444 lines
75 KiB
XML
444 lines
75 KiB
XML
<document id="B447B1CFA18BF21B812CF5AD85A0EA88" ID-CLB-Dataset="54025" ID-DOI="10.11646/zootaxa.5231.1.3" ID-GBIF-Dataset="e44954f0-eb71-4c39-95ad-c71a4214eb27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7574093" ID-ZooBank="9C938B40-376D-4AB0-A5E8-B267DD2D5725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674712506927" checkinUser="plazi" docAuthor="Soares, Matheus M. M., Capellari, Renato S. & Ale-Rocha, Rosaly" docDate="2023" docId="039987AEFF80FFEDE4D9FBFAFA25FCBD" docLanguage="en" docName="zootaxa.5231.1.3.pdf" docOrigin="Zootaxa 5231 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraclius vulcanoae Soares & Capellari & Ale-Rocha 2023, sp. nov." docType="treatment" docUuid="094E4DF3-76FA-49E2-9FC3-FC43334A14A5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="49" masterDocId="FFA0FFD6FF87FFE1E44EFFC3FFB9FF84" masterDocTitle="Species of Paraclius Loew (Diptera: Dolichopodidae) with bi-lamellate antennal stylus: new combination, new synonym, and two new species" masterLastPageNumber="51" masterPageNumber="37" pageNumber="44" updateTime="1699474335796" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="90F65DE8BAAA9DC38AEC0D60AE095CB5">Species of Paraclius Loew (Diptera: Dolichopodidae) with bi-lamellate antennal stylus: new combination, new synonym, and two new species</mods:title>
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<mods:name id="8F333540D6ACBE2A6B1218F0221FCA65" type="personal">
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<mods:roleTerm id="58EC86A04F9382E81C70BC6527180981">Author</mods:roleTerm>
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<mods:namePart id="C2A7B15D94B72C44692C6084D096FEC1">Soares, Matheus M. M.</mods:namePart>
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<mods:nameIdentifier id="FA85BDF3A22750665E42A7750899FB2A" type="ORCID">0000-0002-2355-1441</mods:nameIdentifier>
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<mods:affiliation id="D358D89C1406F4B0B4B37F230F7DFC94">Graduate Program in Entomology (PPG-Ent), Instituto Nacional de Pesquisas da Amazônia (INPA). & matheusmmsoares @ gmail. com; https: // orcid. org / 0000 - 0002 - 2355 - 1441</mods:affiliation>
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<mods:nameIdentifier id="63CCAAE7EBEB2563C12C4AC5D2A671FC" type="email">matheusmmsoares@gmail.com</mods:nameIdentifier>
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<mods:name id="D52655E70C10FA0716269D7967D11528" type="personal">
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<mods:roleTerm id="709140513745216B2C86FE5172B96CFD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1302E14A876949E923A6F135E24BAEDE">Capellari, Renato S.</mods:namePart>
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<mods:nameIdentifier id="A54DC8B47169ED96A4E5B8B70ECBC812" type="ORCID">0000-0002-8410-9235</mods:nameIdentifier>
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<mods:affiliation id="6BED3463D0EBCA8CD90F135726AF6B39">Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064 - 790, Brazil. & rscapellari @ gmail. com; https: // orcid. org / 000 - 0002 - 8410 - 9235</mods:affiliation>
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<mods:nameIdentifier id="939F9F63D2266FC8C550CE6F89175A61" type="email">rscapellari@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="E59DF99A48BC87F0360493C45097BBBD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="746754742EC108190C2C1BCDEAD28469">Ale-Rocha, Rosaly</mods:namePart>
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<mods:nameIdentifier id="1DA7DBD725666157C65EF454BBFCEDE8" type="ORCID">0000-0001-9874-9770</mods:nameIdentifier>
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<mods:affiliation id="594B4FFA1436D308506083D1E94F16BA">Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067 - 375, Manaus, Amazonas, Brazil. & alerocha @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 9874 - 9770</mods:affiliation>
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<mods:nameIdentifier id="1AEE25DBE265E029EE05069F99F51A2E" type="email">alerocha@inpa.gov.br</mods:nameIdentifier>
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<mods:title id="312D2363FCD1CDAAED41105FE7802C84">Zootaxa</mods:title>
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<mods:part id="49B198C88999117A34B0781EB098372B">
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<mods:date id="B6488F4E33781425B6967A20B7BCAB2B">2023</mods:date>
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<mods:number id="A6D1261BAC025015C749C3772F6B0616">2023-01-26</mods:number>
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<mods:url id="AE6CE4E2A659ED9A21F5E2572BD785A9">http://dx.doi.org/10.11646/zootaxa.5231.1.3</mods:url>
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<mods:classification id="9991DDBAD720B512FE353C3667B3F2BA">journal article</mods:classification>
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<mods:identifier id="431B09521294503F02393AC747377EA2" type="DOI">10.11646/zootaxa.5231.1.3</mods:identifier>
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<treatment id="039987AEFF80FFEDE4D9FBFAFA25FCBD" ID-DOI="http://doi.org/10.5281/zenodo.7574117" ID-GBIF-Taxon="209104899" ID-Zenodo-Dep="7574117" LSID="urn:lsid:zoobank.org:act:094E4DF3-76FA-49E2-9FC3-FC43334A14A5" httpUri="http://treatment.plazi.org/id/039987AEFF80FFEDE4D9FBFAFA25FCBD" lastPageId="12" lastPageNumber="49" pageId="7" pageNumber="44">
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<subSubSection id="C32A6533FF80FFE6E4D9FBFAFE4AFBD7" box="[151,499,1081,1108]" pageId="7" pageNumber="44" type="nomenclature">
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<paragraph id="8B8F36B8FF80FFE6E4D9FBFAFE4AFBD7" blockId="7.[151,499,1081,1144]" box="[151,499,1081,1108]" pageId="7" pageNumber="44">
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<heading id="D0C781D4FF80FFE6E4D9FBFAFE4AFBD7" bold="true" box="[151,499,1081,1108]" fontSize="11" level="1" pageId="7" pageNumber="44" reason="1">
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<emphasis id="B944EAAAFF80FFE6E4D9FBFAFE4AFBD7" bold="true" box="[151,499,1081,1108]" pageId="7" pageNumber="44">
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<taxonomicName id="4C304D3BFF80FFE6E4D9FBFAFE37FBD0" ID-CoL="BDKG5" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[151,398,1081,1108]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
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<emphasis id="B944EAAAFF80FFE6E4D9FBFAFE37FBD0" bold="true" box="[151,398,1081,1108]" italics="true" pageId="7" pageNumber="44">Paraclius vulcanoae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27757D1FF80FFE6E5DBFBFAFE4AFBD7" box="[405,499,1081,1107]" pageId="7" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32A6533FF80FFE6E4D9FB9EFEFAFBF3" box="[151,323,1117,1144]" pageId="7" pageNumber="44" type="description">
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<paragraph id="8B8F36B8FF80FFE6E4D9FB9EFEFAFBF3" blockId="7.[151,499,1081,1144]" box="[151,323,1117,1144]" pageId="7" pageNumber="44">
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(
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<figureCitation id="130B2A3DFF80FFE6E4D1FB9EFF51FBF3" box="[159,232,1117,1143]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Figs 4</figureCitation>
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,
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<figureCitation id="130B2A3DFF80FFE6E4BAFB9EFEBDFBFC" box="[244,260,1117,1144]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="7" pageNumber="44">5</figureCitation>
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,
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<figureCitation id="130B2A3DFF80FFE6E55EFB9EFE99FBF3" box="[272,288,1117,1143]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="7" pageNumber="44">6</figureCitation>
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,
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<figureCitation id="130B2A3DFF80FFE6E562FB9EFE82FBF3" box="[300,315,1117,1143]" captionStart="FIGURE 7" captionStartId="12.[151,250,1643,1667]" captionTargetBox="[151,1435,849,1620]" captionTargetId="figure-338@12.[151,1435,849,1620]" captionTargetPageId="12" captionText="FIGURE 7. Geographic distribution of Paraclius Loew with bi-lamellate arista-like stylus." figureDoi="http://doi.org/10.5281/zenodo.7574115" httpUri="https://zenodo.org/record/7574115/files/figure.png" pageId="7" pageNumber="44">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A6533FF80FFE6E4D9FB63FD5DFAA2" pageId="7" pageNumber="44" type="diagnosis">
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<paragraph id="8B8F36B8FF80FFE6E4D9FB63FD5DFAA2" blockId="7.[151,1437,1184,2039]" pageId="7" pageNumber="44">
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<emphasis id="B944EAAAFF80FFE6E4D9FB63FED6FB3E" bold="true" box="[151,367,1184,1210]" pageId="7" pageNumber="44">Diagnosis (males).</emphasis>
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Clypeus protruding, longer than face and pointed below, ending close to lower eye margin (
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<figureCitation id="130B2A3DFF80FFE6E125FB63FF05FB5A" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Figs 4A</figureCitation>
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,
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<figureCitation id="130B2A3DFF80FFE6E489FB07FEB2FB5A" box="[199,267,1220,1247]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="7" pageNumber="44">5A, B</figureCitation>
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). Postpedicel wider than long, shorter than scape and pedicel combined; arista-like stylus one segmented, 2X longer than eye height, with lamella at middle and apex, bare, except for dorsal pubescence on lamellae, apical lamella wider than long, truncated at apex (
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<figureCitation id="130B2A3DFF80FFE6E633FACFFD61FAA2" box="[637,728,1292,1318]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Fig. 4A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A6533FF80FFE9E489FAF3FECBF865" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="description">
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<paragraph id="8B8F36B8FF80FFE9E489FAF3FECBF865" blockId="7.[151,1437,1184,2039]" lastBlockId="8.[151,1437,151,2017]" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
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<emphasis id="B944EAAAFF80FFE6E489FAF3FE25FACE" bold="true" box="[199,412,1328,1354]" pageId="7" pageNumber="44">Description. Male</emphasis>
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(
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<figureCitation id="130B2A3DFF80FFE6E5E4FAF3FDB9FACE" box="[426,512,1328,1354]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Fig. 4B</figureCitation>
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,
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<typeStatus id="548B881AFF80FFE6E645FAF3FDCBFACE" box="[523,626,1328,1354]" pageId="7" pageNumber="44" type="holotype">holotype</typeStatus>
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). Body length:
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<quantity id="4CC89B5DFF80FFE6E754FAF3FCCBFACF" box="[794,882,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="44" unit="mm" value="5.2">5.2 mm</quantity>
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; wing:
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<quantity id="4CC89B5DFF80FFE6E78DFAF3FBA1FACF" box="[963,1048,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="44" unit="mm" value="5.6">5.6 mm</quantity>
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long, wide:
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<quantity id="4CC89B5DFF80FFE6E0EFFAF3FB5DFACF" box="[1185,1252,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="44" unit="mm" value="2.0">2 mm</quantity>
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.
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||
<emphasis id="B944EAAAFF80FFE6E0A3FAF3FA94FACE" bold="true" box="[1261,1325,1328,1354]" pageId="7" pageNumber="44">Head</emphasis>
|
||
(
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<figureCitation id="130B2A3DFF80FFE6E175FAF3FA21FACE" box="[1339,1432,1328,1354]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Figs 4A</figureCitation>
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,
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<figureCitation id="130B2A3DFF80FFE6E4D9FA97FF66FAEA" box="[151,223,1364,1391]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="7" pageNumber="44">5A, B</figureCitation>
|
||
). Distinctly wider than high. Postocular setae white, ending in 1 stronger seta below, 6 upper-most black. Frons rectangular, about 4X wider than long, dark metallic green, obscured by dense silvery pruinosity. Face about 1.25X higher than wide, wider than ocellar tubercle; yellowish gray, covered with silvery pruinosity, mostly evident on sides of face, and yellowish pruinosity at middle. Clypeus protruding, 2X longer than wide and 2X longer than face, lower margin pointed, ending close to lower eye margin; clypeus yellow, covered with yellow pruinosity, clypeal suture inconspicuous. Palpus yellow, subtriangular, with central region bare, shiny, and ventral edge covered with a few yellow setae, lacking apical stronger seta. Proboscis yellow, labellum with a few short pale setae, and 1 stronger seta at apex. One pair of divergent strong ocellar setae and 1 pair of tiny postocellar setae; 1 pair of strong vertical setae; 1 pair of short paravertical setae, slightly longer than upper-most postocular seta. Postcranium dark green, covered with silvery pruinosity, lower margin of postcranium with 6 strong white setae, 1 strong white seta and a few slender white hairs below occiput. Antenna inserted on top of head, orangish yellow, except arista brown; scape 2X longer than pedicel, with apical projection curved apically, dorsal surface covered with short black setae; pedicel shorter than postpedicel, arising dorsally from scape, with crown of setae; postpedicel about 2X wider than long, with rounded apex, covered with short whitish pubescence; arista-like stylus dorsal, arising from upper-edge of postpedicel at middle, one segmented, 2X longer than eye height, with lamella at middle and apex, bare, except for dorsal pubescence on lamellae, apical lamella wider than long, truncated, with whitish pubescence at apex.
|
||
<emphasis id="B944EAAAFF80FFE6E12DF8B3FF06F82A" bold="true" pageId="7" pageNumber="44">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF80FFE6E481F857FE92F82A" box="[207,299,1940,1966]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="7" pageNumber="44">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="130B2A3DFF80FFE6E576F857FEE5F82B" box="[312,348,1940,1967]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="7" pageNumber="44">5C</figureCitation>
|
||
). Metallic dark green, with weak coppery reflections, except for postpronotal lobe, region above notopleuron and extending to postalar callus brown, mesonotum covered with weak silvery pruinosity, dense on anterior 0.5/6 of mesonotum, lateral margins of postpronotal lobe and entire notopleuron. Scutellum concolorous with mesonotum. Pleura mostly brownish gray, with greenish reflections and covered by silvery pruinosity, except metepisternum (around posterior spiracle yellow); metepimeron gray.
|
||
<emphasis id="B944EAAAFF8FFFE9E7F2FF7AFB85FF57" box="[956,1084,185,211]" italics="true" pageId="8" pageNumber="45">Chaetotaxy</emphasis>
|
||
: pronotum with row of strong black setae; anterior 0.5/6 of mesonotum covered with short black setae; acrostichals biseriate, ending at fourth dorsocentral setae; 6 pairs of dorsocentral setae increasing in length posteriorly; 1 pre-, 1 sutural and 1 postsutural intra-alars; 2 strong supra-alar setae, 1 positioned between postpronotal lobe and notopleuron and 1 near postalar callus; 1 strong and 2 short postpronotals; 2 strong notopleurals, 1 at middle of lower edge and 1 at posterior margin of notopleuron; 1 strong postalar; scutellum with 1 pair of strong medial setae, 1 pair of smaller setae laterad about 1/4 as long as medial scutellars; upper and lower surface of proepisternum with a few short, slender pale setae and lower surface with 1 stronger black seta directed anteroventrally; metepisternum with row of a few slender pale setae.
|
||
<emphasis id="B944EAAAFF8FFFE9E491FE0DFE99FE6C" bold="true" box="[223,288,462,488]" pageId="8" pageNumber="45">Wing</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8FFFE9E57FFE0CFE34FE6D" box="[305,397,462,489]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="8" pageNumber="45">Fig. 5D</figureCitation>
|
||
). Membrane light brown, mostly dark on anterior half of wing, veins brown. Costa ending close wing apex, at vein M
|
||
<subScript id="17B434FDFF8FFFE9E589FDC7FE76FD96" attach="left" box="[455,463,516,530]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
; R
|
||
<subScript id="17B434FDFF8FFFE9E5BFFDC7FE40FD96" attach="left" box="[497,505,516,530]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
ending at basal 3/8 of wing; R
|
||
<subScript id="17B434FDFF8FFFE9E71AFDC7FCD4FD96" attach="left" box="[852,877,516,530]" fontSize="6" pageId="8" pageNumber="45">2+3</subScript>
|
||
nearly straight, R
|
||
<subScript id="17B434FDFF8FFFE9E079FDC7FBE8FD96" attach="left" box="[1079,1105,516,530]" fontSize="6" pageId="8" pageNumber="45">4+5</subScript>
|
||
slightly curved posteriorly at apex; last section of vein M
|
||
<subScript id="17B434FDFF8FFFE9E59DFDE5FE62FDB0" attach="left" box="[467,475,550,564]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
straight at base, gradually curved towards R
|
||
<subScript id="17B434FDFF8FFFE9E798FDE5FC49FDB0" attach="left" box="[982,1008,550,564]" fontSize="6" pageId="8" pageNumber="45">4+5</subScript>
|
||
at apical 6/8; maximum width of cell r
|
||
<subScript id="17B434FDFF8FFFE9E4EFFD8AFF03FDD3" attach="right" box="[161,186,585,599]" fontSize="6" pageId="8" pageNumber="45">4+5</subScript>
|
||
/length of dm-m: 0.9; bm+dm ending at apical 5/8 of wing, dm-m straight; M
|
||
<subScript id="17B434FDFF8FFFE9E057FD8AFB98FDD3" attach="left" box="[1049,1057,585,599]" fontSize="6" pageId="8" pageNumber="45">4</subScript>
|
||
and CuA+CuP not reaching wing margin; length of crossvein dm-m/last part of M
|
||
<subScript id="17B434FDFF8FFFE9E6F9FDAFFD06FDFE" attach="left" box="[695,703,620,634]" fontSize="6" pageId="8" pageNumber="45">4</subScript>
|
||
(“CuAx ratio”): 1.75. Lower calypter yellow with blackish cilia; halter yellow.
|
||
<emphasis id="B944EAAAFF8FFFE9E578FDBFFED7FD12" bold="true" box="[310,366,636,662]" pageId="8" pageNumber="45">Legs</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8FFFE9E533FDBFFE6CFD12" box="[381,469,636,662]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="8" pageNumber="45">Fig. 4B</figureCitation>
|
||
). Yellow, except lateral surface of coxa II, apex of femur III with narrow spot dorsally, tarsus I from apex of It
|
||
<subScript id="17B434FDFF8FFFE9E5DBFD72FE24FD3B" attach="left" box="[405,413,689,703]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
, tarsus II from apical 1/3 of IIt
|
||
<subScript id="17B434FDFF8FFFE9E6BCFD72FD43FD3B" attach="left" box="[754,762,689,703]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
and entirely tarsus III brownish.
|
||
<emphasis id="B944EAAAFF8FFFE9E024FD5CFB16FD3D" bold="true" box="[1130,1199,671,697]" pageId="8" pageNumber="45">Leg I.</emphasis>
|
||
Podomere ratios: 60, 55, 31/12/10/8/7. Anterior surface of coxa I covered by silvery pruinosity, with short pale setae, outer edge with 2 strong setae near apex, apical edge with 1 strong and 3–4 less conspicuous setae. Femur I covered by vestiture of short black setae, except ventral surface mostly bare, one anteroventral row of subequally long setae at apical 1/2, 3 conspicuous posterodorsal preapical setae, apicalmost longer than preceding setae. Tibia I with 1 pair of antero- and posterodorsal setae at 2/6, 1 anterodorsal seta at 3/6, 1 dorsal at 3.5/6 and 1 dorsal and 1 posterodorsal preapical. It
|
||
<subScript id="17B434FDFF8FFFE9E1DAFC9DFA25FCE8" attach="left" box="[1428,1436,862,876]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
with 1 short basiventral seta, ventral surface from apex of It
|
||
<subScript id="17B434FDFF8FFFE9E761FC42FC8EFC0B" attach="left" box="[815,823,897,911]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
to It
|
||
<subScript id="17B434FDFF8FFFE9E725FC42FCCAFC0B" attach="left" box="[875,883,897,911]" fontSize="6" pageId="8" pageNumber="45">4</subScript>
|
||
with pile of whitish setae (MSSC).
|
||
<emphasis id="B944EAAAFF8FFFE9E14CFCACFAEAFC0D" bold="true" box="[1282,1363,879,905]" pageId="8" pageNumber="45">Leg II.</emphasis>
|
||
Podomere ratios: 80, 80, 32/20/15/10/8. Anterior surface of coxa II covered by silvery pruinosity and a few short pale setae, outer edge with one row of setae increasing in length from basal 1/3 ending in 2 strong setae, 1 at middle and 1 near apex, apical edge with 3 conspicuous setae. Femur II covered by short vestiture of black setae, except ventral surface mostly bare, with anteroventral row of short setae decreasing in length from base to 3/6, 1 short anteroventral preapical seta, and 1 stronger anterior seta at 5/6, 2 short posteroventral preapical setae. Tibia II with 3 pairs of antero- and posterodorsal setae at 1/6 (about half length of second pair), 1.5/6, and 3.5/6, 1 anterodorsal seta at 2/6, 2 anteroventral setae at 2.5/6 and 4/6, 1 posteroventral seta at 4.5/6, one crown of setae at apex: 1 antero- and 1 posterodorsal, 1 dorsal, 1 antero- and 1 posteroventral. IIt
|
||
<subScript id="17B434FDFF8FFFE9E751FB55FC9EFB20" attach="left" box="[799,807,1174,1188]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
with 1 short basiventral seta.
|
||
<emphasis id="B944EAAAFF8FFFE9E034FB47FB6FFB1A" bold="true" box="[1146,1238,1156,1182]" pageId="8" pageNumber="45">Leg III.</emphasis>
|
||
Podomere ratios: 80, 98, 33, 35, 22, 14, 10. Apical edge of coxa III with 3–4 longer pale setae, lateral surface with 1 long seta at middle and 1 short at apex. Femur III 1.3X wider than femur II at broadest point, femur III covered by short vestiture of black setae, except ventral surface mostly bare, 1 stronger anterodorsal seta at 4.5/6, and 1 short anteroventral preapical seta, one short ventral to anteroventral row of pale short setae from basal 1/6 to apical 4/6, 2 short posteroventral preapical setae. Tibia III with 3 pairs of antero- and posterodorsal setae, 1 short, about 1/3 as long as second pair at 1/6, 1 at 1.5/6 and 1 at 2.5/6, 1 dorsal seta at 3/6, 1 anterodorsal at 3.5/6, 1 dorsal, 1 ventral and 1 anterodorsal preapical seta, 1 ventral seta at 4.5/6, one row of short, sparse and erected setae from basal 1.5/6 to apical 4.5/6. IIIt
|
||
<subScript id="17B434FDFF8FFFE9E4F5FA6FFF7AFA3E" attach="left" box="[187,195,1452,1466]" fontSize="6" pageId="8" pageNumber="45">1</subScript>
|
||
with 1 short basiventral seta.
|
||
<emphasis id="B944EAAAFF8FFFE9E65BFA5AFD33FA37" bold="true" box="[533,650,1433,1459]" pageId="8" pageNumber="45">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8FFFE9E6D7FA5AFD4BFA37" box="[665,754,1433,1459]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="8" pageNumber="45">Fig. 4B</figureCitation>
|
||
). About 1.5X longer than thorax. Dark brown, with greenish reflections. Tergites 1–5 setose, posterior margin with long and strong setae, with wide lateral patches of silvery pruinosity. Tergite 6 bare. Segment 7 brown, bare, short, well sclerotized. Sternite 8 brown, homogeneously covered with long black setae.
|
||
<emphasis id="B944EAAAFF8FFFE9E5D7F9C2FD8CF99F" bold="true" box="[409,565,1537,1563]" pageId="8" pageNumber="45">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8FFFE9E60BF9C2FD32F99F" box="[581,651,1537,1563]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6</figureCitation>
|
||
). Epandrium dark brown, subrectangular, about 2X longer than wide (
|
||
<figureCitation id="130B2A3DFF8FFFE9E4D1F9E7FF43F9BA" box="[159,250,1572,1598]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6A</figureCitation>
|
||
). Basiventral epandrial lobe long, with rounded apex, and 1 short medioventral seta (
|
||
<figureCitation id="130B2A3DFF8FFFE9E0FDF9E7FA94F9BA" box="[1203,1325,1572,1598]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6A, B</figureCitation>
|
||
). Apicoventral epandrial lobe wider than long, somewhat fan-shaped, slightly shorter than cercus, outer edge with 2 short setae (
|
||
<figureCitation id="130B2A3DFF8FFFE9E4AFF9AAFED9F907" box="[225,352,1641,1667]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6B, C</figureCitation>
|
||
). Hypandrium divided in two asymmetrical short arms, left arm curved, with well sclerotized hook-shaped apex, right arm sinuous, slightly narrowing at apex, both shorter than phallus (
|
||
<figureCitation id="130B2A3DFF8FFFE9E0EDF94FFAA7F922" box="[1187,1310,1676,1702]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6B, C</figureCitation>
|
||
). Basal 2/3 of phallus tubular, widening apically, with 1 long right lateral process, branched at apex (
|
||
<figureCitation id="130B2A3DFF8FFFE9E025F96CFABAF94D" box="[1131,1283,1711,1737]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6B, C, G</figureCitation>
|
||
). Ventral lobe of surstylus longer than dorsal lobe, somewhat digitiform, narrowing at apex, with 1 short dorsal seta at basal 1/3 and 1 long apical seta (
|
||
<figureCitation id="130B2A3DFF8FFFE9E5D8F937FDB3F88A" box="[406,522,1780,1806]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6C, F</figureCitation>
|
||
). Dorsal lobe of surstylus with 1 short dorsal digitiform projection near base, apex bifurcated, ventral projection short and rounded, dorsal projection long and mostly membranous (
|
||
<figureCitation id="130B2A3DFF8FFFE9E096F8D4FAE9F8B5" box="[1240,1360,1815,1841]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6C, E</figureCitation>
|
||
). Postgonite plain, shorter than ventral lobe of surstylus, slightly curved ventrally and weakly sclerotized at apex (
|
||
<figureCitation id="130B2A3DFF8FFFE9E10FF8FAFF13F8F2" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6C, F</figureCitation>
|
||
). Sperm pump shorter than epandrium, but as long as ejaculatory apodeme, almost tubular, folded back on itself (
|
||
<figureCitation id="130B2A3DFF8FFFE9E4D1F8BCFEAEF81D" box="[159,279,1919,1945]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6B, C</figureCitation>
|
||
). Ejaculatory apodeme sinuous, well sclerotized and laterally flattened (
|
||
<figureCitation id="130B2A3DFF8FFFE9E076F8BCFB09F81D" box="[1080,1200,1919,1945]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6B, C</figureCitation>
|
||
). Cercus yellow with apical margin dark brown, subrectangular, with short apical projection, covered with yellowish setae (
|
||
<figureCitation id="130B2A3DFF8FFFE9E15CF860FA29F839" box="[1298,1424,1955,1981]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="8" pageNumber="45">Fig. 6A, D</figureCitation>
|
||
).
|
||
<emphasis id="B944EAAAFF8FFFE9E4D9F804FF4CF865" bold="true" box="[151,245,1991,2017]" pageId="8" pageNumber="45">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F6630FF8EFFE8E4D9F841FA2EF81E" ID-DOI="http://doi.org/10.5281/zenodo.7574105" ID-Zenodo-Dep="7574105" box="[151,1431,1922,1946]" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="9" pageNumber="46" startId="9.[151,250,1922,1946]" targetBox="[151,1435,181,1898]" targetPageId="9">
|
||
<paragraph id="8B8F36B8FF8EFFE8E4D9F841FA2EF81E" blockId="9.[151,1431,1922,1946]" box="[151,1431,1922,1946]" pageId="9" pageNumber="46">
|
||
<emphasis id="B944EAAAFF8EFFE8E4D9F841FEAAF81E" bold="true" box="[151,275,1922,1946]" pageId="9" pageNumber="46">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C304D3BFF8EFFE8E557F841FE53F81E" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[281,490,1922,1946]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
|
||
<emphasis id="B944EAAAFF8EFFE8E557F841FE53F81E" box="[281,490,1922,1946]" italics="true" pageId="9" pageNumber="46">Paraclius vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8EFFE8E5BFF841FDF8F81E" bold="true" box="[497,577,1922,1946]" pageId="9" pageNumber="46">
|
||
<taxonomicNameLabel id="A27757D1FF8EFFE8E5BFF841FDF8F81E" box="[497,577,1922,1946]" pageId="9" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male holotype.
|
||
<emphasis id="B944EAAAFF8EFFE8E6A6F841FCB9F81E" bold="true" box="[744,768,1922,1946]" pageId="9" pageNumber="46">A.</emphasis>
|
||
Head, anterior view;
|
||
<emphasis id="B944EAAAFF8EFFE8E792F841FC4DF81E" bold="true" box="[988,1012,1922,1946]" pageId="9" pageNumber="46">B.</emphasis>
|
||
Habitus, lateral view (before dissection).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6630FF8DFFEBE4D9F854FC09F87C" ID-DOI="http://doi.org/10.5281/zenodo.7574109" ID-Zenodo-Dep="7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="10" pageNumber="47" startId="10.[151,250,1943,1967]" targetBox="[156,1431,181,1920]" targetPageId="10">
|
||
<paragraph id="8B8F36B8FF8DFFEBE4D9F854FA25F834" blockId="10.[151,1436,1943,2046]" box="[151,1436,1943,1968]" pageId="10" pageNumber="47">
|
||
<emphasis id="B944EAAAFF8DFFEBE4D9F854FEADF834" bold="true" box="[151,276,1943,1968]" pageId="10" pageNumber="47">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C304D3BFF8DFFEBE552F85BFE57F834" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[284,494,1944,1968]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
|
||
<emphasis id="B944EAAAFF8DFFEBE552F85BFE57F834" box="[284,494,1944,1968]" italics="true" pageId="10" pageNumber="47">Paraclius vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8DFFEBE5B8F85BFDF1F834" bold="true" box="[502,584,1944,1968]" pageId="10" pageNumber="47">
|
||
<taxonomicNameLabel id="A27757D1FF8DFFEBE5B8F85BFDF1F834" box="[502,584,1944,1968]" pageId="10" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male holotype.
|
||
<emphasis id="B944EAAAFF8DFFEBE6BCF854FC90F82B" bold="true" box="[754,809,1943,1967]" pageId="10" pageNumber="47">A–B.</emphasis>
|
||
Head, anterior and lateral views;
|
||
<emphasis id="B944EAAAFF8DFFEBE0C9F854FB26F834" bold="true" box="[1159,1183,1943,1968]" pageId="10" pageNumber="47">C.</emphasis>
|
||
Thorax, dorsal view;
|
||
<emphasis id="B944EAAAFF8DFFEBE1CAF85BFA25F834" bold="true" box="[1412,1436,1944,1968]" pageId="10" pageNumber="47">D.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8F36B8FF8DFFEBE4D9F87FFC09F87C" blockId="10.[151,1436,1943,2046]" pageId="10" pageNumber="47">
|
||
Wing. Abbreviations: dm-m = discal medial crossvein; M
|
||
<subScript id="17B434FDFF8DFFEBE6AFF808FD51F85D" attach="left" box="[737,744,1995,2009]" fontSize="6" pageId="10" pageNumber="47">1</subScript>
|
||
= first branch of media; M
|
||
<subScript id="17B434FDFF8DFFEBE7B0F808FBBCF85D" attach="left" box="[1022,1029,1995,2009]" fontSize="6" pageId="10" pageNumber="47">4</subScript>
|
||
= fourth branch of media; R
|
||
<subScript id="17B434FDFF8DFFEBE164F808FA88F85D" attach="left" box="[1322,1329,1995,2009]" fontSize="6" pageId="10" pageNumber="47">1</subScript>
|
||
= anterior branch of radius; R
|
||
<subScript id="17B434FDFF8DFFEBE514F82CFEC8F879" attach="left" box="[346,369,2031,2045]" fontSize="6" pageId="10" pageNumber="47">2+3</subScript>
|
||
= second branch of radius; R
|
||
<subScript id="17B434FDFF8DFFEBE6D2F833FD0AF87A" attach="left" box="[668,691,2032,2046]" fontSize="6" pageId="10" pageNumber="47">4+5</subScript>
|
||
= third branch of radius.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6630FF8CFFEAE4D9F94EFD9EF8DD" ID-DOI="http://doi.org/10.5281/zenodo.7574111" ID-Zenodo-Dep="7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="11" pageNumber="48" startId="11.[151,250,1677,1701]" targetBox="[151,1435,181,1653]" targetPageId="11">
|
||
<paragraph id="8B8F36B8FF8CFFEAE4D9F94EFD9EF8DD" blockId="11.[151,1436,1677,1881]" pageId="11" pageNumber="48">
|
||
<emphasis id="B944EAAAFF8CFFEAE4D9F94EFEAFF921" bold="true" box="[151,278,1677,1701]" pageId="11" pageNumber="48">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C304D3BFF8CFFEAE551F94EFE4AF921" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[287,499,1677,1701]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
|
||
<emphasis id="B944EAAAFF8CFFEAE551F94EFE4AF921" box="[287,499,1677,1701]" italics="true" pageId="11" pageNumber="48">Paraclius vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8CFFEAE5B3F94EFDE9F921" bold="true" box="[509,592,1677,1701]" pageId="11" pageNumber="48">
|
||
<taxonomicNameLabel id="A27757D1FF8CFFEAE5B3F94EFDE9F921" box="[509,592,1677,1701]" pageId="11" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male holotype.
|
||
<emphasis id="B944EAAAFF8CFFEAE74EF94EFC8FF921" bold="true" box="[768,822,1677,1701]" pageId="11" pageNumber="48">A–B.</emphasis>
|
||
Hypopygium, not dissected and dissected respectively;
|
||
<emphasis id="B944EAAAFF8CFFEAE1CAF94EFA25F921" bold="true" box="[1412,1436,1677,1701]" pageId="11" pageNumber="48">C.</emphasis>
|
||
Internal appendages, left lateral view;
|
||
<emphasis id="B944EAAAFF8CFFEAE66FF972FD83F94D" bold="true" box="[545,570,1713,1737]" pageId="11" pageNumber="48">D.</emphasis>
|
||
Right cercus, outer view;
|
||
<emphasis id="B944EAAAFF8CFFEAE705F972FCDBF94D" bold="true" box="[843,866,1713,1737]" pageId="11" pageNumber="48">E.</emphasis>
|
||
Dorsal lobe of surstylus, inner view;
|
||
<emphasis id="B944EAAAFF8CFFEAE0A6F972FB42F94D" bold="true" box="[1256,1275,1713,1737]" pageId="11" pageNumber="48">F.</emphasis>
|
||
Ventral lobe of surstylus and postgonite, outer view;
|
||
<emphasis id="B944EAAAFF8CFFEAE655F916FD8CF969" bold="true" box="[539,565,1749,1773]" pageId="11" pageNumber="48">G.</emphasis>
|
||
Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A6533FF8CFFEDE489F844FC7FFF7D" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48" type="materials_examined">
|
||
<paragraph id="8B8F36B8FF8CFFEDE489F844FC7FFF7D" blockId="11.[151,1437,1927,2026]" lastBlockId="12.[151,1437,151,825]" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48">
|
||
<emphasis id="B944EAAAFF8CFFEAE489F844FD9FF825" bold="true" box="[199,550,1927,1953]" pageId="11" pageNumber="48">
|
||
Type material.
|
||
<typeStatus id="548B881AFF8CFFEAE5C9F844FD9FF825" box="[391,550,1927,1953]" pageId="11" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
♁, labelled: “[
|
||
<collectingCountry id="F3277628FF8CFFEAE697F844FCFFF825" box="[729,838,1927,1953]" name="Brazil" pageId="11" pageNumber="48">BRAZIL</collectingCountry>
|
||
]
|
||
<collectingRegion id="49F4F85AFF8CFFEAE714F844FC69F825" box="[858,976,1927,1953]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="48">São Paulo</collectingRegion>
|
||
| Campos do Jordão [ca
|
||
<geoCoordinate id="EE04507FFF8CFFEAE149F84AFA25F820" box="[1287,1436,1929,1956]" degrees="22" direction="south" minutes="44" orientation="latitude" pageId="11" pageNumber="48" precision="1" seconds="17.3" value="-22.738138">22°44′17.3″S</geoCoordinate>
|
||
<geoCoordinate id="EE04507FFF8CFFEAE4D9F86EFE82F84C" box="[151,315,1965,1992]" degrees="45" direction="west" minutes="36" orientation="longitude" pageId="11" pageNumber="48" precision="1" seconds="35.7" value="-45.609917">45°36′35.7″W</geoCoordinate>
|
||
] |
|
||
<date id="FF8E1078FF8CFFEAE514F868FE60F841" box="[346,473,1963,1989]" pageId="11" pageNumber="48" value="1944-12-29">29.xii.1944</date>
|
||
| F. Lane col.”; “25657”; “
|
||
<taxonomicName id="4C304D3BFF8CFFEAE75BF868FC2BF841" authorityName="Meigen" authorityYear="1824" box="[789,914,1963,1989]" class="Insecta" family="Dolichopodidae" genus="Sybistroma" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAAAFF8CFFEAE75BF868FC2BF841" box="[789,914,1963,1989]" italics="true" pageId="11" pageNumber="48">Sybistroma</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C304D3BFF8CFFEAE7E5F86FFB9AF841" authorityName="Schiner. MMMS" authorityYear="1868" box="[939,1059,1964,1989]" class="Insecta" family="Dolichopodidae" genus="Sybistroma" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis id="B944EAAAFF8CFFEAE7E5F86FFB9AF841" box="[939,1059,1964,1989]" italics="true" pageId="11" pageNumber="48">americana</emphasis>
|
||
</taxonomicName>
|
||
| Schin.” [handwritten]; “O gen.
|
||
<emphasis id="B944EAAAFF8CFFEAE4D9F80CFF6BF86D" box="[151,210,1999,2025]" italics="true" pageId="11" pageNumber="48">Sybis</emphasis>
|
||
|
|
||
<emphasis id="B944EAAAFF8CFFEAE4A7F813FE92F86D" box="[233,299,2000,2025]" italics="true" pageId="11" pageNumber="48">troma</emphasis>
|
||
é | Europeu | A espécie | america- | na deve | ser um no- | vo gênero” [The genus
|
||
<taxonomicName id="4C304D3BFF8CFFEAE09DF80CFAE9F86D" authorityName="Meigen" authorityYear="1824" box="[1235,1360,1999,2025]" class="Insecta" family="Dolichopodidae" genus="Sybistroma" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAAAFF8CFFEAE09DF80CFAE9F86D" box="[1235,1360,1999,2025]" italics="true" pageId="11" pageNumber="48">Sybistroma</emphasis>
|
||
</taxonomicName>
|
||
is European, and the species
|
||
<taxonomicName id="4C304D3BFF8BFFEDE5E4FF54FD9BFF34" authorityName="Schiner. MMMS" authorityYear="1868" box="[426,546,151,176]" class="Insecta" family="Dolichopodidae" genus="Sybistroma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis id="B944EAAAFF8BFFEDE5E4FF54FD9BFF34" box="[426,546,151,176]" italics="true" pageId="12" pageNumber="49">americana</emphasis>
|
||
</taxonomicName>
|
||
should be a new genus] [handwritten]; “
|
||
<typeStatus id="548B881AFF8BFFEDE7B8FF54FB34FF35" box="[1014,1165,151,177]" pageId="12" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
||
|
|
||
<taxonomicName id="4C304D3BFF8BFFEDE0EAFF54FA34FF35" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[1188,1421,151,177]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="vulcanoae">
|
||
<emphasis id="B944EAAAFF8BFFEDE0EAFF54FA34FF35" box="[1188,1421,151,177]" italics="true" pageId="12" pageNumber="49">Paraclius vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
| Soares, Capellari & Ale-Rocha [red label]” (
|
||
<collectionCode id="ED21AE7DFF8BFFEDE6C7FF78FD54FF51" box="[649,749,187,213]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="49">MZUSP</collectionCode>
|
||
).
|
||
<typeStatus id="548B881AFF8BFFEDE6B1FF78FCDEFF51" box="[767,871,187,213]" pageId="12" pageNumber="49" type="holotype">Holotype</typeStatus>
|
||
in good condition, terminalia dissected and stored in microvial on the same pin, both wings damaged, right IIIt
|
||
<subScript id="17B434FDFF8BFFEDE774FF32FCFBFF7B" attach="left" box="[826,834,241,255]" fontSize="6" pageId="12" pageNumber="49">5</subScript>
|
||
broken off.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6533FF8BFFEDE489FEC0FE8AFD2D" pageId="12" pageNumber="49" type="discussion">
|
||
<paragraph id="8B8F36B8FF8BFFEDE489FEC0FE8AFD2D" blockId="12.[151,1437,151,825]" pageId="12" pageNumber="49">
|
||
<emphasis id="B944EAAAFF8BFFEDE489FEC0FE82FE99" bold="true" box="[199,315,259,285]" pageId="12" pageNumber="49">Remarks.</emphasis>
|
||
This species is closely related to
|
||
<taxonomicName id="4C304D3BFF8BFFEDE68CFEC0FC02FE98" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[706,955,259,285]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus" status="comb. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE68CFEC0FC02FE98" box="[706,955,259,285]" italics="true" pageId="12" pageNumber="49">Paraclius americanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE78AFEC0FBFEFE99" bold="true" box="[964,1095,259,285]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE78AFEC0FBFEFE99" box="[964,1095,259,285]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as discussed above (see “Remarks” under
|
||
<taxonomicName id="4C304D3BFF8BFFEDE57DFEE4FE6DFEC4" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[307,468,295,321]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus" status="comb. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE57DFEE4FEFDFEC4" box="[307,324,295,320]" italics="true" pageId="12" pageNumber="49">P</emphasis>
|
||
.
|
||
<emphasis id="B944EAAAFF8BFFEDE51FFEE4FE6DFEC4" box="[337,468,295,320]" italics="true" pageId="12" pageNumber="49">americanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE594FEE4FDE0FEC5" bold="true" box="[474,601,295,321]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE594FEE4FDE0FEC5" box="[474,601,295,321]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). The new species can be differentiated by lower margin of clypeus pointed (
|
||
<figureCitation id="130B2A3DFF8BFFEDE4D1FE88FF45FEE1" box="[159,252,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1922,1946]" captionTargetBox="[151,1435,181,1898]" captionTargetId="figure-18@9.[151,1435,181,1898]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius vulcanoae sp. nov. Male holotype. A. Head, anterior view; B. Habitus, lateral view (before dissection)." figureDoi="http://doi.org/10.5281/zenodo.7574105" httpUri="https://zenodo.org/record/7574105/files/figure.png" pageId="12" pageNumber="49">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="130B2A3DFF8BFFEDE549FE89FE93FEE1" box="[263,298,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[156,1431,181,1920]" captionTargetId="figure-18@10.[156,1431,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Paraclius vulcanoae sp. nov. Male holotype. A–B. Head, anterior and lateral views; C. Thorax, dorsal view; D. Wing. Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R1 = anterior branch of radius; R2+3 = second branch of radius; R 4+5 = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.7574109" httpUri="https://zenodo.org/record/7574109/files/figure.png" pageId="12" pageNumber="49">5A</figureCitation>
|
||
) (lower margin of clypeus straight in
|
||
<taxonomicName id="4C304D3BFF8BFFEDE681FE88FCC8FEE0" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[719,881,331,357]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus" status="comb. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE681FE88FD59FEE0" box="[719,736,331,356]" italics="true" pageId="12" pageNumber="49">P</emphasis>
|
||
.
|
||
<emphasis id="B944EAAAFF8BFFEDE6A0FE88FCC8FEE0" box="[750,881,331,356]" italics="true" pageId="12" pageNumber="49">americanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE739FE88FC4EFEE1" bold="true" box="[887,1015,331,357]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE739FE88FC4EFEE1" box="[887,1015,331,357]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8BFFEDE048FE88FBE5FEE1" box="[1030,1116,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1841,1865]" captionTargetBox="[151,1435,416,1818]" captionTargetId="figure-111@2.[151,1435,416,1818]" captionTargetPageId="2" captionText="FIGURE 1. Paraclius americanus (Schiner) comb. nov. A, B, D, F Male lectotype (NHMW). C, E, G Female paralectotype (NHMW). A. Male habitus, lateral view; B. Male labels; C. Female habitus, lateral view; D. Male head, anterior view; E. Female head, anterior view; F. Male arista-like stylus; G. Female arista-like stylus. Original images lacked scale bars. Photographs by Ģnther Ŵss, NHMW." figureDoi="http://doi.org/10.5281/zenodo.7574097" httpUri="https://zenodo.org/record/7574097/files/figure.png" pageId="12" pageNumber="49">Fig. 1D</figureCitation>
|
||
)), basiventral epandrial lobe rounded at apex, apicoventral epandrial lobe wider than long, somewhat fan-shaped (
|
||
<figureCitation id="130B2A3DFF8BFFEDE000FEACFB7DFE0D" box="[1102,1220,367,393]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="12" pageNumber="49">Fig. 6B, C</figureCitation>
|
||
) (basiventral epandrial lobe hook-shaped and apicoventral epandrial lobe longer than wide in
|
||
<taxonomicName id="4C304D3BFF8BFFEDE7B8FE50FB23FE28" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[1014,1178,403,429]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus" status="comb. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE7B8FE50FBBEFE28" box="[1014,1031,403,428]" italics="true" pageId="12" pageNumber="49">P</emphasis>
|
||
.
|
||
<emphasis id="B944EAAAFF8BFFEDE059FE50FB23FE28" box="[1047,1178,403,428]" italics="true" pageId="12" pageNumber="49">americanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE0ECFE50FA9DFE29" bold="true" box="[1186,1316,403,429]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE0ECFE50FA9DFE29" box="[1186,1316,403,429]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA14B49FF8BFFEDE17AFE50FF6BFE55" author="Capellari, R. S." pageId="12" pageNumber="49" pagination="293 - 296" refId="ref8937" refString="Capellari, R. S. (2013) A remarkable new species of Paraclius Loew from Brazil (Diptera: Dolichopodidae). Zootaxa, 3681 (3), 293 - 296. https: // doi. org / 10.11646 / zootaxa. 3681.3.10" type="journal article" year="2013">Capellari 2013</bibRefCitation>
|
||
, fig. 5)). In addition, the two species share a sperm pump folded over itself, but evidently shorter in
|
||
<taxonomicName id="4C304D3BFF8BFFEDE174FE74FF74FE70" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE174FE74FF74FE70" italics="true" pageId="12" pageNumber="49">P. vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE49BFE18FE89FE71" bold="true" box="[213,304,475,501]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE49BFE18FE89FE71" box="[213,304,475,501]" pageId="12" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3DFF8BFFEDE50EFE18FE20FE71" box="[320,409,475,501]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="12" pageNumber="49">Fig. 6C</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFA14B49FF8BFFEDE5E3FE18FDCFFE71" author="Grichanov, I. Ya." box="[429,630,475,501]" pageId="12" pageNumber="49" pagination="183 - 190" refId="ref9162" refString="Grichanov, I. Ya. (1998) Afrotropical species of the genus Acropsilus Mik (Diptera: Dolichopodidae). International Journal of Dipterological Research, 9 (3), 183 - 190." type="journal article" year="1998">Grichanov (1998)</bibRefCitation>
|
||
suggested that differences in the orientation of the hypandrium-phallus apparatus in species of
|
||
<taxonomicName id="4C304D3BFF8BFFEDE5D4FE3CFDFEFD9D" authority="Mik" authorityName="Mik" authorityYear="1878" box="[410,583,511,537]" class="Insecta" family="Dolichopodidae" genus="Acropsilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAAAFF8BFFEDE5D4FE3CFDA8FD9D" box="[410,529,511,537]" italics="true" pageId="12" pageNumber="49">Acropsilus</emphasis>
|
||
Mik
|
||
</taxonomicName>
|
||
might be due by copulation (hypandrium and phallus would lay perpendicularly to each other after copulation). In a similar way, the shorter sperm pump in
|
||
<taxonomicName id="4C304D3BFF8BFFEDE052FDE0FB13FDB9" authority="Soares & Capellari & Ale-Rocha, 2023" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" box="[1052,1194,547,573]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="vulcanoae" status="sp. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE052FDE0FB13FDB9" box="[1052,1194,547,573]" italics="true" pageId="12" pageNumber="49">P. vulcanoae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE0FFFDE0FAB2FDB9" bold="true" box="[1201,1291,547,573]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE0FFFDE0FAB2FDB9" box="[1201,1291,547,573]" pageId="12" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
compared to that of
|
||
<taxonomicName id="4C304D3BFF8BFFEDE4A4FD84FE33FDE4" authorityName="Soares & Capellari & Ale-Rocha" authorityYear="2023" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[234,394,583,608]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="americanus" status="comb. nov.">
|
||
<emphasis id="B944EAAAFF8BFFEDE4A4FD84FE33FDE4" box="[234,394,583,608]" italics="true" pageId="12" pageNumber="49">P. americanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944EAAAFF8BFFEDE5DCFD84FDADFDE5" bold="true" box="[402,532,583,609]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="A27757D1FF8BFFEDE5DCFD84FDADFDE5" box="[402,532,583,609]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
could be a by-product of copulation, including the perpendicular orientation of hypandrium and phallus seen in the
|
||
<typeStatus id="548B881AFF8BFFEDE663FDA8FD36FD01" box="[557,655,619,645]" pageId="12" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130B2A3DFF8BFFEDE6D1FDA8FCA0FD01" box="[671,793,619,645]" captionStart="FIGURE 6" captionStartId="11.[151,250,1677,1701]" captionTargetBox="[151,1435,181,1653]" captionTargetId="figure-18@11.[151,1435,181,1653]" captionTargetPageId="11" captionText="FIGURE 6. Paraclius vulcanoae sp. nov. Male holotype. A–B. Hypopygium, not dissected and dissected respectively; C. Internal appendages, left lateral view; D. Right cercus, outer view; E. Dorsal lobe of surstylus, inner view; F. Ventral lobe of surstylus and postgonite, outer view; G. Phallus, ventral view. Abbreviations: apv lb = apicoventral lobe of epandrium; bv s = basiventral epandrial seta; bv lb = basiventral lobe of epandrium; cerc = cercus; d sur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; ph = phallus; ph process = lateral process of phallus; sp = sperm pump; v sur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.7574111" httpUri="https://zenodo.org/record/7574111/files/figure.png" pageId="12" pageNumber="49">Fig. 6A, B</figureCitation>
|
||
), as presumed by
|
||
<bibRefCitation id="EFA14B49FF8BFFEDE7ABFDA8FB15FD01" author="Grichanov, I. Ya." box="[997,1196,619,645]" pageId="12" pageNumber="49" pagination="183 - 190" refId="ref9162" refString="Grichanov, I. Ya. (1998) Afrotropical species of the genus Acropsilus Mik (Diptera: Dolichopodidae). International Journal of Dipterological Research, 9 (3), 183 - 190." type="journal article" year="1998">Grichanov (1998)</bibRefCitation>
|
||
for mated specimens of
|
||
<taxonomicName id="4C304D3BFF8BFFEDE4F8FD4CFE94FD2D" authorityName="Mik" authorityYear="1878" box="[182,301,655,681]" class="Insecta" family="Dolichopodidae" genus="Acropsilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAAAFF8BFFEDE4F8FD4CFE94FD2D" box="[182,301,655,681]" italics="true" pageId="12" pageNumber="49">Acropsilus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6533FF8BFFEDE489FD70FB5AFC91" pageId="12" pageNumber="49" type="etymology">
|
||
<paragraph id="8B8F36B8FF8BFFEDE489FD70FB5AFC91" blockId="12.[151,1437,151,825]" pageId="12" pageNumber="49">
|
||
<emphasis id="B944EAAAFF8BFFEDE489FD70FEF7FD49" bold="true" box="[199,334,691,717]" pageId="12" pageNumber="49">Etymology.</emphasis>
|
||
The new species is named after Maria Aparecida Vulcano (
|
||
<emphasis id="B944EAAAFF8BFFEDE048FD70FB24FD48" box="[1030,1181,691,716]" italics="true" pageId="12" pageNumber="49">in memoriam</emphasis>
|
||
), who was one of the precursors of the MZUSP
|
||
<taxonomicName id="4C304D3BFF8BFFEDE586FD14FDA5FD75" box="[456,540,727,753]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="12" pageNumber="49" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
|
||
collection during the 1940s. She wrote the handwritten label on the
|
||
<typeStatus id="548B881AFF8BFFEDE174FD14FA25FD75" box="[1338,1436,727,753]" pageId="12" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
mentioned above and the patronym acknowledges her insight while first identifying the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6533FF8BFFEDE489FCDDFA25FCBD" box="[199,1436,798,825]" pageId="12" pageNumber="49" type="distribution">
|
||
<paragraph id="8B8F36B8FF8BFFEDE489FCDDFA25FCBD" blockId="12.[151,1437,151,825]" box="[199,1436,798,825]" pageId="12" pageNumber="49">
|
||
<emphasis id="B944EAAAFF8BFFEDE489FCDDFED8FCBC" bold="true" box="[199,353,798,824]" pageId="12" pageNumber="49">Distribution.</emphasis>
|
||
<collectingCountry id="F3277628FF8BFFEDE52BFCDCFE10FCBD" box="[357,425,799,825]" name="Brazil" pageId="12" pageNumber="49">Brazil</collectingCountry>
|
||
(state of S„o Paulo).This species is known to occur only in the Atlantic Forest biome (
|
||
<figureCitation id="130B2A3DFF8BFFEDE102FCDCFA37FCBD" box="[1356,1422,799,825]" captionStart="FIGURE 7" captionStartId="12.[151,250,1643,1667]" captionTargetBox="[151,1435,849,1620]" captionTargetId="figure-338@12.[151,1435,849,1620]" captionTargetPageId="12" captionText="FIGURE 7. Geographic distribution of Paraclius Loew with bi-lamellate arista-like stylus." figureDoi="http://doi.org/10.5281/zenodo.7574115" httpUri="https://zenodo.org/record/7574115/files/figure.png" pageId="12" pageNumber="49">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |