507 lines
60 KiB
XML
507 lines
60 KiB
XML
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<mods:title id="37EF2EFCDE313DE6117BF1F058330E86">A first account of Chyromyidae (Diptera: Acalyptratae) from continental Central and South America and some Caribbean Islands, with descriptions of new species</mods:title>
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<mods:namePart id="607FAF63D4961B6B3959891DF9586EA7">Ebejer, Martin J.</mods:namePart>
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<mods:title id="096733EDE54A82AAA8530759321A200A">Zootaxa</mods:title>
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<mods:date id="19A8B11EEC8F4AF136828847C522C44A">2023</mods:date>
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<taxonomicName id="4C2C97CCFFE8FF9A37DEF89EFE58701A" authority="Ebejer, 2009" authorityName="Ebejer" authorityYear="2009" box="[151,458,1834,1861]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFE8FF9A37DEF89EFE58701A" bold="true" box="[151,458,1834,1861]" pageId="11" pageNumber="312">
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<emphasis id="B958305DFFE8FF9A37DEF89EFEB5701B" bold="true" box="[151,295,1834,1861]" italics="true" pageId="11" pageNumber="312">Tethysimyia</emphasis>
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<bibRefCitation id="EFBD91BEFFE8FF9A3667F89EFE58701A" author="Ebejer, M. J." box="[302,458,1834,1860]" pageId="11" pageNumber="312" pagination="321 - 434" refId="ref14102" refString="Ebejer, M. J. (2009) A revision of Afrotropical Chyromyidae (excluding Gymnochiromyia Hendel) (Diptera: Schizophora), with the recognition of two subfamilies and the description of new genera. African Invertebrates, 50 (2), 321 - 434. https: // doi. org / 10.5733 / afin. 050.0208" type="journal article" year="2009">Ebejer, 2009</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C336BFC4FFE8FF9D37DEF8C4FC0772C3" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312" type="materials_examined">
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<paragraph id="8B93EC4FFFE8FF9A37DEF8C4FC0B70D9" blockId="11.[151,921,1903,1928]" box="[151,921,1903,1928]" pageId="11" pageNumber="312">
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<typeStatus id="549752EDFFE8FF9A37DEF8C4FF5B70D6" box="[151,201,1904,1928]" pageId="11" pageNumber="312">Type</typeStatus>
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species:
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<taxonomicName id="4C2C97CCFFE8FF9A366FF8DBFD0C70D9" authority="Ebejer, 1996" authorityName="Ebejer" authorityYear="1996" box="[294,670,1903,1927]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="deemingi">
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<emphasis id="B958305DFFE8FF9A366FF8DBFD8770D9" box="[294,533,1903,1927]" italics="true" pageId="11" pageNumber="312">Aphaniosoma deemingi</emphasis>
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<bibRefCitation id="EFBD91BEFFE8FF9A3552F8DBFD0C70D9" author="Ebejer, M. J." box="[539,670,1903,1927]" pageId="11" pageNumber="312" pagination="280 - 299" refId="ref14014" refString="Ebejer, M. J. (1996) Chyromyidae (Diptera: Schizophora) from the Arabian Peninsula with descriptions of twelve new species. Fauna of Saudi Arabia, 15, 280 - 299." type="journal article" year="1996">Ebejer, 1996</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="8B93EC4FFFE8FF9D37DEF801FE6E74FB" blockId="11.[151,1436,1973,2036]" lastBlockId="12.[151,1437,151,1437]" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
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With the availability of more material it is now possible to give some more detail on characters of
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<taxonomicName id="4C2C97CCFFE8FF9A3259F801FA077091" authorityName="Ebejer" authorityYear="2009" box="[1296,1429,1973,1999]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFE8FF9A3259F801FA077091" box="[1296,1429,1973,1999]" italics="true" pageId="11" pageNumber="312">Tethysimyia</emphasis>
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: fronto-orbital setae usually reduced in size and number or absent, paravertical setulae reduced to a minute size, they can be convergent or divergent, parallel or absent; 1 dorsocentral seta and usually a short one in front of this; acrostichals always in at least 6 rows, usually in 8–10 and somewhat irregular; intrapostalar and prescutellar setae/setulae present; proepisternal seta/setula absent; epandrium small with surstylus fused, aedeagus simple, of uncomplicated structure, with distiphallus more or less straight, weakly sclerotized and mostly translucent; pregonites and postgonites though usually present are often not clearly discernible. The hypandrium is narrow with almost parallel sides and elongated; phallapodeme long rod-like, extending anteriorly in the abdomen. None of the species has the wing veins
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<collectionCode id="ED3D748AFFEFFF9D3627FEDBFE1376D7" box="[366,385,367,393]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="313">R</collectionCode>
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<subScript id="17A8EE0AFFEFFF9D36C8FE35FE0976D1" attach="left" box="[385,411,385,399]" fontSize="6" pageId="12" pageNumber="313">4+5</subScript>
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and M
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<subScript id="17A8EE0AFFEFFF9D36AEFE35FE7D76D1" attach="left" box="[487,495,385,399]" fontSize="6" pageId="12" pageNumber="313">1</subScript>
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diverging towards the apex of the wing or the complex development of the sternites and gonites typical of
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<taxonomicName id="4C2C97CCFFEFFF9D36C4FE27FDB576F3" authorityName="Becker" authorityYear="1903" box="[397,551,403,429]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D36C4FE27FDB576F3" box="[397,551,403,429]" italics="true" pageId="12" pageNumber="313">Aphaniosoma</emphasis>
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</taxonomicName>
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. In
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<taxonomicName id="4C2C97CCFFEFFF9D351BFE27FD4576F3" authorityName="Ebejer" authorityYear="2009" box="[594,727,403,429]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D351BFE27FD4576F3" box="[594,727,403,429]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
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</taxonomicName>
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there is sometimes a row of stronger setulae along the length of the hind tibia, which may end in a longer preapical seta. Species of
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<taxonomicName id="4C2C97CCFFEFFF9D3435FE03FB93768F" authorityName="Ebejer" authorityYear="2009" box="[892,1025,439,465]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D3435FE03FB93768F" box="[892,1025,439,465]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
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</taxonomicName>
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differ from
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<taxonomicName id="4C2C97CCFFEFFF9D33CEFE03FF5D76AB" authority="Ebejer, 2009" authorityName="Ebejer" authorityYear="2009" class="Insecta" family="Chyromyidae" genus="Paraphaniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D33CEFE03FADA768F" box="[1159,1352,439,465]" italics="true" pageId="12" pageNumber="313">Paraphaniosoma</emphasis>
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||
<bibRefCitation id="EFBD91BEFFEFFF9D3204FE03FF5D76AB" author="Ebejer, M. J." pageId="12" pageNumber="313" pagination="321 - 434" refId="ref14102" refString="Ebejer, M. J. (2009) A revision of Afrotropical Chyromyidae (excluding Gymnochiromyia Hendel) (Diptera: Schizophora), with the recognition of two subfamilies and the description of new genera. African Invertebrates, 50 (2), 321 - 434. https: // doi. org / 10.5733 / afin. 050.0208" type="journal article" year="2009">
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Ebejer,
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||
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</bibRefCitation>
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in the broader frons that projects above and in front of the eye, in a more developed facial carina and much broader gena.
