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<document id="1EFD2B964D4626B8C2C62965ABDAF4F3" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario &amp; Michat, Mariano C." docDate="2015" docId="03EE87FEFFE1FFE791A6FB1CFE6EA683" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus improvidus Toledo &amp; Michat, 2015, sp. n." docType="treatment" docVersion="9" lastPageNumber="320" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="319" updateTime="1698611165842" updateUser="plazi">
<mods:mods id="B1DF823AFE6AF873C4634FFCB66D4399" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B0B67C751EB2E2902039E3A648680A01">
<mods:title id="99EFE2573ADE21FD44208A10292C03D8">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
</mods:titleInfo>
<mods:name id="45F4C77EFA446C8B3441176F1AFB944A" type="personal">
<mods:role id="24CAC3936379CB53682CC0E3EB532BAE">
<mods:roleTerm id="5FEC087666D9E44DBA8BDEF315FE6375">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="971CD8953ECA014AA2DF46C38C1F8FAD">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="5A070A06B294F6C6CFB9CEA941CD5DAF" type="personal">
<mods:role id="13E6753F2E282940BF49A9361E34B7BD">
<mods:roleTerm id="D99A68A4EE2D540D2340DFF7CB318DB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6DCA7C2F352F60D9B4535D56AAE1531A">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DA136AEB9BD2934098F6A387EAB0FB14">text</mods:typeOfResource>
<mods:relatedItem id="2F53EA20134C8840AA01845544BCFB26" type="host">
<mods:titleInfo id="5A95D0481EA30D65AF5AEC3B81B565BF">
<mods:title id="7F2E40371C609FEE25F21928368D7DF0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F20DF71937BB78553F8863C98A873DD6">
<mods:date id="C5574E25BD7462F19418184F0B97298E">2015</mods:date>
<mods:detail id="CEBBFB261DD24AA36BE8B9F6B81470C3" type="volume">
<mods:number id="363A1D3FDA8646BCA09572CA6E2AD12F">3990</mods:number>
</mods:detail>
<mods:detail id="E62BAD87FC6D3192D28D5F7BE295E48F" type="issue">
<mods:number id="A8CAFD6B7262497817DEDD2A198F84F6">3</mods:number>
</mods:detail>
<mods:extent id="5AA286E4E7A92203584BFD8A1E00BA7B" unit="page">
<mods:start id="5C88F4F9359A37D84204CBC75C4A359A">301</mods:start>
<mods:end id="5B0AF51F9199CD6E217D1AF79754B1E7">354</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9483145FFD865376EE382D21E7A6BCC9">journal article</mods:classification>
<mods:identifier id="023DA5654E2E83E08D83A00995B24473" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
<mods:identifier id="2C456ED8506282ADA381B35DE9A4034B" type="GBIF-Dataset">cf747765-b9bf-42ed-8479-35355e712db0</mods:identifier>
<mods:identifier id="FADDAB9B32C4CBF849F329E83C525469" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8BBEB8E35F7872FBABA6BDE9AE2E3DF1" type="Zenodo-Dep">244301</mods:identifier>
<mods:identifier id="EC89A8345E3339EAD2795EB7EA448667" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
</mods:mods>
<treatment id="03EE87FEFFE1FFE791A6FB1CFE6EA683" ID-DOI="http://doi.org/10.5281/zenodo.5692054" ID-GBIF-Taxon="119638446" ID-Zenodo-Dep="5692054" LSID="urn:lsid:plazi:treatment:03EE87FEFFE1FFE791A6FB1CFE6EA683" httpUri="http://treatment.plazi.org/id/03EE87FEFFE1FFE791A6FB1CFE6EA683" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<subSubSection id="C35D6563FFE1FFE691A6FB1CFE5DA4E2" pageId="18" pageNumber="319" type="nomenclature">
<paragraph id="8BF836E8FFE1FFE691A6FB1CFDF1A485" blockId="18.[151,521,1185,1245]" box="[151,521,1185,1211]" pageId="18" pageNumber="319">
<heading id="D0B08184FFE1FFE691A6FB1CFDF1A485" bold="true" box="[151,521,1185,1211]" fontSize="11" level="1" pageId="18" pageNumber="319" reason="1">
<emphasis id="B933EAFAFFE1FFE691A6FB1CFDF1A485" bold="true" box="[151,521,1185,1211]" pageId="18" pageNumber="319">
<taxonomicName id="4C474D6BFFE1FFE691A6FB1CFE38A485" ID-CoL="8LXJX" box="[151,448,1185,1211]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="improvidus" status="sp. nov.">
<emphasis id="B933EAFAFFE1FFE691A6FB1CFE38A485" bold="true" box="[151,448,1185,1211]" italics="true" pageId="18" pageNumber="319">Laccomimus improvidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2005781FFE1FFE690F6FB1CFDF1A485" box="[455,521,1185,1211]" pageId="18" pageNumber="319" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836E8FFE1FFE691A6FB79FE5DA4E2" blockId="18.[151,521,1185,1245]" box="[151,421,1220,1245]" pageId="18" pageNumber="319">
(
<figureCitation id="137C2A6DFFE1FFE691AEFB79FF0EA4E2" box="[159,246,1220,1245]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="18" pageNumber="319">Figs 89</figureCitation>
; 112; 129; 133)
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE1FFE791A6FAB1FC77A122" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319" type="materials_examined">
<paragraph id="8BF836E8FFE1FFE691A6FAB1FEB4A576" blockId="18.[151,1437,1292,2037]" pageId="18" pageNumber="319">
<emphasis id="B933EAFAFFE1FFE691A6FAB1FECDA51B" bold="true" box="[151,309,1292,1317]" pageId="18" pageNumber="319">
<typeStatus id="54FC884AFFE1FFE691A6FAB1FF2AA51B" box="[151,210,1292,1317]" pageId="18" pageNumber="319">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3507678FFE1FFE6900AFAB1FE49A51B" box="[315,433,1292,1317]" name="Venezuela" pageId="18" pageNumber="319">Venezuela</collectingCountry>
, Guarico State,
<geoCoordinate id="EE73502FFFE1FFE69350FAB1FD1BA51B" box="[609,739,1292,1317]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="1" value="8.528417">8°31.705'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE693DFFAB1FC7AA51B" box="[750,898,1292,1317]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="1" value="-66.3767">66°22.602'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE1FFE692BDFAB1FC2DA51A" box="[908,981,1292,1317]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="319" unit="m" value="145.0">145 m</quantity>
, ca.
<quantity id="4CBF9B0DFFE1FFE69537FAB1FBB5A51B" box="[1030,1101,1292,1317]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="319" unit="km" value="65.0">65 km</quantity>
S Las Mercedes, large lagoon with vegetation.
</paragraph>
<paragraph id="8BF836E8FFE1FFE691F6FAE9FD36A5C3" blockId="18.[151,1437,1292,2037]" pageId="18" pageNumber="319">
<emphasis id="B933EAFAFFE1FFE691F6FAE9FE82A553" bold="true" box="[199,378,1364,1389]" pageId="18" pageNumber="319">
<typeStatus id="54FC884AFFE1FFE691F6FAE9FEFAA553" box="[199,258,1364,1389]" pageId="18" pageNumber="319">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54FC884AFFE1FFE690B7FAE9FE16A553" box="[390,494,1364,1389]" pageId="18" pageNumber="319" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED56AE2DFFE1FFE69374FAE9FD6BA552" LSID="urn:lsid:biocol.org:col:33834" box="[581,659,1364,1388]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
): dissected by us; last two metaventrites and genitaia included in
<collectionCode id="ED56AE2DFFE1FFE691A6FAC5FF10A5B1" box="[151,232,1400,1423]" pageId="18" pageNumber="319">DMHF</collectionCode>
on plastic label, pinned under the
<typeStatus id="54FC884AFFE1FFE69340FACAFD2BA5AE" box="[625,723,1399,1424]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
“Guarico State,
<geoCoordinate id="EE73502FFFE1FFE692A1FACAFBEAA5AE" box="[912,1042,1399,1424]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="1" value="8.528417">8°31.705'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE6952FFACAFB4AA5AE" box="[1054,1202,1399,1424]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="1" value="-66.3767">66°22.602'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE1FFE6958FFACAFAF0A5B1" box="[1214,1288,1399,1424]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="319" unit="m" value="145.0">145 m</quantity>
, ca.
<quantity id="4CBF9B0DFFE1FFE6940CFACAFA7DA5AE" box="[1341,1413,1399,1424]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="319" unit="km" value="65.0">65 km</quantity>
S Las Mercedes, large lagoon with vegetation,
<date id="FFF91028FFE1FFE69391FA21FD07A58A" box="[672,767,1436,1460]" pageId="18" pageNumber="319" value="2009-01-09">9.I.2009</date>
, leg. Short, Miller, Garcia &amp; Camacho, VZ09-0109-
<date id="FFF91028FFE1FFE69453FA21FA69A58A" box="[1378,1425,1436,1460]" pageId="18" pageNumber="319" value="2001-10">01X</date>
” [printed], “
<collectionCode id="ED56AE2DFFE1FFE69023FA7DFE46A5E6" box="[274,446,1472,1496]" pageId="18" pageNumber="319">KUNHM-ENT</collectionCode>
SM0845078” [printed, together with barcode], “
<typeStatus id="54FC884AFFE1FFE692D8FA7DFB78A5E6" box="[1001,1152,1472,1496]" pageId="18" pageNumber="319" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C474D6BFFE1FFE695B8FA7DFA64A5E6" box="[1161,1436,1472,1496]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="improvidus">
<emphasis id="B933EAFAFFE1FFE695B8FA7DFA64A5E6" box="[1161,1436,1472,1496]" italics="true" pageId="18" pageNumber="319">Laccomimus improvidus</emphasis>
</taxonomicName>
Toledo &amp; Michat” [red, rectangular, printed label].
</paragraph>
<paragraph id="8BF836E8FFE1FFE791F6F9B5FC77A122" blockId="18.[151,1437,1292,2037]" lastBlockId="19.[151,1437,151,1725]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<typeStatus id="54FC884AFFE1FFE691F6F9B5FECCA61E" box="[199,308,1544,1568]" pageId="18" pageNumber="319" type="paratype">Paratypes</typeStatus>
(26 exs, all bearing red, printed label “
<typeStatus id="54FC884AFFE1FFE69231F9B5FC76A61E" box="[768,910,1544,1568]" pageId="18" pageNumber="319" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C474D6BFFE1FFE692A8F9B5FB55A61E" box="[921,1197,1544,1568]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="improvidus">
<emphasis id="B933EAFAFFE1FFE692A8F9B5FB55A61E" box="[921,1197,1544,1568]" italics="true" pageId="18" pageNumber="319">Laccomimus improvidus</emphasis>
</taxonomicName>
Toledo &amp; Michat”):
<emphasis id="B933EAFAFFE1FFE691A6F991FEE0A67B" bold="true" box="[151,280,1580,1605]" pageId="18" pageNumber="319">
<collectingCountry id="F3507678FFE1FFE691A6F991FEECA67B" box="[151,276,1580,1605]" name="Venezuela" pageId="18" pageNumber="319">Venezuela</collectingCountry>
:
</emphasis>
same data of
<typeStatus id="54FC884AFFE1FFE69084F991FDEFA67B" box="[437,535,1580,1605]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D41FD61FFE1FFE69318F991FD8AA67B" box="[553,626,1580,1605]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE6934BF991FD45A67A" box="[634,701,1580,1604]" pageId="18" pageNumber="319">VAIP</collectionCode>
); “Monagas State,
<geoCoordinate id="EE73502FFFE1FFE692A1F991FBEAA67B" box="[912,1042,1580,1605]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="1" value="9.60985">9°36.591'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE6952CF991FB5BA67B" box="[1053,1187,1580,1605]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="1" value="-63.13825">63°8.295'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE1FFE6959FF991FB12A67A" box="[1198,1258,1580,1605]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="319" unit="m" value="45.0">45 m</quantity>
, S. of Marturin, vegetated river”, “morichal margin,
<date id="FFF91028FFE1FFE69307F9EDFD67A656" box="[566,671,1616,1640]" pageId="18" pageNumber="319" value="2010-02-02">2.II.2010</date>
, leg. Short &amp; Garcia, VZ10-0202-01B” (
<specimenCount id="9D41FD61FFE1FFE6954AF9EDFB30A656" box="[1147,1224,1615,1640]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE695E0F9EDFAECA656" box="[1233,1300,1616,1640]" pageId="18" pageNumber="319">VAIP</collectionCode>
); “Monagas State,
<geoCoordinate id="EE73502FFFE1FFE691D3F9C9FE9CA6B3" box="[226,356,1652,1677]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="1" value="8.773983">8°46.439'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE69044F9C9FDF0A6B3" box="[373,520,1652,1677]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="1" value="-62.238434">62°14.306'W</geoCoordinate>
,
<quantity id="4CBF9B0DFFE1FFE69328F9C9FDA1A6B2" box="[537,601,1652,1676]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="18" pageNumber="319" unit="m" value="19.0">19 m</quantity>
, between Tucupita and Temblador, pond margin,
<date id="FFF91028FFE1FFE69598F9C9FAEAA6B2" box="[1193,1298,1652,1676]" pageId="18" pageNumber="319" value="2010-02-03">3.II.2010</date>
, leg. Short, Garcia &amp; Joly, VZ10-0203-02A” (
<specimenCount id="9D41FD61FFE1FFE6931AF925FD80A68E" box="[555,632,1687,1712]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE693B2F925FD3EA68E" box="[643,710,1688,1712]" pageId="18" pageNumber="319">VAIP</collectionCode>
); each one of these two latter specimens bearing second label (
<collectionCode id="ED56AE2DFFE1FFE691AFF901FEB3A6EA" box="[158,331,1724,1748]" pageId="18" pageNumber="319">KUNHM-ENT</collectionCode>
) with its own barcode and catalogue number; “Km 88 El Dorado,
<collectingCountry id="F3507678FFE1FFE69546F901FAFAA6EB" box="[1143,1282,1724,1749]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="18" pageNumber="319">Santa Elena</collectingCountry>
BO,
<quantity id="4CBF9B0DFFE1FFE69478F901FA6FA6EA" box="[1353,1431,1724,1749]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="319" unit="m" value="160.0">160 m</quantity>
,
<date id="FFF91028FFE1FFE691A6F962FEE1A6C6" box="[151,281,1759,1784]" pageId="18" pageNumber="319" value="1966-11-14">14.XI.1966</date>
, J. &amp; B. Bechyne &amp; E. Osuna” (
<specimenCount id="9D41FD61FFE1FFE69382F95DFCFDA6C6" box="[691,773,1759,1784]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
,
<specimenCount id="9D41FD61FFE1FFE69227F95DFC70A6C6" box="[790,904,1759,1784]" pageId="18" pageNumber="319" type="female">3 females</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE692A5F95DFC19A6C6" LSID="urn:lsid:biocol.org:col:33834" box="[916,993,1760,1784]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
); “Bolivar, Guri, Rio Caroni,
<quantity id="4CBF9B0DFFE1FFE69479F95DFA6FA6C9" box="[1352,1431,1760,1784]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="319" unit="m" value="100.0">100 m</quantity>
,
<date id="FFF91028FFE1FFE691A6F8B9FEE1A723" box="[151,281,1796,1821]" pageId="18" pageNumber="319" value="1966-11-16">16.XI.1966</date>
, J. &amp; B. Bechyne &amp; E. Osuna” (
<specimenCount id="9D41FD61FFE1FFE69397F8B9FCF1A723" box="[678,777,1796,1821]" pageId="18" pageNumber="319" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE69224F8B9FC9AA722" LSID="urn:lsid:biocol.org:col:33834" box="[789,866,1796,1820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
); “Bolivar, Ana[illegible letter]oco,
<quantity id="4CBF9B0DFFE1FFE69437F8B9FABDA722" box="[1286,1349,1796,1821]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="319" unit="m" value="60.0">60 m</quantity>
, 6°5',N
<geoCoordinate id="EE73502FFFE1FFE691A6F89AFF14A77E" box="[151,236,1831,1856]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="925" value="-61.133335">61°8'W</geoCoordinate>
,
<date id="FFF91028FFE1FFE691C8F895FE4EA77E" box="[249,438,1832,1856]" pageId="18" pageNumber="319" value="1979-08-10" valueMax="1979-08-30" valueMin="1979-08-10">10–30.VIII.1979</date>
” (
<specimenCount id="9D41FD61FFE1FFE690E4F895FDD8A77E" box="[469,544,1831,1856]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE69318F895FD8EA77E" LSID="urn:lsid:biocol.org:col:33834" box="[553,630,1832,1856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
); “Aragua, El Limon,
<quantity id="4CBF9B0DFFE1FFE69245F89AFC47A701" box="[884,959,1831,1856]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="319" unit="m" value="450.0">450 m</quantity>
,
<date id="FFF91028FFE1FFE692FDF895FBA0A77E" box="[972,1112,1832,1856]" pageId="18" pageNumber="319" value="1977-07-13">
13.
<collectionCode id="ED56AE2DFFE1FFE692DAF895FBEFA77E" box="[1003,1047,1832,1856]" pageId="18" pageNumber="319">VII</collectionCode>
.1977
</date>
, leg. B. Bechyne” (
<specimenCount id="9D41FD61FFE1FFE6940AF895FA64A77E" box="[1339,1436,1831,1856]" pageId="18" pageNumber="319" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE691A6F8F1FF1CA75A" LSID="urn:lsid:biocol.org:col:33834" box="[151,228,1868,1892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
); “Coje, Galeras del Pao,
<date id="FFF91028FFE1FFE69332F8F1FD7DA75B" box="[515,645,1868,1893]" pageId="18" pageNumber="319" value="1963-06-28">28.VI.1963</date>
, leg. C. J. Rosales &amp; A. Perez” (
<specimenCount id="9D41FD61FFE1FFE692C0F8F1FBA6A75B" box="[1009,1118,1868,1893]" pageId="18" pageNumber="319" type="female">2 females</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE69554F8F1FB4AA75A" LSID="urn:lsid:biocol.org:col:33834" box="[1125,1202,1868,1892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
); all these specimens in
<collectionCode id="ED56AE2DFFE1FFE69185F8CDFF06A7B6" LSID="urn:lsid:biocol.org:col:33834" box="[180,254,1904,1928]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
, bearing an additional printed green label “
<collectionCode id="ED56AE2DFFE1FFE693EEF8CDFCD6A7B6" LSID="urn:lsid:biocol.org:col:33834" box="[735,814,1904,1928]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="18" pageNumber="319">MIZA</collectionCode>
”.
<emphasis id="B933EAFAFFE1FFE69272F8D2FC38A7B6" bold="true" box="[835,960,1903,1928]" pageId="18" pageNumber="319">
<collectingCountry id="F3507678FFE1FFE69272F8D2FC44A7B6" box="[835,956,1903,1928]" name="Suriname" pageId="18" pageNumber="319">Suriname</collectingCountry>
:
</emphasis>
“Paramaribo, Guest House of Museum, at light,
<date id="FFF91028FFE1FFE691ECF829FE38A793" box="[221,448,1940,1965]" pageId="18" pageNumber="319" value="1969-08-20" valueMax="1969-09-23" valueMin="1969-08-20">20.VIII–23.IX.1969</date>
, leg. N. Nieser” (
<specimenCount id="9D41FD61FFE1FFE693ABF829FD13A793" box="[666,747,1940,1965]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
,
<specimenCount id="9D41FD61FFE1FFE693CDF829FC95A793" box="[764,877,1940,1965]" pageId="18" pageNumber="319" type="female">2 females</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE69248F829FC3EA792" box="[889,966,1940,1964]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="319">NMW</collectionCode>
); idem except for “
<date id="FFF91028FFE1FFE6959EF829FA96A793" box="[1199,1390,1940,1965]" pageId="18" pageNumber="319" value="1969-08-21" valueMax="1969-08-25" valueMin="1969-08-21">21–25.VIII.1969</date>
” (
<specimenCount id="9D41FD61FFE1FFE694A1F829FF1BA7EF" pageId="18" pageNumber="319" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE691C5F805FEB9A7EE" box="[244,321,1976,2000]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="319">NMW</collectionCode>
); idem except for “
<date id="FFF91028FFE1FFE69371F805FCE7A7EF" box="[576,799,1976,2001]" pageId="18" pageNumber="319" value="1969-07-26" valueMax="1969-08-07" valueMin="1969-07-26">
26.
<collectionCode id="ED56AE2DFFE1FFE69350F805FD73A7EF" box="[609,651,1976,2001]" pageId="18" pageNumber="319">VII</collectionCode>
–7.VIII.1969
</date>
” (
<specimenCount id="9D41FD61FFE1FFE69276F805FC48A7EF" box="[839,944,1976,2001]" pageId="18" pageNumber="319" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE692F3F805FBF7A7EE" box="[962,1039,1976,2000]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="319">NMW</collectionCode>
); “
<geoCoordinate id="EE73502FFFE1FFE69506F805FB64A7EF" box="[1079,1180,1976,2001]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="925" value="-55.1">55°06'W</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE69580F805FAE9A7EF" box="[1201,1297,1976,2001]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="925" value="5.75">05°45'N</geoCoordinate>
, Oost-West Verbinding,
<quantity id="4CBF9B0DFFE1FFE69013F861FE8FA7CB" box="[290,375,2012,2037]" metricMagnitude="3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="18" pageNumber="319" unit="km" value="7.5" valueMax="8.0" valueMin="7.0">7–8 km</quantity>
E Meerzorg,
<date id="FFF91028FFE1FFE69320F861FD83A7CB" box="[529,635,2012,2037]" pageId="18" pageNumber="319" value="1969-10-02">2.X.1969</date>
, leg N. Nieser (SN 157)” (
<specimenCount id="9D41FD61FFE1FFE6929BF861FC0BA7CB" box="[938,1011,2012,2037]" pageId="18" pageNumber="319" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE1FFE692CAF861FBB0A7CA" box="[1019,1096,2012,2036]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="319">NMW</collectionCode>
); “
<geoCoordinate id="EE73502FFFE1FFE69557F861FB31A7CB" box="[1126,1225,2012,2037]" direction="west" orientation="longitude" pageId="18" pageNumber="319" precision="925" value="-55.183334">55°11'W</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE1FFE695E4F861FACBA7CB" box="[1237,1331,2012,2037]" direction="north" orientation="latitude" pageId="18" pageNumber="319" precision="925" value="5.5166664">05°31'N</geoCoordinate>
, N banks of Coropinakreek,
<date id="FFF91028FFE0FFE79040FF2AFE0BA08E" box="[369,499,151,176]" pageId="19" pageNumber="320" value="1969-09-25">25.IX.1969</date>
, leg N. Nieser (SN 156A)” (
<specimenCount id="9D41FD61FFE0FFE7927DFF25FC48A08E" box="[844,944,151,176]" pageId="19" pageNumber="320" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE0FFE7928AFF25FBF0A08E" box="[955,1032,152,176]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="320">NMW</collectionCode>
).
<emphasis id="B933EAFAFFE0FFE7952CFF2AFB7DA08E" bold="true" box="[1053,1157,151,176]" pageId="19" pageNumber="320">
<collectingCountry id="F3507678FFE0FFE7952CFF2AFB78A08E" box="[1053,1152,151,176]" name="Guyana" pageId="19" pageNumber="320">Guyana</collectingCountry>
:
</emphasis>
“Karanambo,
<geoCoordinate id="EE73502FFFE0FFE79400FF2AFA6FA08E" box="[1329,1431,151,176]" direction="north" orientation="latitude" pageId="19" pageNumber="320" precision="92" value="3.7516665">3°45.1'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE0FFE791A6FF01FEF7A0EB" box="[151,271,188,213]" direction="west" orientation="longitude" pageId="19" pageNumber="320" precision="92" value="-59.31">59°18.6'W</geoCoordinate>
, blacklight on bank of Rupununi,
<date id="FFF91028FFE0FFE793A4FF01FCEEA0EA" box="[661,790,188,212]" pageId="19" pageNumber="320" value="1994-03-31">31.III.1994</date>
, P.J. Spangler, colln #2” (
<specimenCount id="9D41FD61FFE0FFE7950EFF01FB67A0EB" box="[1087,1183,188,213]" pageId="19" pageNumber="320" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE0FFE79599FF01FB06A0EA" LSID="urn:lsid:biocol.org:col:34871" box="[1192,1278,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="320">USNM</collectionCode>
); idem except for “Buffalo Pond,
<date id="FFF91028FFE0FFE7904BFF5DFE00A0C6" box="[378,504,224,248]" pageId="19" pageNumber="320" value="1994-04-24">24.IV.1994</date>
, colln #11” (
<specimenCount id="9D41FD61FFE0FFE793A7FF5DFD1CA0C6" box="[662,740,223,248]" pageId="19" pageNumber="320" type="male">1 male</specimenCount>
<collectionCode id="ED56AE2DFFE0FFE793DFFF5DFCBCA0C6" LSID="urn:lsid:biocol.org:col:34871" box="[750,836,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="320">USNM</collectionCode>
); “Pirara Ranch,
<geoCoordinate id="EE73502FFFE0FFE7953FFF5DFB8CA0C6" box="[1038,1140,224,248]" direction="north" orientation="latitude" pageId="19" pageNumber="320" precision="92" value="3.5349998">3°32.1'N</geoCoordinate>
,
<geoCoordinate id="EE73502FFFE0FFE795B2FF62FB04A0C6" box="[1155,1276,223,248]" direction="west" orientation="longitude" pageId="19" pageNumber="320" precision="92" value="-59.675">59°40.5'W</geoCoordinate>
, Pirara River,
<date id="FFF91028FFE0FFE791A6FEB9FEEDA123" box="[151,277,260,285]" pageId="19" pageNumber="320" value="1995-04-24">24.IV.1995</date>
, P.J. Spangler &amp; A. Perry, colln #18” (
<specimenCount id="9D41FD61FFE0FFE793F6FEB9FCDEA123" box="[711,806,260,285]" pageId="19" pageNumber="320" type="female">1 female</specimenCount>
<collectionCode id="ED56AE2DFFE0FFE7921FFEB9FC7CA122" LSID="urn:lsid:biocol.org:col:34871" box="[814,900,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="320">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE0FFE791F6FE9AFA92A1EE" pageId="19" pageNumber="320" type="diagnosis">
<paragraph id="8BF836E8FFE0FFE791F6FE9AFA92A1EE" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FE9AFEB9A17E" bold="true" box="[199,321,295,320]" pageId="19" pageNumber="320">Diagnosis.</emphasis>
Externally indistinguishable from other species of
<taxonomicName id="4C474D6BFFE0FFE792B2FE95FBE8A101" box="[899,1040,296,319]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B933EAFAFFE0FFE792B2FE95FBE8A101" box="[899,1040,296,319]" italics="true" pageId="19" pageNumber="320">Laccomimus</emphasis>
</taxonomicName>
, in particular from dark specimens of
<taxonomicName id="4C474D6BFFE0FFE79187FEF0FEC7A15A" box="[182,319,332,356]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE79187FEF0FEC7A15A" box="[182,319,332,356]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D6BFFE0FFE79046FEF0FE11A15A" box="[375,489,332,356]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFE0FFE79046FEF0FE11A15A" box="[375,489,332,356]" italics="true" pageId="19" pageNumber="320">L. malkini</emphasis>
</taxonomicName>
with marked elytral pattern. Examination of female genitalia is also not diagnostic, except from
<taxonomicName id="4C474D6BFFE0FFE79014FECCFE60A1B6" box="[293,408,368,392]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="malkini">
<emphasis id="B933EAFAFFE0FFE79014FECCFE60A1B6" box="[293,408,368,392]" italics="true" pageId="19" pageNumber="320">L. malkini</emphasis>
</taxonomicName>
; male genitalia are distinctive, however, with median lobe more straight and robust than in
<taxonomicName id="4C474D6BFFE0FFE791A6FE28FEF3A192" box="[151,267,404,428]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFE0FFE791A6FE28FEF3A192" box="[151,267,404,428]" italics="true" pageId="19" pageNumber="320">L. pumilio</emphasis>
</taxonomicName>
, with base and distal portion forming a more obtuse angle and apex ending in a wider expansion (
<figureCitation id="137C2A6DFFE0FFE7945EFE29FF4DA1EE" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="19" pageNumber="320">Fig. 89</figureCitation>
a, b); left paramere similar to that of
<taxonomicName id="4C474D6BFFE0FFE79361FE04FD2FA1EE" box="[592,727,440,464]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE79361FE04FD2FA1EE" box="[592,727,440,464]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
, although not or very poorly thickened apically (
<figureCitation id="137C2A6DFFE0FFE795CDFE0AFAA8A1EE" box="[1276,1360,439,464]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="19" pageNumber="320">Fig. 89</figureCitation>
c).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE0FFE791F6FE61FC3CA4D6" pageId="19" pageNumber="320" type="description">
<paragraph id="8BF836E8FFE0FFE791F6FE61FDB6A203" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FE61FEA1A1CB" bold="true" box="[199,345,476,501]" pageId="19" pageNumber="320">Description.</emphasis>
TL: 2.0–
<quantity id="4CBF9B0DFFE0FFE790F2FE61FDD2A1CA" box="[451,554,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="19" pageNumber="320" unit="mm" value="2.25">2.25 mm</quantity>
; MW: 1.0–
<quantity id="4CBF9B0DFFE0FFE79398FE61FD07A1CA" box="[681,767,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="320" unit="mm" value="1.2">1.2 mm</quantity>
(
<typeStatus id="54FC884AFFE0FFE7923FFE61FC8EA1CB" box="[782,886,476,501]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
: TL:
<quantity id="4CBF9B0DFFE0FFE79282FE61FBF4A1CA" box="[947,1036,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="320" unit="mm" value="2.2">2.2 mm</quantity>
; MW:
<quantity id="4CBF9B0DFFE0FFE7956BFE61FB4DA1CA" box="[1114,1205,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="320" unit="mm" value="1.1">1.1 mm</quantity>
); A-TL:
<quantity id="4CBF9B0DFFE0FFE79429FE61FA8AA1CA" box="[1304,1394,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="320" unit="mm" value="2.2">2.2 mm</quantity>
; A- MW:
<quantity id="4CBF9B0DFFE0FFE791EBFDBDFECAA229" box="[218,306,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="320" unit="mm" value="1.1">1.1 mm</quantity>
(NUM: 19 exs); drop-shaped, regularly tapering posteriorly; elytral subapical sinuation missing or hardly visible. Iridescence well visible.
</paragraph>
<paragraph id="8BF836E8FFE0FFE791F6FDF5FEB8A32B" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FDF5FEEDA25E" box="[199,277,584,608]" italics="true" pageId="19" pageNumber="320">Colour</emphasis>
. Head and pronotum markedly paler than elytra. Head uniformly reddish-yellow, at most slightly darkened behind eyes; pronotum of same colour as head, with two brown, rectangular markings along both sides of hind margin. Elytra chestnut-brown, with vague yellow subbasal markings normally fragmented in spots; rarely missing. Apical fourth more or less gradually paler; a poorly defined paler area is often visible on lateral side. Underside uniformly testaceous to reddish-yellow. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker.
</paragraph>
<paragraph id="8BF836E8FFE0FFE791F6FC9DFABDA3F9" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FC9DFE26A306" box="[199,478,800,824]" italics="true" pageId="19" pageNumber="320">Structures and sculpture</emphasis>
. Head smooth, shiny, with fine microreticulation of rounded cells. Pronotum with microreticulation as on head, shiny, with faint iridescence. Smooth, lacking punctures except along hind and fore margins. Elytra apically not markedly pointed and subapical sinuation poorly or not pronounced; surface smooth, with punctures very scarce or lacking and often with few, very shallow, gnarled scratches; reticulation poorly impressed; iridescence well visible. Prosternal process and lobes of metacoxal processes as in
<taxonomicName id="4C474D6BFFE0FFE79586FC0CFAC6A3F6" box="[1207,1342,944,968]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE79586FC0CFAC6A3F6" box="[1207,1342,944,968]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF836E8FFE0FFE791F6FC69FE0AA49E" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FC69FEF9A3D2" box="[199,257,980,1004]" italics="true" pageId="19" pageNumber="320">Male</emphasis>
. Last abdominal ventrite tectiform, with antero-lateral sides markedly sinuate. Outer claw of forelegs scimitar-shaped. Median lobe of aedeagus very similar to that of
<taxonomicName id="4C474D6BFFE0FFE7924AFC44FC17A42E" box="[891,1007,1016,1040]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFE0FFE7924AFC44FC17A42E" box="[891,1007,1016,1040]" italics="true" pageId="19" pageNumber="320">L. pumilio</emphasis>
</taxonomicName>
except as follows: in left lateral view (
<figureCitation id="137C2A6DFFE0FFE791AEFBA1FF0AA40A" box="[159,242,1052,1077]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="19" pageNumber="320">Fig. 89</figureCitation>
a, b) more straight and robust, with base and distal portion forming a more obtuse angle; apex ending in a wider expansion, laterally flattened; in dorsal view (
<figureCitation id="137C2A6DFFE0FFE793C4FB82FCAFA466" box="[757,855,1087,1112]" captionStart="FIGURES 107 – 115" captionStartId="46.[151,264,1951,1973]" captionTargetBox="[184,1390,196,1925]" captionTargetId="figure@46.[170,1417,193,1930]" captionTargetPageId="46" captionText="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri="https://zenodo.org/record/244319/files/figure.png" pageId="19" pageNumber="320">Fig. 112</figureCitation>
) more gently attenuate at tip, almost straight, and dorsal lamina more strongly sinuate. Left paramere (
<figureCitation id="137C2A6DFFE0FFE793DBFBD9FCC4A442" box="[746,828,1124,1149]" captionStart="FIGURES 88 – 90" captionStartId="41.[151,264,1919,1941]" captionTargetBox="[225,1366,198,1894]" captionTargetId="figure@41.[204,1370,193,1898]" captionTargetPageId="41" captionText="FIGURES 88 – 90. Male genitalia in Laccomimus species, left lateral view: 88) L. amazonas, a) median lobe, b) left paramere, c) right paramere; 89) L. improvidus, a) median lobe, b) magnification of apex of median lobe, c) left paramere; 90) L. youngi, a) median lobe (holotype), b) magnification of apex of median lobe (holotype), c) magnification of apex of median lobe (different specimen from type locality), d) left paramere (holotype), e) left paramere (different specimen from type locality)." httpUri="https://zenodo.org/record/244314/files/figure.png" pageId="19" pageNumber="320">Fig. 89</figureCitation>
c) very similar to that of
<taxonomicName id="4C474D6BFFE0FFE79566FBD8FB18A442" box="[1111,1248,1124,1148]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE79566FBD8FB18A442" box="[1111,1248,1124,1148]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
, although not or very poorly thickened apically.
</paragraph>
<paragraph id="8BF836E8FFE0FFE791F6FB11FC3CA4D6" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FB11FEE3A4FA" box="[199,283,1196,1220]" italics="true" pageId="19" pageNumber="320">Female</emphasis>
. Elytra duller than in males, with more evident iridescence. Last abdominal ventrite triangular, very narrowly rounded to subtruncate at apex. Genital pieces as in
<taxonomicName id="4C474D6BFFE0FFE7927BFB6CFC45A4D6" box="[842,957,1232,1256]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFE0FFE7927BFB6CFC45A4D6" box="[842,957,1232,1256]" italics="true" pageId="19" pageNumber="320">L. pumilio</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE0FFE791F6FB49FDCEA56A" pageId="19" pageNumber="320" type="distribution">
<paragraph id="8BF836E8FFE0FFE791F6FB49FCCFA532" blockId="19.[151,1437,151,1725]" box="[199,823,1268,1293]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FB49FE98A533" bold="true" box="[199,352,1268,1293]" pageId="19" pageNumber="320">Distribution.</emphasis>
<collectingCountry id="F3507678FFE0FFE79056FB49FE25A533" box="[359,477,1268,1293]" name="Venezuela" pageId="19" pageNumber="320">Venezuela</collectingCountry>
,
<collectingCountry id="F3507678FFE0FFE790D9FB49FDBAA532" box="[488,578,1268,1292]" name="Guyana" pageId="19" pageNumber="320">Guyana</collectingCountry>
,
<collectingCountry id="F3507678FFE0FFE7937CFB49FD40A533" box="[589,696,1268,1293]" name="Suriname" pageId="19" pageNumber="320">Suriname</collectingCountry>
(
<figureCitation id="137C2A6DFFE0FFE793F9FB49FCD2A532" box="[712,810,1268,1293]" captionStart="FIGURES 126 – 129" captionStartId="49.[151,264,1953,1975]" captionTargetBox="[157,1432,197,1931]" captionTargetId="figure@49.[151,1436,193,1932]" captionTargetPageId="49" captionText="FIGURES 126 – 129. Distribution maps of Laccomimus species: 126) L. bordoni in South America; 127) L. bolivari (full circles), L. youngi (single open square in Brazil); 128) L. pumilio (open circles), L. alvarengi in Central America (single triangle in Panamá), L. spangleri (stars), L. bordoni in Central America (asterisks); 129) L. improvidus." httpUri="https://zenodo.org/record/244322/files/figure.png" pageId="19" pageNumber="320">Fig. 129</figureCitation>
).
</paragraph>
<paragraph id="8BF836E8FFE0FFE791F6FAAAFDCEA56A" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FAAAFED2A50E" bold="true" box="[199,298,1303,1328]" pageId="19" pageNumber="320">Habitat.</emphasis>
In Guarico State (
<collectingCountry id="F3507678FFE0FFE79338FAAAFD78A50E" box="[521,640,1303,1328]" name="Venezuela" pageId="19" pageNumber="320">Venezuela</collectingCountry>
) some specimens were collected in a large lagoon with vegetation, exposed to direct sunlight (
<figureCitation id="137C2A6DFFE0FFE790F6FA81FDD1A56A" box="[455,553,1340,1365]" captionStart="FIGURES 132 – 133" captionStartId="51.[151,264,1953,1975]" captionTargetBox="[151,1436,193,1930]" captionTargetId="figure@51.[151,1436,193,1932]" captionTargetPageId="51" captionText="FIGURES 132 – 133. Habitats of Laccomimus species: 132) L. distinctus, ditch at El Cachapé refuge, Chaco Province, Argentina (photo M. C. Michat); 133) L. improvidus and L. bolivari, large lagoon with vegetation ca. 65 km S Las Mercedes, Guarico State, Venezuela (photo A. E. Z. Short)." httpUri="https://zenodo.org/record/244323/files/figure.png" pageId="19" pageNumber="320">Fig. 133</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE0FFE791F6FAE2FF17A5A3" pageId="19" pageNumber="320" type="etymology">
<paragraph id="8BF836E8FFE0FFE791F6FAE2FF17A5A3" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FAE2FEB6A546" bold="true" box="[199,334,1375,1400]" pageId="19" pageNumber="320">Etymology.</emphasis>
Latin adjective for “surprising”, “unexpected”; quite late, in fact, we recognised it as a separate species.
</paragraph>
</subSubSection>
<subSubSection id="C35D6563FFE0FFE791F6FA1AFE6EA683" pageId="19" pageNumber="320" type="discussion">
<paragraph id="8BF836E8FFE0FFE791F6FA1AFE6EA683" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="320">
<emphasis id="B933EAFAFFE0FFE791F6FA1AFEC3A5FE" bold="true" box="[199,315,1447,1472]" pageId="19" pageNumber="320">Remarks.</emphasis>
This species is close to
<taxonomicName id="4C474D6BFFE0FFE79378FA14FD44A5FE" box="[585,700,1448,1472]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B933EAFAFFE0FFE79378FA14FD44A5FE" box="[585,700,1448,1472]" italics="true" pageId="19" pageNumber="320">L. pumilio</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D6BFFE0FFE793C2FA14FC82A5FE" box="[755,890,1448,1472]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE793C2FA14FC82A5FE" box="[755,890,1448,1472]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
. At first, after dissecting the first male specimen (
<collectingCountry id="F3507678FFE0FFE791AFFA71FEF6A5DB" box="[158,270,1484,1509]" name="Suriname" pageId="19" pageNumber="320">Suriname</collectingCountry>
, Paramaribo), we were convinced that it was a dark specimen of
<taxonomicName id="4C474D6BFFE0FFE792C6FA70FB78A5DA" box="[1015,1152,1484,1508]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE792C6FA70FB78A5DA" box="[1015,1152,1484,1508]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
with an aberrant median lobe. We also considered that it could represent a possible case of hybridisation between
<taxonomicName id="4C474D6BFFE0FFE79588FA4CFABDA636" box="[1209,1349,1520,1544]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE79588FA4CFABDA636" box="[1209,1349,1520,1544]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D6BFFE0FFE794B4FA4CFF08A612" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B933EAFAFFE0FFE794B4FA4CFF08A612" italics="true" pageId="19" pageNumber="320">L. bolivari</emphasis>
</taxonomicName>
since the median lobe appears as a broader tip added to an
<taxonomicName id="4C474D6BFFE0FFE792B6F9A8FBF6A612" box="[903,1038,1556,1580]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="alvarengi">
<emphasis id="B933EAFAFFE0FFE792B6F9A8FBF6A612" box="[903,1038,1556,1580]" italics="true" pageId="19" pageNumber="320">L. alvarengi</emphasis>
</taxonomicName>
median lobe. Both species occur at that locality. Additional material found in USNM and NMW, however, made it clear that the specimens belong to an undescribed species, very uniform in elytral colouration and in aedeagal features. Apparently it is a rare species, at present known only from few specimens, often found together with other species. The females are impossible to identify with certainty.
</paragraph>
</subSubSection>
</treatment>
</document>
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