602AA87EFFF90D381309FE399332" LSID="urn:lsid:plazi:treatment:03A587E7A87FFFFB0CBA1554FB8A910A:8ED3602AA87EFFF90D381309FE399332" box="[277,472,333,359]" municipality="Calabozo" name="Hato Masaguaral" pageId="29" pageNumber="30" stateProvince="Pando">Hato Masaguaral</location>
,
<date id="FFB21031A87EFFF90DCF1308FD869332" box="[482,615,332,359]" pageId="29" pageNumber="30" value="1986-03-05">
<collectingDate id="EFF6E9D9A87EFFF90DCF1308FD869332" box="[482,615,332,359]" pageId="29" pageNumber="30" value="1986-03-05">5 Mar 1986</collectingDate>
</date>
, PJSpangler, colln#24 (23
<collectionCode id="ED1DAE34A87EFFF90FBF1309FC0C9332" box="[914,1005,333,359]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF90FD21308FA789332" ID-GBIF-Occurrence="2521263833" box="[1023,1433,332,359]" collectionCode="SEMC" location="Sucre" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Sucre">
<emphasis id="B978EAE3A87EFFF90FD21308FBAD9332" bold="true" box="[1023,1100,332,359]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90FD21308FBA69332" box="[1023,1095,332,359]" country="Venezuela" name="Sucre" pageId="29" pageNumber="30">Sucre</collectingRegion>
:
</emphasis>
VZ10-0129-04A (24
<collectionCode id="ED1DAE34A87EFFF9096D1309FA709332" box="[1344,1425,333,359]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87EFFF90CBA1334FDDC93DE" ID-GBIF-Occurrence="2521263828" box="[151,573,368,395]" collectionCode="SEMC" location="Tachira" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Tachira">
<emphasis id="B978EAE3A87EFFF90CBA1334FF1C93DE" bold="true" box="[151,253,368,395]" pageId="29" pageNumber="30">
<collectingRegion id="49C8F813A87EFFF90CBA1334FF1993DE" box="[151,248,368,395]" country="Venezuela" name="Tachira" pageId="29" pageNumber="30">Tachira</collectingRegion>
:
</emphasis>
VZ12-0126-02A (1
<collectionCode id="ED1DAE34A87EFFF90DCE1335FDD593DE" box="[483,564,369,395]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="29" pageNumber="30" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF736679A87EFFF90CBA1589FC3A95B3" ID-DOI="http://doi.org/10.5281/zenodo.3587207" ID-Zenodo-Dep="3587207" box="[151,987,1997,2022]" httpUri="https://zenodo.org/record/3587207/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1997,2022]" targetBox="[234,1373,450,1944]" targetPageId="29">
<paragraph id="8BB336F1A87EFFF90CBA1589FC3A95B3" blockId="29.[151,987,1997,2022]" box="[151,987,1997,2022]" pageId="29" pageNumber="30">
<emphasis id="B978EAE3A87EFFF90CBA1589FEC195B3" bold="true" box="[151,288,1997,2022]" pageId="29" pageNumber="30">FIGURE 13.</emphasis>
<taxonomicName id="4C0C4D72A87EFFF90D0B158AFDE695B3" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[294,519,1998,2022]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sagittarius" status="sp. nov.">
<emphasis id="B978EAE3A87EFFF90D0B158AFDE695B3" box="[294,519,1998,2022]" italics="true" pageId="29" pageNumber="30">Hydrochus sagittarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A87EFFF90E20158AFDA795B0" box="[525,582,1998,2021]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<caption id="DF736679A87DFFFA0CBA14B9FC869543" ID-DOI="http://doi.org/10.5281/zenodo.3587209" ID-Zenodo-Dep="3587209" box="[151,871,1789,1814]" httpUri="https://zenodo.org/record/3587209/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1789,1814]" targetBox="[190,1404,210,1750]" targetPageId="30">
<paragraph id="8BB336F1A87DFFFA0CBA14B9FC869543" blockId="30.[151,871,1789,1814]" box="[151,871,1789,1814]" pageId="30" pageNumber="31">
<emphasis id="B978EAE3A87DFFFA0CBA14B9FEC19543" bold="true" box="[151,288,1789,1814]" pageId="30" pageNumber="31">FIGURE 14.</emphasis>
<taxonomicName id="4C0C4D72A87DFFFA0D0B14BAFDAD9543" authority="Knisch" authorityName="Knisch" box="[294,588,1789,1814]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B978EAE3A87DFFFA0D0B14BAFE759543" box="[294,404,1790,1814]" italics="true" pageId="30" pageNumber="31">Hydrochus</emphasis>
<emphasis id="B978EAE3A87DFFFA0DB614BAFE1D9543" box="[411,508,1790,1814]" italics="true" pageId="30" pageNumber="31">variabilis</emphasis>
Knisch
</taxonomicName>
, habitus and male genitalia.
</paragraph>
</caption>
<paragraph id="8BB336F1A87DFFFA0CEA1507FA7A95F3" blockId="30.[151,1437,1859,2031]" pageId="30" pageNumber="31">
<emphasis id="B978EAE3A87DFFFA0CEA1507FE30950B" bold="true" box="[199,465,1859,1886]" pageId="30" pageNumber="31">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of moderate size (ca.
<quantity id="4CF49B14A87DFFFA09701500FF1E95D6" metricMagnitude="-3" metricUnit="m" metricValue="3.29" metricValueMax="3.66" metricValueMin="2.92" pageId="30" pageNumber="31" unit="mm" value="3.29" valueMax="3.66" valueMin="2.92">2.92– 3.66 mm</quantity>
), the moderately low and relatively impunctate pronotal reliefs, and the male genitalia (
<figureCitation id="13372A74A87DFFFA08FF152CFAC495D7" box="[1234,1317,1896,1922]" captionStart="FIGURE 18" captionStartId="36.[151,250,1885,1910]" captionTargetBox="[226,1371,295,1841]" captionTargetId="figure@36.[204,1384,273,1860]" captionTargetPageId="36" captionText="FIGURE 18. Hydrochus obscurus Sharp, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587219" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="30" pageNumber="31">Fig. 18</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C0C4D72A87DFFFA0E6615C8FD2495F3" authorityName="Leach" authorityYear="1817" box="[587,709,1932,1958]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A87DFFFA0E6615C8FD2495F3" box="[587,709,1932,1958]" italics="true" pageId="30" pageNumber="31">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87DFFFB0CEA15EBFD5590A6" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="8BB336F1A87DFFFB0CEA15EBFC63934A" blockId="30.[151,1437,1859,2031]" lastBlockId="31.[151,1437,153,863]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="B978EAE3A87DFFFA0CEA15EBFEB0959F" bold="true" box="[199,337,1967,1994]" pageId="30" pageNumber="31">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A87DFFFA0E7B15F4FD49959F" box="[598,680,1968,1994]" captionStart="FIGURE 18" captionStartId="36.[151,250,1885,1910]" captionTargetBox="[226,1371,295,1841]" captionTargetId="figure@36.[204,1384,273,1860]" captionTargetPageId="36" captionText="FIGURE 18. Hydrochus obscurus Sharp, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587219" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="30" pageNumber="31">Fig. 18</figureCitation>
). Size of illustrated male (length/width, mm): body (length to elytral apices) 3.14/1.17; head width 0.77; pronotum l/w 0.71/0.71; PA 0.69; PB 0.63; elytra 1.94 /1.17. Body size moderate (ca.
<quantity id="4CF49B14A87CFFFB0CEA12DDFE8D92E6" box="[199,364,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.29" metricValueMax="3.66" metricValueMin="2.92" pageId="31" pageNumber="32" unit="mm" value="3.29" valueMax="3.66" valueMin="2.92">2.92–3.66 mm</quantity>
, Venezuelan specimens). Dorsum grey, with slight to moderately strong iridescence, elytra with black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–
<date id="FFB21031A87CFFFB084012F9FB669282" box="[1133,1159,189,215]" pageId="31" pageNumber="32">3x</date>
that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
<superScript id="7C799BB9A87CFFFB0F0412A4FCD792BB" attach="left" box="[809,822,224,238]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A87CFFFB093112A4FAC892BB" attach="left" box="[1308,1321,224,238]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C799BB9A87CFFFB0D801340FE5B9347" attach="left" box="[429,442,260,274]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
moderately raised, moderately elongate.
</paragraph>
<paragraph id="8BB336F1A87CFFFB0CEA136DFA7393DE" blockId="31.[151,1437,153,863]" pageId="31" pageNumber="32">Pronotum length and width equal, or only very slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep, much more densely punctate than low reliefs between depressions, latter almost entirely impunctate.</paragraph>
<paragraph id="8BB336F1A87CFFFB0CBA13D1FCBE9386" blockId="31.[151,1437,153,863]" pageId="31" pageNumber="32">
Elytra with spaces between strial punctures ca. 0.5–
<date id="FFB21031A87CFFFB0EC513D0FCE193FA" box="[744,768,404,431]" pageId="31" pageNumber="32">1x</date>
their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.
</paragraph>
<paragraph id="8BB336F1A87CFFFB0CEA1399FB1E93A2" blockId="31.[151,1437,153,863]" box="[199,1279,477,503]" pageId="31" pageNumber="32">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BB336F1A87CFFFB0CEA1045FDFA90D2" blockId="31.[151,1437,153,863]" pageId="31" pageNumber="32">
Male genitalia (
<figureCitation id="13372A74A87CFFFB0D5B1045FE27904E" box="[374,454,513,539]" captionStart="FIGURE 18" captionStartId="36.[151,250,1885,1910]" captionTargetBox="[226,1371,295,1841]" captionTargetId="figure@36.[204,1384,273,1860]" captionTargetPageId="36" captionText="FIGURE 18. Hydrochus obscurus Sharp, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587219" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
) general characters: paramere tips in dorsal and ventral views very weakly sub-sagittate, plsa rounded, lateral margins slightly emarginate, medial margins arcuate toward midline, in lateral view tips narrow and tapering; aedeagus basically lanceolate shaped; basal piece with orifice sclerotized rim produced in small projection on apicoventral margin.
</paragraph>
<paragraph id="8BB336F1A87CFFFB0CEA10D5FB6290FE" blockId="31.[151,1437,153,863]" box="[199,1155,657,683]" pageId="31" pageNumber="32">Dorsal surface: adtl at about distal 1/3; adbl moderately size; pdmm overlapping alm.</paragraph>
<paragraph id="8BB336F1A87CFFFB0CEA10F1FD5590A6" blockId="31.[151,1437,153,863]" pageId="31" pageNumber="32">Ventral surface: aedeagus ventral transition line (avtl) at about proximal 1/3. Paramere medial margins (pvmm) closely approximating aedeagus lateral margins.</paragraph>
</subSubSection>
<subSubSection id="C316657AA87CFFFB0CEA10B9FB8A910A" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BB336F1A87CFFFB0CEA10B9FB8A910A" blockId="31.[151,1437,153,863]" pageId="31" pageNumber="32">
<emphasis id="B978EAE3A87CFFFB0CEA10B9FEDA9142" bold="true" box="[199,315,765,791]" pageId="31" pageNumber="32">Remarks.</emphasis>
The specimen imaged (
<figureCitation id="13372A74A87CFFFB0E6610B9FD7F9142" box="[587,670,765,791]" captionStart="FIGURE 18" captionStartId="36.[151,250,1885,1910]" captionTargetBox="[226,1371,295,1841]" captionTargetId="figure@36.[204,1384,273,1860]" captionTargetPageId="36" captionText="FIGURE 18. Hydrochus obscurus Sharp, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587219" httpUri="https://zenodo.org/record/3587219/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
) is from locality VZ12-0126-01B. I have examined the
<typeStatus id="54B78853A87CFFFB093610B9FA9C9142" box="[1307,1405,765,791]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0C4D72A87CFFFB0CBA1165FD58916E" authority="Makhan 1992" authorityName="Makhan" authorityYear="1992" box="[151,697,801,827]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanbergehenegouweni">
<emphasis id="B978EAE3A87CFFFB0CBA1165FDF2916E" box="[151,531,801,827]" italics="true" pageId="31" pageNumber="32">Hydrochus vanbergehenegouweni</emphasis>
<bibRefCitation id="EF9D4B00A87CFFFB0E361165FD58916E" author="Makhan, D." box="[539,697,801,827]" pageId="31" pageNumber="32" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992</bibRefCitation>
</taxonomicName>
, including the male genitalia. I concur with
<bibRefCitation id="EF9D4B00A87CFFFB08841165FAD89169" author="Oliva, A." box="[1193,1337,801,828]" pageId="31" pageNumber="32" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
that
<taxonomicName id="4C0C4D72A87CFFFB095F1166FDB4910A" authority="Makhan" authorityName="Makhan" authorityYear="1992" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanbergehenegouweni">
<emphasis id="B978EAE3A87CFFFB095F1166FE13910A" italics="true" pageId="31" pageNumber="32">Hydrochus vanbergehenegouweni</emphasis>
Makhan
</taxonomicName>
is conspecific with
<taxonomicName id="4C0C4D72A87CFFFB0F181101FB8A910A" authority="Sharp." authorityName="Sharp." box="[821,1131,837,863]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B978EAE3A87CFFFB0F181101FBFA910A" box="[821,1051,837,863]" italics="true" pageId="31" pageNumber="32">Hydrochus obscurus</emphasis>
Sharp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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