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<taxonomicName id="4C2C97CCFFEFFF9D367DFE4BFE677547" authorityName="Ebejer" authorityYear="2009" box="[308,501,511,537]" class="Insecta" family="Chyromyidae" genus="Paraphaniosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D367DFE4BFE677547" box="[308,501,511,537]" italics="true" pageId="12" pageNumber="313">Paraphaniosoma</emphasis>
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</taxonomicName>
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has a postsutural prealar seta, a more complex male hypopygium with modifications also of the pregenital sternites, and a surstylus articulated with the epandrium. Female
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<taxonomicName id="4C2C97CCFFEFFF9D332CFD97FB787563" authorityName="Ebejer" authorityYear="2009" box="[1125,1258,547,573]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D332CFD97FB787563" box="[1125,1258,547,573]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
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</taxonomicName>
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have the paired spermathecae flat and disc-shaped with slight cupping on one side (except in the Arabian species
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<taxonomicName id="4C2C97CCFFEFFF9D325EFDF3FE0675DB" authority="Ebejer, 1996" authorityName="Ebejer" authorityYear="1996" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="deemingi">
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<emphasis id="B958305DFFEFFF9D325EFDF3FF6C75DB" italics="true" pageId="12" pageNumber="313">Tethysimyia deemingi</emphasis>
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<bibRefCitation id="EFBD91BEFFEFFF9D364DFDDFFE0675DB" author="Ebejer, M. J." box="[260,404,619,645]" pageId="12" pageNumber="313" pagination="280 - 299" refId="ref14014" refString="Ebejer, M. J. (1996) Chyromyidae (Diptera: Schizophora) from the Arabian Peninsula with descriptions of twelve new species. Fauna of Saudi Arabia, 15, 280 - 299." type="journal article" year="1996">Ebejer, 1996</bibRefCitation>
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</taxonomicName>
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). Among the species so far examined these structures are relatively uniform. In
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<taxonomicName id="4C2C97CCFFEFFF9D324BFDDFFA0E75DB" authorityName="Becker" authorityYear="1903" box="[1282,1436,619,645]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B958305DFFEFFF9D324BFDDFFA0E75DB" box="[1282,1436,619,645]" italics="true" pageId="12" pageNumber="313">Aphaniosoma</emphasis>
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||
</taxonomicName>
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the spermathecae tend to be spherical with only some irregularity on one side.
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<taxonomicName id="4C2C97CCFFEFFF9D3348FD3BFB1475F7" authorityName="Ebejer" authorityYear="2009" box="[1025,1158,655,681]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B958305DFFEFFF9D3348FD3BFB1475F7" box="[1025,1158,655,681]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
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||
</taxonomicName>
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has some of the smallest species among the Aphaniosominae. The New World species differ from the characters originally described for the Aphaniosominae in the following: the anepisternal seta is placed at the middle of the posterior margin and not on the junction of the upper and middle third; the humeral crossvein is evanescent in most specimens; the dorsal surface of the costa sometimes bears a few spinose setulae as in the
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<taxonomicName id="4C2C97CCFFEFFF9D3452FCABFC257467" box="[795,951,799,825]" class="Insecta" family="Chyromyidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="subTribe" subTribe="Chyromyinae">Chyromyinae</taxonomicName>
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; some females have mature ova easily seen in a distended abdomen—a feature more typical of the
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<taxonomicName id="4C2C97CCFFEFFF9D35B4FCF7FC0B7403" box="[765,921,835,861]" class="Insecta" family="Chyromyidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="subTribe" subTribe="Chyromyinae">Chyromyinae</taxonomicName>
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. These characters suggest that the Neotropical species might belong in another genus, but this will require stronger evidence, examination of more species, and support from molecular studies.
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</paragraph>
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<paragraph id="8B93EC4FFFEFFF9D378EFC1BFE4373B7" blockId="12.[151,1437,151,1437]" pageId="12" pageNumber="313">
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Based on the above characteristics, the species described in
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<taxonomicName id="4C2C97CCFFEFFF9D3434FC1BFB857497" authorityName="Becker" authorityYear="1903" box="[893,1047,943,969]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
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<emphasis id="B958305DFFEFFF9D3434FC1BFB857497" box="[893,1047,943,969]" italics="true" pageId="12" pageNumber="313">Aphaniosoma</emphasis>
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</taxonomicName>
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from
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<collectingRegion id="49E822ADFFEFFF9D3329FC1BFB237497" box="[1120,1201,943,969]" country="United States of America" name="Hawaii" pageId="12" pageNumber="313">Hawaii</collectingRegion>
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(Hardy, in
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<bibRefCitation id="EFBD91BEFFEFFF9D327EFC1BFEDB74B3" author="Hardy, D. E. & Delfinado, M. D." pageId="12" pageNumber="313" refId="ref14282" refString="Hardy, D. E. & Delfinado, M. D. (1980) Chyromyidae. In: Insects of Hawaii. Vol. 13. Diptera: Cyclorrhapha III. Series Schizophora. Section Acalypterae, exclusive of Family Drosophilidae. University of Hawaii Press, Honolulu, pp. 172 - 178." type="book" year="1980">Hardy & Delfinado 1980</bibRefCitation>
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||
) and the
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<collectingRegion id="49E822ADFFEFFF9D36FFFC67FDBE74B3" box="[438,556,979,1005]" country="Ecuador" name="Galapagos" pageId="12" pageNumber="313">Galapagos</collectingRegion>
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(Wheeler, in
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<bibRefCitation id="EFBD91BEFFEFFF9D3583FC67FC7A74B3" author="Wheeler, T. A. & Sinclair, B. J." box="[714,1000,979,1005]" pageId="12" pageNumber="313" pagination="440 - 453" refId="ref14497" refString="Wheeler, T. A. & Sinclair, B. J. (1994) Chyromyidae (Diptera) from the Galapagos Islands, Ecuador: three new species of Aphaniosoma Becker. Proceedings of the Entomological Society of Washington, 96 (3), 440 - 453." type="journal article" year="1994">Wheeler & Sinclair 1994</bibRefCitation>
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) should be transferred to
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<taxonomicName id="4C2C97CCFFEFFF9D3259FC67FA0774B3" authorityName="Ebejer" authorityYear="2009" box="[1296,1429,979,1005]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958305DFFEFFF9D3259FC67FA0774B3" box="[1296,1429,979,1005]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
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</taxonomicName>
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. The description of
|
||
<taxonomicName id="4C2C97CCFFEFFF9D363CFC43FD33734F" authority="Wheeler, 1961" authorityName="Wheeler" authorityYear="1961" box="[373,673,1015,1041]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="aldrichi">
|
||
<emphasis id="B958305DFFEFFF9D363CFC43FE63734F" box="[373,497,1015,1041]" italics="true" pageId="12" pageNumber="313">A. aldrichi</emphasis>
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D36B3FC43FD33734F" author="Wheeler, M. R." box="[506,673,1015,1041]" pageId="12" pageNumber="313" pagination="86 - 91" refId="ref14458" refString="Wheeler, M. R. (1961) New species of southwestern acalyptrate Diptera. The SouthWestern Naturalist, 6 (2), 86 - 91. https: // doi. org / 10.2307 / 3669590" type="journal article" year="1961">Wheeler, 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
includes the following details: “Cheeks very broad, covered with short pale hairs …. One pair of dorsocentrals, acrostichals in about 8 irregular rows …. Wings …. 2L and 3L parallel at tip ….” (2L and 3L refer to veins
|
||
<collectionCode id="ED3D748AFFEFFF9D35DDFB8BFD357307" box="[660,679,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="313">R</collectionCode>
|
||
<subScript id="17A8EE0AFFEFFF9D35EEFBE5FD527301" attach="left" box="[679,704,1105,1119]" fontSize="6" pageId="12" pageNumber="313">2+3</subScript>
|
||
and
|
||
<collectionCode id="ED3D748AFFEFFF9D35BFFB8BFC9B7307" box="[758,777,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="313">R</collectionCode>
|
||
<subScript id="17A8EE0AFFEFFF9D3440FBE5FCB17301" attach="left" box="[777,803,1105,1119]" fontSize="6" pageId="12" pageNumber="313">4+5</subScript>
|
||
respectively). These characters also place
|
||
<taxonomicName id="4C2C97CCFFEFFF9D324CFB8BFAEC7307" authorityName="Wheeler" authorityYear="1961" box="[1285,1406,1087,1113]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="aldrichi">
|
||
<emphasis id="B958305DFFEFFF9D324CFB8BFAEC7307" box="[1285,1406,1087,1113]" italics="true" pageId="12" pageNumber="313">A. aldrichi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C2C97CCFFEFFF9D37DEFBD7FE8E7323" authorityName="Ebejer" authorityYear="2009" box="[151,284,1123,1149]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958305DFFEFFF9D37DEFBD7FE8E7323" box="[151,284,1123,1149]" italics="true" pageId="12" pageNumber="313">Tethysimyia</emphasis>
|
||
</taxonomicName>
|
||
. Thus, the following new combinations are proposed:
|
||
<taxonomicName id="4C2C97CCFFEFFF9D34D7FBD7FADE7323" authority="(Wheeler, 1961)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1961" box="[926,1356,1123,1149]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="aldrichi" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D34D7FBD7FB157323" box="[926,1159,1123,1149]" italics="true" pageId="12" pageNumber="313">Tethysimyia aldrichi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D33D0FBD7FAD67323" author="Wheeler, M. R." box="[1177,1348,1123,1149]" pageId="12" pageNumber="313" pagination="86 - 91" refId="ref14458" refString="Wheeler, M. R. (1961) New species of southwestern acalyptrate Diptera. The SouthWestern Naturalist, 6 (2), 86 - 91. https: // doi. org / 10.2307 / 3669590" type="journal article" year="1961">Wheeler, 1961</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D321EFBD7FF5B73FF" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2C97CCFFEFFF9D379DFB33FDEC73FF" authority="(Hardy, 1980)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Hardy" baseAuthorityYear="1980" box="[212,638,1159,1185]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="macalpinei" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D379DFB33FE4973FF" box="[212,475,1159,1185]" italics="true" pageId="12" pageNumber="313">Tethysimyia macalpinei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D36A3FB33FDE473FF" author="Hardy, D. E. & Delfinado, M. D." box="[490,630,1159,1185]" pageId="12" pageNumber="313" refId="ref14282" refString="Hardy, D. E. & Delfinado, M. D. (1980) Chyromyidae. In: Insects of Hawaii. Vol. 13. Diptera: Cyclorrhapha III. Series Schizophora. Section Acalypterae, exclusive of Family Drosophilidae. University of Hawaii Press, Honolulu, pp. 172 - 178." type="book" year="1980">Hardy, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D35CCFB33FC9073FF" box="[645,770,1159,1185]" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2C97CCFFEFFF9D3444FB33FB1B73FF" authority="(Hardy, 1980)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Hardy" baseAuthorityYear="1980" box="[781,1161,1159,1185]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="minuta" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D3444FB33FC7473FE" box="[781,998,1159,1185]" italics="true" pageId="12" pageNumber="313">Tethysimyia minuta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D34BDFB33FB1273FF" author="Hardy, D. E. & Delfinado, M. D." box="[1012,1152,1159,1185]" pageId="12" pageNumber="313" refId="ref14282" refString="Hardy, D. E. & Delfinado, M. D. (1980) Chyromyidae. In: Insects of Hawaii. Vol. 13. Diptera: Cyclorrhapha III. Series Schizophora. Section Acalypterae, exclusive of Family Drosophilidae. University of Hawaii Press, Honolulu, pp. 172 - 178." type="book" year="1980">Hardy, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D33C6FB33FA9F73FF" box="[1167,1293,1159,1185]" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2C97CCFFEFFF9D325EFB33FE2E739B" authority="(Wheeler, 1994)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1994" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="arenicola" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D325EFB33FE93739B" italics="true" pageId="12" pageNumber="313">Tethysimyia arenicola</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D3646FB1FFE26739B" author="Wheeler, T. A. & Sinclair, B. J." box="[271,436,1195,1221]" pageId="12" pageNumber="313" pagination="440 - 453" refId="ref14497" refString="Wheeler, T. A. & Sinclair, B. J. (1994) Chyromyidae (Diptera) from the Galapagos Islands, Ecuador: three new species of Aphaniosoma Becker. Proceedings of the Entomological Society of Washington, 96 (3), 440 - 453." type="journal article" year="1994">Wheeler, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D368BFB1FFDAD739B" box="[450,575,1195,1221]" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2C97CCFFEFFF9D3500FB1FFB81739B" authority="(Wheeler, 1994)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1994" box="[585,1043,1195,1221]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="galamarilla" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D3500FB1FFCCA739B" box="[585,856,1195,1221]" italics="true" pageId="12" pageNumber="313">Tethysimyia galamarilla</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D342CFB1FFB99739B" author="Wheeler, T. A. & Sinclair, B. J." box="[869,1035,1195,1221]" pageId="12" pageNumber="313" pagination="440 - 453" refId="ref14497" refString="Wheeler, T. A. & Sinclair, B. J. (1994) Chyromyidae (Diptera) from the Galapagos Islands, Ecuador: three new species of Aphaniosoma Becker. Proceedings of the Entomological Society of Washington, 96 (3), 440 - 453." type="journal article" year="1994">Wheeler, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D3350FB1FFB06739B" box="[1049,1172,1195,1221]" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C2C97CCFFEFFF9D338EFB1FFEDC73B7" authority="(Wheeler, 1994)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1994" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="rabida" status="comb. nov.">
|
||
<emphasis id="B958305DFFEFFF9D338EFB1FFA0F739B" box="[1223,1437,1195,1221]" italics="true" pageId="12" pageNumber="313">Tethysimyia rabida</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D37D6FB7BFED773B7" author="Wheeler, T. A. & Sinclair, B. J." box="[159,325,1231,1257]" pageId="12" pageNumber="313" pagination="440 - 453" refId="ref14497" refString="Wheeler, T. A. & Sinclair, B. J. (1994) Chyromyidae (Diptera) from the Galapagos Islands, Ecuador: three new species of Aphaniosoma Becker. Proceedings of the Entomological Society of Washington, 96 (3), 440 - 453." type="journal article" year="1994">Wheeler, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D361CFB7BFE4373B7" box="[341,465,1231,1257]" pageId="12" pageNumber="313" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEFFF9D378EFB47FC0772C3" blockId="12.[151,1437,151,1437]" pageId="12" pageNumber="313">
|
||
<bibRefCitation id="EFBD91BEFFEFFF9D378EFB47FE617253" author="Wheeler, T. A. & Sinclair, B. J." box="[199,499,1267,1293]" pageId="12" pageNumber="313" pagination="440 - 453" refId="ref14497" refString="Wheeler, T. A. & Sinclair, B. J. (1994) Chyromyidae (Diptera) from the Galapagos Islands, Ecuador: three new species of Aphaniosoma Becker. Proceedings of the Entomological Society of Washington, 96 (3), 440 - 453." type="journal article" year="1994">Wheeler & Sinclair (1994)</bibRefCitation>
|
||
illustrated the terminalia of the three species they described. Their interpretation of some structures differs from that of this author. In their figures, the structure depicted as arising equally from the caudal end of the hypandrium as it is from the anteroventral part of the epandrium is labelled as surstylus, but this is more likely to be the pregonite and their bacilliform sclerite the postgonite. Their figures also depict the long phallapodeme and hypandrium as in the new species here described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336BFC4FFEFFF9C37DEFA5BFB017309" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313" type="key">
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEFA5BFA177172" blockId="12.[151,1437,1518,1581]" pageId="12" pageNumber="313">
|
||
<emphasis id="B958305DFFEFFF9D37DEFA5BFD577156" bold="true" box="[151,709,1518,1545]" pageId="12" pageNumber="313">Key to Central and South American species of</emphasis>
|
||
<emphasis id="B958305DFFEFFF9D3583FA5BFCCF7157" box="[714,861,1519,1545]" italics="true" pageId="12" pageNumber="313">
|
||
<taxonomicName id="4C2C97CCFFEFFF9D3583FA5BFCCA7157" authorityName="Ebejer" authorityYear="2009" box="[714,856,1519,1545]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">Tethysimyia</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B958305DFFEFFF9D342BFA5AFA107156" bold="true" box="[866,1410,1518,1545]" pageId="12" pageNumber="313">
|
||
including the Oceanian species from
|
||
<collectingRegion id="49E822ADFFEFFF9D3268FA5AFAEF7156" box="[1313,1405,1518,1544]" country="United States of America" name="Hawaii" pageId="12" pageNumber="313">Hawaii</collectingRegion>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C2C97CCFFEFFF9D32CEFA5BFF667173" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1961" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="aldrichi">
|
||
<emphasis id="B958305DFFEFFF9D32CEFA5BFF667173" italics="true" pageId="12" pageNumber="313">T. aldrichi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B958305DFFEFFF9D37B2F9A6FA177172" bold="true" box="[251,1413,1554,1581]" pageId="12" pageNumber="313">is excluded as no material has been seen to examine it for detail missing from the description.</emphasis>
|
||
</paragraph>
|
||
<key id="FFBDBB7EFFEFFF9C37DEF9E5FA0F7256" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
|
||
<keyStep id="30D8F4EAFFEFFF9D37DEF9E5FA0E7185" pageId="12" pageNumber="313">
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF9E5FA0F71DA" blockId="12.[151,1437,1617,1991]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF9E5FA0F71DA" pageId="12" pageNumber="313">
|
||
1 Head longer than high; frons not projecting; eye long oval, long diameter about twice length of short diameter; wing veins R
|
||
<subScript id="17A8EE0AFFEFFF9D32CFF9D4FA0E7132" attach="left" box="[1414,1436,1632,1644]" fontSize="5" pageId="12" pageNumber="313">4+5</subScript>
|
||
and M 1 parallel to apex of wing.......................................................
