treatments-xml/data/54/42/4E/54424E035AC3DAF2C040C4D72C1C1DF4.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.769.25258" ID-GBIF-Dataset="7b27f7ff-2920-4834-b08c-754a4c0327c8" ID-PMC="PMC6030179" ID-Pensoft-Pub="1313-2970-769-13" ID-Pensoft-UUID="9C3DFFE3003056499148056FFF8CFFBF" ID-PubMed="29988743" ID-Zenodo-Dep="1304503" ID-ZooBank="C31B0F6BD3C242CDBAED8CE9D5769E8A" ModsDocID="1313-2970-769-13" checkinTime="1530090487889" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander &amp; Feher, Zoltan" docDate="2018" docId="54424E035AC3DAF2C040C4D72C1C1DF4" docLanguage="en" docName="ZooKeys 769: 13-48" docOrigin="ZooKeys 769" docPubDate="2018-06-26" docSource="http://dx.doi.org/10.3897/zookeys.769.25258" docTitle="Spelaeodiscus Brusina 1886" docType="treatment" docVersion="4" id="9C3DFFE3003056499148056FFF8CFFBF" lastPageNumber="13" masterDocId="9C3DFFE3003056499148056FFF8CFFBF" masterDocTitle="Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)" masterLastPageNumber="48" masterPageNumber="13" pageNumber="13" updateTime="1668165931740" updateUser="ExternalLinkService">
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<mods:title>Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deli, Tamas</mods:namePart>
<mods:affiliation>Moricz Zsigmond u. 2, Gyomaendrod, H- 5500, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eross, Zoltan Peter</mods:namePart>
<mods:affiliation>Bem u. 36., Budapest, H- 1151, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Peter L.</mods:namePart>
<mods:affiliation>Puechhaimg. 52, A- 3580, Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimg. 52, A- 3580, Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Feher, Zoltan</mods:namePart>
<mods:affiliation>Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H- 1088, Hungary</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2018</mods:date>
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<mods:number>2018-06-26</mods:number>
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<mods:detail type="volume">
<mods:number>769</mods:number>
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<mods:start>13</mods:start>
<mods:end>48</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.769.25258</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.769.25258</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1304503</mods:identifier>
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<treatment ID-GBIF-Taxon="182229754" LSID="urn:lsid:plazi:treatment:54424E035AC3DAF2C040C4D72C1C1DF4" httpUri="http://treatment.plazi.org/id/54424E035AC3DAF2C040C4D72C1C1DF4" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
Genus 
<taxonomicName authority="Brusina, 1886" authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">Spelaeodiscus Brusina, 1886</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="reference_group">
<paragraph pageId="0" pageNumber="13">
<taxonomicName authority="" class="Gastropoda" family="Spelaeodiscidae" genus="Patula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patula (Spelaeodiscus)" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subGenus" subGenus="Spelaeodiscus">Patula (Spelaeodiscus)</taxonomicName>
Brusina, 1886: 37.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type species">
<paragraph pageId="0" pageNumber="13">Type species.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis italics="true" pageId="0" pageNumber="13">
<taxonomicName class="Gastropoda" family="Spelaeodiscidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">Helix</taxonomicName>
Hauffeni
</emphasis>
Schmidt, 1855.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="distribution">
<paragraph pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="13">
The genus 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
has a disjunct distribution. One species (
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="hauffeni">
<emphasis italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
) is only known from Slovenia, whereas the rest of the genus is distributed in the vicinity of the Skadar Lake Basin (also known as 
<normalizedToken originalValue="Shkodër">Shkoder</normalizedToken>
Lake or Skutari Lake) in Montenegro and northern Albania (Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the genus Spelaeodiscus Brusina, 1886." figureDoi="10.3897/zookeys.769.25258.figure2" httpUri="https://binary.pensoft.net/fig/211836" pageId="0" pageNumber="13">2</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1304589" doi="10.3897/zookeys.769.25258.figure2" httpUri="https://binary.pensoft.net/fig/211836" pageId="0" pageNumber="13" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 2.</emphasis>
Distribution of the genus 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
Brusina, 1886.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="included taxa">
<paragraph pageId="0" pageNumber="13">Included taxa.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName authority="albanicus" authorityName="albanicus (A. J. Wagner" authorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus subsp. albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="albanicus" subSpecies="albanicus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus albanicus albanicus</emphasis>
</taxonomicName>
(A. J. Wagner, 1914), 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. albanicus subsp. edentatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="subSpecies" species="albanicus" subSpecies="edentatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. albanicus edentatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; P. L. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
, ssp. n., 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. dejongi" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="dejongi">
<emphasis italics="true" pageId="0" pageNumber="13">S. dejongi</emphasis>
</taxonomicName>
Gittenberger, 1969, 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. densecostatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="densecostatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. densecostatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; A. 
<normalizedToken originalValue="Reischütz">Reischuetz</normalizedToken>
, sp. n., 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="hauffeni">
<emphasis italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
(Schmidt, 1855) 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. hunyadii" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="hunyadii">
<emphasis italics="true" pageId="0" pageNumber="13">S. hunyadii</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Deli, sp. n., 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="latecostatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. latecostatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken originalValue="Erőss">Eross</normalizedToken>
, sp. n., 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. obodensis" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="obodensis">
<emphasis italics="true" pageId="0" pageNumber="13">S. obodensis</emphasis>
</taxonomicName>
Bole, 1965. 
<taxonomicName authority="unidentatus" family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. unidentatus subsp. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="subSpecies" species="unidentatus" subSpecies="unidentatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. unidentatus unidentatus</emphasis>
</taxonomicName>
Bole, 1961, 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. unidentatus subsp. acutus" order="Stylommatophora" pageId="0" pageNumber="13" rank="subSpecies" species="unidentatus" subSpecies="acutus">
<emphasis italics="true" pageId="0" pageNumber="13">S. unidentatus acutus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
, ssp. n., 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="virpazarioides">
<emphasis italics="true" pageId="0" pageNumber="13">S. virpazarioides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; 
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
, sp. n. For key traits see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Number of ribs on the body whorl, shell size, and key traits of Spelaeodiscus species." httpUri="http://table.plazi.org/id/AE7CCB10728A6F0F311DC97F022F808D" pageId="0" pageNumber="13" tableUuid="AE7CCB10728A6F0F311DC97F022F808D">1</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="AE7CCB10728A6F0F311DC97F022F808D" httpUri="http://table.plazi.org/id/AE7CCB10728A6F0F311DC97F022F808D" pageId="0" pageNumber="13" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Table 1.</emphasis>
Number of ribs on the body whorl, shell size, and key traits of 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
<table pageId="0" pageNumber="13">
<tr pageId="0" pageNumber="13">
<th colspan="1" pageId="0" pageNumber="13" rowspan="1">(Sub)species</th>
<th colspan="1" pageId="0" pageNumber="13" rowspan="1">No. of ribs</th>
<th colspan="1" pageId="0" pageNumber="13" rowspan="1">Shell diameter (in mm)</th>
<th colspan="1" pageId="0" pageNumber="13" rowspan="1">Key traits</th>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="13">
<taxonomicName baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="albanicus">Spelaeodiscus albanicus</taxonomicName>
<taxonomicName baseAuthorityName="A. J. Wagner" baseAuthorityYear="1914" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus albanicus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="albanicus">Spelaeodiscus albanicus</taxonomicName>
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">43-93</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">3.6-4.3</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">matte protoconch, weak palatal and two weak basal teeth</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus edentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="edentatus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus albanicus edentatus</emphasis>
</taxonomicName>
ssp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">35-54</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">3.6-4.2</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">widely-spaced ribs, glossy protoconch glossy, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus dejongi" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="dejongi">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus dejongi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">57-112</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">1.9-3.4</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">dense, low ribs, smooth protoconch, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; A. Reischuetz" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus densecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="densecostatus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus densecostatus</emphasis>
</taxonomicName>
sp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">116</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">very low and dense ribs, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName baseAuthorityName="Schmidt" baseAuthorityYear="1855" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hauffeni">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus hauffeni</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">41-52</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.8-3.5</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">widely spaced, strong ribs, rounded, toothless aperture, finely granular protoconch</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; Eross" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="latecostatus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus latecostatus</emphasis>
</taxonomicName>
sp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">42</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">strong, very widely spaced ribs, glossy protoconch, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; Eross" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus latecostatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="latecostatus">Spelaeodiscus cf. latecostatus</taxonomicName>
sp. n. (2017/005)
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">47-54</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">strong, widely spaced ribs, glossy protoconch, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; Deli" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus hunyadii" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="hunyadii">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus hunyadii</emphasis>
</taxonomicName>
sp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">42-48</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.1-2.2</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">widely spaced, strong ribs, glossy protoconch, strongly oblique, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Páll-Gergely &amp; Deli &amp; Erőss &amp; Reischütz &amp; Reischütz &amp; Fehér" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus obodensis" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="obodensis">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus obodensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">43-76</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.6-3.0</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">elevated spire, roughly sculptured protoconch, strong ribs, toothless aperture</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="13">
<taxonomicName authorityName="Bole" authorityYear="1961" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">Spelaeodiscus unidentatus</taxonomicName>
<taxonomicName authorityName="Bole" authorityYear="1961" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="unidentatus">Spelaeodiscus unidentatus</taxonomicName>
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">74-118</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.4-3.2</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">low basal tooth; palatal part of peristome with strong incision</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus unidentatus subsp. acutus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="unidentatus" subSpecies="acutus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus unidentatus acutus</emphasis>
</taxonomicName>
ssp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">64-91</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">2.9-3.5</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">pointed basal tooth; palatal part of peristome with shallow incision</td>
</tr>
<tr pageId="0" pageNumber="13">
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">
<taxonomicName authorityName="Pall-Gergely &amp; Feher" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus virpazarioides" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="species" species="virpazarioides">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus virpazarioides</emphasis>
</taxonomicName>
sp. n.
</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">40-70</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">3.3-3.6</td>
<td colspan="1" pageId="0" pageNumber="13" rowspan="1">spiral sculpture, thickened callus</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="delimitation of this genus">
<paragraph pageId="0" pageNumber="13">Delimitation of this genus.</paragraph>
<paragraph pageId="0" pageNumber="13">
The reproductive anatomy of 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
is characterized by a short penial caecum, a well-developed penial appendix, sometimes an epiphallic caecum, and a bursa copulatrix without a diverticulum. The retractor muscle is divided into two bounds, one inserting on the penial appendix, whereas the other at the base of the penial caecum. Examining the anatomical descriptions and drawings of 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
) and 
<taxonomicName authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
(
<bibRefCitation author="Hudec, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="13" pagination="157 - 165" refId="B19" refString="Hudec, V, 1965. Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna. Archiv fuer Molluskenkunde 94: 157 - 165" title="Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna." volume="94" year="1965">Hudec 1965</bibRefCitation>
, 
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="13" pagination="263 - 277" refId="B14" refString="Gittenberger, E, 1975. Beitraege zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277" title="Beitraege zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag." volume="48" year="1975">Gittenberger 1975</bibRefCitation>
, 
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="13" pagination="1 - 127" refId="B38" refString="Schileyko, AA, 1998. Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. Ruthenica, Supplement 2 (1): 1 - 127" title="Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae." volume="2" year="1998">Schileyko 1998</bibRefCitation>
, 
<bibRefCitation author="Subai, P" journalOrPublisher="Basteria" pageId="0" pageNumber="13" pagination="111 - 118" refId="B44" refString="Subai, P, Dedov, I, 2008. A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov. Basteria 72: 111 - 118" title="A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov." volume="72" year="2008">Subai and Dedov 2008</bibRefCitation>
), we were unable to find characters that would constantly differ between the two groups. For example, the penial caecum was long and slender in 
<taxonomicName lsidName="A. tatrica" pageId="0" pageNumber="13" rank="species" species="tatrica">
<emphasis italics="true" pageId="0" pageNumber="13">A. tatrica</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Hudec, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="13" pagination="157 - 165" refId="B19" refString="Hudec, V, 1965. Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna. Archiv fuer Molluskenkunde 94: 157 - 165" title="Neue Erkenntnisse ueber den Geschlechtsapparat von Argna bielzi (Rossmaessler) und Bemerkungen zur systematischen Stellung der Gattung Argna." volume="94" year="1965">Hudec 1965</bibRefCitation>
) and 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="hauffeni">
<emphasis italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
(see 
<bibRefCitation DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
), but was short and conical in 
<taxonomicName lsidName="A. triaria" pageId="0" pageNumber="13" rank="species" species="triaria">
<emphasis italics="true" pageId="0" pageNumber="13">A. triaria</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Subai, P" journalOrPublisher="Basteria" pageId="0" pageNumber="13" pagination="111 - 118" refId="B44" refString="Subai, P, Dedov, I, 2008. A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov. Basteria 72: 111 - 118" title="A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov." volume="72" year="2008">Subai and Dedov 2008</bibRefCitation>
) and 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="unidentatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. unidentatus</emphasis>
</taxonomicName>
(see 
<bibRefCitation DOI="https://doi.org/10.5038/1827-806X.1.3.9" author="Bole, J" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="13" pagination="349 - 356" refId="B5" refString="Bole, J, 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356, DOI: https://doi.org/10.5038/1827-806X.1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien." url="https://doi.org/10.5038/1827-806X.1.3.9" volume="1" year="1965">Bole 1965</bibRefCitation>
). Also, the shape of the bursa and the position of the starting point of the penial appendix was greatly variable across genera. Clear epiphallic caecum was only found in 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. hauffeni" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="hauffeni">
<emphasis italics="true" pageId="0" pageNumber="13">S. hauffeni</emphasis>
</taxonomicName>
, but some thickening was visible in 
<taxonomicName family="Spelaeodiscidae" kingdom="Animalia" lsidName="S. unidentatus" order="Stylommatophora" pageId="0" pageNumber="13" rank="species" species="unidentatus">
<emphasis italics="true" pageId="0" pageNumber="13">S. unidentatus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. triaria" pageId="0" pageNumber="13" rank="species" species="triaria">
<emphasis italics="true" pageId="0" pageNumber="13">A. triaria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="13">
As for shell characters, 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
is characterized by a mostly colourless shell that is smaller than 4.3 mm (majority of species are even smaller than 3.5 mm), the spire is relatively low (height of body whorl at least two third of the height of the entire shell), the body whorl is evenly rounded, the edge of the parietal callus is straight, and the peristome is only slightly expanded. In contrast, 
<taxonomicName authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
shells are brownish, larger than 4.3 mm, they have higher spire (height of body whorl is approximately half of the height of the entire shell), the shell is shape reverse trapezoid from standard apertural view, the callus is heart-shaped, and the basal part of the peristome is strongly expanded.
</paragraph>
<paragraph pageId="0" pageNumber="13">
The habitat was the only 
<normalizedToken originalValue="“trait”">&quot;trait&quot;</normalizedToken>
mentioned by 
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="13" pagination="287 - 306" refId="B13" refString="Gittenberger, E, 1969. Beitraege zur Kenntnis der Pupillacea I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306" title="Beitraege zur Kenntnis der Pupillacea I. Die Spelaeodiscinae." volume="43" year="1969">Gittenberger (1969)</bibRefCitation>
as difference between the two groups. Namely, 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
is subterranean, whereas 
<taxonomicName authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
can be found on rock surfaces and among leaf litter at the base of limestone rocks. In the lack of sound molecular data it is difficult to infer their relationship, but based only on ecological, conchological, and biogeographical differences it seems reasonable to keep 
<taxonomicName authorityName="Westerlund" authorityYear="1889" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspasita" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Aspasita</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Brusina" authorityYear="1886" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" kingdom="Animalia" lsidName="Spelaeodiscus" order="Stylommatophora" pageId="0" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Spelaeodiscus</emphasis>
</taxonomicName>
as distinct genera.
</paragraph>
</subSubSection>
</treatment>
</document>