322 lines
36 KiB
XML
322 lines
36 KiB
XML
<document id="0E57E35E19717C87CEE9E611315BAE98" ID="10.11646/zootaxa.4139.3.1" ID-DOI="10.11646/zootaxa.4139.3.1" ID-GBIF-Dataset="ae883579-c6f5-4ee8-9c4e-22a4c13552b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="262893" ID-ZooBank="7D131A9B-0DF4-4B80-97FF-AAB8966CC7EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469091365757" checkinUser="plazi" docAuthor="Pirani, Gabriela & Amorim, Dalton De Souza" docDate="2016" docId="039987F43C12A201FF669181FC92F805" docLanguage="en" docName="zootaxa.4139.3.1.pdf" docOrigin="Zootaxa 4139 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladochaeta nebulosa" docType="treatment" docVersion="7" lastPageNumber="338" masterDocId="FFA0FF8C3C33A224FFF19036FFFFFF94" masterDocTitle="Going beyond the tip of the Drosophilidae iceberg: New Cladochaeta Coquillett, 1900 (Diptera: Drosophilidae) from Brazil" masterLastPageNumber="344" masterPageNumber="301" pageNumber="334" updateTime="1698365442733" updateUser="plazi">
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<mods:mods id="D8507CC35CBB13DA7C8FD7030592FC74" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AA67E8044B142377B547BC83FA5A3515">
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<mods:title id="89363DFFBF41B5E865C70ADB79C3C1B5">Going beyond the tip of the Drosophilidae iceberg: New Cladochaeta Coquillett, 1900 (Diptera: Drosophilidae) from Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="8A923774CFAC8AF10938EB291861974A" type="personal">
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<mods:roleTerm id="87BF464B168986B438B46A777E063247">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7FD332A8413282D044BBCF4C3F0FE5E2">Pirani, Gabriela</mods:namePart>
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</mods:name>
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<mods:name id="42929A4CC70F2B0A5B8673C35E300580" type="personal">
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<mods:role id="80E6BC8E0E5357850C7314741F8884DC">
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<mods:roleTerm id="CA8B14AE9FB9FAF198B700571AAA616E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DE70A68A3AA10B9957E94E8F4BC97C2A">Amorim, Dalton De Souza</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="913F98DE514C3067055169423E29AE86">text</mods:typeOfResource>
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<mods:titleInfo id="D6A038CDCB3AEFD7F977E0F246CA2D9D">
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<mods:title id="EAFC6DA738CFCDAB925B7C91A6011F8D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FB584FC8AE4077B218D1BF874600D2AA">
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<mods:date id="50FFE24F22C889738BE0D4E2DF637A98">2016</mods:date>
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<mods:detail id="664234C6DA319D228B46DAF99AF79AF3" type="volume">
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<mods:number id="C79E4507C4FB9DDD7EF3F97491676078">4139</mods:number>
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</mods:detail>
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<mods:detail id="E9E86D474BFC26EB231663F94C6CFF98" type="issue">
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<mods:number id="18E41543D8726D5B19DE3D7C6135E7D4">3</mods:number>
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</mods:detail>
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<mods:start id="A4EE516A422593761C4689583FB21DEC">301</mods:start>
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<mods:end id="29C11425D3D036ACE9231DA104C9ABC1">344</mods:end>
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</mods:extent>
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<mods:classification id="287E2A5E7561A95E9D76D8BEEBD88480">journal article</mods:classification>
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<mods:identifier id="7C533AFF8A226DD4E4C7AFEF63E83936" type="DOI">10.11646/zootaxa.4139.3.1</mods:identifier>
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<mods:identifier id="992EDEFD77DB9C2EAD74DD2FFE07179E" type="GBIF-Dataset">ae883579-c6f5-4ee8-9c4e-22a4c13552b6</mods:identifier>
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<mods:identifier id="AFF91FA88451C469BA04DE35AFCA2ABC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A42AC8B35C8445C0ADF25DA5A1439C4B" type="Zenodo-Dep">262893</mods:identifier>
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<mods:identifier id="754BF30C480B90F46A584CCF237882A7" type="ZooBank">7D131A9B-0DF4-4B80-97FF-AAB8966CC7EA</mods:identifier>
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</mods:mods>
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<treatment id="039987F43C12A201FF669181FC92F805" ID-DOI="http://doi.org/10.5281/zenodo.6083807" ID-GBIF-Taxon="123198535" ID-Zenodo-Dep="6083807" LSID="urn:lsid:plazi:treatment:039987F43C12A201FF669181FC92F805" httpUri="http://treatment.plazi.org/id/039987F43C12A201FF669181FC92F805" lastPageId="37" lastPageNumber="338" pageId="33" pageNumber="334">
