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<mods:title id="A11981E0374771322E518F7965D4563D">Macropodidae</mods:title>
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<mods:namePart id="38F301906D3B8FB00F6431710B0B877C">Russell A. Mittermeier</mods:namePart>
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<mods:title id="0301EB986BFF57FE6DEBDF16407D64D7">Handbook of the Mammals of the World Volume 5 Monotremes and Marsupials</mods:title>
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<treatment id="039504399654FFB16ABAF4F2F8D33192" ID-DOI="http://doi.org/10.5281/zenodo.6722388" ID-GBIF-Taxon="196400521" ID-Zenodo-Dep="6722388" LSID="urn:lsid:plazi:treatment:039504399654FFB16ABAF4F2F8D33192" httpUri="http://treatment.plazi.org/id/039504399654FFB16ABAF4F2F8D33192" lastPageNumber="700" pageId="9" pageNumber="700">
<subSubSection id="C326E6A49654FFB16ABAF4F2FF373C04" box="[177,231,2841,2887]" pageId="9" pageNumber="700" type="multiple">
<paragraph id="8B83B52F9654FFB16ABAF4F2FF373C04" blockId="9.[172,1150,2841,2969]" box="[177,231,2841,2887]" pageId="9" pageNumber="700">
<heading id="D0CB02439654FFB16ABAF4F2FF373C04" box="[177,231,2841,2887]" pageId="9" pageNumber="700">
<figureCitation id="1307A9AA9654FFB16ABAF4F2FF373C04" box="[177,231,2841,2887]" captionStart="Plate 37: Macropodidae" captionStartId="12.[93,123,3416,3441]" captionTargetBox="[13,2734,13,3644]" captionTargetPageId="7" captionText="10. Rufous-bellied Pademelon (Thylogale billardieru), 11. Red-necked Pademelon (Thylogale thetis), 12. Red-legged Pademelon (Thylogale stigmatica), 13. New Guinea Pademelon (Thylogale browni), 14. Dusky Pademelon (Thylogale brunii), 15. Calabys Pademelon (Thylogale calabyi), 16. Bennett's Tree Kangaroo (Dendrolagus bennettianus), 17. Lumholtzs Tree Kangaroo (Dendrolagus lumholtzi), 18. Grizzled Tree Kangaroo (Dendrolagus inustus), 19. Vogelkop Tree Kangaroo (Dendrolagus ursinus), 20. Dingiso (Dendrolagus mbaiso), 21. Dorias. 9 Tree Kangaroo Dendrolagus (- dorianus), 22. Scotts Tree Kangaroo (Dendrolagus scottae), 23. Goodfellows Tree Kangaroo (Dendrolagus goodfellowi), 24. Huon Tree Kangaroo (Dendrolagus matschie), 25. Lowland Tree Kangaroo (Dendrolagus spadix)" figureDoi="http://doi.org/10.5281/zenodo.6723980" httpUri="https://zenodo.org/record/6723980/files/figure.png" pageId="9" pageNumber="700">13.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C326E6A49654FFB16AF3F4F2FD243C04" box="[248,756,2841,2887]" pageId="9" pageNumber="700" type="vernacular_names">
<paragraph id="8B83B52F9654FFB16AF3F4F2FD243C04" blockId="9.[172,1150,2841,2969]" box="[248,756,2841,2887]" pageId="9" pageNumber="700">
<heading id="D0CB02439654FFB16AF3F4F2FD243C04" box="[248,756,2841,2887]" pageId="9" pageNumber="700">
<vernacularName id="053FC5019654FFB16AF3F4F2FD243C04" box="[248,756,2841,2887]" pageId="9" pageNumber="700">New Guinea Pademelon</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16935F4F2FBB73C04" box="[830,1127,2841,2887]" pageId="9" pageNumber="700" type="nomenclature">
<paragraph id="8B83B52F9654FFB16935F4F2FBB73C04" blockId="9.[172,1150,2841,2969]" box="[830,1127,2841,2887]" pageId="9" pageNumber="700">
<heading id="D0CB02439654FFB16935F4F2FBB73C04" box="[830,1127,2841,2887]" pageId="9" pageNumber="700">
<taxonomicName id="4C3CCEAC9654FFB16935F4F2FBB73C04" ID-CoL="56Q7J" baseAuthorityName="Ramsay" baseAuthorityYear="1877" box="[830,1127,2841,2887]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="browni">
<emphasis id="B948693D9654FFB16935F4F2FBB73C04" box="[830,1127,2841,2887]" italics="true" pageId="9" pageNumber="700">Thylogale browni</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16AA6F4B2FC773CD5" pageId="9" pageNumber="700" type="vernacular_names">
<paragraph id="8B83B52F9654FFB16AA6F4B2FBAD3C2D" blockId="9.[172,1150,2841,2969]" box="[173,1149,2905,2926]" pageId="9" pageNumber="700">
<heading id="D0CB02439654FFB16AA6F4B2FBAD3C2D" box="[173,1149,2905,2926]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16AA6F4B2FF293C2D" bold="true" box="[173,249,2905,2926]" pageId="9" pageNumber="700">French:</emphasis>
<vernacularName id="053FC5019654FFB16AF5F4B2FE143C2D" box="[254,452,2905,2926]" pageId="9" pageNumber="700">Thylogale de Brown</vernacularName>
/
<emphasis id="B948693D9654FFB16BD1F4B2FDE43C2D" bold="true" box="[474,564,2905,2926]" pageId="9" pageNumber="700">German:</emphasis>
<vernacularName id="053FC5019654FFB16834F4B2FD323C2D" box="[575,738,2905,2926]" pageId="9" pageNumber="700">Brauner Filander</vernacularName>
/
<emphasis id="B948693D9654FFB168FDF4B2FC823C2D" bold="true" box="[758,850,2905,2926]" pageId="9" pageNumber="700">Spanish:</emphasis>
<vernacularName id="053FC5019654FFB16950F4B2FBAD3C2D" box="[859,1149,2905,2926]" pageId="9" pageNumber="700">Pademelon de Nueva Guinea</vernacularName>
</heading>
</paragraph>
<paragraph id="8B83B52F9654FFB16AA6F46AFC773CD5" blockId="9.[172,1150,2841,2969]" box="[173,935,2945,2966]" pageId="9" pageNumber="700">
<heading id="D0CB02439654FFB16AA6F46AFC773CD5" box="[173,935,2945,2966]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16AA6F46AFE743CD5" bold="true" box="[173,420,2945,2966]" pageId="9" pageNumber="700">Other common names:</emphasis>
<vernacularName id="053FC5019654FFB16BA5F46AFDA53CD5" box="[430,629,2945,2966]" pageId="9" pageNumber="700">Brown's Pademelon</vernacularName>
,
<vernacularName id="053FC5019654FFB1688FF46AFC773CD5" box="[644,935,2945,2966]" pageId="9" pageNumber="700">Mountain Pademelon (/anata)</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB1691BF423FC8A3B4F" pageId="9" pageNumber="700" type="reference_group">
<paragraph id="8B83B52F9654FFB1691BF423FC8A3B4F" blockId="9.[784,1379,3016,3438]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB1691BF423FC7C3CA6" bold="true" box="[784,940,3016,3045]" pageId="9" pageNumber="700">Taxonomy.</emphasis>
<taxonomicName id="4C3CCEAC9654FFB169C1F423FC863B4F" authority="Ramsay, 1877" authorityName="Ramsay" authorityYear="1877" class="Mammalia" family="Macropodidae" genus="Halmaturus" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="browni">Halmaturus browni: Ramsay, 1877</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB1696EF404FAE73B4F" box="[869,1335,3055,3084]" pageId="9" pageNumber="700" type="reference_group">
<paragraph id="8B83B52F9654FFB1696EF404FAE73B4F" blockId="9.[784,1379,3016,3438]" box="[869,1335,3055,3084]" pageId="9" pageNumber="700">
<materialsCitation id="3B54BF729654FFB1696EF404FAE73B4F" ID-GBIF-Occurrence="3819448344" box="[869,1335,3055,3084]" pageId="9" pageNumber="700">New Ireland, Papua New Guinea.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16919F3F9FDFB3AD7" pageId="9" pageNumber="700" type="discussion">
<paragraph id="8B83B52F9654FFB16919F3F9FA8D3A2D" blockId="9.[784,1379,3016,3438]" pageId="9" pageNumber="700">
Formerly regarded as a subspecies of 1 bruni, but was reinstated as a species in 1992. It has been suggested that lanata be treated as a full species; recent molecular studies, however, do not support this, nor do they support the distinction between
<taxonomicName id="4C3CCEAC9654FFB1691EF315FC5E3A5C" authorityName="Ramsay" authorityYear="1877" baseAuthorityName="Ramsay" baseAuthorityYear="1877" box="[789,910,3326,3359]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="browni">T. browni</taxonomicName>
and either of the two other endemic New Guinea
<taxonomicName id="4C3CCEAC9654FFB16E38F2CDFB7A3A04" box="[1075,1194,3366,3399]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="genus">Thylogale</taxonomicName>
taxa (
<taxonomicName id="4C3CCEAC9654FFB16F01F2CDFCE53A2D" authorityName="Schreber" authorityYear="1778" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="brunii">T. brunii</taxonomicName>
,
<taxonomicName id="4C3CCEAC9654FFB16944F2BAFC033A2D" authorityName="Flannery" authorityYear="1992" box="[847,979,3409,3438]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="calabyi">T. calabyi</taxonomicName>
). Further studies required.
</paragraph>
<paragraph id="8B83B52F9654FFB16AA7F298FDFB3AD7" blockId="9.[172,555,3443,3476]" box="[172,555,3443,3476]" pageId="9" pageNumber="700">Two subspecies recognized.</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA5FECEF6AA36FE" pageId="9" pageNumber="700" type="synonymic_list">
<caption id="DF43E5A79654FFB16FA5FECEF6AA36FE" ID-DOI="http://doi.org/10.5281/zenodo.6723762" ID-Zenodo-Dep="6723762" httpUri="https://zenodo.org/record/6723762/files/figure.png" inLine="true" pageId="9" pageNumber="700" targetBox="[170,761,3022,3435]" targetPageId="9">
<paragraph id="8B83B52F9654FFB16FA5FECEF8EE3605" blockId="9.[1450,2661,293,1751]" box="[1454,1854,293,326]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA5FECEF8EE3605" bold="true" box="[1454,1854,293,326]" pageId="9" pageNumber="700">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="8B83B52F9654FFB16FB9FEA6F7EC362D" blockId="9.[1450,2661,293,1751]" box="[1458,2108,333,366]" pageId="9" pageNumber="700">
<taxonomicName id="4C3CCEAC9654FFB16FB9FEA6F8D8362D" authority="Ramsay, 1877" authorityName="Ramsay" authorityYear="1877" box="[1458,1800,333,366]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="subSpecies" species="browni" subSpecies="browni">T. b. browni Ramsay, 1877</taxonomicName>
— N &amp; E New Guinea.
</paragraph>
<paragraph id="8B83B52F9654FFB16FB9FE9FF6BC36D6" blockId="9.[1450,2661,293,1751]" box="[1458,2412,372,405]" pageId="9" pageNumber="700">
<taxonomicName id="4C3CCEAC9654FFB16FB9FE9FF8DA36D6" authorityName="Thomas" authorityYear="1922" box="[1458,1802,372,405]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="subSpecies" species="browni" subSpecies="lanata">T.b. lanata Thomas 1922</taxonomicName>
— Huon Peninsula, NE Papua New Guinea.
</paragraph>
<paragraph id="8B83B52F9654FFB16FA5FE77F6AA36FE" blockId="9.[1450,2661,293,1751]" box="[1454,2426,412,445]" pageId="9" pageNumber="700">Introduced into Yapen I and several islands in Bismarck Archipelago.</paragraph>
</caption>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA6FE2CF85335BB" pageId="9" pageNumber="700" type="description">
<paragraph id="8B83B52F9654FFB16FA6FE2CF85335BB" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA6FE2CF97A36A7" bold="true" box="[1453,1706,455,484]" pageId="9" pageNumber="700">Descriptive notes.</emphasis>
Head—body 53:5-66.7 cm (males) and 48.7-56 cm (females), tail 36.6-52 cm (males) and 30-42.5 cm (females); weight 4.9-1 kg (males) and 3.5-9 kg (females). Light to dark brown dorsally, paler ventrally (gray to white, underfur gray). Fur typically short, but can be long in specimens from high elevations. Limbs same color as body, and largely hairless on inside. Face may be darker. Tail dark dorsally, but thinly furred ventrally and distally; a pale tail tip occasionally present. In 7. b. lanata, fur is long and woolly,tail shorter and better furred, and nape paler. Diploid chromosome number of
<taxonomicName id="4C3CCEAC9654FFB16CA8FD3CF8E835BB" authority="Ramsay, 1877" authorityName="Ramsay" authorityYear="1877" box="[1699,1848,727,760]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="browni">T. b. browni</taxonomicName>
is 22.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA6FD14F86134D5" pageId="9" pageNumber="700" type="biology_ecology">
<paragraph id="8B83B52F9654FFB16FA6FD14F86134D5" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA6FD14F9CC3463" bold="true" box="[1453,1564,767,800]" pageId="9" pageNumber="700">Habitat.</emphasis>
<taxonomicName id="4C3CCEAC9654FFB16C20FD14F9113463" authorityName="Ramsay" authorityYear="1877" box="[1579,1729,767,800]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="700" phylum="Chordata" rank="species" species="browni">T. b. browni</taxonomicName>
inhabits primary and secondary lowland and lower montane rainforest, adjacent grassland and abandoned gardens, below 2100 m elevation; may prefer disturbed areas or ecotones. 7. b. lanata occurs in subalpine grassland and adjacent montane rainforest, at 3000-3800 m.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA6FC76F95034A7" pageId="9" pageNumber="700" type="food_feeding">
<paragraph id="8B83B52F9654FFB16FA6FC76F95034A7" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA6FC76F96634FD" bold="true" box="[1453,1718,925,958]" pageId="9" pageNumber="700">Food and Feeding.</emphasis>
Poorly known. Reported to feed on grass, herbs, browse, and occasionally fruit.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA6FC00F9103370" pageId="9" pageNumber="700" type="breeding">
<paragraph id="8B83B52F9654FFB16FA6FC00F9103370" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA6FC00F9E3334F" bold="true" box="[1453,1587,1003,1036]" pageId="9" pageNumber="700">Breeding.</emphasis>
Poorly known. Females reportedly produce a single young and are likely to breed continuously.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA1FBD1F85A33C1" pageId="9" pageNumber="700" type="activity">
<paragraph id="8B83B52F9654FFB16FA1FBD1F85A33C1" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA1FBD1F9453318" bold="true" box="[1450,1685,1082,1115]" pageId="9" pageNumber="700">Activity patterns.</emphasis>
Likely to be nocturnal or crepuscular, but specific activity patterns of this species are virtually unknown.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA7FB62F9503393" pageId="9" pageNumber="700" type="biology_ecology">
<paragraph id="8B83B52F9654FFB16FA7FB62F9503393" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA7FB62F7A333E9" bold="true" box="[1452,2163,1161,1194]" pageId="9" pageNumber="700">Movements, Home range and Social organization.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA7FB33F8113118" pageId="9" pageNumber="700" type="conservation">
<paragraph id="8B83B52F9654FFB16FA7FB33F8113118" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA7FB33F8DA33BA" bold="true" box="[1452,1802,1240,1273]" pageId="9" pageNumber="700">Status and Conservation.</emphasis>
Treated as two species on the IUCN Red List: “1. brown?” classified as Vulnerable and “7. lanatus” classified as Endangered. This pademelon has declined in parts of its range and continues to be threatened by hunting. It is a favored subsistence game animal and is susceptible to overhunting by local people using dogs. The species was introduced to islands around northern New Guinea (e.g. Bismarck Archipelago, Yapen) in prehistory, most likely as a source of food. Subsequently, several of these populations have become extinct and others (e.g. New Ireland) are experiencing severe declines owing to hunting pressure. Additional research on taxonomy, abundance, general ecology, and impact of threats is required; the implementation of hunting regulations may be necessary.
</paragraph>
</subSubSection>
<subSubSection id="C326E6A49654FFB16FA7F981F8D33192" pageId="9" pageNumber="700" type="bibRefCitation_list">
<paragraph id="8B83B52F9654FFB16FA7F981F8D33192" blockId="9.[1450,2661,293,1751]" pageId="9" pageNumber="700">
<emphasis id="B948693D9654FFB16FA7F981F99531C0" bold="true" box="[1452,1605,1642,1667]" pageId="9" pageNumber="700">Bibliography.</emphasis>
Flannery (1992, 1995a, 1995b), Flannery et al. (1988), Groves (2005b), Hayman (1989), Heinsohn (2003, 2005a), Leary, Seri, Flannery, Wright, Hamilton, Helgen, Singadan, Menzies, Allison &amp; James (2008a, 2008b), Macqueen et al. (2011).
</paragraph>
</subSubSection>
</treatment>
</document>