treatments-xml/data/03/EE/87/03EE87A6A025FFD639FC6E2BFD71B6CA.xml

<document ID-DOI="10.1590/S1679-62252008000300006" ID-ISSN="1982-0224" ID-Zenodo-Dep="5419740" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1630375433146" checkinUser="carolina" docAuthor="Carvalho, Tiago Pinto, A., Pablo Lehmann &amp; Reis, Roberto E." docDate="2008" docId="03EE87A6A025FFD639FC6E2BFD71B6CA" docLanguage="en" docName="NeotropIchthyol.2008.31.pdf" docOrigin="Neotropical Ichthyology 6 (3)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252008000300006&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Gymnotocinclus anosteos Carvalho &amp; A. &amp; Reis 2008, new species" docType="treatment" docVersion="2" lastPageId="7" lastPageNumber="336" masterDocId="FFD7FFDEA024FFD13A516C39FF91B633" masterDocTitle="Gymnotocinclus anosteos, a new uniquely-plated genus and species of loricariid catfish (Teleostei: Siluriformes) from the upper rio Tocantins basin, central Brazil" masterLastPageNumber="338" masterPageNumber="329" pageId="1" pageNumber="330" updateTime="1630683736786" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Gymnotocinclus anosteos, a new uniquely-plated genus and species of loricariid catfish (Teleostei: Siluriformes) from the upper rio Tocantins basin, central Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carvalho, Tiago Pinto</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>A., Pablo Lehmann</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reis, Roberto E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>329</mods:start>
<mods:end>338</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252008000300006&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252008000300006</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">5419740</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03EE87A6A025FFD639FC6E2BFD71B6CA" httpUri="http://treatment.plazi.org/id/03EE87A6A025FFD639FC6E2BFD71B6CA" lastPageId="7" lastPageNumber="336" pageId="1" pageNumber="330">
<subSubSection pageId="1" pageNumber="330" type="nomenclature">
<paragraph blockId="1.[941,1364,530,585]" pageId="1" pageNumber="330">
<emphasis bold="true" box="[941,1364,530,554]" pageId="1" pageNumber="330">
<taxonomicName authority="Carvalho &amp; A. &amp; Reis, 2008" authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" box="[941,1223,530,554]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="330" phylum="Chordata" rank="species" species="anosteos" status="new species">
<emphasis bold="true" box="[941,1223,530,554]" italics="true" pageId="1" pageNumber="330">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1234,1364,530,553]" pageId="1" pageNumber="330" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation box="[1107,1198,562,585]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[158,197,1920,1943]" captionStartId-1="3.[151,190,962,985]" captionTargetBox-0="[158,1481,538,1896]" captionTargetBox-1="[171,1462,199,909]" captionTargetId-0="figure-194@2.[158,1481,538,1896]" captionTargetId-1="figure-159@3.[151,1475,194,945]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Gymnotocinclus anosteos, holotype, MCN 18624, male, 44.3 mm SL, ribeirão das Cobras, near highway GO-327, tributary of rio Tocantinzinho, rio Tocantins Basin, Alto Paraíso de Goiás, Goiás, Brazil." captionText-1="Fig. 2. Gymnotocinclus anosteos, cleared and stained paratype, MCP 41726, male, 40.5 mm SL, collected with the holotype." figureDoi-0="http://doi.org/10.5281/zenodo.5419742" figureDoi-1="http://doi.org/10.5281/zenodo.5419744" httpUri-0="https://zenodo.org/record/5419742/files/figure.png" httpUri-1="https://zenodo.org/record/5419744/files/figure.png" pageId="1" pageNumber="330">Figs. 1-2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="330" type="materials_examined">
<paragraph blockId="1.[831,1474,626,992]" pageId="1" pageNumber="330">
<materialsCitation accessionNumber="MCN 18624" collectingDate="1996-11-09" collectionCode="MCN, R" collectorName="das Cobras &amp; Koch &amp; K. M. Grosser." country="Brazil" latitude="-14.161944" location="rio Tocantinzinho" longLatPrecision="21" longitude="-47.6325" municipality="Alto Paraiso" pageId="1" pageNumber="330" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
<emphasis bold="true" box="[831,932,626,647]" pageId="1" pageNumber="330">
<typeStatus box="[831,928,626,647]" pageId="1" pageNumber="330">Holotype</typeStatus>
.
</emphasis>
<accessionNumber box="[941,1068,626,647]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCN18624" pageId="1" pageNumber="330">
<collectionCode box="[941,996,626,647]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">MCN</collectionCode>
18624
</accessionNumber>
, 
<quantity box="[1081,1171,626,647]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" pageId="1" pageNumber="330" unit="mm" value="44.3">44.3 mm</quantity>
SL, male, 
<collectingCountry box="[1285,1347,626,647]" name="Brazil" pageId="1" pageNumber="330">Brazil</collectingCountry>
, 
<collectingRegion box="[1360,1419,626,647]" country="Brazil" name="Goias" pageId="1" pageNumber="330">Goiás</collectingRegion>
, 
<collectingMunicipality pageId="1" pageNumber="330">Alto Paraíso</collectingMunicipality>
de 
<collectingRegion box="[937,996,654,675]" country="Brazil" name="Goias" pageId="1" pageNumber="330">Goiás</collectingRegion>
, ribeirão 
<collectorName box="[1088,1198,654,675]" pageId="1" pageNumber="330">das Cobras</collectorName>
, near to road GO-239, tributary to 
<location LSID="urn:lsid:plazi:treatment:03EE87A6A025FFD639FC6E2BFD71B6CA:8E98606BA025FFD039C26E92FBC7B4F3" box="[915,1110,683,704]" country="Brazil" latitude="-14.161944" longLatPrecision="21" longitude="-47.6325" municipality="Alto Paraiso" name="rio Tocantinzinho" pageId="1" pageNumber="330" stateProvince="Goias">rio Tocantinzinho</location>
, rio 
<collectingRegion box="[1165,1268,683,704]" country="Brazil" name="Tocantins" pageId="1" pageNumber="330">Tocantins</collectingRegion>
basin, 
<geoCoordinate box="[1352,1474,683,704]" degrees="14" direction="south" minutes="09" orientation="latitude" pageId="1" pageNumber="330" precision="15" seconds="43" value="-14.161944">14º09’43&quot;S</geoCoordinate>
<geoCoordinate box="[831,952,712,733]" degrees="47" direction="west" minutes="37" orientation="longitude" pageId="1" pageNumber="330" precision="15" seconds="57" value="-47.6325">47º37’57&quot;W</geoCoordinate>
, 
<date box="[963,1081,712,733]" pageId="1" pageNumber="330" value="1996-11-09">
<collectingDate box="[963,1081,712,733]" pageId="1" pageNumber="330" value="1996-11-09">9 Nov 1996</collectingDate>
</date>
, W. 