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<subSubSection id="C32A65693C12A205FF669181FE03FD8C" pageId="33" pageNumber="334" type="nomenclature">
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<paragraph id="8B8F36E23C12A205FF669181FD00FE46" blockId="33.[151,767,439,466]" box="[151,767,439,466]" pageId="33" pageNumber="334">
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<heading id="D0C7818E3C12A205FF669181FD00FE46" bold="true" box="[151,767,439,466]" fontSize="11" level="1" pageId="33" pageNumber="334" reason="1">
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<emphasis id="B944EAF03C12A205FF669181FD00FE46" bold="true" box="[151,767,439,466]" pageId="33" pageNumber="334">
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||
<taxonomicName id="4C304D613C12A205FF669181FEFBFE45" ID-CoL="5YWCG" box="[151,260,439,465]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="nebulosa">
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<emphasis id="B944EAF03C12A205FF669181FEFBFE45" bold="true" box="[151,260,439,465]" italics="true" pageId="33" pageNumber="334">nebulosa</emphasis>
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</taxonomicName>
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species group
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<bibRefCitation id="EFA14B133C12A205FE4A918EFD00FE46" author="Grimaldi" box="[443,767,440,466]" pageId="33" pageNumber="334" refString="Grimaldi, D. A. & Nguyen, T. (1999) Monograph on the spittlebug flies, genus Cladochaeta (Diptera: Drosophilidae: Cladochaetini). Bulletin of the American Museum of Natural History, 241, 1 - 326." type="journal article" year="1999">Grimaldi & Nguyen, 1999</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F36E23C12A205FF6691C8FE03FD8C" blockId="33.[151,623,510,570]" box="[151,508,510,536]" pageId="33" pageNumber="334">
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<emphasis id="B944EAF03C12A205FF6691C8FE64FD8C" bold="true" box="[151,411,510,536]" italics="true" pageId="33" pageNumber="334">
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<taxonomicName id="4C304D613C12A205FF6691C8FED1FD8C" box="[151,302,510,536]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">Cladochaeta</taxonomicName>
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stigmata
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</emphasis>
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<emphasis id="B944EAF03C12A205FE5291C9FE03FD8C" bold="true" box="[419,508,511,536]" pageId="33" pageNumber="334">nov. sp.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A65693C12A205FF669217FD90FDAD" box="[151,623,545,570]" pageId="33" pageNumber="334" type="description">
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||
<paragraph id="8B8F36E23C12A205FF669217FD90FDAD" blockId="33.[151,623,510,570]" box="[151,623,545,570]" pageId="33" pageNumber="334">
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||
(
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<figureCitation id="130B2A673C12A205FF6E9217FF05FDAD" box="[159,250,545,570]" captionStart="FIGURE 30" captionStartId="33.[151,250,1962,1984]" captionTargetBox="[157,1423,958,1930]" captionTargetId="figure@33.[150,1437,946,1941]" captionTargetPageId="33" captionText="FIGURE 30. Cladochaeta stigmata, nov. sp., holotype. A. Head, dorsal view. B. Head, lateral view. C. Head, frontal view. D. Thorax, lateral view. E. Thorax, dorsal view. F. Abdomen, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/262923/files/figure.png" pageId="33" pageNumber="334">Figs. 30</figureCitation>
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A–F; 31 A–F; 17 B; 34 D; 36 C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A65693C12A205FF66925FFE5CFD7A" pageId="33" pageNumber="334" type="diagnosis">
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<paragraph id="8B8F36E23C12A205FF66925FFE5CFD7A" blockId="33.[151,1437,617,894]" pageId="33" pageNumber="334">
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<emphasis id="B944EAF03C12A205FF66925FFEEEFD16" bold="true" box="[151,273,617,642]" pageId="33" pageNumber="334">Diagnosis.</emphasis>
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Thorax light yellow, abdomen ochre; wing with infuscation over costal region and dm-cu crossvein; legs pale yellow to whitish; arista with only dorsal basal branch; phallus only partially sclerotized: broad bulbous membranous portion between two thin asymmetrical, sclerotized arms, membrane projected anteriorly, reaching phallapodeme medially.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A65693C12A205FF3692CFFDFBFCEA" pageId="33" pageNumber="334" type="materials_examined">
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<paragraph id="8B8F36E23C12A205FF3692CFFDFBFCEA" blockId="33.[151,1437,617,894]" pageId="33" pageNumber="334">
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<emphasis id="B944EAF03C12A205FF3692CFFDDAFC86" bold="true" box="[199,549,761,786]" pageId="33" pageNumber="334">
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Material examined.
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<typeStatus id="548B88403C12A205FE4992CFFDDAFC86" box="[440,549,761,786]" pageId="33" pageNumber="334" type="holotype">Holotype</typeStatus>
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</emphasis>
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Ƌ: (Left wing and antenna slide-mounted, terminalia on vial),
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<collectingCountry id="F32776723C12A205FB1C92CCFAAAFC86" box="[1261,1365,762,786]" name="Brazil" pageId="33" pageNumber="334">BRAZIL</collectingCountry>
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,
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<emphasis id="B944EAF03C12A205FA9092CFFEC7FCA2" bold="true" pageId="33" pageNumber="334">
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State of
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<collectingRegion id="49F4F8003C12A205FF4A932BFEC7FCA2" box="[187,312,797,822]" country="Brazil" name="Sao Paulo" pageId="33" pageNumber="334">São Paulo</collectingRegion>
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</emphasis>
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, Gália, E.E. Caetetus,
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<geoCoordinate id="EE0450253C12A205FDA19328FD2DFCA2" box="[592,722,798,822]" direction="south" orientation="latitude" pageId="33" pageNumber="334" precision="15" value="-22.291388">22º17’29”S</geoCoordinate>
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<geoCoordinate id="EE0450253C12A205FD2E9328FC93FCA2" box="[735,876,798,822]" direction="west" orientation="longitude" pageId="33" pageNumber="334" precision="15" value="-49.552776">49º33’10”W</geoCoordinate>
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,
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<date id="FF8E10223C12A205FC8C932BFC06FCA2" box="[893,1017,797,822]" pageId="33" pageNumber="334" value="2010-03-12">12.iii.2010</date>
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, Sweeping, Bertoni col. (
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<collectionCode id="ED21AE273C12A205FADC9328FA6EFCA2" box="[1325,1425,798,822]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="33" pageNumber="334">MZUSP</collectionCode>
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).
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<emphasis id="B944EAF03C12A205FF669377FEFBFCCE" bold="true" box="[151,260,833,858]" pageId="33" pageNumber="334">
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<typeStatus id="548B88403C12A205FF669377FEFBFCCE" box="[151,260,833,858]" pageId="33" pageNumber="334" type="paratype">Paratype</typeStatus>
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</emphasis>
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Ƌ: (terminalia on vial),
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<collectingCountry id="F32776723C12A205FDD79374FD71FCCE" box="[550,654,834,858]" name="Brazil" pageId="33" pageNumber="334">BRAZIL</collectingCountry>
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,
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<emphasis id="B944EAF03C12A205FD6D9377FC82FCCE" bold="true" box="[668,893,833,858]" pageId="33" pageNumber="334">
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State of
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<collectingRegion id="49F4F8003C12A205FCF29377FC82FCCE" box="[771,893,833,858]" country="Brazil" name="Sao Paulo" pageId="33" pageNumber="334">São Paulo</collectingRegion>
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</emphasis>
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, Serra do Japi,
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<date id="FF8E10223C12A205FBC99377FB4CFCCE" box="[1080,1203,833,858]" pageId="33" pageNumber="334" value="2010-03-18">18.iii.2010</date>
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, Malaise trap, N.W. Perioto and team coll. (
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<collectionCode id="ED21AE273C12A205FE6A9350FE03FCEA" box="[411,508,870,894]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="33" pageNumber="334">MZUSP</collectionCode>
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)
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</paragraph>
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</subSubSection>
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<caption id="DF4F666A3C12A205FF66979CFBD0F84B" httpUri="https://zenodo.org/record/262923/files/figure.png" pageId="33" pageNumber="334" targetBox="[157,1423,958,1930]" targetPageId="33">
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<paragraph id="8B8F36E23C12A205FF66979CFBD0F84B" blockId="33.[151,1436,1962,2015]" pageId="33" pageNumber="334">
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<emphasis id="B944EAF03C12A205FF66979CFEDFF82B" bold="true" box="[151,288,1962,1984]" pageId="33" pageNumber="334">FIGURE 30.</emphasis>
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<emphasis id="B944EAF03C12A205FED7979CFDFAF854" box="[294,517,1962,1984]" italics="true" pageId="33" pageNumber="334">
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<taxonomicName id="4C304D613C12A205FED7979CFE58F854" box="[294,423,1962,1984]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">Cladochaeta</taxonomicName>
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stigmata
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</emphasis>
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,
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<emphasis id="B944EAF03C12A205FDE3979DFD9BF854" bold="true" box="[530,612,1963,1984]" pageId="33" pageNumber="334">nov. sp.</emphasis>
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, holotype.
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<emphasis id="B944EAF03C12A205FD27979CFD11F82B" bold="true" box="[726,750,1962,1983]" pageId="33" pageNumber="334">A.</emphasis>
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Head, dorsal view.
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<emphasis id="B944EAF03C12A205FC49979DFC30F854" bold="true" box="[952,975,1963,1984]" pageId="33" pageNumber="334">B.</emphasis>
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Head, lateral view.
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<emphasis id="B944EAF03C12A205FB6A979CFB4CF854" bold="true" box="[1179,1203,1962,1984]" pageId="33" pageNumber="334">C.</emphasis>
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Head, frontal view.
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<emphasis id="B944EAF03C12A205FA72979DFA63F854" bold="true" box="[1411,1436,1963,1984]" pageId="33" pageNumber="334">D.</emphasis>
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Thorax, lateral view.
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<emphasis id="B944EAF03C12A205FE9F97FFFE79F84A" bold="true" box="[366,390,1993,2014]" pageId="33" pageNumber="334">E.</emphasis>
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Thorax, dorsal view.
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<emphasis id="B944EAF03C12A205FD9397FFFD8CF84A" bold="true" box="[610,627,1993,2014]" pageId="33" pageNumber="334">F.</emphasis>
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Abdomen, dorsal view. Scale bars: 0.1mm.
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</paragraph>
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</caption>
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<caption id="DF4F666A3C11A206FF6694BFFECAFB29" httpUri="https://zenodo.org/record/262924/files/figure.png" pageId="34" pageNumber="335" targetBox="[163,1422,205,1125]" targetPageId="34">
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||
<paragraph id="8B8F36E23C11A206FF6694BFFECAFB29" blockId="34.[151,1436,1161,1213]" pageId="34" pageNumber="335">
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<emphasis id="B944EAF03C11A206FF6694BFFEE0FB0A" bold="true" box="[151,287,1161,1183]" pageId="34" pageNumber="335">FIGURE 31.</emphasis>
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<emphasis id="B944EAF03C11A206FED494BFFDFCFB0B" box="[293,515,1161,1183]" italics="true" pageId="34" pageNumber="335">
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<taxonomicName id="4C304D613C11A206FED494BFFE59FB0B" box="[293,422,1161,1183]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="genus">Cladochaeta</taxonomicName>
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stigmata
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</emphasis>
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,
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<emphasis id="B944EAF03C11A206FDE094BFFD9DFB0A" bold="true" box="[529,610,1161,1182]" pageId="34" pageNumber="335">nov. sp.</emphasis>
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, holotype, male terminalia.
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<emphasis id="B944EAF03C11A206FC8F94BFFC4AFB0A" bold="true" box="[894,949,1161,1182]" pageId="34" pageNumber="335">A, D.</emphasis>
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Terminal view.
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<emphasis id="B944EAF03C11A206FBA894BFFB71FB0A" bold="true" box="[1113,1166,1161,1182]" pageId="34" pageNumber="335">B, E.</emphasis>
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Terminal oblique view.
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<emphasis id="B944EAF03C11A206FA7294BFFF57FB28" bold="true" pageId="34" pageNumber="335">C, F.</emphasis>
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Lateral view.
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</paragraph>
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</caption>
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<subSubSection id="C32A65693C11A206FF3694D2FB3BF819" pageId="34" pageNumber="335" type="description">
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||
<paragraph id="8B8F36E23C11A206FF3694D2FB3BF819" blockId="34.[151,1437,1252,2006]" pageId="34" pageNumber="335">
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<emphasis id="B944EAF03C11A206FF3694D2FE5EFB69" bold="true" box="[199,417,1252,1277]" pageId="34" pageNumber="335">Description. Head</emphasis>
|
||
(
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<figureCitation id="130B2A673C11A206FE4294D2FDEFFB69" box="[435,528,1252,1277]" captionStart="FIGURE 30" captionStartId="33.[151,250,1962,1984]" captionTargetBox="[157,1423,958,1930]" captionTargetId="figure@33.[150,1437,946,1941]" captionTargetPageId="33" captionText="FIGURE 30. Cladochaeta stigmata, nov. sp., holotype. A. Head, dorsal view. B. Head, lateral view. C. Head, frontal view. D. Thorax, lateral view. E. Thorax, dorsal view. F. Abdomen, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/262923/files/figure.png" pageId="34" pageNumber="335">Figs. 30</figureCitation>
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A–C). Higher than wide. Eyes bare, light red. Pedicel ochre, first flagellomere light ochre; arista curved downwards, with only basal dorsal branch (
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<figureCitation id="130B2A673C11A206FC6C953EFC12FAB5" box="[925,1005,1288,1313]" captionStart="FIGURE 17" captionStartId="20.[151,250,818,840]" captionTargetBox="[180,1375,241,777]" captionTargetId="figure@20.[149,1437,175,815]" captionTargetPageId="20" captionText="FIGURE 17. Arista conditions in Cladochaeta. A. C. chauliodactyla nov. sp., holotype. B. C. stigmata nov. sp., holotype. Scale bars: 0.1 mm. Abbreviations: Ap. f.: apical fork; d- 1: first dorsal branch; d- 2: second dorsal branch; d- 3: third dorsal branch; d- 4: forth dorsal branch; dors. bas. br.: dorsal basal branch; vent. br.: ventral branch." httpUri="https://zenodo.org/record/262910/files/figure.png" pageId="34" pageNumber="335">Fig. 17</figureCitation>
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B). Frons pale yellow, ocellar triangle concolor. Fronto-orbital setae: anterior reclinate short, barely distinguishable from other fronto-orbitals, posterior reclinate slightly smaller than proclinate, closer to inner vertical than to proclinate; anterior reclinate slightly closer to proclinate than to posterior reclinate. Face flat, gena and face light ochre. Proboscis and palpus concolor with gena.
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<emphasis id="B944EAF03C11A206FF2B95AEFECCFA25" bold="true" box="[218,307,1432,1457]" pageId="34" pageNumber="335">Thorax</emphasis>
|
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(
|
||
<figureCitation id="130B2A673C11A206FEB395AEFE61FA25" box="[322,414,1432,1457]" captionStart="FIGURE 30" captionStartId="33.[151,250,1962,1984]" captionTargetBox="[157,1423,958,1930]" captionTargetId="figure@33.[150,1437,946,1941]" captionTargetPageId="33" captionText="FIGURE 30. Cladochaeta stigmata, nov. sp., holotype. A. Head, dorsal view. B. Head, lateral view. C. Head, frontal view. D. Thorax, lateral view. E. Thorax, dorsal view. F. Abdomen, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/262923/files/figure.png" pageId="34" pageNumber="335">Figs. 30</figureCitation>
|
||
D–E). Light yellow. Anterior dorsocentral ca. ½ length of posterior dorsocentral. Posterior dorsocentral closer to scutoscutellar suture than to anterior dorsocentral. Acrostichals in 6 even rows. Basal and apical scutellars convergent. Three postpronotal setae, median longer than other two. Legs uniformily pale yellow.
|
||
<emphasis id="B944EAF03C11A206FF669633FF27F98A" bold="true" box="[151,216,1541,1566]" pageId="34" pageNumber="335">Wing</emphasis>
|
||
(
|
||
<figureCitation id="130B2A673C11A206FF189633FEC3F98A" box="[233,316,1541,1566]" captionStart="FIGURE 36" captionStartId="39.[151,250,867,889]" captionTargetBox="[164,1420,213,833]" captionTargetId="figure@39.[152,1435,193,846]" captionTargetPageId="39" captionText="FIGURE 36. Wings of Cladochaeta species. A. C. periotoi nov. sp. (holoype). B. C. phallotrixa nov. sp. (holotype). C. C. stigmata nov. sp. (holotype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/262929/files/figure.png" pageId="34" pageNumber="335">Fig. 36</figureCitation>
|
||
C). Length:
|
||
<quantity id="4CC89B073C11A206FE229633FDD1F989" box="[467,558,1541,1566]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="34" pageNumber="335" unit="mm" value="1.6">1.6 mm</quantity>
|
||
. Membrane infuscate, darker area extending from r2+3 cell to mid of r4+5 cell, gradually fading; small, rounded and diffuse cloud over r-m and a small darker cloud over dm-cu barely extending to CuA1. Tip of R2+3 curved towards wing margin. R4+5 and M parallel, tips slightly divergent. Crossvein dm-cu slightly sinuous, perpendicular to CuA1. Wing tip slightly acute. Halter whitish.
|
||
<emphasis id="B944EAF03C11A206FBEA9640FB6FF91B" bold="true" box="[1051,1168,1654,1679]" pageId="34" pageNumber="335">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="130B2A673C11A206FB519640FB0FF91B" box="[1184,1264,1654,1679]" captionStart="FIGURE 30" captionStartId="33.[151,250,1962,1984]" captionTargetBox="[157,1423,958,1930]" captionTargetId="figure@33.[150,1437,946,1941]" captionTargetPageId="33" captionText="FIGURE 30. Cladochaeta stigmata, nov. sp., holotype. A. Head, dorsal view. B. Head, lateral view. C. Head, frontal view. D. Thorax, lateral view. E. Thorax, dorsal view. F. Abdomen, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/262923/files/figure.png" pageId="34" pageNumber="335">Fig. 30</figureCitation>
|
||
F). All tergites ochre, posterior margins darker.
