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<document id="6F424B892EF5C7CE2DB2D04D66474E02" ID-CLB-Dataset="20038" ID-DOI="10.5281/zenodo.5318986" ID-GBIF-Dataset="3b197df1-d7f8-4513-9a9b-e5c3654415e4" ID-ISSN="0374-1036" ID-Zenodo-Dep="5318986" ID-ZooBank="A159251E-8C8E-4509-8C0C-34FBD0A79279M" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630107039701" checkinUser="marcus" docAuthor="Löbl, Ivan" docDate="2015" docId="03998786FF8D5F2AFE62FCFC0A1BFC49" docLanguage="en" docName="ActaEntMusNatPra.2015.55.1.129-144.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Scaphisoma apparatum Löbl 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="131" masterDocId="FFA0FFFEFF8C5F28FFCBFFED082FFFFB" masterDocTitle="Notes on Scaphisoma of Kalimantan (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="144" masterPageNumber="129" pageNumber="130" updateTime="1699156701922" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A0822A324F3692D6765194AD76B0C1BA">Notes on Scaphisoma of Kalimantan (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
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<mods:namePart id="7F2381B5428CF053555B3CF2745ADB65">Löbl, Ivan</mods:namePart>
<mods:affiliation id="568BD44EB5ECF028557290FF453085DA">Muséum dhistoire naturelle, Route de Malagnou 1, CH- 1208 Genève, Switzerland</mods:affiliation>
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<mods:title id="9D176AF70AD3D8EFB09280EB7EA5F545">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="D031B4398E54817E88DFDFE82DA5DB09">2015</mods:date>
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<treatment id="03998786FF8D5F2AFE62FCFC0A1BFC49" ID-DOI="http://doi.org/10.5281/zenodo.6402987" ID-GBIF-Taxon="194318098" ID-Zenodo-Dep="6402987" LSID="urn:lsid:plazi:treatment:03998786FF8D5F2AFE62FCFC0A1BFC49" httpUri="http://treatment.plazi.org/id/03998786FF8D5F2AFE62FCFC0A1BFC49" lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130">
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<paragraph id="8B8F3690FF8D5F29FE62FCFC0B26FCD2" blockId="1.[425,777,785,809]" box="[425,777,785,809]" pageId="1" pageNumber="130">
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<emphasis id="B944EA82FF8D5F29FE62FCFC0B26FCD2" bold="true" box="[425,777,785,809]" pageId="1" pageNumber="130">
<taxonomicName id="4C304D13FF8D5F29FE62FCFC0A82FCD3" ID-CoL="6Y5NJ" authority="Löbl, 2015" authorityName="Löbl" authorityYear="2015" box="[425,685,785,809]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="apparatum" status="sp. nov.">
<emphasis id="B944EA82FF8D5F29FE62FCFC0A82FCD3" bold="true" box="[425,685,785,809]" italics="true" pageId="1" pageNumber="130">Scaphisoma apparatum</emphasis>
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<taxonomicNameLabel id="A27757F9FF8D5F29FD7FFCFF0B26FCD2" box="[692,777,786,809]" pageId="1" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B8F3690FF8D5F29FFBAFCD20911FC96" blockId="1.[113,1089,831,983]" pageId="1" pageNumber="130">
<emphasis id="B944EA82FF8D5F29FFBAFCD208C4FCA9" bold="true" box="[113,235,831,850]" pageId="1" pageNumber="130">
<typeStatus id="548B8832FF8D5F29FFBAFCD208B1FCA9" box="[113,158,831,850]" pageId="1" pageNumber="130">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3277600FF8D5F29FF3AFCD20969FCA9" box="[241,326,831,850]" name="Indonesia" pageId="1" pageNumber="130">Indonesia</collectingCountry>
,
<collectingRegion id="49F4F872FF8D5F29FE84FCD209CFFCA9" box="[335,480,831,850]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="130">East Kalimantan</collectingRegion>
, ca
<quantity id="4CC89B75FF8D5F29FDC9FCD20A16FCA9" box="[514,569,831,850]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="130" unit="km" value="55.0">55 km</quantity>