|
||
<taxonomicName id="4C2C97CCFFEFFF9D33E5F9DAFA9171DA" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[1196,1283,1646,1668]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis id="B958305DFFEFFF9D33E5F9DAFA9171DA" bold="true" box="[1196,1283,1646,1668]" italics="true" pageId="12" pageNumber="313">chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEFFF9D3241F9DBFAC171DA" box="[1288,1363,1647,1668]" pageId="12" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry id="F33BACDFFFEFFF9D3216F9DAFA0471DA" box="[1375,1430,1646,1668]" name="Chile" pageId="12" pageNumber="313">Chile</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF938FA0E7185" blockId="12.[151,1437,1617,1991]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF938FA0E7185" pageId="12" pageNumber="313">
|
||
- Head spherical, or a little higher than long; frons projecting beyond eye margin above and anteriorly; eye short oval with long diameter only about 1.3 times as long as short diameter; wing veins R
|
||
<subScript id="17A8EE0AFFEFFF9D34CCF90CFC08719A" attach="left" box="[901,922,1720,1732]" fontSize="5" pageId="12" pageNumber="313">4+5</subScript>
|
||
and M
|
||
<subScript id="17A8EE0AFFEFFF9D3494F90CFC76719A" attach="left" box="[989,996,1720,1732]" fontSize="5" pageId="12" pageNumber="313">1</subScript>
|
||
convergent towards apex of wing, sometimes only slightly so...................................................................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEFFF9D37DEF8B5FA0E7030" pageId="12" pageNumber="313">
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF8B5FA0E706A" blockId="12.[151,1437,1617,1991]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF8B5FA0E706A" pageId="12" pageNumber="313">
|
||
2 Frontal setulae reclinate; scutum grey anteriorly; acrostichals in 8–10 rows; abdominal tergites grey with pale posterior margins....................................................................
|
||
<taxonomicName id="4C2C97CCFFEFFF9D335DF8AAFB54706A" authority="Wheeler" authorityName="Wheeler" box="[1044,1222,1822,1844]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="B958305DFFEFFF9D335DF8AAFBFD706D" box="[1044,1135,1822,1843]" italics="true" pageId="12" pageNumber="313">arenicola</emphasis>
|
||
Wheeler
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F33BACDFFFEFFF9D339BF8AAFAB7706A" box="[1234,1317,1822,1844]" name="Ecuador" pageId="12" pageNumber="313">Ecuador</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E822ADFFEFFF9D3266F8AAFA04706A" box="[1327,1430,1822,1844]" country="Ecuador" name="Galapagos" pageId="12" pageNumber="313">Galapagos</collectingRegion>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF888FA0E7030" blockId="12.[151,1437,1617,1991]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF888FA0E7030" pageId="12" pageNumber="313">- Frontal setulae proclinate; scutum yellow; acrostichals in 6–8 rows; abdominal tergites yellow, at most with narrow, pale brown bands anterodorsally.................................................................................. 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEFFF9D37DEF820FA0E7098" pageId="12" pageNumber="313">
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF820FA0E70F7" blockId="12.[151,1437,1617,1991]" box="[151,1436,1940,1962]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF820FA0E70F7" box="[151,1436,1940,1962]" pageId="12" pageNumber="313">
|
||
<specimenCount id="9D2A27C6FFEFFF9D37DEF820FE8F70F4" box="[151,285,1940,1962]" pageId="12" pageNumber="313" type="male">3 Males</specimenCount>
|
||
.............................................................................................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEFFF9D37DEF805FA0E7098" blockId="12.[151,1437,1617,1991]" box="[151,1436,1969,1991]" pageId="12" pageNumber="313">
|
||
<keyLead id="30DD4F7AFFEFFF9D37DEF805FA0E7098" box="[151,1436,1969,1991]" pageId="12" pageNumber="313">- Females............................................................................................ 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFF2CFA0E7795" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFF2CFA0F77F3" blockId="13.[151,1437,151,1288]" box="[151,1437,151,173]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFF2CFA0F77F3" box="[151,1437,151,173]" pageId="13" pageNumber="314">
|
||
4 Frons, face and basal flagellomere black (
|
||
<figureCitation id="1317F0CAFFEEFF9C353AFF23FD2A77F3" box="[627,696,151,173]" captionStart="FIGURE 15" captionStartId="23.[151,250,1884,1908]" captionTargetBox="[189,1398,993,1861]" captionTargetId="figure-326@23.[189,1398,993,1861]" captionTargetPageId="23" captionText="FIGURE 15. Tethysimyia nigrifacies sp. nov., holotype, male. A. habitus, lateral view. B. dorsal oblique view. C. head, anterolateral view. D. same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8202879" httpUri="https://zenodo.org/record/8202879/files/figure.png" pageId="13" pageNumber="314">Fig. 15</figureCitation>
|
||
).................................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3314FF23FB5477F3" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[1117,1222,151,173]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="nigrifacies" status="sp. nov.">
|
||
<emphasis id="B958305DFFEEFF9C3314FF23FB5477F3" bold="true" box="[1117,1222,151,173]" italics="true" pageId="13" pageNumber="314">nigrifacies</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEEFF9C3385FF2CFA8577F3" box="[1228,1303,152,173]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(West Indies)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFF01FA0E7795" blockId="13.[151,1437,151,1288]" box="[151,1436,181,203]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFF01FA0E7795" box="[151,1436,181,203]" pageId="13" pageNumber="314">– Frons, face and basal flagellomere yellow................................................................. 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFF5BFA0E761E" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFF5BFA0E767D" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFF5BFA0E767D" pageId="13" pageNumber="314">
|
||
5 Hind basitarsus distinctly modified: broad and flattened dorsoventrally; hind tibia with prominent anterior setae
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3251FF44FA0E765B" authority="Hardy" authorityName="Hardy" box="[1304,1436,239,261]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="minuta">
|
||
<emphasis id="B958305DFFEEFF9C3251FF44FAC9765B" box="[1304,1371,240,261]" italics="true" pageId="13" pageNumber="314">minuta</emphasis>
|
||
Hardy
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="49E822ADFFEEFF9C37A1FEB9FEA2767D" box="[232,304,269,291]" country="United States of America" name="Hawaii" pageId="13" pageNumber="314">Hawaii</collectingRegion>
|
||
) &
|
||
<taxonomicName id="4C2C97CCFFEEFF9C361FFEB9FD8C767D" authority="Wheeler" authorityName="Wheeler" box="[342,542,269,291]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="galamarilla">
|
||
<emphasis id="B958305DFFEEFF9C361FFEB9FE55767C" box="[342,455,269,290]" italics="true" pageId="13" pageNumber="314">galamarilla</emphasis>
|
||
Wheeler
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="49E822ADFFEEFF9C3562FEB9FD01767D" box="[555,659,269,291]" country="Ecuador" name="Galapagos" pageId="13" pageNumber="314">Galapagos</collectingRegion>
|
||
)................(distinguishable by the characters in the male hypopygium)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFE9FFA0E761E" blockId="13.[151,1437,151,1288]" box="[151,1436,298,320]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFE9FFA0E761E" box="[151,1436,298,320]" pageId="13" pageNumber="314">- Hind basitarsus not modified; tibia without prominent anterior setae............................................. 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFED1FA0E76EB" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFED1FA0E76C6" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFED1FA0E76C6" pageId="13" pageNumber="314">
|
||
6 Mediotergite black; scutum usually with pale brown vittae; abdominal tergites dark shiny brown with narrow pale hind margins; eyes with microtrichia between every 1 or 2 ommatidia........
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3429FE36FC2176C6" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[864,947,386,408]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="litophila" status="sp. nov.">
|
||
<emphasis id="B958305DFFEEFF9C3429FE36FC2176C6" bold="true" box="[864,947,386,408]" italics="true" pageId="13" pageNumber="314">litophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEEFF9C34F0FE37FB9176C6" box="[953,1027,387,408]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Pacific coasts of
|
||
<collectingCountry id="F33BACDFFFEEFF9C33FDFE36FA8E76C6" box="[1204,1308,386,408]" name="Costa Rica" pageId="13" pageNumber="314">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BACDFFFEEFF9C3202FE36FA0476C6" box="[1355,1430,386,408]" name="Mexico" pageId="13" pageNumber="314">Mexico</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFE14FA0E76EB" blockId="13.[151,1437,151,1288]" box="[151,1436,415,437]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFE14FA0E76EB" box="[151,1436,415,437]" pageId="13" pageNumber="314">- Without the above combination of characters............................................................... 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFE6FFA0E7553" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFE6FFA0F76AE" blockId="13.[151,1437,151,1288]" box="[151,1437,474,496]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFE6FFA0F76AE" box="[151,1437,474,496]" pageId="13" pageNumber="314">
|
||
7 Paravertical setulae present; acrostichals in 6–8 rows..................................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C33C3FE6EFAB376AE" authority="Wheeler" authorityName="Wheeler" box="[1162,1313,474,496]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rabida">
|
||
<emphasis id="B958305DFFEEFF9C33C3FE6EFB5876B1" box="[1162,1226,474,495]" italics="true" pageId="13" pageNumber="314">rabida</emphasis>
|
||
Wheeler
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="49E822ADFFEEFF9C3267FE6EFA0476AE" box="[1326,1430,474,496]" country="Ecuador" name="Galapagos" pageId="13" pageNumber="314">Galapagos</collectingRegion>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFE4CFA0E7553" blockId="13.[151,1437,151,1288]" box="[151,1436,503,525]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFE4CFA0E7553" box="[151,1436,503,525]" pageId="13" pageNumber="314">- Paravertical setulae absent; acrostichals in 8–10 rows........................................................ 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFD86FA0F753B" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFD86FA0F7516" blockId="13.[151,1437,151,1288]" box="[151,1437,562,584]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFD86FA0F7516" box="[151,1437,562,584]" pageId="13" pageNumber="314">
|
||
8 acrostichals in 10 rows at level of transverse scutal suture................................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C33DDFD86FAD37516" authority="Hardy" authorityName="Hardy" box="[1172,1345,562,584]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="macalpinei">
|
||
<emphasis id="B958305DFFEEFF9C33DDFD86FB6D7519" box="[1172,1279,562,583]" italics="true" pageId="13" pageNumber="314">macalpinei</emphasis>
|
||
Hardy
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="49E822ADFFEEFF9C3207FD86FA047516" box="[1358,1430,562,584]" country="United States of America" name="Hawaii" pageId="13" pageNumber="314">Hawaii</collectingRegion>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFDE4FA0F753B" blockId="13.[151,1437,151,1288]" box="[151,1437,591,613]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFDE4FA0F753B" box="[151,1437,591,613]" pageId="13" pageNumber="314">
|
||
- acrostichals in 8 rows at level of transverse scutal suture...............................
|
||
<emphasis id="B958305DFFEEFF9C3331FDFBFB78753B" box="[1144,1258,591,613]" italics="true" pageId="13" pageNumber="314">
|
||
<emphasis id="B958305DFFEEFF9C3331FDFBFB73753B" bold="true" box="[1144,1249,591,613]" italics="true" pageId="13" pageNumber="314">bonairensi</emphasis>
|
||
s
|
||
</emphasis>
|
||
sp. nov. (
|
||
<collectingCountry id="F33BACDFFFEEFF9C320EFDFBFA04753B" box="[1351,1430,591,613]" name="Bonaire, Sint Eustatius and Saba" pageId="13" pageNumber="314">Bonaire</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFD3EFA0E75E3" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFD3EFA0E75C1" blockId="13.[151,1437,151,1288]" box="[151,1436,650,672]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFD3EFA0E75C1" box="[151,1436,650,672]" pageId="13" pageNumber="314">
|
||
9 Paravertical setulae absent (
|
||
<figureCitation id="1317F0CAFFEEFF9C36A7FD3EFDD775C1" box="[494,581,650,672]" captionStart="FIGURE 19" captionStartId="26.[151,250,1815,1839]" captionTargetBox="[204,1384,273,1791]" captionTargetId="figure-79@26.[204,1384,273,1792]" captionTargetPageId="26" captionText="FIGURE 19. Tethysimyia spp. females, A. species A, head, posterolateral view, scale bar = 0.25mm. B. same species (another specimen), lateral view. C. species C, head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8202887" httpUri="https://zenodo.org/record/8202887/files/figure.png" pageId="13" pageNumber="314">Fig. 19A</figureCitation>
|
||
).................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFD1CFA0E75E3" blockId="13.[151,1437,151,1288]" box="[151,1436,679,701]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFD1CFA0E75E3" box="[151,1436,679,701]" pageId="13" pageNumber="314">- Paravertical setulae present............................................................................ 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFD56FA0F746D" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFD56FA0F75A6" blockId="13.[151,1437,151,1288]" box="[151,1437,738,760]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFD56FA0F75A6" box="[151,1437,738,760]" pageId="13" pageNumber="314">
|
||
10 Entirely yellow species........................................................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3331FD56FB7975A6" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[1144,1259,738,760]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="bonairensis" status="sp. nov.">
|
||
<emphasis id="B958305DFFEEFF9C3331FD56FB7975A6" bold="true" box="[1144,1259,738,760]" italics="true" pageId="13" pageNumber="314">bonairensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEEFF9C33B9FD57FAA975A6" box="[1264,1339,739,760]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry id="F33BACDFFFEEFF9C320EFD56FA0475A6" box="[1351,1430,738,760]" name="Bonaire, Sint Eustatius and Saba" pageId="13" pageNumber="314">Bonaire</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFCB4FA0F746D" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFCB4FA0F746D" pageId="13" pageNumber="314">
|
||
- Brown transverse bands present on abdominal tergites 3 and 4; lateral spots present on abdominal tergites 2, 5, 6 (
|
||
<figureCitation id="1317F0CAFFEEFF9C326FFD4BFAEF744B" box="[1318,1405,767,789]" captionStart="FIGURE 20" captionStartId="27.[151,250,1961,1985]" captionTargetBox="[425,1162,417,1938]" captionTargetId="figure-70@27.[425,1162,417,1938]" captionTargetPageId="27" captionText="FIGURE 20. Tethysimyia spp. female specimens from Bonaire, A. species A. B. species B. C. species C." figureDoi="http://doi.org/10.5281/zenodo.8202889" httpUri="https://zenodo.org/record/8202889/files/figure.png" pageId="13" pageNumber="314">Fig. 20A</figureCitation>
|
||
)........................................................................... undescribed species A (
|
||
<collectingCountry id="F33BACDFFFEEFF9C320EFCA9FA04746D" box="[1351,1430,797,819]" name="Bonaire, Sint Eustatius and Saba" pageId="13" pageNumber="314">Bonaire</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFCECFA0E749B" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFCECFA0F74F6" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFCECFA0F74F6" pageId="13" pageNumber="314">
|
||
11 3 reclinate fronto-orbital setae; tibia and tarsus of anterior and middle leg pale brown; abdominal tergites with narrow dark bands on segments 2–6, and segments 3–5 with dark spot or a widening at midpoint of band (
|
||
<figureCitation id="1317F0CAFFEEFF9C332BFCC1FB2A74D4" box="[1122,1208,885,907]" captionStart="FIGURE 20" captionStartId="27.[151,250,1961,1985]" captionTargetBox="[425,1162,417,1938]" captionTargetId="figure-70@27.[425,1162,417,1938]" captionTargetPageId="27" captionText="FIGURE 20. Tethysimyia spp. female specimens from Bonaire, A. species A. B. species B. C. species C." figureDoi="http://doi.org/10.5281/zenodo.8202889" httpUri="https://zenodo.org/record/8202889/files/figure.png" pageId="13" pageNumber="314">Fig. 20C</figureCitation>
|
||
)........................................................................................... undescribed species C (
|
||
<collectingCountry id="F33BACDFFFEEFF9C320EFC26FA0474F6" box="[1351,1430,914,936]" name="Bonaire, Sint Eustatius and Saba" pageId="13" pageNumber="314">Bonaire</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFC04FA0E749B" blockId="13.[151,1437,151,1288]" box="[151,1436,943,965]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFC04FA0E749B" box="[151,1436,943,965]" pageId="13" pageNumber="314">- 2 reclinate fronto-orbital setae; legs all yellow; abdominal tergite with different pattern if bands are present............ 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFC5EFA0E732B" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFC5EFB017309" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFC5EFB017309" pageId="13" pageNumber="314">
|
||
12 Mediotergite yellow; anterior declivity of scutum yellow; abdomen with transverse bands on tergites 1–5, if present, at most pale brown; tergite 6 with pair of small pale brown spots; eyes bare....................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3314FBB3FB547343" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[1117,1222,1031,1053]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="nigrifacies" status="sp. nov.">