<collectionCode box="[1124,1143,712,733]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="330">R</collectionCode>
. 
<collectorName box="[1152,1205,712,733]" pageId="1" pageNumber="330">Koch</collectorName>
&amp; 
<collectorName box="[1237,1382,712,733]" pageId="1" pageNumber="330">K. M. Grosser.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="1.[831,1474,626,992]" pageId="1" pageNumber="330">
<materialsCitation accessionNumber="MCN 13543, MCP 41726, MCN 13154, MCN 13155, MCN 13538" collectingDate="1996-08-04" collectingDateMax="1996-11-09" collectingDateMin="1996-08-04" collectionCode="MCN, MCP, R" collectorName="Koch, P. C. C &amp; K. M. Grosser. &amp; Koch &amp; K. M. Grosser" country="Brazil" county="All" latitude="-14.139444" location="rio Tocantinzinho" longLatPrecision="21" longitude="-47.662502" municipality="Alto Paraiso" pageId="1" pageNumber="330" specimenCode="12, 5, 7, 6, ANSP 187156, 1" specimenCount="2" stateProvince="Goias" typeStatus="paratype">
<emphasis bold="true" box="[831,941,741,762]" pageId="1" pageNumber="330">
<typeStatus box="[831,937,741,762]" pageId="1" pageNumber="330">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty box="[949,980,741,762]" pageId="1" pageNumber="330">All</collectingCounty>
from 
<collectingCountry box="[1043,1105,741,762]" name="Brazil" pageId="1" pageNumber="330">Brazil</collectingCountry>
, 
<collectingRegion box="[1117,1176,741,762]" country="Brazil" name="Goias" pageId="1" pageNumber="330">Goiás</collectingRegion>
, 
<collectingMunicipality box="[1186,1310,741,762]" pageId="1" pageNumber="330">Alto Paraíso</collectingMunicipality>
de 
<collectingRegion box="[1348,1408,741,762]" country="Brazil" name="Goias" pageId="1" pageNumber="330">Goiás</collectingRegion>
. 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MCN13543" pageId="1" pageNumber="330">
<collectionCode box="[1419,1474,741,762]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">MCN</collectionCode>
13543
</accessionNumber>
, 
<specimenCode box="[903,929,770,791]" collectionCode="MCN" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">12</specimenCode>
, 
<quantity box="[939,1075,770,791]" metricMagnitude="-2" metricUnit="m" metricValue="3.255" metricValueMax="3.9299999999999997" metricValueMin="2.58" pageId="1" pageNumber="330" unit="mm" value="32.55" valueMax="39.3" valueMin="25.8">25.8-39.3 mm</quantity>
SL and 
<accessionNumber box="[1153,1272,770,791]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP41726" pageId="1" pageNumber="330">
<collectionCode box="[1153,1204,770,791]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="1" pageNumber="330">MCP</collectionCode>
41726
</accessionNumber>
, 
<specimenCode box="[1281,1295,770,791]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="1" pageNumber="330">5</specimenCode>
, 
<quantity box="[1304,1441,770,791]" metricMagnitude="-2" metricUnit="m" metricValue="3.185" metricValueMax="4.16" metricValueMin="2.21" pageId="1" pageNumber="330" unit="mm" value="31.85" valueMax="41.6" valueMin="22.1">22.1-41.6 mm</quantity>
SL + 3 c&amp;s, 
<quantity box="[931,1073,798,819]" metricMagnitude="-2" metricUnit="m" metricValue="3.145" metricValueMax="4.05" metricValueMin="2.2399999999999998" pageId="1" pageNumber="330" unit="mm" value="31.45" valueMax="40.5" valueMin="22.4">22.4-40.5 mm</quantity>
SL, collected with the 
<typeStatus box="[1317,1406,798,819]" pageId="1" pageNumber="330">holotype</typeStatus>
. 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MCN13154" pageId="1" pageNumber="330">
<collectionCode box="[1419,1474,798,819]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">MCN</collectionCode>
13154
</accessionNumber>
, 
<specimenCode box="[903,917,827,848]" collectionCode="MCN" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">7</specimenCode>
, 
<quantity box="[926,1061,827,848]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" metricValueMax="3.4299999999999997" metricValueMin="1.63" pageId="1" pageNumber="330" unit="mm" value="25.299999999999997" valueMax="34.3" valueMin="16.3">16.3-34.3 mm</quantity>
SL, 
<accessionNumber box="[1105,1228,827,848]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCN13155" pageId="1" pageNumber="330">
<collectionCode box="[1105,1160,827,848]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">MCN</collectionCode>
13155
</accessionNumber>
, 
<specimenCode box="[1236,1250,827,848]" collectionCode="MCN" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">6</specimenCode>
, 19.0- 
<quantity box="[1308,1394,827,848]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="1" pageNumber="330" unit="mm" value="31.6">31.6 mm</quantity>
SL, and 
<specimenCode box="[831,975,856,877]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1" pageNumber="330">ANSP 187156</specimenCode>
, 
<specimenCode box="[986,1000,856,877]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1" pageNumber="330">5</specimenCode>
, 
<quantity box="[1012,1150,856,877]" metricMagnitude="-2" metricUnit="m" metricValue="3.165" metricValueMax="4.08" metricValueMin="2.25" pageId="1" pageNumber="330" unit="mm" value="31.65" valueMax="40.8" valueMin="22.5">22.5-40.8 mm</quantity>
SL, same locality as holotype, 
<date pageId="1" pageNumber="330" value="1996-08-04">
<collectingDate pageId="1" pageNumber="330" value="1996-08-04">4 Aug 1996</collectingDate>
</date>
, W. 