|
||
<emphasis id="B944EAF03C11A206FDF396ABFD3DF922" bold="true" box="[514,706,1693,1718]" pageId="34" pageNumber="335">Male terminalia</emphasis>
|
||
(
|
||
<figureCitation id="130B2A673C11A206FD2096ABFCD2F922" box="[721,813,1693,1718]" captionStart="FIGURE 31" captionStartId="34.[151,250,1161,1183]" captionTargetBox="[163,1422,205,1125]" captionTargetId="figure@34.[151,1436,193,1139]" captionTargetPageId="34" captionText="FIGURE 31. Cladochaeta stigmata, nov. sp., holotype, male terminalia. A, D. Terminal view. B, E. Terminal oblique view. C, F. Lateral view." httpUri="https://zenodo.org/record/262924/files/figure.png" pageId="34" pageNumber="335">Figs. 31</figureCitation>
|
||
A–F). Epandrium inverted U-shaped, almost as high as wide, with 7 long setae on ventrolateral halves; ventral tip of ventrolateral halves gradually tapering, curved inwards terminally. Cercus with ventrolateral edges rounded, not extended. Phallus partially sclerotized: membranous portion broadly bulbous, placed between two thin asymmetrical sclerotized arms; membranous area projected anteriorly, reaching the phallapodeme medially; right sclerotized arm long, left shorter, both connected by a thin dorsal bridge. Phallapodeme thin, sinuous, heavily sclerotized. Surstylus simple, clavate; setae present on entire dorsal and dorsolateral surfaces of surstylus. Hypandrium Y-shaped, ventral keel present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A65693C11A206FF3697AFFD65F842" pageId="34" pageNumber="335" type="etymology">
|
||
<paragraph id="8B8F36E23C11A206FF3697AFFD65F842" blockId="34.[151,1437,1252,2006]" pageId="34" pageNumber="335">
|
||
<emphasis id="B944EAF03C11A206FF3697AFFEB9F826" bold="true" box="[199,326,1945,1970]" pageId="34" pageNumber="335">Etymology</emphasis>
|
||
. The specific epithet comes from the Greek root
|
||
<emphasis id="B944EAF03C11A206FC4597ACFC00F825" box="[948,1023,1946,1969]" italics="true" pageId="34" pageNumber="335">stigma</emphasis>
|
||
(mark), refering to the clouds of infuscation typical of the wing of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F666A3C10A207FF669798FC90F877" httpUri="https://zenodo.org/record/262925/files/figure.png" pageId="35" pageNumber="336" targetBox="[216,1373,213,1914]" targetPageId="35">
|
||
<paragraph id="8B8F36E23C10A207FF669798FC90F877" blockId="35.[151,1435,1966,2019]" pageId="35" pageNumber="336">
|
||
<emphasis id="B944EAF03C10A207FF669798FEDEF850" bold="true" box="[151,289,1966,1988]" pageId="35" pageNumber="336">FIGURE 32.</emphasis>
|
||
Lateral habitus of
|
||
<taxonomicName id="4C304D613C10A207FE159798FD9AF850" box="[484,613,1966,1988]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAF03C10A207FE159798FD9AF850" box="[484,613,1966,1988]" italics="true" pageId="35" pageNumber="336">Cladochaeta</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="B944EAF03C10A207FD329798FD23F857" bold="true" box="[707,732,1966,1987]" pageId="35" pageNumber="336">A.</emphasis>
|
||
<taxonomicName id="4C304D613C10A207FD129799FCB4F851" box="[739,843,1967,1989]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="armata">
|
||
<emphasis id="B944EAF03C10A207FD129799FCB4F851" box="[739,843,1967,1989]" italics="true" pageId="35" pageNumber="336">C. armata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B944EAF03C10A207FCA99799FC8FF850" bold="true" box="[856,880,1967,1988]" pageId="35" pageNumber="336">B.</emphasis>
|
||
<taxonomicName id="4C304D613C10A207FC869799FBF9F851" box="[887,1030,1967,1989]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="armatopsis">
|
||
<emphasis id="B944EAF03C10A207FC869799FBF9F851" box="[887,1030,1967,1989]" italics="true" pageId="35" pageNumber="336">C. armatopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C10A207FBFC9799FB9FF850" bold="true" box="[1037,1120,1967,1988]" pageId="35" pageNumber="336">nov. sp.</emphasis>
|
||
(holotype).
|
||
<emphasis id="B944EAF03C10A207FB2F9798FB09F850" bold="true" box="[1246,1270,1966,1988]" pageId="35" pageNumber="336">C.</emphasis>
|
||
<taxonomicName id="4C304D613C10A207FB0F9799FA97F850" box="[1278,1384,1966,1988]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="balbiae">
|
||
<emphasis id="B944EAF03C10A207FB0F9799FA97F850" box="[1278,1384,1966,1988]" italics="true" pageId="35" pageNumber="336">C. balbiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C10A207FA9E9799FF4AF877" bold="true" pageId="35" pageNumber="336">nov. sp.</emphasis>
|
||
(holotype).
|
||
<emphasis id="B944EAF03C10A207FEC097F8FEB5F877" bold="true" box="[305,330,1998,2019]" pageId="35" pageNumber="336">D.</emphasis>
|
||
<taxonomicName id="4C304D613C10A207FEA197F8FE26F877" box="[336,473,1998,2019]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="paraitinga">
|
||
<emphasis id="B944EAF03C10A207FEA197F8FE26F877" box="[336,473,1998,2019]" italics="true" pageId="35" pageNumber="336">C. paraitinga</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C10A207FE2E97F8FDCFF877" bold="true" box="[479,560,1998,2019]" pageId="35" pageNumber="336">nov. sp.</emphasis>
|
||
(holotype). Scale bars: 0.5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F666A3C17A200FF6697B1FA63F828" httpUri="https://zenodo.org/record/262926/files/figure.png" pageId="36" pageNumber="337" targetBox="[165,1416,209,1891]" targetPageId="36">
|
||
<paragraph id="8B8F36E23C17A200FF6697B1FA63F828" blockId="36.[151,1436,1927,1980]" pageId="36" pageNumber="337">
|
||
<emphasis id="B944EAF03C17A200FF6697B1FEDFF808" bold="true" box="[151,288,1927,1949]" pageId="36" pageNumber="337">FIGURE 33.</emphasis>
|
||
Lateral habitus of
|
||
<taxonomicName id="4C304D613C17A200FE1097B1FD9CF809" box="[481,611,1927,1949]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAF03C17A200FE1097B1FD9CF809" box="[481,611,1927,1949]" italics="true" pageId="36" pageNumber="337">Cladochaeta</emphasis>
|
||
</taxonomicName>
|
||
species, holotypes.
|
||
<emphasis id="B944EAF03C17A200FCDF97B1FCB8F808" bold="true" box="[814,839,1927,1948]" pageId="36" pageNumber="337">A.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FCBF97BEFC27F809" box="[846,984,1927,1949]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="bomplandi">
|
||
<emphasis id="B944EAF03C17A200FCBF97BEFC27F809" box="[846,984,1927,1949]" italics="true" pageId="36" pageNumber="337">C. bomplandi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B944EAF03C17A200FC1497BEFC03F809" bold="true" box="[997,1020,1928,1949]" pageId="36" pageNumber="337">B.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FBF297BEFB6FF809" box="[1027,1168,1927,1949]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="arthrostyla">
|
||
<emphasis id="B944EAF03C17A200FBF297BEFB6FF809" box="[1027,1168,1927,1949]" italics="true" pageId="36" pageNumber="337">C. arthrostyla</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B944EAF03C17A200FB6F97B1FB49F809" bold="true" box="[1182,1206,1927,1949]" pageId="36" pageNumber="337">C.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FB4C97BEFADBF809" box="[1213,1316,1927,1949]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="asapha">
|
||
<emphasis id="B944EAF03C17A200FB4C97BEFADBF809" box="[1213,1316,1927,1949]" italics="true" pageId="36" pageNumber="337">C. asapha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C17A200FADA97BEFA63F809" bold="true" box="[1323,1436,1928,1949]" pageId="36" pageNumber="337">nov. sp. D.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FF669791FEB5F828" box="[151,330,1958,1980]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="chauliodactyla">
|
||
<emphasis id="B944EAF03C17A200FF669791FEB5F828" box="[151,330,1958,1980]" italics="true" pageId="36" pageNumber="337">C. chauliodactyla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C17A200FEA19791FE41F828" bold="true" box="[336,446,1959,1980]" pageId="36" pageNumber="337">nov. sp. E.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FE359791FD8FF828" box="[452,624,1958,1980]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="conicophallus">
|
||
<emphasis id="B944EAF03C17A200FE359791FD8FF828" box="[452,624,1958,1980]" italics="true" pageId="36" pageNumber="337">C. conicophallus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C17A200FD879791FD21F828" bold="true" box="[630,734,1959,1980]" pageId="36" pageNumber="337">nov. sp. F.</emphasis>
|
||
<emphasis id="B944EAF03C17A200FD179791FC75F828" box="[742,906,1958,1980]" italics="true" pageId="36" pageNumber="337">
|
||
<taxonomicName id="4C304D613C17A200FD179791FC7AF828" box="[742,901,1958,1980]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="dicrophallus">C. dicrophallus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B944EAF03C17A200FC7F9791FC06F828" bold="true" box="[910,1017,1958,1980]" pageId="36" pageNumber="337">nov. sp. G.</emphasis>
|
||
<taxonomicName id="4C304D613C17A200FBF39791FB82F828" box="[1026,1149,1958,1980]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B944EAF03C17A200FBF39791FB82F828" box="[1026,1149,1958,1980]" italics="true" pageId="36" pageNumber="337">C. grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C17A200FB759791FB2AF828" bold="true" box="[1156,1237,1959,1980]" pageId="36" pageNumber="337">nov. sp.</emphasis>
|
||
Scale bars: 0.5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F666A3C16A201FF669655FCE6F90C" httpUri="https://zenodo.org/record/262927/files/figure.png" pageId="37" pageNumber="338" targetBox="[163,1418,209,1600]" targetPageId="37">
|
||
<paragraph id="8B8F36E23C16A201FF669655FCE6F90C" blockId="37.[151,1436,1635,1688]" pageId="37" pageNumber="338">
|
||
<emphasis id="B944EAF03C16A201FF669655FEDBF9EC" bold="true" box="[151,292,1635,1657]" pageId="37" pageNumber="338">FIGURE 34.</emphasis>
|
||
Lateral habitus of
|
||
<taxonomicName id="4C304D613C16A201FE069655FD86F9ED" box="[503,633,1635,1657]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAF03C16A201FE069655FD86F9ED" box="[503,633,1635,1657]" italics="true" pageId="37" pageNumber="338">Cladochaeta</emphasis>
|
||
</taxonomicName>
|
||
species, holotypes.
|
||
<emphasis id="B944EAF03C16A201FCA29655FC94F9EC" bold="true" box="[851,875,1635,1656]" pageId="37" pageNumber="338">A.</emphasis>
|
||
<taxonomicName id="4C304D613C16A201FC879652FC0CF9ED" box="[886,1011,1635,1657]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="atlantica">
|
||
<emphasis id="B944EAF03C16A201FC879652FC0CF9ED" box="[886,1011,1635,1657]" italics="true" pageId="37" pageNumber="338">C. atlantica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C16A201FC0F9652FB89F9ED" bold="true" box="[1022,1142,1636,1657]" pageId="37" pageNumber="338">nov. sp. B.</emphasis>
|
||
<taxonomicName id="4C304D613C16A201FB739652FB0BF9ED" box="[1154,1268,1636,1657]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="periotoi">
|
||
<emphasis id="B944EAF03C16A201FB739652FB0BF9ED" box="[1154,1268,1636,1657]" italics="true" pageId="37" pageNumber="338">C. periotoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C16A201FAF19652FA86F9ED" bold="true" box="[1280,1401,1635,1657]" pageId="37" pageNumber="338">nov. sp. C.</emphasis>
|
||
<taxonomicName id="4C304D613C16A201FA759652FEF7F90C" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="phallotrixa">
|
||
<emphasis id="B944EAF03C16A201FA759652FEF7F90C" italics="true" pageId="37" pageNumber="338">C. phallotrixa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAF03C16A201FEFC96B4FE81F903" bold="true" box="[269,382,1666,1687]" pageId="37" pageNumber="338">nov. sp. D.</emphasis>
|
||
<emphasis id="B944EAF03C16A201FE7596B5FE07F90C" box="[388,504,1667,1688]" italics="true" pageId="37" pageNumber="338">C. stigmata</emphasis>
|
||
<emphasis id="B944EAF03C16A201FE0E96B4FDAFF903" bold="true" box="[511,592,1666,1687]" pageId="37" pageNumber="338">nov. sp.</emphasis>
|
||
Scale bars: 0.5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A65693C16A201FF3696F2FC92F805" pageId="37" pageNumber="338" type="discussion">
|
||
<paragraph id="8B8F36E23C16A201FF3696F2FC92F805" blockId="37.[151,1436,1732,1937]" pageId="37" pageNumber="338">
|
||
<emphasis id="B944EAF03C16A201FF3696F2FEBBF949" bold="true" box="[199,324,1732,1757]" pageId="37" pageNumber="338">Comment.</emphasis>
|
||
This species was placed in the
|
||
<taxonomicName id="4C304D613C16A201FD5C96F3FCEDF949" box="[685,786,1733,1757]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="nebulosa">
|
||
<emphasis id="B944EAF03C16A201FD5C96F3FCEDF949" box="[685,786,1733,1757]" italics="true" pageId="37" pageNumber="338">nebulosa</emphasis>
|
||
</taxonomicName>
|
||
species group due to the arista with a single dorsal basal branch, the clouded wing membrane, and the membranous, laterally flattened, median portion of the phallus, with a pair of asymmetrical, long and thin sclerotized arms. It closely resembles
|
||
<taxonomicName id="4C304D613C16A201FC10973BFA63F8B1" authority="Grimaldi & Nguyen, 1999" authorityName="Grimaldi & Nguyen" authorityYear="1999" box="[993,1436,1805,1830]" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="genuinus">
|
||
<emphasis id="B944EAF03C16A201FC10973BFB97F8B1" box="[993,1128,1805,1829]" italics="true" pageId="37" pageNumber="338">C. genuinus</emphasis>
|
||
<bibRefCitation id="EFA14B133C16A201FB81973BFA63F8B1" author="Grimaldi" box="[1136,1436,1805,1830]" pageId="37" pageNumber="338" refString="Grimaldi, D. A. & Nguyen, T. (1999) Monograph on the spittlebug flies, genus Cladochaeta (Diptera: Drosophilidae: Cladochaetini). Bulletin of the American Museum of Natural History, 241, 1 - 326." type="journal article" year="1999">Grimaldi & Nguyen, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
on the morphology of phallus, clouding pattern of the wing and the sinuous dm-cu crossvein. However,
|
||
<taxonomicName id="4C304D613C16A201FA739707FF03F8F9" class="Insecta" family="Drosophilidae" genus="Cladochaeta" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="genuinus">
|
||
<emphasis id="B944EAF03C16A201FA739707FF03F8F9" italics="true" pageId="37" pageNumber="338">C. genuinus</emphasis>
|
||
</taxonomicName>
|
||
has a long surstylus ventral lobe, absent in
|
||
<emphasis id="B944EAF03C16A201FD049763FCF7F8F8" box="[757,776,1877,1900]" italics="true" pageId="37" pageNumber="338">C</emphasis>
|
||
.
|
||
<emphasis id="B944EAF03C16A201FCE99760FC87F8F9" box="[792,888,1878,1901]" italics="true" pageId="37" pageNumber="338">stigmata</emphasis>
|
||
,
|
||
<emphasis id="B944EAF03C16A201FC799763FC1BF8FA" bold="true" box="[904,996,1877,1902]" pageId="37" pageNumber="338">nov. sp.</emphasis>
|
||
and the proportional length of the left sclerotized arm of the phallus is different in both species as well.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |