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<document id="0A6E92961D325DCF2FBA99C5CAC31306" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario &amp; Michat, Mariano C." docDate="2015" docId="03EE87FEFFE5FFE391A6FB54FB71A6DE" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus bolivari Toledo &amp; Michat, 2015, sp. n." docType="treatment" docVersion="9" lastPageNumber="324" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="323" updateTime="1698611165842" updateUser="plazi">
<mods:mods id="B51AD46B2D0C7EAC3DA96C8B80DEE399" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="761EB6F1DB2B52F5D6B88B785A2DB54D">
<mods:title id="616216E4E53955A95D9BD6018CDD8B8D">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
</mods:titleInfo>
<mods:name id="513E8626F8419B9F3FC62F740042D9DF" type="personal">
<mods:role id="EC3B5B959156089E4438CE97ACB2644C">
<mods:roleTerm id="BF2310E83C5292B9D3A60FCBCDB0142B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5947772EE987F257DC8CFA7FF52554FD">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="6C2CFCD478C1B9B5FCEA40EE875AAD22" type="personal">
<mods:role id="2AFEFA4E64EFB2B04ABB9F7288EC2E75">
<mods:roleTerm id="897CBCD3B14E8E266B6F26E7F0FF0B24">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8542D0B6D5F26455A9F12BE3E35B1361">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="579CDF5D6F0341E262064AC34F3920A6">text</mods:typeOfResource>
<mods:relatedItem id="49554EDBDF0E018B67072ED670EF7BB6" type="host">
<mods:titleInfo id="E514B3A0EA39FEA7035091199DC61AA0">
<mods:title id="70520EDF8D46C0F0CCD83159929D2C5B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E5F7C687F8A99E44D2D362B1372CC7C3">
<mods:date id="29143DF1A522734D975222A36907BC21">2015</mods:date>
<mods:detail id="C968F8AFEC9B004BDF40985BEAB5716C" type="volume">
<mods:number id="7E8E3B0D6F442B63A23347D3047C7EFD">3990</mods:number>
</mods:detail>
<mods:detail id="720C2E39D221663B4AEC3ADFBDB41F62" type="issue">
<mods:number id="8B9FA821118EB8BC087FE05DD6C6F083">3</mods:number>
</mods:detail>
<mods:extent id="3677D97CB7677F3E27806816D5247F51" unit="page">
<mods:start id="814F4E92C1DEBE8849A8F00A83DCF6A5">301</mods:start>
<mods:end id="0E83E1A6ED1F0C5BD9D8EE2544A0E910">354</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="048F78315F5F6F744968DFAF6E5995BB">journal article</mods:classification>
<mods:identifier id="12A9A6396BF246397B1F6DE1BFA338A4" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
<mods:identifier id="A85A9D853F6D378A3F56E57B9CCCEEC7" type="GBIF-Dataset">cf747765-b9bf-42ed-8479-35355e712db0</mods:identifier>
<mods:identifier id="B240206CCC156D5CEC14154C6A29770E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="895F8A843B0ADD19EB2765EFD4E4A108" type="Zenodo-Dep">244301</mods:identifier>
<mods:identifier id="45BA63767BEB2DF08F8B434645998AD7" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
</mods:mods>
<treatment id="03EE87FEFFE5FFE391A6FB54FB71A6DE" ID-DOI="http://doi.org/10.5281/zenodo.5692060" ID-GBIF-Taxon="119638443" ID-Zenodo-Dep="5692060" LSID="urn:lsid:plazi:treatment:03EE87FEFFE5FFE391A6FB54FB71A6DE" httpUri="http://treatment.plazi.org/id/03EE87FEFFE5FFE391A6FB54FB71A6DE" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
<subSubSection id="C35D6563FFE5FFE291A6FB54FCDCA51B" pageId="22" pageNumber="323" type="nomenclature">
<paragraph id="8BF836E8FFE5FFE291A6FB54FE27A53D" blockId="22.[151,804,1257,1317]" box="[151,479,1257,1283]" pageId="22" pageNumber="323">
<heading id="D0B08184FFE5FFE291A6FB54FE27A53D" bold="true" box="[151,479,1257,1283]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
<emphasis id="B933EAFAFFE5FFE291A6FB54FE27A53D" bold="true" box="[151,479,1257,1283]" pageId="22" pageNumber="323">
<taxonomicName id="4C474D6BFFE5FFE291A6FB54FE6EA53D" ID-CoL="8LXJV" box="[151,406,1257,1283]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="bolivari" status="sp. nov.">
<emphasis id="B933EAFAFFE5FFE291A6FB54FE6EA53D" bold="true" box="[151,406,1257,1283]" italics="true" pageId="22" pageNumber="323">Laccomimus bolivari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005781FFE5FFE290ACFB54FE27A53D" box="[413,479,1257,1283]" pageId="22" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836E8FFE5FFE291A6FAB1FCDCA51B" blockId="22.[151,804,1257,1317]" box="[151,804,1292,1317]" pageId="22" pageNumber="323">
(
<figureCitation id="137C2A6DFFE5FFE291AEFAB1FF1EA51A" box="[159,230,1292,1317]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="22" pageNumber="323">Figs 1</figureCitation>
; 13; 41; 58; 74; 105–106; 115; 127; 133–134; 136)
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE5FFE391A6FAE9FA6BA122" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323" type="materials_examined">
<paragraph id="8BF836E8FFE5FFE291A6FAE9FB11A552" blockId="22.[151,1437,1364,2037]" box="[151,1257,1364,1389]" pageId="22" pageNumber="323">
<emphasis id="B933EAFAFFE5FFE291A6FAE9FECDA553" bold="true" box="[151,309,1364,1389]" pageId="22" pageNumber="323">
<typeStatus id="54FC884AFFE5FFE291A6FAE9FF2AA553" box="[151,210,1364,1389]" pageId="22" pageNumber="323">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3507678FFE5FFE2900DFAE9FE4AA553" box="[316,434,1364,1389]" name="Venezuela" pageId="22" pageNumber="323">Venezuela</collectingCountry>
, Bolivar State, Los Pijiguaos,
<geoCoordinate id="EE73502FFFE5FFE29234FAE9FC7FA553" box="[773,903,1364,1389]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="6.5936165">6°35.617'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE292A3FAE9FBDEA553" box="[914,1062,1364,1389]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-66.82063">66°49.238'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29500FAE9FB92A552" box="[1073,1130,1364,1388]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="323" unit="m" value="80.0">80 m</quantity>
(Fig. 134).
</paragraph>
<paragraph id="8BF836E8FFE5FFE291F6FACAFE47A61E" blockId="22.[151,1437,1364,2037]" pageId="22" pageNumber="323">
<emphasis id="B933EAFAFFE5FFE291F6FACAFE82A5AE" bold="true" box="[199,378,1399,1424]" pageId="22" pageNumber="323">
<typeStatus id="54FC884AFFE5FFE291F6FACAFEFAA5AE" box="[199,258,1399,1424]" pageId="22" pageNumber="323">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54FC884AFFE5FFE290B7FACAFE16A5AE" box="[390,494,1399,1424]" pageId="22" pageNumber="323" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED56AE2DFFE5FFE29374FAC5FD68A5AE" LSID="urn:lsid:biocol.org:col:33834" box="[581,656,1400,1424]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="22" pageNumber="323">MIZA</collectionCode>
) dissected by us; genitalia glued on the same rectangular card as
<typeStatus id="54FC884AFFE5FFE291A6FA21FF01A58B" box="[151,249,1436,1461]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
“
<collectingCountry id="F3507678FFE5FFE2903DFA21FE7CA58B" box="[268,388,1436,1461]" name="Venezuela" pageId="22" pageNumber="323">Venezuela</collectingCountry>
, Bolivar State,
<geoCoordinate id="EE73502FFFE5FFE29304FA21FD4FA58B" box="[565,695,1436,1461]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="6.5936165">6°35.617'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE293F5FA21FCA0A58B" box="[708,856,1436,1461]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-66.82063">66°49.238'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29254FA21FC59A58A" box="[869,929,1436,1460]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="323" unit="m" value="80.0">80 m</quantity>
, Los Pijiguaos, outcrop/morichal,
<date id="FFF91028FFE5FFE2941BFA21FA60A58A" box="[1322,1432,1436,1460]" pageId="22" pageNumber="323" value="2009-01-12">12.I.2009</date>
, leg. Short, Garcia, Camacho, Miller &amp; Joly, VZ09-0112-01A, morichal margin” [printed], “
<collectionCode id="ED56AE2DFFE5FFE295C1FA7DFA64A5E6" box="[1264,1436,1472,1496]" pageId="22" pageNumber="323">KUNHM-ENT</collectionCode>
<collectionCode id="ED56AE2DFFE5FFE291A6FA59FF2AA5C3" box="[151,210,1508,1533]" httpUri="http://grbio.org/cool/dhkd-089i" name="Sedgwick Museum" pageId="22" pageNumber="323">SMC</collectionCode>
0856789” [printed, together with barcode], “
<typeStatus id="54FC884AFFE5FFE293D1FA59FC8FA5C2" box="[736,887,1508,1532]" pageId="22" pageNumber="323" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C474D6BFFE5FFE292B2FA58FB8DA5C2" box="[899,1141,1508,1532]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFE5FFE292B2FA58FB8DA5C2" box="[899,1141,1508,1532]" italics="true" pageId="22" pageNumber="323">Laccomimus bolivari</emphasis>
</taxonomicName>
Toledo &amp; Michat” [red, rectangular, printed label].
</paragraph>
<paragraph id="8BF836E8FFE5FFE291F6F990FA65A7CA" blockId="22.[151,1437,1364,2037]" pageId="22" pageNumber="323">
<typeStatus id="54FC884AFFE5FFE291F6F990FECCA67B" box="[199,308,1581,1605]" pageId="22" pageNumber="323" type="paratype">Paratypes</typeStatus>
(87 exs, all bearing red, printed label “
<typeStatus id="54FC884AFFE5FFE29226F991FC5DA67A" box="[791,933,1580,1604]" pageId="22" pageNumber="323" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C474D6BFFE5FFE29283F990FB5DA67A" box="[946,1189,1580,1604]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFE5FFE29283F990FB5DA67A" box="[946,1189,1580,1604]" italics="true" pageId="22" pageNumber="323">Laccomimus bolivari</emphasis>
</taxonomicName>
Toledo &amp; Michat”):
<emphasis id="B933EAFAFFE5FFE291A6F9F2FEE0A656" bold="true" box="[151,280,1615,1640]" pageId="22" pageNumber="323">
<collectingCountry id="F3507678FFE5FFE291A6F9F2FEECA656" box="[151,276,1615,1640]" name="Venezuela" pageId="22" pageNumber="323">Venezuela</collectingCountry>
:
</emphasis>
same data as
<typeStatus id="54FC884AFFE5FFE290E5F9F2FDCEA656" box="[468,566,1615,1640]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
(4 exs
<collectionCode id="ED56AE2DFFE5FFE29391F9EDFD1BA656" box="[672,739,1616,1640]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “Bolivar State,
<geoCoordinate id="EE73502FFFE5FFE2928AF9F2FBB0A656" box="[955,1096,1615,1640]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="5.847">5°50'49.2&quot;N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE2956DF9F2FB16A656" box="[1116,1262,1615,1640]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-61.467335">61°28'2.4&quot;W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29430F9EDFA98A659" box="[1281,1376,1616,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="22" pageNumber="323" unit="m" value="1340.0">1340 m</quantity>
, Rio Aponowao at Hwy 10,
<date id="FFF91028FFE5FFE29094F9C9FDC8A6B2" box="[421,560,1652,1676]" pageId="22" pageNumber="323" value="2008-07-31">
31.
<collectionCode id="ED56AE2DFFE5FFE290F5F9C9FE17A6B2" box="[452,495,1652,1676]" pageId="22" pageNumber="323">VII</collectionCode>
.2008
</date>
, leg. A. Short, M. Garcia, AS-08-060a, small vegetated pool” (1 ex.
<collectionCode id="ED56AE2DFFE5FFE2947FF9C9FA69A6B2" box="[1358,1425,1652,1676]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “
<taxonomicName id="4C474D6BFFE5FFE29190F925FEE2A68E" box="[161,282,1688,1712]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
State,
<geoCoordinate id="EE73502FFFE5FFE2905CF92AFE0DA68E" box="[365,501,1687,1712]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="5.3419">5°20.514&quot;N</geoCoordinate>
, 67°.45,315&quot;W,
<quantity id="4CBF9B0DFFE5FFE29385F925FD0AA691" box="[692,754,1688,1712]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="22" pageNumber="323" unit="m" value="87.0">87 m</quantity>
, S- Communidad Porvenir,
<date id="FFF91028FFE5FFE29509F92AFB5EA68E" box="[1080,1190,1687,1712]" pageId="22" pageNumber="323" value="2009-01-15">15.I.2009</date>
, leg. Miller &amp; Short, VZ09-0115-03B: small streamlet” (19 exs
<collectionCode id="ED56AE2DFFE5FFE29349F901FD4EA6EA" box="[632,694,1724,1748]" pageId="22" pageNumber="323">VAIP</collectionCode>
,
<collectionCode id="ED56AE2DFFE5FFE293F0F901FCF8A6EA" box="[705,768,1724,1748]" pageId="22" pageNumber="323">MTP</collectionCode>
); “
<taxonomicName id="4C474D6BFFE5FFE2922CF901FC6EA6EA" box="[797,918,1724,1748]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
State,
<geoCoordinate id="EE73502FFFE5FFE292D2F901FB9EA6EB" box="[995,1126,1724,1749]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="5.3413167">5°20.479'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE29543F901FAFDA6EB" box="[1138,1285,1724,1749]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-67.75597">67°45.358'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29420F901FAB4A6EA" box="[1297,1356,1724,1748]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="22" pageNumber="323" unit="m" value="72.0">72 m</quantity>
, ca.
<quantity id="4CBF9B0DFFE5FFE294B1F901FF43A6C6" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="323" unit="km" value="12.0">12 km</quantity>
N Samariapo, small stream at rd x-ing,
<date id="FFF91028FFE5FFE293A7F962FD0DA6C6" box="[662,757,1759,1784]" pageId="22" pageNumber="323" value="2006-01-06">6.I.2006</date>
, AS-06-013, leg. A.E.Z. Short” (2 exs
<collectionCode id="ED56AE2DFFE5FFE295F1F95DFAFBA6C6" box="[1216,1283,1760,1784]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “
<taxonomicName id="4C474D6BFFE5FFE29412F95DFA64A6C6" box="[1315,1436,1760,1784]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
State,
<geoCoordinate id="EE73502FFFE5FFE291ECF8B9FEA7A723" box="[221,351,1796,1821]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="5.2773166">5°16.639'N</geoCoordinate>
, 67°.48.044'W,
<quantity id="4CBF9B0DFFE5FFE29320F8B9FDB5A722" box="[529,589,1796,1821]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="323" unit="m" value="60.0">60 m</quantity>
, ca.
<quantity id="4CBF9B0DFFE5FFE293B1F8B9FD41A723" box="[640,697,1796,1821]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="323" unit="km" value="4.0">4 km</quantity>
N Samariapo, small stream at rd x-ing,
<date id="FFF91028FFE5FFE2954CF8B9FB24A723" box="[1149,1244,1796,1821]" pageId="22" pageNumber="323" value="2006-01-06">6.I.2006</date>
, AS-06-012, leg. A.E.Z. Short” (2 exs
<collectionCode id="ED56AE2DFFE5FFE290ABF895FE25A77E" box="[410,477,1832,1856]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “
<taxonomicName id="4C474D6BFFE5FFE290CEF895FD80A77E" box="[511,632,1832,1856]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="354" phylum="Arthropoda" rank="species" species="amazonas">Amazonas</taxonomicName>
State,
<geoCoordinate id="EE73502FFFE5FFE293FEF89AFCA9A77E" box="[719,849,1831,1856]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="5.8069">5°48.414'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE29253F89AFC0DA77E" box="[866,1013,1831,1856]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-67.43855">67°26.313'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29537F895FBBFA701" box="[1030,1095,1832,1856]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="323" unit="m" value="80.0">80 m</quantity>
, nr. Iboruwa: &quot;Tobogancito&quot;,
<date id="FFF91028FFE5FFE291A6F8F1FEFDA75A" box="[151,261,1868,1892]" pageId="22" pageNumber="323" value="2009-01-13">13.I.2009</date>
, leg. Short
<emphasis id="B933EAFAFFE5FFE290B9F8F0FE39A75A" box="[392,449,1868,1892]" italics="true" pageId="22" pageNumber="323">et al.</emphasis>
, VZ09-0113-
<date id="FFF91028FFE5FFE2936DF8F1FD75A75A" box="[604,653,1868,1892]" pageId="22" pageNumber="323" value="2002-10">02X</date>
” (1 ex.
<collectionCode id="ED56AE2DFFE5FFE293D8F8F1FCD4A75A" box="[745,812,1868,1892]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “Guarico State,
<geoCoordinate id="EE73502FFFE5FFE292C7F8F1FB80A75B" box="[1014,1144,1868,1893]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="8.528417">8°31.705'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE295B4F8F1FAE1A75B" box="[1157,1305,1868,1893]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-66.3767">66°22.602'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE29414F8F1FA97A75A" box="[1317,1391,1868,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="22" pageNumber="323" unit="m" value="145.0">145 m</quantity>
, ca.
<quantity id="4CBF9B0DFFE5FFE291A6F8D2FF26A7B6" box="[151,222,1903,1928]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="22" pageNumber="323" unit="km" value="65.0">65 km</quantity>
S Las Mercedes,
<date id="FFF91028FFE5FFE29097F8CDFDFEA7B6" box="[422,518,1904,1928]" pageId="22" pageNumber="323" value="2009-01-09">9.I.2009</date>
, leg Short, Miller, Garcia &amp; Camarcho, large lagoon w/veg., VZ09-0109-
<date id="FFF91028FFE5FFE29472F8CDFA8BA7B6" box="[1347,1395,1904,1928]" pageId="22" pageNumber="323" value="2001-10">01X</date>
” (2 exs
<collectionCode id="ED56AE2DFFE5FFE291F2F829FEFEA792" box="[195,262,1940,1964]" pageId="22" pageNumber="323">VAIP</collectionCode>
); “Guarico State,
<geoCoordinate id="EE73502FFFE5FFE290FAF829FDB4A793" box="[459,588,1940,1965]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="1" value="9.772">9°46.320'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE5FFE29369F829FD14A793" box="[600,748,1940,1965]" direction="west" orientation="longitude" pageId="22" pageNumber="323" precision="1" value="-67.35295">67°21.177'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE5FFE293C6F829FCC7A792" box="[759,831,1940,1964]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="22" pageNumber="323" unit="m" value="280.0">280 m</quantity>
, Rio San Antonio, N Dos Caminos,
<date id="FFF91028FFE5FFE295E5F829FACBA792" box="[1236,1331,1940,1964]" pageId="22" pageNumber="323" value="2009-01-08">8.I.2009</date>
, leg K.B. Miller, mic. Habitats, VZ09-0108-02C” (1 ex.
<collectionCode id="ED56AE2DFFE5FFE29384F805FD0DA7EE" box="[693,757,1976,2000]" pageId="22" pageNumber="323">VAIP</collectionCode>
) [each specimen in
<collectionCode id="ED56AE2DFFE5FFE292D0F804FBE7A7EF" box="[993,1055,1977,2001]" pageId="22" pageNumber="323">VAIP</collectionCode>
bearing second label (
<collectionCode id="ED56AE2DFFE5FFE29416F805FA64A7EE" box="[1319,1436,1976,2000]" pageId="22" pageNumber="323">KUNHM-</collectionCode>
<collectionCode id="ED56AE2DFFE5FFE291A6F861FF37A7CA" LSID="urn:lsid:biocol.org:col:13589" box="[151,207,2012,2036]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="22" pageNumber="323">ENT</collectionCode>
) with its own barcode and catalogue number]; “Guarico,
<quantity id="4CBF9B0DFFE5FFE292A3F861FC18A7CB" box="[914,992,2012,2037]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="323" unit="km" value="12.0">12 km</quantity>
S Calabozo,
<date id="FFF91028FFE5FFE295BAF861FAC9A7CB" box="[1163,1329,2012,2037]" pageId="22" pageNumber="323" value="1969-12-11" valueMax="1969-12-06" valueMin="1969-12-11">11–6.XII.1969</date>
, P. &amp; P.
</paragraph>
<paragraph id="8BF836E8FFE4FFE391A6FF2AFA6BA122" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
Spangler”, “Est. Biologica Los Llanos, black light” (3 exs
<collectionCode id="ED56AE2DFFE4FFE39203FF25FC70A08E" LSID="urn:lsid:biocol.org:col:34871" box="[818,904,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="324">USNM</collectionCode>
).
<emphasis id="B933EAFAFFE4FFE392AAFF2AFBE2A08E" bold="true" box="[923,1050,151,176]" pageId="23" pageNumber="324">
<collectingCountry id="F3507678FFE4FFE392AAFF2AFBECA08E" box="[923,1044,151,176]" name="Suriname" pageId="23" pageNumber="324">Suriname</collectingCountry>
:
</emphasis>
“Brokopondo,
<geoCoordinate id="EE73502FFFE4FFE395F8FF2AFAD4A08E" box="[1225,1324,151,176]" direction="west" orientation="longitude" pageId="23" pageNumber="324" precision="925" value="-55.166668">55°10'W</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE4FFE39409FF2AFA6FA08E" box="[1336,1431,151,176]" direction="north" orientation="latitude" pageId="23" pageNumber="324" precision="925" value="4.9833336">04°59'N</geoCoordinate>
, rd. Brownsberg–Brownsweg,
<date id="FFF91028FFE4FFE390D6FF01FD98A0EB" box="[487,608,188,213]" pageId="23" pageNumber="324" value="1969-10-24">24.X.1969</date>
, leg. N. Nieser (SN 201A)” (1 ex.
<collectionCode id="ED56AE2DFFE4FFE392D6FF01FBCCA0EA" box="[999,1076,188,212]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="324">NMW</collectionCode>
).
<emphasis id="B933EAFAFFE4FFE39577FF01FB41A0EB" bold="true" box="[1094,1209,188,213]" pageId="23" pageNumber="324">
<collectingCountry id="F3507678FFE4FFE39577FF01FB4CA0EB" box="[1094,1204,188,213]" name="Trinidad and Tobago" pageId="23" pageNumber="324">Trinidad</collectingCountry>
:
</emphasis>
“Cumuto,
<collectingCountry id="F3507678FFE4FFE39404FF01FA6FA0EB" box="[1333,1431,188,213]" name="Trinidad and Tobago" pageId="23" pageNumber="324">Trinidad</collectingCountry>
, 1929, Darlington”; all printed label, except for one specimen, handwritten, presumably by Young; two specimens with additional label “
<taxonomicName id="4C474D6BFFE4FFE390A1FEB8FD4FA122" box="[400,695,260,284]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="trinidadensis" status="sp. nov.">
<emphasis id="B933EAFAFFE4FFE390A1FEB8FD4FA122" box="[400,695,260,284]" italics="true" pageId="23" pageNumber="324">Laccomimus trinidadensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005781FFE4FFE3938FFEB8FCF6A122" box="[702,782,261,284]" pageId="23" pageNumber="324" rank="species">sp.nov.</taxonomicNameLabel>
det. F.N. Young” handwritten by Young (51 exs
<collectionCode id="ED56AE2DFFE4FFE39400FEB9FA7FA122" box="[1329,1415,260,284]" pageId="23" pageNumber="324">NMHS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFE391F6FE9AFEE1A193" pageId="23" pageNumber="324" type="diagnosis">
<paragraph id="8BF836E8FFE4FFE391F6FE9AFEE1A193" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FE9AFEB9A17E" bold="true" box="[199,321,295,320]" pageId="23" pageNumber="324">Diagnosis.</emphasis>
Externally hard to tell apart from other members of
<taxonomicName id="4C474D6BFFE4FFE392A9FE95FBDDA101" box="[920,1061,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFE4FFE392A9FE95FBDDA101" box="[920,1061,296,319]" italics="true" pageId="23" pageNumber="324">Laccomimus</emphasis>
</taxonomicName>
. Aedeagal characters distinctive, median lobe with angle between base and distal portion almost indistinct, apex ending in wide and flattened expansion (
<figureCitation id="137C2A6DFFE4FFE39026FED2FE84A1B6" box="[279,380,367,392]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="23" pageNumber="324">Figs 105</figureCitation>
a; 106a); left paramere straight (
<figureCitation id="137C2A6DFFE4FFE393D3FED2FCBBA1B6" box="[738,835,367,392]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="23" pageNumber="324">Fig. 105</figureCitation>
c). Females impossible to identify without association with males.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFE391F6FE0AFC69A4D6" pageId="23" pageNumber="324" type="description">
<paragraph id="8BF836E8FFE4FFE391F6FE0AFC73A229" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FE0AFEA1A1EE" bold="true" box="[199,345,439,464]" pageId="23" pageNumber="324">Description.</emphasis>
<emphasis id="B933EAFAFFE4FFE39051FE05FE41A1EE" box="[352,441,440,464]" italics="true" pageId="23" pageNumber="324">Habitus</emphasis>
(
<figureCitation id="137C2A6DFFE4FFE390F9FE0AFDF2A1EE" box="[456,522,439,464]" captionStart="FIGURES 1 – 7. 1" captionStartId="29.[151,264,1905,1927]" captionTargetBox="[169,1412,219,1874]" captionTargetId="figure@29.[169,1413,193,1876]" captionTargetPageId="29" captionText="FIGURES 1 – 7. 1) Habitus of Laccomimus bolivari, dorsal view (Photo A. E. Z. Short); 2) habitus of Laccodytes apalodes, dorsal view (Photo A. E. Z. Short); 3 – 7) SEMs of Laccomimus species (females): 3) habitus of L. alvarengi, dorsal view; 4) habitus of L. bordoni, dorsal view; 5) pronotum of L. alvarengi, dorsal view; 6) pronotum of L. bordoni, dorsal view; 7) pronotum of L. bordoni, dorsolateral view." httpUri="https://zenodo.org/record/244302/files/figure.png" pageId="23" pageNumber="324">Fig. 1</figureCitation>
). TL: 2.0–
<quantity id="4CBF9B0DFFE4FFE393B7FE05FD18A1F1" box="[646,736,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="324" unit="mm" value="2.4">2.4 mm</quantity>
; MW: 1.0–
<quantity id="4CBF9B0DFFE4FFE3926EFE05FC4DA1F1" box="[863,949,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="324" unit="mm" value="1.2">1.2 mm</quantity>
(
<typeStatus id="54FC884AFFE4FFE392F5FE0AFBD4A1EE" box="[964,1068,439,464]" pageId="23" pageNumber="324" type="holotype">holotype</typeStatus>
: TL:
<quantity id="4CBF9B0DFFE4FFE39559FE05FB3AA1F1" box="[1128,1218,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="324" unit="mm" value="2.3">2.3 mm</quantity>
; MW:
<quantity id="4CBF9B0DFFE4FFE39421FE05FA92A1EE" box="[1296,1386,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="324" unit="mm" value="1.2">1.2 mm</quantity>
); A- TL:
<quantity id="4CBF9B0DFFE4FFE391FBFE61FEDCA1CA" box="[202,292,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="324" unit="mm" value="2.1">2.1 mm</quantity>
; A-MW:
<quantity id="4CBF9B0DFFE4FFE390A3FE61FE12A1CA" box="[402,490,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="324" unit="mm" value="1.1">1.1 mm</quantity>
(NUM: 35 exs); very similar to
<taxonomicName id="4C474D6BFFE4FFE39250FE60FC12A1CA" box="[865,1002,476,500]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE4FFE39250FE60FC12A1CA" box="[865,1002,476,500]" italics="true" pageId="23" pageNumber="324">L. alvarengi</emphasis>
</taxonomicName>
in body shape and colouration; elytra markedly sinuate subapically; ventral surface uniformly testaceous.
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6FD99FEB8A2F3" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FD99FEEDA202" box="[199,277,548,572]" italics="true" pageId="23" pageNumber="324">Colour</emphasis>
. Head, pronotum, and elytra as in
<taxonomicName id="4C474D6BFFE4FFE393AEFD98FCDFA202" box="[671,807,548,572]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE4FFE393AEFD98FCDFA202" box="[671,807,548,572]" italics="true" pageId="23" pageNumber="324">L. alvarengi</emphasis>
</taxonomicName>
. Pale markings on elytra always well defined, subbasal band in most cases complete, occasionally fragmented in spots; lateral markings missing or very reduced. Subapical band extending to lateral margin, not reaching elytral apex. Underside testaceous, slightly darker along sutures and apex of abdominal ventrites 3, 4 and 5. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker.
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6FD65FF3BA39A" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FD65FE2EA2CE" box="[199,470,728,752]" italics="true" pageId="23" pageNumber="324">Structures and sculpture</emphasis>
. Head smooth, shiny, with hardly visible microreticulation of rounded cells. Pronotum with quite impressed microreticulation, with iridescence visible as on elytra. Few scattered punctures mostly along fore and hind margins. Elytra lacking subapical sinuation; less narrowed at tip than preceding species; smooth with quite impressed reticulation and intense iridescence. Underside with faint iridescence. Prosternal process quite elongate and visibly carinate, almost acute at tip; lobes of metacoxal processes ending in obtuse angulation (
<figureCitation id="137C2A6DFFE4FFE3945EFCDAFF4EA39A" captionStart="FIGURES 38 – 43" captionStartId="35.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@35.[151,1436,193,1932]" captionTargetPageId="35" captionText="FIGURES 38 – 43. SEMs of Laccomimus species, ventral view. 38 – 41) metacoxal processes: 38) L. amazonas (male); 39) L. alvarengi (female); 40) L. bordoni (female); 41) L. bolivari (female); 42 – 43) abdomen on L. bordoni, showing sexual difference in last ventrite: 42) male (arrow indicates slightly sinuate caudal margins); 43) female." httpUri="https://zenodo.org/record/244308/files/figure.png" pageId="23" pageNumber="324">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6FC0DFE3CA442" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FC0DFEF9A3F6" box="[199,257,944,968]" italics="true" pageId="23" pageNumber="324">Male</emphasis>
. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped (
<figureCitation id="137C2A6DFFE4FFE39067FC69FE52A3D2" box="[342,426,980,1005]" captionStart="FIGURES 67 – 83" captionStartId="39.[151,264,1798,1820]" captionTargetBox="[159,1430,201,1748]" captionTargetId="figure@39.[151,1436,193,1777]" captionTargetPageId="39" captionText="FIGURES 67 – 83. Prosternal process and claws in Laccomimus species. 67 – 69) P rosternal process: 67) L. pumilio (Florida, USA); 68) L. pumilio (Cuba); 69) L. alvarengi (Perú). 70 – 83) M ale left protarsal claws: 70) L. bordoni (Paraguay); 71) L. malkini (Madre de Dios, Perú); 72) L. spangleri (holotype); 73) L. distinctus (holotype); 74) L. bolivari (Trinidad); 75) L. amazonas (holotype); 76) L. variegatus (Colonia Calleria, Perú); 77) L. alvarengi (holotype); 78) L. alvarengi (Loreto, Perú); 79) L. alvarengi (Guayaramerin, Bolivia); 80) L. alvarengi (Esmeraldas, Ecuador); 81) L. alvarengi (Albrook, Panamá); 82) L. pumilio (Cayo District, Belize); 83) L. pumilio (Cayamas, Cuba)." httpUri="https://zenodo.org/record/244312/files/figure.png" pageId="23" pageNumber="324">Fig. 74</figureCitation>
). Median lobe of aedeagus, in left lateral view (
<figureCitation id="137C2A6DFFE4FFE3928BFC69FBE6A3D3" box="[954,1054,980,1005]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="23" pageNumber="324">Figs 105</figureCitation>
a; 106a), elongate, almost straight, with angle between base and distal portion almost indistinct; gradually narrowed to apex, ending in characteristic, wide and flattened expansion. In dorsal view (
<figureCitation id="137C2A6DFFE4FFE3938BFBA1FCE7A40B" box="[698,799,1052,1077]" captionStart="FIGURES 107 – 115" captionStartId="46.[151,264,1951,1973]" captionTargetBox="[184,1390,196,1925]" captionTargetId="figure@46.[170,1417,193,1930]" captionTargetPageId="46" captionText="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri="https://zenodo.org/record/244319/files/figure.png" pageId="23" pageNumber="324">Fig. 115</figureCitation>
), slender, gently but visibly turned to right; ejaculator groove gradually narrowed to apex, closed before apical fifth of median lobe. Left paramere (
<figureCitation id="137C2A6DFFE4FFE395F2FB82FADCA466" box="[1219,1316,1087,1112]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="23" pageNumber="324">Fig. 105</figureCitation>
c) straight, gradually tapering to apex.
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6FB35FC69A4D6" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FB35FEE3A49E" box="[199,283,1160,1184]" italics="true" pageId="23" pageNumber="324">Female</emphasis>
. Iridescence as in male or slightly more intense. Last abdominal ventrite triangular, very narrowly rounded at apex, almost pointed. Lateral tergites thin and partly membranous. Gonocoxal blade with single denticle; ramus as in preceding species, with large and spaced teeth.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFE391F6FB49FDCCA56B" pageId="23" pageNumber="324" type="distribution">
<paragraph id="8BF836E8FFE4FFE391F6FB49FCC7A532" blockId="23.[151,1437,151,1760]" box="[199,831,1268,1293]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FB49FE98A533" bold="true" box="[199,352,1268,1293]" pageId="23" pageNumber="324">Distribution.</emphasis>
<collectingCountry id="F3507678FFE4FFE39056FB49FE25A533" box="[359,477,1268,1293]" name="Venezuela" pageId="23" pageNumber="324">Venezuela</collectingCountry>
,
<collectingCountry id="F3507678FFE4FFE390D9FB49FDB2A533" box="[488,586,1268,1293]" name="Trinidad and Tobago" pageId="23" pageNumber="324">Trinidad</collectingCountry>
,
<collectingCountry id="F3507678FFE4FFE39364FB49FD38A533" box="[597,704,1268,1293]" name="Suriname" pageId="23" pageNumber="324">Suriname</collectingCountry>
(
<figureCitation id="137C2A6DFFE4FFE393E1FB49FCCBA532" box="[720,819,1268,1293]" captionStart="FIGURES 126 – 129" captionStartId="49.[151,264,1953,1975]" captionTargetBox="[157,1432,197,1931]" captionTargetId="figure@49.[151,1436,193,1932]" captionTargetPageId="49" captionText="FIGURES 126 – 129. Distribution maps of Laccomimus species: 126) L. bordoni in South America; 127) L. bolivari (full circles), L. youngi (single open square in Brazil); 128) L. pumilio (open circles), L. alvarengi in Central America (single triangle in Panamá), L. spangleri (stars), L. bordoni in Central America (asterisks); 129) L. improvidus." httpUri="https://zenodo.org/record/244322/files/figure.png" pageId="23" pageNumber="324">Fig. 127</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6FAAAFDCCA56B" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FAAAFED3A50E" bold="true" box="[199,299,1303,1328]" pageId="23" pageNumber="324">Ecology.</emphasis>
VAIP’s field data suggest a wide ecological range, with collections from a morichal, a lagoon and a small streamlet (
<figureCitation id="137C2A6DFFE4FFE39061FA81FE4BA56A" box="[336,435,1340,1365]" captionStart="FIGURES 132 – 133" captionStartId="51.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@51.[151,1436,193,1932]" captionTargetPageId="51" captionText="FIGURES 132 – 133. Habitats of Laccomimus species: 132) L. distinctus, ditch at El Cachapé refuge, Chaco Province, Argentina (photo M. C. Michat); 133) L. improvidus and L. bolivari, large lagoon with vegetation ca. 65 km S Las Mercedes, Guarico State, Venezuela (photo A. E. Z. Short)." httpUri="https://zenodo.org/record/244323/files/figure.png" pageId="23" pageNumber="324">Figs 133</figureCitation>
–134; 136).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFE391F6FAE2FF26A64B" pageId="23" pageNumber="324" type="etymology">
<paragraph id="8BF836E8FFE4FFE391F6FAE2FF37A636" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6FAE2FEB6A546" bold="true" box="[199,334,1375,1400]" pageId="23" pageNumber="324">Etymology.</emphasis>
Young and Spangler named
<emphasis id="B933EAFAFFE4FFE393A1FADDFD0EA546" box="[656,758,1376,1400]" italics="true" pageId="23" pageNumber="324">in litteris</emphasis>
this species
<taxonomicName id="4C474D6BFFE4FFE392B4FADDFB53A546" box="[901,1195,1376,1400]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="trinidadensis">
<emphasis id="B933EAFAFFE4FFE392B4FADDFB53A546" box="[901,1195,1376,1400]" italics="true" pageId="23" pageNumber="324">Laccomimus trinidadensis</emphasis>
</taxonomicName>
, based on material in USNM, in great part collected from
<collectingCountry id="F3507678FFE4FFE39305FA39FD6FA5A3" box="[564,663,1412,1437]" name="Trinidad and Tobago" pageId="23" pageNumber="324">Trinidad</collectingCountry>
. However, it is clear that this species has a much wider distribution. We decided to name this taxon
<taxonomicName id="4C474D6BFFE4FFE39333FA15FDA3A5FE" box="[514,603,1448,1472]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFE4FFE39333FA15FDA3A5FE" box="[514,603,1448,1472]" italics="true" pageId="23" pageNumber="324">bolivari</emphasis>
</taxonomicName>
in reference to the
<typeStatus id="54FC884AFFE4FFE3920FFA14FC96A5FF" box="[830,878,1449,1473]" pageId="23" pageNumber="324">type</typeStatus>
locality and in honour of the Venezuelan patriot Simón José Antonio de la Santísima
<collectingCountry id="F3507678FFE4FFE3931EFA71FD77A5DB" box="[559,655,1484,1509]" name="Trinidad and Tobago" pageId="23" pageNumber="324">Trinidad</collectingCountry>
Bolivar, “El Libertador”. The specific epithet is a noun in the genitive case.
</paragraph>
<paragraph id="8BF836E8FFE4FFE391F6F9A9FF26A64B" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6F9A9FE43A613" bold="true" box="[199,443,1556,1581]" pageId="23" pageNumber="324">Notes on variability.</emphasis>
The only specimen seen from
<collectingCountry id="F3507678FFE4FFE39211F9A9FC73A613" box="[800,907,1556,1581]" name="Suriname" pageId="23" pageNumber="324">Suriname</collectingCountry>
has the median lobe in lateral view thicker in distal portion and more abruptly narrowing to apex in distal fourth; the apical expansion is slightly smaller (
<figureCitation id="137C2A6DFFE4FFE3945EF98AFF3CA64B" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="23" pageNumber="324">Fig. 106</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE4FFE391F6F9C2FB71A6DE" pageId="23" pageNumber="324" type="discussion">
<paragraph id="8BF836E8FFE4FFE391F6F9C2FB71A6DE" blockId="23.[151,1437,151,1760]" pageId="23" pageNumber="324">
<emphasis id="B933EAFAFFE4FFE391F6F9C2FEC3A6A6" bold="true" box="[199,315,1663,1688]" pageId="23" pageNumber="324">Remarks.</emphasis>
This species is close to
<taxonomicName id="4C474D6BFFE4FFE3937EF93CFD3AA6A6" box="[591,706,1664,1688]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFE4FFE3937EF93CFD3AA6A6" box="[591,706,1664,1688]" italics="true" pageId="23" pageNumber="324">L. pumilio</emphasis>
</taxonomicName>
and related species, sharing with them external male characters, similar female genitalia, and a similar shape of the prosternal and metacoxal processes. It is, however, well characterised by aedeagal features, unusual compared to the other species of
<taxonomicName id="4C474D6BFFE4FFE392C5F975FB79A6E1" box="[1012,1153,1736,1759]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFE4FFE392C5F975FB79A6E1" box="[1012,1153,1736,1759]" italics="true" pageId="23" pageNumber="324">Laccomimus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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