<collectionCode box="[979,998,885,906]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="330">R</collectionCode>
. 
<collectorName box="[1009,1151,885,906]" pageId="1" pageNumber="330">Koch, P. C. C</collectorName>
. Milani &amp; 
<collectorName box="[1262,1412,885,906]" pageId="1" pageNumber="330">K. M. Grosser.</collectorName>
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MCN13538" pageId="1" pageNumber="330">
<collectionCode box="[1419,1474,885,906]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">MCN</collectionCode>
13538
</accessionNumber>
, 
<specimenCode box="[908,920,914,935]" collectionCode="MCN" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="1" pageNumber="330" type="Herbarium">1</specimenCode>
, 
<quantity box="[936,1026,914,935]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="1" pageNumber="330" unit="mm" value="28.1">28.1 mm</quantity>
SL, creek on road GO-239, tributary to 
<location LSID="urn:lsid:plazi:treatment:03EE87A6A025FFD639FC6E2BFD71B6CA:8E98606BA025FFD03FF66FABFC5DB5F0" country="Brazil" county="All" latitude="-14.139444" longLatPrecision="21" longitude="-47.662502" municipality="Alto Paraiso" name="rio Tocantinzinho" pageId="1" pageNumber="330" stateProvince="Goias">rio Tocantinzinho</location>
, rio 
<collectingRegion box="[1011,1104,942,963]" country="Brazil" name="Tocantins" pageId="1" pageNumber="330">Tocantins</collectingRegion>
basin, 
<geoCoordinate box="[1171,1282,942,963]" degrees="14" direction="south" minutes="08" orientation="latitude" pageId="1" pageNumber="330" precision="15" seconds="22" value="-14.139444">14º08’22&quot;S</geoCoordinate>
<geoCoordinate box="[1287,1407,942,963]" degrees="47" direction="west" minutes="39" orientation="longitude" pageId="1" pageNumber="330" precision="15" seconds="45" value="-47.662502">47º39’45&quot;W</geoCoordinate>
, 
<date pageId="1" pageNumber="330" value="1996-11-09">
<collectingDate pageId="1" pageNumber="330" value="1996-11-09">9 Nov 1996</collectingDate>
</date>
, W. 
<collectionCode box="[926,945,971,992]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="330">R</collectionCode>
. 
<collectorName box="[955,1008,971,992]" pageId="1" pageNumber="330">Koch</collectorName>
&amp; 
<collectorName box="[1040,1183,971,992]" pageId="1" pageNumber="330">K. M. Grosser</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[831,1158,1029,1052]" pageId="1" pageNumber="330" type="diagnosis">
<paragraph blockId="1.[831,1158,1029,1052]" box="[831,1158,1029,1052]" pageId="1" pageNumber="330">
<emphasis bold="true" box="[831,949,1029,1052]" pageId="1" pageNumber="330">Diagnosis.</emphasis>
Same as for genus.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="335" pageId="1" pageNumber="330" type="description">
<paragraph blockId="1.[831,1475,1087,1974]" pageId="1" pageNumber="330">
<emphasis bold="true" box="[831,972,1087,1110]" pageId="1" pageNumber="330">Description.</emphasis>
Morphometrics and meristics given in 
<tableCitation box="[1385,1473,1087,1110]" captionStart="Table 1" captionStartId="4.[839,904,632,655]" captionText="Table 1. Descriptive morphometrics of Gymnotocinclus anosteos. Values are given for holotype (H) and ranges of 20 specimens including the holotype. SD = standard deviation." pageId="1" pageNumber="330">Tables 1</tableCitation>
and 
<tableCitation box="[877,890,1119,1142]" captionStart="Table 2" captionStartId="6.[158,221,197,220]" captionText="Table 2. Frequency distribution for meristics of Gymnotocinclus anosteos. Holotype values are marked with an asterisk. N = number of specimens." pageId="1" pageNumber="330">2</tableCitation>
respectively. Dorsal profile gently curved from snout tip to parieto-supraoccipital bone, almost straight from that point to dorsal-fin origin. Slightly concave to straight at dorsal-fin base. Straight from end of dorsal-fin base to caudal-fin origin. Ventral profile straight to slightly convex from snout tip to posterior border of lip, straight and posteroventrally sloped from that point to origin of pelvic fin, straight and posterodorsally inclined from pelvic fin to caudal-fin origin. Body flattened at predorsal region. Ventral surface of trunk flattened at base of paired fins. Caudal peduncle round in cross section. Greatest body depth at dorsal-fin origin. Least body depth at posterior end of caudal peduncle. Head and snout broad, snout anterior margin rounded in dorsal view. Body progressively narrowing posteriorly from cleithrum. Head flattened at parieto-supraoccipital, interorbital region straight. Upper margin of orbit not elevated. Eye small, dorsolaterally placed. Iris operculum small, less than half pupil diameter.
</paragraph>
<paragraph blockId="1.[831,1475,1087,1974]" lastBlockId="2.[839,1482,194,505]" lastPageId="2" lastPageNumber="331" pageId="1" pageNumber="330">
Pectoral fin I,6, posterior margin slightly rounded; when depressed reaching to middle of adpressed pelvic fin. Posterior margin of pectoral-fin spine smooth. Soft portion of pectoral spine large, about one-fourth of spine length. Pectoralfin axillary slit absent. Coracoid and cleithrum exposed laterally, supporting odontodes only in restricted lateral area near base of pectoral fin; covered by skin medially (
<figureCitation box="[1324,1389,1855,1878]" captionStart="Fig" captionStartId="4.[158,197,1955,1978]" captionTargetBox="[171,1481,1284,1902]" captionTargetId="figure-141@4.[158,1482,1275,1940]" captionTargetPageId="4" captionText="Fig. 6. Pectoral girdle in ventral view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419752" httpUri="https://zenodo.org/record/5419752/files/figure.png" pageId="1" pageNumber="330">Fig. 6</figureCitation>
). Pelvic fin i,5, reaching anal-fin origin when depressed. Pelvic girdle with slender lateropterygium, club shaped, expanded proximally and thinner distally. Basipterygium ossified anteriorly, lacking anterior fenestrae (
<figureCitation box="[453,522,194,217]" captionStart="Fig" captionStartId="5.[151,190,961,984]" captionTargetBox="[358,1288,193,902]" captionTargetId="figure-11@5.[336,1315,185,917]" captionTargetPageId="5" captionText="Fig. 7. Pelvic girdle in dorsal view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419754" httpUri="https://zenodo.org/record/5419754/files/figure.png" pageId="2" pageNumber="331">Fig. 7</figureCitation>
). Dorsal fin II,7; spinelet present, reduced in size, quadrangular in shape. Nuchal plate present, triangular in shape. Dorsal-fin origin located posterior to vertical through pelvic-fin origin.Anal fin i,5, first analfin pterygiophore not exposed on ventral surface anterior to first anal-fin unbranched ray. Bifid hemal spines absent from abdominal vertebrae posterior to first anal-fin proximal radial. Caudal fin i,14,i. Compound hypurals 1 and 2 almost completely fused to the compound hypurals 3-5, posterior margin of caudal-fin skeleton with slight median notch. Total vertebrae 28-29 (3 c&amp;s). Adipose fin absent, single series of three to six platelets at adipose fin position.
</paragraph>
<paragraph blockId="2.[839,1482,194,505]" lastBlockId="3.[831,1474,1729,1976]" lastPageId="3" lastPageNumber="332" pageId="2" pageNumber="331">
Head plates present, including postrostral plates, infraorbital series, cheek plates, and small scattered plates anterior to nostril. Body almost completely naked, without dermal plates (
<figureCitation box="[918,986,354,377]" captionStart="Fig" captionStartId="3.[151,190,962,985]" captionTargetBox="[171,1462,199,909]" captionTargetId="figure-159@3.[151,1475,194,945]" captionTargetPageId="3" captionText="Fig. 2. Gymnotocinclus anosteos, cleared and stained paratype, MCP 41726, male, 40.5 mm SL, collected with the holotype." figureDoi="http://doi.org/10.5281/zenodo.5419744" httpUri="https://zenodo.org/record/5419744/files/figure.png" pageId="2" pageNumber="331">Fig. 2</figureCitation>
), except for small platelets scattered all over body dorsal surface, not forming regular series; small platelets at lateral line composing median series; about six large plates on anterior portion of mid-ventral series; and last plates of ventral and dorsal series from adipose fin region to posterior end of caudal peduncle. Abdomen with scattered platelets irregularly arranged between posterior margin of coracoid and region anterior to anal-fin origin; abdominal platelets absent near pelvic-fin base. Fourth infraorbital not expanded ventrally, naked area below infraorbitals four and five and anterior to preopercle (
<figureCitation box="[510,579,1826,1849]" captionStart="Fig" captionStartId="5.[151,190,1954,1977]" captionTargetBox="[195,1416,1087,1919]" captionTargetId="figure-29@5.[151,1475,1067,1933]" captionTargetPageId="5" captionText="Fig. 8. Skull in lateral view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419756" httpUri="https://zenodo.org/record/5419756/files/figure.png" pageId="3" pageNumber="332">Fig. 8</figureCitation>
). Infraorbital canal entering infraorbital series via sphenotic. Canal-bearing cheek plate not extending medially in ventral view. Lateral line complete, without midbody gap. Raised odontodes absent from parieto-supraoccipital in adults, smaller juveniles with three raised crests of odontodes on posterior portion of parieto-supraoccipital. Odontodes on trunk with uniform size and distribution, not arranged into distinct rows, except for larger odontodes on ventral surface of pectoral-fin spine, and first pelvic- and anal-fin unbranched rays. Odontodes of pelvic fin turned mesially.Tip of odontodes on last lateral plates curved dorsally or anteriorly (
<figureCitation box="[1060,1123,1921,1944]" captionStart="Fig" captionStartId="3.[831,870,1497,1520]" captionTargetBox="[834,1476,1007,1476]" captionTargetId="figure-200@3.[834,1476,1007,1476]" captionTargetPageId="3" captionText="Fig. 4. Scanning electron microscope image of dermal plates at posterior end of caudal peduncle. Gymnotocinclus anosteos, paratype, MCP 41726, male, 40.5 mm SL. LLP = lateral line plates, VPS = ventral plate series, DPS = dorsal plate series.Arrow indicates the modified odontode tips of the last plates on caudal peduncle." figureDoi="http://doi.org/10.5281/zenodo.5419746" httpUri="https://zenodo.org/record/5419746/files/figure.png" pageId="3" pageNumber="332">Fig. 4</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419742" ID-Zenodo-Dep="5419742" httpUri="https://zenodo.org/record/5419742/files/figure.png" pageId="2" pageNumber="331" startId="2.[158,197,1920,1943]" targetBox="[158,1481,538,1896]" targetPageId="2">
<paragraph blockId="2.[158,1481,1920,1975]" pageId="2" pageNumber="331">
<emphasis bold="true" box="[158,230,1920,1943]" pageId="2" pageNumber="331">Fig. 1.</emphasis>
<taxonomicName authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" box="[238,515,1920,1943]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="331" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[238,515,1920,1943]" italics="true" pageId="2" pageNumber="331">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
, holotype, MCN 18624, male, 44.3 mm SL, ribeirão das Cobras, near highway GO-327, tributary of rio Tocantinzinho, rio Tocantins Basin, Alto Paraíso de Goiás, Goiás, Brazil.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419744" ID-Zenodo-Dep="5419744" box="[151,1468,962,985]" httpUri="https://zenodo.org/record/5419744/files/figure.png" pageId="3" pageNumber="332" startId="3.[151,190,962,985]" targetBox="[171,1462,199,909]" targetPageId="3">
<paragraph blockId="3.[151,1468,962,985]" box="[151,1468,962,985]" pageId="3" pageNumber="332">
<emphasis bold="true" box="[151,220,962,985]" pageId="3" pageNumber="332">Fig. 2.</emphasis>
<taxonomicName authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" box="[226,500,962,985]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="332" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[226,500,962,985]" italics="true" pageId="3" pageNumber="332">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
, cleared and stained paratype, MCP 41726, male, 40.5 mm SL, collected with the holotype.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419746" ID-Zenodo-Dep="5419746" httpUri="https://zenodo.org/record/5419746/files/figure.png" pageId="3" pageNumber="332" startId="3.[831,870,1497,1520]" targetBox="[834,1476,1007,1476]" targetPageId="3">
<paragraph blockId="3.[831,1474,1497,1680]" pageId="3" pageNumber="332">
<emphasis bold="true" box="[831,900,1497,1520]" pageId="3" pageNumber="332">Fig. 4.</emphasis>
Scanning electron microscope image of dermal plates at posterior end of caudal peduncle. 
<taxonomicName authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="332" phylum="Chordata" rank="species" species="anosteos">
<emphasis italics="true" pageId="3" pageNumber="332">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
, paratype, MCP 41726, male, 40.5 mm SL. LLP = lateral line plates, VPS = ventral plate series, DPS = dorsal plate series. Arrow indicates the modified odontode tips of the last plates on caudal peduncle.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419748" ID-Zenodo-Dep="5419748" httpUri="https://zenodo.org/record/5419748/files/figure.png" pageId="3" pageNumber="332" startId="3.[151,191,1550,1573]" targetBox="[178,768,1018,1513]" targetPageId="3">
<paragraph blockId="3.[151,793,1550,1637]" pageId="3" pageNumber="332">
<emphasis bold="true" box="[151,227,1550,1573]" pageId="3" pageNumber="332">Fig. 3.</emphasis>
Posterior abdominal vertebrae of 
<taxonomicName authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="332" phylum="Chordata" rank="species" species="anosteos">
<emphasis italics="true" pageId="3" pageNumber="332">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
lacking bifid hemal spines. MCP 41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<paragraph blockId="3.[831,1474,1729,1976]" lastBlockId="4.[839,1482,194,601]" lastPageId="4" lastPageNumber="333" pageId="3" pageNumber="332">
Anterior margin of mesethmoid rounded, not expanded laterally. Ventral surface of mesethmoid anterior to ventral articular condyle simple, without anteroventral lamina. Vertical lateral ethmoid ridge elevated, posterior portion contacting metapterygoid. Lateral ethmoid encapsulating nasal organ, with small longitudinal opening at lateral border of nasal capsule (
<figureCitation box="[846,913,354,377]" captionStart="Fig" captionStartId="4.[158,200,1113,1136]" captionTargetBox="[203,787,198,1039]" captionTargetId="figure-121@4.[156,805,158,1079]" captionTargetPageId="4" captionText="Fig. 5. Left side of neurocranium in ventral view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419750" httpUri="https://zenodo.org/record/5419750/files/figure.png" pageId="4" pageNumber="333">Fig. 5</figureCitation>
). Lateral ethmoid exposed on dorsal surface of head posterior to nostrils supporting one series of odontodes (
<figureCitation captionStart="Fig" captionStartId="5.[151,190,1954,1977]" captionTargetBox="[195,1416,1087,1919]" captionTargetId="figure-29@5.[151,1475,1067,1933]" captionTargetPageId="5" captionText="Fig. 8. Skull in lateral view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419756" httpUri="https://zenodo.org/record/5419756/files/figure.png" pageId="4" pageNumber="333">Fig. 8</figureCitation>
). Hyomandibula articular condyle contacting prootic. Compound pterotic not enlarged posterolaterally, posterior portion somewhat rounded, not pointed. Anteroventral margin of compound pterotic with irregularly arranged perforations of median to large size. Swimbladder capsule relatively enlarged, its posterior margin extending to joint of Weberian complex with
</paragraph>
<caption pageId="4" pageNumber="333" startId="4.[839,904,632,655]">
<paragraph blockId="4.[839,1482,632,719]" pageId="4" pageNumber="333">
<emphasis bold="true" box="[839,935,632,655]" pageId="4" pageNumber="333">Table 1.</emphasis>
Descriptive morphometrics of 
<taxonomicName authorityName="Carvalho &amp; A. &amp; Reis" authorityYear="2008" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="anosteos">
<emphasis italics="true" pageId="4" pageNumber="333">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
. Values are given for holotype (H) and ranges of 20 specimens including the holotype. SD = standard deviation.
</paragraph>
</caption>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1460,749,768]" pageId="4" pageNumber="333">Character H N Low High Mean SD</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" pageId="4" pageNumber="333">Standard length (mm) 44.3 20 28.8 44.3 35.7 Percentages of standard length</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,823,842]" pageId="4" pageNumber="333">Head length 33.0 20 32.5 34.4 33.3 0.56</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,848,867]" pageId="4" pageNumber="333">Predorsal length 45.7 20 43.5 48.4 46.1 1.03</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,873,892]" pageId="4" pageNumber="333">Dorsal-fin spine length 20.7 20 19.2 22.6 20.9 0.86</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,897,916]" pageId="4" pageNumber="333">Anal-fin unbranched ray length 17.8 20 16.3 20.3 18.0 1.02</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,921,940]" pageId="4" pageNumber="333">Pectoral-fin spine length 20.6 19 16.6 23.7 20.3 1.72</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,946,965]" pageId="4" pageNumber="333">Pelvic-fin unbranched ray length 20.0 20 15.7 20.5 18.0 1.47</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,970,989]" pageId="4" pageNumber="333">Cleithral width 26.7 20 24.6 27.0 25.8 0.71</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,995,1014]" pageId="4" pageNumber="333">Trunk length 17.0 20 15.2 17.3 16.2 0.71</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1020,1039]" pageId="4" pageNumber="333">Abdominal length 22.8 20 19.9 23.2 21.6 1.06</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1044,1063]" pageId="4" pageNumber="333">Body depth at dorsal-fin origin 19.1 20 16.4 19.7 17.5 0.86</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1069,1088]" pageId="4" pageNumber="333">Caudal-peduncle length 32.0 20 28.6 35.6 32.2 1.52</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" pageId="4" pageNumber="333">Caudal-peduncle depth 10.5 20 8.5 10.5 9.7 0.47 Percentages of head length</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419750" ID-Zenodo-Dep="5419750" httpUri="https://zenodo.org/record/5419750/files/figure.png" pageId="4" pageNumber="333" startId="4.[158,200,1113,1136]" targetBox="[203,787,198,1039]" targetPageId="4">
<paragraph blockId="4.[158,801,1113,1200]" pageId="4" pageNumber="333">
<emphasis bold="true" box="[158,239,1113,1136]" pageId="4" pageNumber="333">Fig. 5.</emphasis>
Left side of neurocranium in ventral view of 
<taxonomicName authority=", MCP" authorityName="MCP" box="[158,502,1145,1168]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[158,432,1145,1168]" italics="true" pageId="4" pageNumber="333">Gymnotocinclus anosteos</emphasis>
, MCP
</taxonomicName>
41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1142,1161]" pageId="4" pageNumber="333">Snout length 58.6 20 54.7 62.0 57.9 1.87</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1167,1186]" pageId="4" pageNumber="333">Orbital diameter 14.0 20 12.6 16.7 14.5 1.04</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1191,1210]" pageId="4" pageNumber="333">Interorbital width 41.5 20 37.8 42.8 39.7 1.34</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1216,1235]" pageId="4" pageNumber="333">Head depth 50.8 20 43.4 50.9 48.1 1.75</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1240,1259]" pageId="4" pageNumber="333">Suborbital depth 28.5 20 20.7 28.5 24.7 2.18</paragraph>
<paragraph blockId="4.[843,1469,749,1284]" box="[843,1469,1265,1284]" pageId="4" pageNumber="333">Mandibular ramus 13.3 20 10.3 14.8 12.7 1.34</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419752" ID-Zenodo-Dep="5419752" box="[158,1271,1955,1978]" httpUri="https://zenodo.org/record/5419752/files/figure.png" pageId="4" pageNumber="333" startId="4.[158,197,1955,1978]" targetBox="[171,1481,1284,1902]" targetPageId="4">
<paragraph blockId="4.[158,1271,1955,1978]" box="[158,1271,1955,1978]" pageId="4" pageNumber="333">
<emphasis bold="true" box="[158,227,1955,1978]" pageId="4" pageNumber="333">Fig. 6.</emphasis>
Pectoral girdle in ventral view of 
<taxonomicName authority=", MCP" authorityName="MCP" box="[581,920,1955,1978]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[581,852,1955,1978]" italics="true" pageId="4" pageNumber="333">Gymnotocinclus anosteos</emphasis>
, MCP
</taxonomicName>
41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419754" ID-Zenodo-Dep="5419754" box="[151,1234,961,984]" httpUri="https://zenodo.org/record/5419754/files/figure.png" pageId="5" pageNumber="334" startId="5.[151,190,961,984]" targetBox="[358,1288,193,902]" targetPageId="5">
<paragraph blockId="5.[151,1234,961,984]" box="[151,1234,961,984]" pageId="5" pageNumber="334">
<emphasis bold="true" box="[151,219,961,984]" pageId="5" pageNumber="334">Fig. 7.</emphasis>
Pelvic girdle in dorsal view of 
<taxonomicName authority=", MCP" authorityName="MCP" box="[544,883,961,984]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="334" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[544,815,961,984]" italics="true" pageId="5" pageNumber="334">Gymnotocinclus anosteos</emphasis>
, MCP
</taxonomicName>
41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419756" ID-Zenodo-Dep="5419756" box="[151,1154,1954,1977]" httpUri="https://zenodo.org/record/5419756/files/figure.png" pageId="5" pageNumber="334" startId="5.[151,190,1954,1977]" targetBox="[195,1416,1087,1919]" targetPageId="5">
<paragraph blockId="5.[151,1154,1954,1977]" box="[151,1154,1954,1977]" pageId="5" pageNumber="334">
<emphasis bold="true" box="[151,220,1954,1977]" pageId="5" pageNumber="334">Fig. 8.</emphasis>
Skull in lateral view of 
<taxonomicName authority=", MCP" authorityName="MCP" box="[466,804,1954,1977]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="334" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[466,737,1954,1977]" italics="true" pageId="5" pageNumber="334">Gymnotocinclus anosteos</emphasis>
, MCP
</taxonomicName>
41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<caption pageId="6" pageNumber="335" startId="6.[158,221,197,220]">
<paragraph blockId="6.[158,1481,197,252]" pageId="6" pageNumber="335">
<emphasis bold="true" box="[158,247,197,220]" pageId="6" pageNumber="335">Table 2.</emphasis>
Frequency distribution for meristics of 
<taxonomicName box="[670,945,197,220]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="335" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[670,945,197,220]" italics="true" pageId="6" pageNumber="335">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
. Holotype values are marked with an asterisk. N = number of specimens.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="335">
<table box="[325,1306,274,441]" gridcols="4" gridrows="4" pageId="6" pageNumber="335">
<tr box="[325,1306,274,293]" gridrow="0" pageId="6" pageNumber="335">
<td box="[325,531,274,293]" gridcol="0" gridrow="0" pageId="6" pageNumber="335">Character</td>
<td box="[573,1141,274,293]" gridcol="1" gridrow="0" pageId="6" pageNumber="335">FrequencyDistribution</td>
<td box="[1199,1221,274,293]" gridcol="2" gridrow="0" pageId="6" pageNumber="335">N</td>
<td box="[1258,1306,274,293]" gridcol="3" gridrow="0" pageId="6" pageNumber="335">Mean</td>
</tr>
<tr box="[325,1306,300,368]" gridrow="1" pageId="6" pageNumber="335">
<td box="[325,531,300,368]" gridcol="0" gridrow="1" pageId="6" pageNumber="335">Left perforate plates Right perforate plates Left premaxillary teeth</td>
<td box="[573,1141,300,368]" gridcol="1" gridrow="1" pageId="6" pageNumber="335">23(5) 24(12*) 25(3) 23(8*) 24(7) 25(5) 25(1) 26(1) 27(2) 28(5*) 29(3) 30(1) 31(2) 32(2) 35(1)</td>
<td box="[1199,1221,300,368]" gridcol="2" gridrow="1" pageId="6" pageNumber="335">20 20 18</td>
<td box="[1258,1306,300,368]" gridcol="3" gridrow="1" pageId="6" pageNumber="335">24.0 23.8 29.1</td>
</tr>
<tr box="[325,1306,373,417]" gridrow="2" pageId="6" pageNumber="335">
<td box="[325,531,373,417]" gridcol="0" gridrow="2" pageId="6" pageNumber="335">Right premaxillary teeth Left dentary teeth</td>
<td box="[573,1141,373,417]" gridcol="1" gridrow="2" pageId="6" pageNumber="335">25(1) 26(2*) 27(4) 28(3) 29(1) 30(1) 31(1) 32(1) 33(4) 34(1) 23(1) 25(2) 26(3) 27(3*) 28(4) 29(2) 30(2) 33(2)</td>
<td box="[1199,1221,373,417]" gridcol="2" gridrow="2" pageId="6" pageNumber="335">19 19</td>
<td box="[1258,1306,373,417]" gridcol="3" gridrow="2" pageId="6" pageNumber="335">29.3 27.8</td>
</tr>
<tr box="[325,1306,422,441]" gridrow="3" pageId="6" pageNumber="335">
<td box="[325,531,422,441]" gridcol="0" gridrow="3" pageId="6" pageNumber="335">Right dentary teeth</td>
<td box="[573,1141,422,441]" gridcol="1" gridrow="3" pageId="6" pageNumber="335">22(1) 25(3) 26(3) 27(2*) 28(1) 29(4) 30(1) 31(1) 32(1) 33(1) 35(1)</td>
<td box="[1199,1221,422,441]" gridcol="2" gridrow="3" pageId="6" pageNumber="335">19</td>
<td box="[1258,1306,422,441]" gridcol="3" gridrow="3" pageId="6" pageNumber="335">28.0</td>
</tr>
</table>
</paragraph>
<paragraph blockId="6.[158,802,482,1145]" pageId="6" pageNumber="335">
sixth centrum. Metapterygoid channel present, deep. Levator crest present, well developed, its extension not reaching dorsal portion of hyomandibula. Hyomandibula adductor crest present, extending from posterior corner of preopercle to dorsal articular condyle. Suture between metapterigoid and hyomandibula elongate, beginning almost on dorsal margin of bones (
<figureCitation box="[232,294,674,697]" captionStart="Fig" captionStartId="6.[158,197,1954,1977]" captionTargetBox="[332,1323,1322,1906]" captionTargetId="figure-415@6.[297,1338,1305,1925]" captionTargetPageId="6" captionText="Fig. 9. Suspensorium in lateral view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419758" httpUri="https://zenodo.org/record/5419758/files/figure.png" pageId="6" pageNumber="335">Fig. 9</figureCitation>
). Preopercle laterosensorial canal straight, posterior ramus contacting hyomandibula adductor crest. Preopercle ventral margin straight, not reflected medially.
</paragraph>
<paragraph blockId="6.[158,802,482,1145]" pageId="6" pageNumber="335">Teeth slender, bifid; major (medial) cusp slightly rounded and approximately three times longer than minor (lateral) cusp, minor cusp minute and pointed. Accessory teeth absent in premaxilla and dentary. Lips roundish, papillose, and comparatively large; anterior portion of lips well developed and fleshy, posterior portion almost reaching pectoral girdle. Maxillary barbel adnate, completely attached to lower lip.</paragraph>
<paragraph blockId="6.[158,802,482,1145]" pageId="6" pageNumber="335">
Upper pharyngeal toothplate larger than fourth pharyngobranchial, which partially supports it.Accessory ceratobranchial flange present and well developed, almost same size of first ceratobranchial. Basibranchial 2 and 4 present. Basibranchial 2 partially ossified. Four branchiostegal rays (
<figureCitation box="[645,724,1122,1145]" captionStart="Fig" captionStartId="7.[151,190,1954,1977]" captionTargetBox="[165,1419,1177,1926]" captionTargetId="figure-516@7.[151,1475,1157,1939]" captionTargetPageId="7" captionText="Fig. 10. Branchial skeleton in dorsal view of Gymnotocinclus anosteos, MCP 41726, paratype, male, 40.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5419760" httpUri="https://zenodo.org/record/5419760/files/figure.png" pageId="6" pageNumber="335">Fig. 10</figureCitation>
).
</paragraph>
<paragraph blockId="6.[158,801,1186,1273]" lastBlockId="6.[839,1482,482,953]" pageId="6" pageNumber="335">
<emphasis bold="true" box="[158,345,1186,1209]" pageId="6" pageNumber="335">Color in alcohol.</emphasis>
Ground color of dorsal surface of head and body light to dark brown. Dorsal surface of head darker than body. Ventrolateral margins of head and trunk yellowish. Ventral surface yellowish and almost unpigmented. Wide longitudinal dark lateral stripe. Lateral line unpigmented, forming thin light longitudinal line in middle of dark lateral stripe. Four transverse dark saddles on dorsum, beginning at dark lateral longitudinal stripe and extending to middorsal line. Most anterior saddle inconspicuous, at dorsal-fin origin; second saddle beginning at end of dorsal-fin base and reaching vertical through second branched anal-fin ray; third saddle at typical adiposefin region; and posterior saddle at end of caudal peduncle. Pectoral and pelvic fins mostly hyaline, with few scattered chromatophores. Dorsal and anal fins with chromatophores forming two transverse dark bands, most conspicuous on dorsal fin. Caudal fin mostly dark, except for transverse hyaline band at middle and posterior end of rays. In some specimens, middle caudal rays darker forming longitudinal stripe.
</paragraph>
<paragraph blockId="6.[839,1481,994,1145]" pageId="6" pageNumber="335">
<emphasis bold="true" box="[839,1070,994,1017]" pageId="6" pageNumber="335">Sexual dimorphism.</emphasis>
Males with urogenital papilla just behind the anal opening. Adult males have a fleshy flap along the dorsal margin of the pelvic-fin thickened ray, which is wider basally and progressively narrowing distally. The flap is absent in juveniles and females.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="335" type="distribution">
<paragraph blockId="6.[839,1481,1186,1273]" pageId="6" pageNumber="335">
<emphasis bold="true" box="[839,987,1186,1209]" pageId="6" pageNumber="335">Distribution.</emphasis>
<taxonomicName box="[992,1267,1186,1209]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="335" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[992,1267,1186,1209]" italics="true" pageId="6" pageNumber="335">Gymnotocinclus anosteos</emphasis>
</taxonomicName>
is known only from two creeks, both tributaries to the rio Tocantinzinho, in the headwaters of the rio 
<collectingRegion box="[1063,1166,1250,1273]" country="Brazil" name="Tocantins" pageId="6" pageNumber="335">Tocantins</collectingRegion>
basin, 
<collectingRegion box="[1239,1304,1250,1273]" country="Brazil" name="Goias" pageId="6" pageNumber="335">Goiás</collectingRegion>
, 
<collectingCountry box="[1313,1378,1250,1273]" name="Brazil" pageId="6" pageNumber="335">Brazil</collectingCountry>
(
<figureCitation box="[1392,1468,1250,1273]" captionStart="Fig" captionStartId="8.[158,197,1875,1898]" captionTargetBox="[198,793,404,1838]" captionTargetId="figure-568@8.[198,793,404,1838]" captionTargetPageId="8" captionText="Fig. 11. Geographic distribution of Gymnotocinclus anosteos in the upper rio Tocantins drainage. Circles represent collecting sites, and T is the type-locality." figureDoi="http://doi.org/10.5281/zenodo.5419762" httpUri="https://zenodo.org/record/5419762/files/figure.png" pageId="6" pageNumber="335">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5419758" ID-Zenodo-Dep="5419758" box="[158,1261,1954,1977]" httpUri="https://zenodo.org/record/5419758/files/figure.png" pageId="6" pageNumber="335" startId="6.[158,197,1954,1977]" targetBox="[332,1323,1322,1906]" targetPageId="6">
<paragraph blockId="6.[158,1261,1954,1977]" box="[158,1261,1954,1977]" pageId="6" pageNumber="335">
<emphasis bold="true" box="[158,227,1954,1977]" pageId="6" pageNumber="335">Fig. 9.</emphasis>
Suspensorium in lateral view of 
<taxonomicName authority=", MCP" authorityName="MCP" box="[570,910,1954,1977]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="335" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[570,842,1954,1977]" italics="true" pageId="6" pageNumber="335">Gymnotocinclus anosteos</emphasis>
, MCP
</taxonomicName>
41726, paratype, male, 40.5 mm.
</paragraph>
</caption>
<subSubSection pageId="7" pageNumber="336" type="etymology">
<paragraph blockId="7.[151,794,194,249]" pageId="7" pageNumber="336">
<emphasis bold="true" box="[151,281,194,217]" pageId="7" pageNumber="336">Etymology.</emphasis>
From the Greek 
<taxonomicName box="[461,555,194,217]" class="Actinopterygii" family="Loricariidae" genus="Gymnotocinclus" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="336" phylum="Chordata" rank="species" species="anosteos">
<emphasis box="[461,555,194,217]" italics="true" pageId="7" pageNumber="336">anosteos</emphasis>
</taxonomicName>
, meaning boneless, in allusion to the absence of the lateral connecting bone.
</paragraph>
</subSubSection>
</treatment>
</document>