W of Balikpapan, PT Fajar Surya Swadaya area,
<geoCoordinate id="EE045057FF8D5F29FC23FCD20C13FCA9" box="[1000,1084,831,850]" degrees="01" direction="south" minutes="13.3" orientation="latitude" pageId="1" pageNumber="130" precision="92" value="-1.2216667">01°13.3S</geoCoordinate>
,
<geoCoordinate id="EE045057FF8D5F29FFBAFCB708FDFC96" box="[113,210,858,877]" degrees="116" direction="east" minutes="21.0" orientation="longitude" pageId="1" pageNumber="130" precision="92" value="116.35">116°21.0E</geoCoordinate>
,
<quantity id="4CC89B75FF8D5F29FF10FCB7093EFC96" box="[219,273,858,877]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="130" unit="m" value="100.0">100 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B8F3690FF8D5F29FFBAFC990A11FC2C" blockId="1.[113,1089,831,983]" pageId="1" pageNumber="130">
<emphasis id="B944EA82FF8D5F29FFBAFC9908DAFC7C" bold="true" box="[113,245,884,903]" pageId="1" pageNumber="130">Type material.</emphasis>
<materialsCitation id="3B583CCDFF8D5F29FF32FC990C24FC46" ID-GBIF-Occurrence="3744299316" collectingDate="2011-11-24" collectingDateMax="2011-11-29" collectingDateMin="2011-11-24" collectingMethod="individual collecting, + light trap'" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" elevation="100" latitude="-1.2216667" location="border of Acacia mangium plantation and primary rainforest, stream and waterfall, puddles" longLatPrecision="130" longitude="116.35" pageId="1" pageNumber="130" specimenCount="1" specimenCount-female="1" stateProvince="E. Kalimantan" typeStatus="holotype">
<typeStatus id="548B8832FF8D5F29FF32FC990963FC7D" box="[249,332,884,903]" pageId="1" pageNumber="130" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D36FD19FF8D5F29FE9DFC830949FC7C" box="[342,358,878,903]" pageId="1" pageNumber="130" type="female"></specimenCount>
,
<collectingCountry id="F3277600FF8D5F29FEB8FC9909E3FC7C" box="[371,460,884,903]" name="Indonesia" pageId="1" pageNumber="130">Indonesia</collectingCountry>
,
<collectingRegion id="49F4F872FF8D5F29FE18FC990A7FFC7C" box="[467,592,884,903]" pageId="1" pageNumber="130">E. Kalimantan</collectingRegion>
,
<locationDeviation id="9A3D2FB2FF8D5F29FD9CFC990C6FFC7C" box="[599,1088,884,903]" pageId="1" pageNumber="130">
ca
<quantity id="4CC89B75FF8D5F29FDA5FC990A8CFC7C" box="[622,675,884,903]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="130" unit="km" value="55.0">55 km</quantity>
W of Balikpapan PT Fajar Surya Swadaya [area]
</locationDeviation>
<geoCoordinate id="EE045057FF8D5F29FFBAFC6208E7FC59" box="[113,200,911,930]" degrees="01" direction="south" minutes="13.3" orientation="latitude" pageId="1" pageNumber="130" precision="92" value="-1.2216667">01°13.3S</geoCoordinate>
<geoCoordinate id="EE045057FF8D5F29FF06FC62091FFC59" box="[205,304,911,930]" degrees="116" direction="east" minutes="21.0" orientation="longitude" pageId="1" pageNumber="130" precision="92" value="116.35">116°21.0E</geoCoordinate>
,
<quantity id="4CC89B75FF8D5F29FEF2FC620946FC59" box="[313,361,911,930]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="130" unit="m" value="100.0">
<elevation id="001DD1A3FF8D5F29FEF2FC620946FC59" box="[313,361,911,930]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="130" unit="m" value="100.0">100m</elevation>
</quantity>
<collectorName id="26C55346FF8D5F29FEA4FC620998FC59" box="[367,439,911,930]" pageId="1" pageNumber="130">J. Hájek</collectorName>
,
<collectorName id="26C55346FF8D5F29FE0BFC620A05FC59" box="[448,554,911,930]" pageId="1" pageNumber="130">J. Schneider</collectorName>
,
<collectorName id="26C55346FF8D5F29FDF9FC620AA6FC59" box="[562,649,911,930]" pageId="1" pageNumber="130">P. Votruba</collectorName>
leg.
<collectingDate id="EFCAE9B8FF8D5F29FD78FC620B63FC59" box="[691,844,911,930]" pageId="1" pageNumber="130">
24-25+
<date id="FF8E1050FF8D5F29FD39FC620B63FC59" box="[754,844,911,930]" pageId="1" pageNumber="130" value="2011-11-29">29.xi.2011</date>
</collectingDate>
/
<location id="8EEF604BFF8D5F29FC97FC620AA0FC46" LSID="urn:lsid:plazi:treatment:03998786FF8D5F2AFE62FCFC0A1BFC49:8EEF604BFF8D5F29FC97FC620AA0FC46" country="Indonesia" latitude="-1.2216667" longLatPrecision="130" longitude="116.35" name="border of Acacia mangium plantation and primary rainforest, stream and waterfall, puddles" pageId="1" pageNumber="130" stateProvince="E. Kalimantan">
border of
<taxonomicName id="4C304D13FF8D5F29FC64FC620C6EFC59" box="[943,1089,911,930]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="130" phylum="Tracheophyta" rank="species" species="mangium">Acacia mangium</taxonomicName>
plantation and primary rainforest, stream and waterfall, puddles
</location>
;
<collectingMethod id="52714E87FF8D5F29FD53FC470B9AFC46" box="[664,949,938,957]" pageId="1" pageNumber="130">individual collecting, + light trap</collectingMethod>
(
<collectionCode id="ED21AE55FF8D5F29FC74FC470C2FFC46" box="[959,1024,938,957]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="130">NMPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B583CCDFF8D5F29FBC5FC470A11FC2C" ID-GBIF-Occurrence="3744299313" collectingDate="2011-11-24" collectingDateMax="2011-11-29" collectingDateMin="2011-11-24" collectingMethod="individual collecting, + light trap'" collectionCode="MHNG" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" elevation="100" latitude="-1.2216667" location="border of Acacia mangium plantation and primary rainforest, stream and waterfall, puddles" longLatPrecision="130" longitude="116.35" pageId="1" pageNumber="130" specimenCount="1" specimenCount-female="1" stateProvince="E. Kalimantan" typeStatus="paratype">
<typeStatus id="548B8832FF8D5F29FBC5FC4708B9FC2D" pageId="1" pageNumber="130" type="paratype">
PARA
<subScript id="17B434D5FF8D5F29FBF2FC4708B9FC2D" attach="none" fontSize="6" pageId="1" pageNumber="130">TYPE</subScript>
</typeStatus>
:
<specimenCount id="9D36FD19FF8D5F29FF69FC53089DFC2C" box="[162,178,958,983]" pageId="1" pageNumber="130" type="female"></specimenCount>
, with the same data as the holotype (
<collectionCode id="ED21AE55FF8D5F29FE26FC290A1CFC2C" box="[493,563,964,983]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="130" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF8D5F2AFFBAFC1D0A36FD89" lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130" type="description">
<paragraph id="8B8F3690FF8D5F2AFFBAFC1D0A36FD89" blockId="1.[113,1090,1008,1575]" lastBlockId="2.[151,1127,219,946]" lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130">
<emphasis id="B944EA82FF8D5F29FFBAFC1D08D4FBFC" bold="true" box="[113,251,1008,1031]" pageId="1" pageNumber="130">Description.</emphasis>
Length
<quantity id="4CC89B75FF8D5F29FE84FC1D0981FBFC" box="[335,430,1008,1031]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="130" unit="mm" value="1.9">1.90 mm</quantity>
, width
<quantity id="4CC89B75FF8D5F29FE33FC1D0A77FBFC" box="[504,600,1008,1031]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="1" pageNumber="130" unit="mm" value="1.36">1.36 mm</quantity>
. Head and body dark reddish-brown, pronotum darkened along basal margin, elytra darkened in middle, between basal and apical fourth. Femora, tibiae and abdominal ventrites lighter, tarsi and antennae much lighter than pronotum. Pronotum without microsculpture, elytra with conspicuous punctulate microsculpture, dull. Antennae with length/width ratio of antennomeres as follows: III 17/11: IV 33/10: V 45/10: VI 56/11: VII 62/14: VIII 52/11: IX 67/15: X 55/15: XI 77/15. Pronotum with lateral margins rounded, lateral margin carinae visible in dorsal view except near angles, discal punctation fairly fine and dense, well delimited, puncture intervals mostly about twice as large to as large as puncture diameters, pubescence distinct. Apex of scutellum exposed. Elytra dull, with conspicuous microsculpture consisting of punctures and striae, and very shallowly wrinkled; lateral margin carinae exposed in dorsal view, apical margins slightly rounded, inner apical angle not prominent, situated about in level with outer angles, sutural margin not raised, sutural striae shallow, curved at base, extended along basal margins to form basal striae reaching outer third of elytral basal width (dorsal view), parallel between level of scutellum and mid-length, converging from mid-length apically, adsutural areas flat, with puncture row, combined about
<quantity id="4CC89B75FF8D5F29FE31FA3D0A75FA1C" box="[506,602,1488,1511]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="130" unit="mm" value="0.15">0.15 mm</quantity>
wide at mid-length, with very fine, irregular puncture row. Elytral punctation very fine and sparse, hardly visible at 50× magnification, punctures margining sutural striae distinct. Punctation fairly distinct and dense on basal part of pygidium, becoming finer toward pygidial apex. Hypomera not microsculptured, very finely punctate. Mesepimeres shorter than interval to mesocoxae and 4 times as long as wide. Metaventrite not microsculptured, convex in middle, flattened apicomesally, without apicomedian impressions, with punctation very fine between mesocoxae, becoming coarse and dense toward apical intercoxal process, with coarse punctures well delimited and to part larger than puncture intervals. Lateral parts of metaventrite very finely and sparsely punctate, without antecoxal puncture row. Submesocoxal lines parallel, margined by conspicuously coarse puncture; submesocoxal areas
<quantity id="4CC89B75FF8E5F2AFDFFFE560AB7FE29" box="[564,664,443,466]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="131" unit="mm" value="0.05">0.05 mm</quantity>
, about as fourth of interval to metacoxae. Metanepisterna convex,
<quantity id="4CC89B75FF8E5F2AFE68FE360A2AFE09" box="[419,517,475,498]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="131" unit="mm" value="0.12">0.12 mm</quantity>
wide, moderately narrowed anteriad, with inner suture almost straight, deeply impressed. Tibiae straight. Abdomen with strigulate microsculpture, ventrites extremely finely and sparsely punctate. Submetacoxale lines of ventrite 1 subparallel, conspicuously coarsely punctate, submetacoxal areas about
<quantity id="4CC89B75FF8E5F2AFCD8FDD60B5EFDA9" box="[787,881,571,594]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="131" unit="mm" value="0.05">0.05 mm</quantity>
long, almost as fifth of interval to apical margin of ventrite.
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF8E5F2AFF5CFD960B48FCA9" pageId="2" pageNumber="131" type="diagnosis">
<paragraph id="8B8F3690FF8E5F2AFF5CFD960B48FCA9" blockId="2.[151,1127,219,946]" pageId="2" pageNumber="131">
<emphasis id="B944EA82FF8E5F2AFF5CFD9609B9FD69" bold="true" box="[151,406,635,658]" pageId="2" pageNumber="131">Differential diagnosis.</emphasis>
This species differs significantly from all Asian congeners, with the notable exception of the Japanese
<taxonomicName id="4C304D13FF8E5F2AFDE7FD770AD2FD49" authority="Lobl, 1982" authorityName="Lobl" authorityYear="1982" box="[556,765,666,690]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B944EA82FF8E5F2AFDE7FD770A51FD49" box="[556,638,666,690]" italics="true" pageId="2" pageNumber="131">S. pecki</emphasis>
<bibRefCitation id="EFA14B61FF8E5F2AFD4EFD760AD2FD49" author="LOBL I." box="[645,765,667,690]" pageId="2" pageNumber="131" pagination="5 - 16" refId="ref8555" refString="LOBL I. 1982: Little known and new Oriental species of the genus Scaphisoma Leach (Coleoptera, Scaphidiidae). Pp. 5 - 16. In: SATO M., HORI Y., ARITA Y. &amp; OKADOME T. (eds): Special issue to the memory of retirement of Emeritus Professor Michio Chujo. Association of the Memorial Issue of Emeritus Professor M. Chujo C / O Biological Laboratory, Nagoya Women's University, Nagoya, Japan, 185 pp." type="book chapter" year="1982">Löbl, 1982</bibRefCitation>
</taxonomicName>
, by its dull, conspicuously microsculptured elytra. It may be easily distinguished from
<taxonomicName id="4C304D13FF8E5F2AFD16FD570B1FFD29" authorityName="Lobl" authorityYear="1982" box="[733,816,698,722]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B944EA82FF8E5F2AFD16FD570B1FFD29" box="[733,816,698,722]" italics="true" pageId="2" pageNumber="131">S. pecki</emphasis>
</taxonomicName>
by the darkened middle part of the elytra, the elytral punctation being hardly visible, much finer than pronotal punctation, the antennomere XI longer than X, while both are equally long in
<taxonomicName id="4C304D13FF8E5F2AFC92FD170B83FCE9" authorityName="Lobl" authorityYear="1982" box="[857,940,762,786]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B944EA82FF8E5F2AFC92FD170B83FCE9" box="[857,940,762,786]" italics="true" pageId="2" pageNumber="131">S. pecki</emphasis>
</taxonomicName>
, the metaventrite lacking impressions and with narrower submesocoxal areas, the presence of strigulate abdominal microsculpture, and the entire ventrite 1 very finely punctate.
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF8E5F2AFF5CFCB60924FC69" pageId="2" pageNumber="131" type="etymology">
<paragraph id="8B8F3690FF8E5F2AFF5CFCB60924FC69" blockId="2.[151,1127,219,946]" pageId="2" pageNumber="131">
<emphasis id="B944EA82FF8E5F2AFF5CFCB60937FC89" bold="true" box="[151,280,859,882]" pageId="2" pageNumber="131">Etymology.</emphasis>
The species epithet is a Latin adjective, referring to the conspicuous distinctive characters.
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF8E5F2AFF5CFC760A1BFC49" box="[151,564,923,946]" pageId="2" pageNumber="131" type="distribution">
<paragraph id="8B8F3690FF8E5F2AFF5CFC760A1BFC49" blockId="2.[151,1127,219,946]" box="[151,564,923,946]" pageId="2" pageNumber="131">
<emphasis id="B944EA82FF8E5F2AFF5CFC760905FC49" bold="true" box="[151,298,923,946]" pageId="2" pageNumber="131">Distribution.</emphasis>
<collectingCountry id="F3277600FF8E5F2AFEFBFC7609B7FC49" box="[304,408,923,946]" name="Indonesia" pageId="2" pageNumber="131">Indonesia</collectingCountry>
(
<collectingRegion id="49F4F872FF8E5F2AFE6DFC760A06FC49" box="[422,553,923,946]" country="Indonesia" name="Kalimantan Utara" pageId="2" pageNumber="131">Kalimantan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>