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<document id="260CCFB3B51709C4D56B8D209D35CEB9" ID-CLB-Dataset="21918" ID-DOI="10.11646/zootaxa.4780.1.1" ID-GBIF-Dataset="af0a7e60-adc6-4896-8544-63725501fa13" ID-ISSN="1175-5326" ID-Zenodo-Dep="3856493" ID-ZooBank="D6CC1683-8BF0-4ABF-ABFE-3EC63E66AE5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590138264658" checkinUser="plazi" docAuthor="Suárez, William" docDate="2020" docId="039EF96AFFE62249ED83FF6EFACAFE33" docLanguage="en" docName="zootaxa.4780.1.1.pdf" docOrigin="Zootaxa 4780 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyto cravesae in Suarez &amp; Olson 2015" docType="treatment" docVersion="7" lastPageNumber="31" masterDocId="FFA78112FFFB2257ED14FFF7FFB9FFB8" masterDocTitle="The fossil avifauna of the tar seeps Las Breas de San Felipe, Matanzas, Cuba" masterLastPageNumber="53" masterPageNumber="1" pageNumber="30" updateTime="1698840883311" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2ED9BE01FD99B2F039EC52CB266E79F7">The fossil avifauna of the tar seeps Las Breas de San Felipe, Matanzas, Cuba</mods:title>
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<treatment id="039EF96AFFE62249ED83FF6EFACAFE33" ID-DOI="http://doi.org/10.5281/zenodo.3856839" ID-GBIF-Taxon="164248753" ID-Zenodo-Dep="3856839" LSID="urn:lsid:plazi:treatment:039EF96AFFE62249ED83FF6EFACAFE33" httpUri="http://treatment.plazi.org/id/039EF96AFFE62249ED83FF6EFACAFE33" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="C32D1BF7FFE6224AED83FF6EFD4CFF6F" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8B88487CFFE6224AED83FF6EFDD8FF0B" blockId="29.[151,757,153,252]" box="[151,609,153,180]" pageId="29" pageNumber="30">
<heading id="D0C0FF10FFE6224AED83FF6EFDD8FF0B" bold="true" box="[151,609,153,180]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B943946EFFE6224AED83FF6EFDD8FF0B" bold="true" box="[151,609,153,180]" pageId="29" pageNumber="30">
<taxonomicName id="4C3733FFFFE6224AEDB2FF6DFDD8FF0B" authority="Suarez &amp; Olson, 2015" authorityName="in Suarez &amp; Olson" authorityYear="2015" box="[166,609,153,180]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cravesae">
<emphasis id="B943946EFFE6224AEDB2FF6DFEF3FF0C" bold="true" box="[166,330,154,180]" italics="true" pageId="29" pageNumber="30">Tyto cravesae</emphasis>
<bibRefCitation id="EFA6358DFFE6224AEC45FF6EFDD8FF0B" author="Suarez, W. &amp; Olson, S. L." box="[337,609,153,180]" pageId="29" pageNumber="30" pagination="533 - 553" refId="ref31508" refString="Suarez, W. &amp; Olson, S. L. (2015) Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa, 4020 (3), 533 - 553. https: // doi. org / 10.11646 / zootaxa. 4020.3.7" type="journal article" year="2015">Suárez &amp; Olson, 2015</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B88487CFFE6224AED83FF4AFD4CFF6F" blockId="29.[151,757,153,252]" box="[151,757,189,216]" pageId="29" pageNumber="30">Cravess Giant Barn Owl; Lechuza Gigante de Craves</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE6224AED83FF16FEE1FF44" box="[151,344,225,252]" pageId="29" pageNumber="30" type="description">
<paragraph id="8B88487CFFE6224AED83FF16FEE1FF44" blockId="29.[151,757,153,252]" box="[151,344,225,252]" pageId="29" pageNumber="30">
(
<figureCitation id="130C54F9FFE6224AED8BFF16FEA8FF43" box="[159,273,225,251]" captionStart="FIGURE 12" captionStartId="29.[151,250,1908,1933]" captionTargetBox="[180,1423,537,1872]" captionTargetId="figure@29.[151,1436,520,1883]" captionTargetPageId="29" captionText="FIGURE 12. Tyto cravesae: Left tarsometatarsus (Paratype, MNHNCu 75.4801) in anterior (A) and distal (B) views. Pulsatrix arredondoi: Left tarsometatarsus (MNHNCu 75.4808) in anterior (C) and proximal (D) views. Ornimegalonyx oteroi: Right tarsometatarsus (MNHNCu 75.4804) in anterior (E), proximal (F), distal (G) and posterior (H) views. Ornimegalonyx sp.: Right femur (MNHNCu 75.4714) in proximal (I) and anterior (J) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856515" httpUri="https://zenodo.org/record/3856515/files/figure.png" pageId="29" pageNumber="30">Figure 12</figureCitation>
: AB)
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE6224AED83FED1FCA3FE86" box="[151,794,293,318]" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="8B88487CFFE6224AED83FED1FCA3FE86" blockId="29.[151,794,293,318]" box="[151,794,293,318]" pageId="29" pageNumber="30">
<taxonomicName id="4C3733FFFFE6224AED83FED1FDB4FE86" authority="Suarez &amp; Olson, 2015" authorityName="in Suarez &amp; Olson" authorityYear="2015" box="[151,525,293,318]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cravesae">
<emphasis id="B943946EFFE6224AED83FED1FE9AFE85" box="[151,291,294,317]" italics="true" pageId="29" pageNumber="30">Tyto cravesae</emphasis>
<bibRefCitation id="EFA6358DFFE6224AEC3DFED1FDB4FE86" author="Suarez, W. &amp; Olson, S. L." box="[297,525,293,318]" pageId="29" pageNumber="30" pagination="533 - 553" refId="ref31508" refString="Suarez, W. &amp; Olson, S. L. (2015) Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa, 4020 (3), 533 - 553. https: // doi. org / 10.11646 / zootaxa. 4020.3.7" type="journal article" year="2015">Suárez &amp; Olson, 2015</bibRefCitation>
</taxonomicName>
, Zootaxa 4020 (3), p. 545.
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE6224AED83FE9CFB22FE3E" box="[151,1179,363,391]" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8B88487CFFE6224AED83FE9CFB22FE3E" blockId="29.[151,1436,363,499]" box="[151,1179,363,391]" pageId="29" pageNumber="30">
<emphasis id="B943946EFFE6224AED83FE9CFECCFE3E" bold="true" box="[151,373,363,390]" pageId="29" pageNumber="30">Referred material.</emphasis>
San Felipe I: Distal end of the left tarsometatarsus, MNHNCu 75.4801.
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE62249EDD3FE78FEB7FF6F" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8B88487CFFE6224AEDD3FE78FC7EFE4A" blockId="29.[151,1436,363,499]" pageId="29" pageNumber="30">
<emphasis id="B943946EFFE6224AEDD3FE78FEE0FE12" bold="true" box="[199,345,399,426]" pageId="29" pageNumber="30">Description.</emphasis>
The specimen MNHNCu 75.4801 has been compared and described by
<bibRefCitation id="EFA6358DFFE6224AE99AFE67FA21FE12" author="Suarez, W. &amp; Olson, S. L." box="[1166,1432,399,426]" pageId="29" pageNumber="30" pagination="533 - 553" refId="ref31508" refString="Suarez, W. &amp; Olson, S. L. (2015) Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa, 4020 (3), 533 - 553. https: // doi. org / 10.11646 / zootaxa. 4020.3.7" type="journal article" year="2015">Suárez &amp; Olson (2015)</bibRefCitation>
, where
<taxonomicName id="4C3733FFFFE6224AEDF7FE42FDA2FE76" authority="Arredondo, 1972" authorityName="Arredondo" authorityYear="1972" box="[227,539,436,462]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="noeli">
<emphasis id="B943946EFFE6224AEDF7FE42FEEBFE76" box="[227,338,436,462]" italics="true" pageId="29" pageNumber="30">Tyto noeli</emphasis>
<bibRefCitation id="EFA6358DFFE6224AEC4CFE43FDA2FE76" author="Arredondo, O." box="[344,539,436,462]" pageId="29" pageNumber="30" pagination="415 - 431" refId="ref24552" refString="Arredondo, O. (1972) Nueva especie de ave fosil (Strigiformes: Tytonidae) del Pleistoceno superior de Cuba. Boletin de la Sociedad Venezolana de Ciencias Naturales, 29, 415 - 431." type="journal article" year="1972">Arredondo, 1972</bibRefCitation>
</taxonomicName>
, is smaller, but
<taxonomicName id="4C3733FFFFE6224AEFD8FE42FC48FE76" authority="Wetmore, 1937" authorityName="Wetmore" authorityYear="1937" box="[716,1009,436,463]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="pollens">
<emphasis id="B943946EFFE6224AEFD8FE42FD65FE76" box="[716,732,437,462]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B943946EFFE6224AEFFEFE43FC82FE76" box="[746,827,436,462]" italics="true" pageId="29" pageNumber="30">pollens</emphasis>
<bibRefCitation id="EFA6358DFFE6224AEE56FE42FC48FE76" author="Wetmore, A." box="[834,1009,436,463]" pageId="29" pageNumber="30" pagination="427 - 441" refId="ref32224" refString="Wetmore, A. (1937) Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bulletin of the Museum of Comparative Zoology, 80, 427 - 441." type="journal article" year="1937">Wetmore, 1937</bibRefCitation>
</taxonomicName>
, much larger. See additional comparisons in the original description of
<taxonomicName id="4C3733FFFFE6224AEF02FE2EFC7BFE4A" authority="in Suarez &amp; Olson (2015)" authorityName="in Suarez &amp; Olson" authorityYear="2015" box="[534,962,471,499]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cravesae">
<emphasis id="B943946EFFE6224AEF02FE2EFD9FFE4A" box="[534,550,473,498]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B943946EFFE6224AEF27FE2EFD2EFE4A" box="[563,663,473,498]" italics="true" pageId="29" pageNumber="30">cravesae</emphasis>
in
<bibRefCitation id="EFA6358DFFE6224AEFAFFE2FFC7BFE4A" author="Suarez, W. &amp; Olson, S. L." box="[699,962,471,498]" pageId="29" pageNumber="30" pagination="533 - 553" refId="ref31508" refString="Suarez, W. &amp; Olson, S. L. (2015) Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa, 4020 (3), 533 - 553. https: // doi. org / 10.11646 / zootaxa. 4020.3.7" type="journal article" year="2015">Suárez &amp; Olson (2015)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="DF4818F4FFE6224AED83F883FC7DF840" ID-DOI="http://doi.org/10.5281/zenodo.3856515" ID-Zenodo-Dep="3856515" httpUri="https://zenodo.org/record/3856515/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1908,1933]" targetBox="[180,1423,537,1872]" targetPageId="29">
<paragraph id="8B88487CFFE6224AED83F883FC7DF840" blockId="29.[151,1436,1908,2041]" pageId="29" pageNumber="30">
<emphasis id="B943946EFFE6224AED83F883FEA6F835" bold="true" box="[151,287,1908,1933]" pageId="29" pageNumber="30">FIGURE 12.</emphasis>
<taxonomicName id="4C3733FFFFE6224AEC31F882FE51F835" authority=": Left" authorityName="Left" box="[293,488,1908,1933]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cravesae">
<emphasis id="B943946EFFE6224AEC31F882FE08F834" box="[293,433,1909,1932]" italics="true" pageId="29" pageNumber="30">Tyto cravesae</emphasis>
: Left
</taxonomicName>
tarsometatarsus (Paratype, MNHNCu 75.4801) in anterior (A) and distal (B) views.
<taxonomicName id="4C3733FFFFE6224AE82BF882FEFBF809" authority=": Left" authorityName="Left" class="Aves" family="Strigidae" genus="Pulsatrix" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="arredondoi">
<emphasis id="B943946EFFE6224AE82BF882FEB1F809" italics="true" pageId="29" pageNumber="30">Pulsatrix arredondoi</emphasis>
: Left
</taxonomicName>
tarsometatarsus (MNHNCu 75.4808) in anterior (C) and proximal (D) views.
<taxonomicName id="4C3733FFFFE6224AE967F86EFA25F809" authority=": Right" authorityName="Right" box="[1139,1436,1944,1969]" class="Aves" family="Strigidae" genus="Ornimegalonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="oteroi">
<emphasis id="B943946EFFE6224AE967F86EFAECF808" box="[1139,1365,1945,1969]" italics="true" pageId="29" pageNumber="30">Ornimegalonyx oteroi</emphasis>
: Right
</taxonomicName>
tarsometatarsus (MNHNCu 75.4804) in anterior (E), proximal (F), distal (G) and posterior (H) views.
<taxonomicName id="4C3733FFFFE6224AE983F84AFA8CF86D" authorityName="Arredondo" authorityYear="1958" box="[1175,1333,1981,2005]" class="Aves" family="Strigidae" genus="Ornimegalonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strigiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE6224AE983F84AFA8CF86D" box="[1175,1333,1981,2005]" italics="true" pageId="29" pageNumber="30">Ornimegalonyx</emphasis>
</taxonomicName>
sp.: Right femur (MNHNCu 75.4714) in proximal (I) and anterior (J) views. Scale = 2 cm.
</paragraph>
</caption>
<paragraph id="8B88487CFFE52249EDD3FF6EFEB7FF6F" blockId="30.[151,1436,152,395]" pageId="30" pageNumber="31">
<emphasis id="B943946EFFE52249EDD3FF6EFECFFF0B" bold="true" box="[199,374,153,179]" pageId="30" pageNumber="31">Measurements</emphasis>
(
<taxonomicName id="4C3733FFFFE52249EC92FF6DFE4DFF0B" authorityName="Arredondo" authorityYear="1972" box="[390,500,153,179]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="noeli">
<emphasis id="B943946EFFE52249EC92FF6DFE4DFF0B" box="[390,500,153,179]" italics="true" pageId="30" pageNumber="31">Tyto noeli</emphasis>
</taxonomicName>
in parentheses, after
<bibRefCitation id="EFA6358DFFE52249EFF5FF6EFC6FFF0B" author="Suarez, W. &amp; Olson, S. L." box="[737,982,152,179]" pageId="30" pageNumber="31" pagination="533 - 553" refId="ref31508" refString="Suarez, W. &amp; Olson, S. L. (2015) Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa, 4020 (3), 533 - 553. https: // doi. org / 10.11646 / zootaxa. 4020.3.7" type="journal article" year="2015">Suárez &amp; Olson 2015</bibRefCitation>
: table 2).
<emphasis id="B943946EFFE52249E923FF6DFBEDFF0B" box="[1079,1108,154,179]" italics="true" pageId="30" pageNumber="31"></emphasis>
Distal width: 18.1 (14.417.3 [15.7] 12).
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE52249EDD3FF17FACAFE33" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8B88487CFFE52249EDD3FF17FACAFE33" blockId="30.[151,1436,152,395]" pageId="30" pageNumber="31">
<emphasis id="B943946EFFE52249EDD3FF17FEF6FF43" bold="true" box="[199,335,224,251]" pageId="30" pageNumber="31">Comments.</emphasis>
The distal end of the tarsometatarsus that represents this large barn owl in Las Breas de San Felipe (
<figureCitation id="130C54F9FFE52249EDB4FEF2FF57FEA7" box="[160,238,261,287]" captionStart="FIGURE 12" captionStartId="29.[151,250,1908,1933]" captionTargetBox="[180,1423,537,1872]" captionTargetId="figure@29.[151,1436,520,1883]" captionTargetPageId="29" captionText="FIGURE 12. Tyto cravesae: Left tarsometatarsus (Paratype, MNHNCu 75.4801) in anterior (A) and distal (B) views. Pulsatrix arredondoi: Left tarsometatarsus (MNHNCu 75.4808) in anterior (C) and proximal (D) views. Ornimegalonyx oteroi: Right tarsometatarsus (MNHNCu 75.4804) in anterior (E), proximal (F), distal (G) and posterior (H) views. Ornimegalonyx sp.: Right femur (MNHNCu 75.4714) in proximal (I) and anterior (J) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856515" httpUri="https://zenodo.org/record/3856515/files/figure.png" pageId="30" pageNumber="31">Fig. 12</figureCitation>
AB) was formerly recorded as
<taxonomicName id="4C3733FFFFE52249EF41FEF1FD3CFEA7" authorityName="Billberg" authorityYear="1828" box="[597,645,262,287]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE52249EF41FEF1FD3CFEA7" box="[597,645,262,287]" italics="true" pageId="30" pageNumber="31">Tyto</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="EFA6358DFFE52249EFADFEF2FC49FEA7" author="Iturralde-Vinent, M. &amp; MacPhee, R. D. E. &amp; Diaz-Franco, S. &amp; Rojas-Consuegra, R. &amp; Suarez, W. &amp; Lomba, A." box="[697,1008,261,287]" pageId="30" pageNumber="31" pagination="300 - 313" refId="ref27570" refString="Iturralde-Vinent, M., MacPhee, R. D. E., Diaz-Franco, S., Rojas-Consuegra, R., Suarez, W. &amp; Lomba, A. (2000) Las Breas de San Felipe, a Quaternary asphalt seep near Marti (Matanzas Province, Cuba). Caribbean Journal of Science, 36, 300 - 313." type="journal article" year="2000">
Iturralde-Vinent
<emphasis id="B943946EFFE52249EE62FEF1FC1EFEA7" box="[886,935,261,287]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 2000
</bibRefCitation>
: table 2), and subsequently designated as the
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of
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.
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(see
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). The assessment that
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.
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was the rarest of the Cuban large barn owls (
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) and that in
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existed four large tytonids with marked anatomical adaptations for terrestrial locomotion (
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), is wrong (see
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, 539540).
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