|
||
<emphasis id="B958305DFFEEFF9C3314FBB3FB547343" bold="true" box="[1117,1222,1031,1053]" italics="true" pageId="13" pageNumber="314">nigrifacies</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEEFF9C3385FBBCFA857343" box="[1228,1303,1032,1053]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(West Indies) (
|
||
<taxonomicName id="4C2C97CCFFEEFF9C37A3FB91FE057365" authority="Hardy" authorityName="Hardy" box="[234,407,1061,1083]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="macalpinei">
|
||
<emphasis id="B958305DFFEEFF9C37A3FB91FEC77364" box="[234,341,1061,1082]" italics="true" pageId="13" pageNumber="314">macalpinei</emphasis>
|
||
Hardy
|
||
</taxonomicName>
|
||
(Hawaii),
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3549FB91FD177365" authority="Hardy" authorityName="Hardy" box="[512,645,1061,1083]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="minuta">
|
||
<emphasis id="B958305DFFEEFF9C3549FB91FDD17364" box="[512,579,1061,1082]" italics="true" pageId="13" pageNumber="314">minuta</emphasis>
|
||
Hardy
|
||
</taxonomicName>
|
||
(Hawaii),
|
||
<taxonomicName id="4C2C97CCFFEEFF9C35A4FB91FC547365" authority="(Wheeler)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1994" box="[749,966,1061,1083]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="galamarilla">
|
||
<emphasis id="B958305DFFEEFF9C35A4FB91FCCC7364" box="[749,862,1061,1082]" italics="true" pageId="13" pageNumber="314">galamarilla</emphasis>
|
||
(Wheeler)
|
||
</taxonomicName>
|
||
(Galapagos),
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3306FB91FB657365" authority="(Wheeler)" authorityName="Ebejer" authorityYear="2023" baseAuthorityName="Wheeler" baseAuthorityYear="1994" box="[1103,1271,1061,1083]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rabida">
|
||
<emphasis id="B958305DFFEEFF9C3306FB91FB1D7364" box="[1103,1167,1061,1082]" italics="true" pageId="13" pageNumber="314">rabida</emphasis>
|
||
(Wheeler)
|
||
</taxonomicName>
|
||
(Galapagos) and undescribed species D (
|
||
<collectingCountry id="F33BACDFFFEEFF9C368CFBF6FDC47306" box="[453,598,1090,1112]" name="French Guiana" pageId="13" pageNumber="314">French Guiana</collectingCountry>
|
||
) are not reliably distinguished at present in the female sex.)
|
||
</keyLead>
|
||
</paragraph>
|
||
<subSubSection id="C336BFC4FFEEFF9C37DEFBD4FA0F7256" pageId="13" pageNumber="314" type="description">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFBD4FA0E732B" blockId="13.[151,1437,151,1288]" box="[151,1436,1119,1141]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFBD4FA0E732B" box="[151,1436,1119,1141]" pageId="13" pageNumber="314">- Mediotergite black; anterior declivity of scutum with dark patch............................................... 13</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30D8F4EAFFEEFF9C37DEFB2EFA0F7256" pageId="13" pageNumber="314">
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFB2EFA0E7393" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFB2EFA0E7393" pageId="13" pageNumber="314">
|
||
13 Abdomen with broad dark brown transverse bands on tergites 1–5; tergite 6 with pair of large brown spots (
|
||
<figureCitation id="1317F0CAFFEEFF9C33AAFB2EFAB573F1" box="[1251,1319,1178,1200]" captionStart="FIGURE 13" captionStartId="21.[151,250,1890,1914]" captionTargetBox="[189,1398,561,1866]" captionTargetId="figure-172@21.[189,1398,561,1866]" captionTargetPageId="21" captionText="FIGURE 13. Tethysimyia litophila sp. nov,. A. holotype, male. B. same head dorsal view. C. same, hypopygium in situ, posterior view. D. paratype, female, abdomen, posterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.8202875" httpUri="https://zenodo.org/record/8202875/files/figure.png" pageId="13" pageNumber="314">Fig. 13</figureCitation>
|
||
); eyes with microtrichia between every 1 or 2 ommatidia................
|
||
<taxonomicName id="4C2C97CCFFEEFF9C3429FB03FC217393" authority="Ebejer, 2023" authorityName="Ebejer" authorityYear="2023" box="[864,947,1207,1229]" class="Insecta" family="Chyromyidae" genus="Tethysimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="litophila" status="sp. nov.">
|
||
<emphasis id="B958305DFFEEFF9C3429FB03FC217393" bold="true" box="[864,947,1207,1229]" italics="true" pageId="13" pageNumber="314">litophila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B8D26FFEEFF9C34F0FB0CFB917393" box="[953,1027,1208,1229]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Pacific coasts of
|
||
<collectingCountry id="F33BACDFFFEEFF9C33FDFB0CFA8E7393" box="[1204,1308,1207,1229]" name="Costa Rica" pageId="13" pageNumber="314">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BACDFFFEEFF9C3202FB03FA047393" box="[1355,1430,1207,1229]" name="Mexico" pageId="13" pageNumber="314">Mexico</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93EC4FFFEEFF9C37DEFB61FA0F7256" blockId="13.[151,1437,151,1288]" pageId="13" pageNumber="314">
|
||
<keyLead id="30DD4F7AFFEEFF9C37DEFB61FA0F7256" pageId="13" pageNumber="314">
|
||
- Abdomen with narrow brown transverse bands on abdominal tergites 3–5; tergites 2 and 6 with small lateral spots (
|
||
<figureCitation id="1317F0CAFFEEFF9C3271FB61FA0373B4" box="[1336,1425,1237,1259]" captionStart="FIGURE 20" captionStartId="27.[151,250,1961,1985]" captionTargetBox="[425,1162,417,1938]" captionTargetId="figure-70@27.[425,1162,417,1938]" captionTargetPageId="27" captionText="FIGURE 20. Tethysimyia spp. female specimens from Bonaire, A. species A. B. species B. C. species C." figureDoi="http://doi.org/10.5281/zenodo.8202889" httpUri="https://zenodo.org/record/8202889/files/figure.png" pageId="13" pageNumber="314">Fig. 20B</figureCitation>
|
||
); eyes bare................................................................... undescribed species B (
|
||
<collectingCountry id="F33BACDFFFEEFF9C320EFB46FA047256" box="[1351,1430,1266,1288]" name="Bonaire, Sint Eustatius and Saba" pageId="13" pageNumber="314">Bonaire</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |