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<document id="F97AAA2BD280D8A4A9A0877EB137A961" ID-CLB-Dataset="294849" ID-DOI="10.11646/zootaxa.5443.4.2" ID-GBIF-Dataset="1e37690a-e157-439e-ac09-6e26fdd1c4c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064278" ID-ZooBank="A91A8D67-A81B-427D-8096-59E41326A8E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1714028852870" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2024" docId="03CB87B7E944FFEBF2CFFC88FD03FA26" docLanguage="en" docName="zootaxa.5443.4.2.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Octavius Fauvel 1873" docType="key" docVersion="2" lastPageNumber="521" masterDocId="FFF2FFCFE95EFFF1F258FF9CFFC6FFD6" masterDocTitle="On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)" masterLastPageNumber="522" masterPageNumber="495" pageNumber="521" updateTime="1714144938831" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="06EAACD905811EF25AB4CC4392532DA7">On Octavius from Maloti-Drakensberg and Golden Gate National Parks, South Africa (Coleoptera: Staphylinidae: Euaesthetinae)</mods:title>
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<subSubSection id="C378652AE944FFEBF2CFFC88FF18FC84" pageId="26" pageNumber="521" type="nomenclature">
<paragraph id="8BDD36A1E944FFEBF2CFFC88FF18FC84" blockId="26.[151,1436,787,850]" pageId="26" pageNumber="521">
<heading id="D09581CDE944FFEBF2CFFC88FF18FC84" bold="true" centered="true" fontSize="11" level="1" pageId="26" pageNumber="521" reason="1">
<emphasis id="B916EAB3E944FFEBF2CFFC88FF18FC84" bold="true" pageId="26" pageNumber="521">
Updated key to the
<taxonomicName id="4C624D22E944FFEBF3DAFC8FFE21FCFB" ID-CoL="8VZT3" authority="Fauvel, 1873" authorityName="Fauvel" authorityYear="1873" box="[386,487,787,813]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B916EAB3E944FFEBF3DAFC8FFE21FCFB" bold="true" box="[386,487,787,813]" italics="true" pageId="26" pageNumber="521">Octavius</emphasis>
</taxonomicName>
species of
<collectingCountry id="F3757631E944FFEBF03FFC8FFD39FCFB" box="[615,767,787,813]" name="South Africa" pageId="26" pageNumber="521">South Africa</collectingCountry>
with one ommatidium and pronotum much longer than broad
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378652AE944FFEBF2CFFCE3FCDDFC68" pageId="26" pageNumber="521" type="discussion">
<paragraph id="8BDD36A1E944FFEBF2CFFCE3FCDDFC68" blockId="26.[151,1436,895,958]" pageId="26" pageNumber="521">
This key updates a part of the key published by
<bibRefCitation id="EFF34B50E944FFEBF081FCE3FCB4FC4F" author="Puthz, V." box="[729,882,895,921]" pageId="26" pageNumber="521" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz (2006)</bibRefCitation>
. Couplets 168 and 7185 remains unchanged, couplets 86 and following were updated by
<bibRefCitation id="EFF34B50E944FFEBF0DAFC3FFCD1FC68" author="Janak, J." box="[642,791,931,958]" pageId="26" pageNumber="521" pagination="195 - 231" refId="ref16165" refString="Janak, J. (2014) New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae). Acta Entomologica Musei Nationalis Pragae, 54 (1), 195 - 231." type="journal article" year="2014">Janák (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378652AE944FFEBF2CFFC74FD03FA26" pageId="26" pageNumber="521" type="key">
<paragraph id="8BDD36A1E944FFEBF2CFFC74FD6AFC2B" blockId="26.[151,1437,1000,1521]" box="[151,684,1000,1022]" pageId="26" pageNumber="521">69 (70) Pronotum much longer than broad (R&gt; 1.1)</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFB9AFA5AFBCA" blockId="26.[151,1437,1000,1521]" box="[151,1436,1030,1052]" pageId="26" pageNumber="521">
69a (69b) Parameres markedly longer than median lobe (
<figureCitation id="13592A24E944FFEBF09DFB9AFD38FBCD" box="[709,766,1030,1052]" captionStart="FIGURES 115" captionStartId="3.[151,264,1794,1818]" captionTargetBox="[168,1419,192,1769]" captionTargetId="figure-15@3.[156,1432,179,1769]" captionText="FIGURES 115. 110. Octavius adriani Janák: 13, 6, 910, Gxaligengwa Forest; 45, 78, Fort Nottingham, paratypes. 1115. Octavius clarkei Janák, sp. nov., 11, 1315, holotype, 12, paratype. 1, 11, habitus; 25, 1213, aedeagus ventral; 68, apex of aedeagus ventral; 9, 14, male sternite VIII; 10, 15, male sternite IX. Scale 1 mm: 1, 11; scales 0.05 mm: 68; 0.1 mm: remaining figures. LI: lateral impression, TI: transverse impression, DI: dorsal impressions, LRP: lateral ridge on pronotum, LIR: lateral internal ridge, LOR: lateral external ridge." figureDoi="http://doi.org/10.5281/zenodo.11074157" httpUri="https://zenodo.org/record/11074157/files/figure.png" pageId="26" pageNumber="521">
Fig.
<quantity id="4C9A9B44E944FFEBF0AAFB9AFCD1FBCA" box="[754,791,1030,1052]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="26" pageNumber="521" unit="in" value="7.0">7 in</quantity>
</figureCitation>
<bibRefCitation id="EFF34B50E944FFEBF144FB9AFC4CFBCA" author="Puthz, V." box="[796,906,1030,1052]" pageId="26" pageNumber="521" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz 2006</bibRefCitation>
)....................
<taxonomicName id="4C624D22E944FFEBF6D4FB9AFA5AFBCA" authority="Puthz, 2006" authorityName="Puthz" authorityYear="2006" box="[1164,1436,1030,1052]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B916EAB3E944FFEBF6D4FB9AFAE2FBCD" box="[1164,1316,1030,1051]" italics="true" pageId="26" pageNumber="521">O. angusticollis</emphasis>
<bibRefCitation id="EFF34B50E944FFEBF772FB9AFA5AFBCA" author="Puthz, V." box="[1322,1436,1030,1052]" pageId="26" pageNumber="521" pagination="1 - 28" refId="ref16421" refString="Puthz, V. (2006) New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 1 - 28." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFBBFFC43FBEF" blockId="26.[151,1437,1000,1521]" box="[151,901,1059,1081]" pageId="26" pageNumber="521">69b (69a) Parameres slightly shorter or at least about as long as median lobe.</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFBDCFA5AFB83" blockId="26.[151,1437,1000,1521]" box="[151,1436,1088,1110]" pageId="26" pageNumber="521">
69c (69d) Aedeagus with internal structure very long, awl shaped (
<figureCitation id="13592A24E944FFEBF17BFBDCFC6AFB83" box="[803,940,1088,1110]" captionStart="FIGURES 99112" captionStartId="21.[151,264,1492,1516]" captionTargetBox="[172,1416,286,1456]" captionTargetId="figure-41@21.[159,1429,273,1469]" captionTargetPageId="21" captionText="FIGURES 99112. 99106: Octavius regius Janák, sp. nov.; 99103, 105106, holotype; 104, paratype. 107112: Octavius subulatus Janák, sp. nov.; 107110, 112113, holotype; 111, paratype. 99, 107, habitus; 100, 108, fore parts; 101, 109, fore parts lateral; 102, detail of eye with omatidia; 103104, 110111, aedeagus ventral; 105, 112, male sternite VIII; 106, 113, male sternite IX. Scale 0.5 mm: 99, 107; scales 0.1 mm: 100101, 108109; 103104; 110111; 105106, 112113. LOR: lateral outer ridge, o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064296" httpUri="https://zenodo.org/record/11064296/files/figure.png" pageId="26" pageNumber="521">Figs. 110111</figureCitation>
).................
<taxonomicName id="4C624D22E944FFEBF6D6FBDDFA80FB80" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1166,1350,1088,1110]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="subulatus" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF6D6FBDDFAC1FB83" box="[1166,1287,1088,1110]" italics="true" pageId="26" pageNumber="521">O. subulatus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E944FFEBF708FBDCFA5AFB83" bold="true" box="[1360,1436,1088,1109]" pageId="26" pageNumber="521">
<taxonomicNameLabel id="A22557C8E944FFEBF708FBDCFA5AFB83" box="[1360,1436,1088,1109]" pageId="26" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFBC2FD5BFBA2" blockId="26.[151,1437,1000,1521]" box="[151,669,1118,1140]" pageId="26" pageNumber="521">69d (69c) Aedeagus with internal structure different.</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFBE7FA5AFB7B" blockId="26.[151,1437,1000,1521]" pageId="26" pageNumber="521">
69e (69f) Median part of aedeagus without internal structures, internal sac only with small dense scales (
<figureCitation id="13592A24E944FFEBF6C9FBE7FAC3FB47" box="[1169,1285,1147,1169]" captionStart="FIGURES 7184" captionStartId="16.[151,264,1389,1413]" captionTargetBox="[193,1394,195,1353]" captionTargetId="figure-29@16.[180,1407,181,1366]" captionTargetPageId="16" captionText="FIGURES 7184. 7177: Octavius aureaportalis Janák, sp. nov.; 7174, 7677, holotype; 75, paratype. 7884: Octavius gracilis Janák, sp. nov.: 7881, 8384, holotype; 82, paratype. 71,78, habitus; 72, 79, fore parts; 73, 80, fore parts lateral; 7475, 8182, aedeagus ventral; 76, 83, male sternite VIII; 77, 84, male sternite IX. Scale 0.5 mm: 71, 78; scales 0.1 mm: 7273, 7980; 7475; 8182; 7677, 8384, o ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064290" httpUri="https://zenodo.org/record/11064290/files/figure.png" pageId="26" pageNumber="521">Figs. 7475</figureCitation>
)..................................................................................
<taxonomicName id="4C624D22E944FFEBF630FB05FA80FB78" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1128,1350,1176,1198]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="aureaportalis" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF630FB05FAC1FB7B" box="[1128,1287,1176,1198]" italics="true" pageId="26" pageNumber="521">O. aureaportalis</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E944FFEBF708FB04FA5AFB7B" bold="true" box="[1360,1436,1176,1197]" pageId="26" pageNumber="521">
<taxonomicNameLabel id="A22557C8E944FFEBF708FB04FA5AFB7B" box="[1360,1436,1176,1197]" pageId="26" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFB2AFCEEFB1D" blockId="26.[151,1437,1000,1521]" box="[151,808,1206,1228]" pageId="26" pageNumber="521">69f (69e) Median part of aedeagus with distinct internal structures.</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFB4FFA5AFB3E" blockId="26.[151,1437,1000,1521]" box="[151,1436,1235,1257]" pageId="26" pageNumber="521">
<quantity id="4C9A9B44E944FFEBF2CFFB4FFF7AFB3F" box="[151,188,1235,1257]" metricMagnitude="-2" metricUnit="kg" metricValue="6.9" pageId="26" pageNumber="521" unit="g" value="69.0">69g</quantity>
(69h) Aedeagus with internal tube and two “V” shaped structures (
<figureCitation id="13592A24E944FFEBF113FB4FFC78FB3E" box="[843,958,1235,1257]" captionStart="FIGURES 7184" captionStartId="16.[151,264,1389,1413]" captionTargetBox="[193,1394,195,1353]" captionTargetId="figure-29@16.[180,1407,181,1366]" captionTargetPageId="16" captionText="FIGURES 7184. 7177: Octavius aureaportalis Janák, sp. nov.; 7174, 7677, holotype; 75, paratype. 7884: Octavius gracilis Janák, sp. nov.: 7881, 8384, holotype; 82, paratype. 71,78, habitus; 72, 79, fore parts; 73, 80, fore parts lateral; 7475, 8182, aedeagus ventral; 76, 83, male sternite VIII; 77, 84, male sternite IX. Scale 0.5 mm: 71, 78; scales 0.1 mm: 7273, 7980; 7475; 8182; 7677, 8384, o ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064290" httpUri="https://zenodo.org/record/11064290/files/figure.png" pageId="26" pageNumber="521">Figs. 8182</figureCitation>
).................
<taxonomicName id="4C624D22E944FFEBF6F8FB4FFA80FB3F" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1184,1350,1235,1257]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF6F8FB4FFAC1FB3E" box="[1184,1287,1235,1256]" italics="true" pageId="26" pageNumber="521">O. gracilis</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E944FFEBF708FB4FFA5AFB3E" bold="true" box="[1360,1436,1235,1256]" pageId="26" pageNumber="521">
<taxonomicNameLabel id="A22557C8E944FFEBF708FB4FFA5AFB3E" box="[1360,1436,1235,1256]" pageId="26" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFB6CFD60FAD0" blockId="26.[151,1437,1000,1521]" box="[151,678,1264,1286]" pageId="26" pageNumber="521">
69h (
<quantity id="4C9A9B44E944FFEBF290FB6CFF28FAD0" box="[200,238,1264,1286]" metricMagnitude="-2" metricUnit="kg" metricValue="6.9" pageId="26" pageNumber="521" unit="g" value="69.0">69g</quantity>
) Aedeagus with different internal structures.
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFA92FA5AFAF5" blockId="26.[151,1437,1000,1521]" box="[151,1436,1293,1316]" pageId="26" pageNumber="521">
69i (69j) Median part of aedeagus with a pair of very long internal structures (
<figureCitation id="13592A24E944FFEBF1C1FA92FBCBFAF5" box="[921,1037,1293,1316]" captionStart="FIGURES 8598" captionStartId="18.[151,264,1364,1388]" captionTargetBox="[179,1409,194,1327]" captionTargetId="figure-29@18.[166,1422,181,1340]" captionTargetPageId="18" captionText="FIGURES 8598. 8591: Octavius gxaligengwanus Janák, sp. nov.; 8588, 9091, holotype; 89, paratype. 9298: Octavius monachicullus Janák, sp. nov.; 9295, 9798, holotype; 96, paratype, Injisuthi. 85, 92, habitus; 86, 93, fore parts; 87, 94, fore parts lateral; 8889, 9596, aedeagus ventral; 90, 97, male sternite VIII; 91, 98, male sternite IX. Scale 0.5 mm: 85, 92; scales 0.1 mm: 8687, 9394; 8889, 9596; 9091, 9798; o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064292" httpUri="https://zenodo.org/record/11064292/files/figure.png" pageId="26" pageNumber="521">Figs. 9596</figureCitation>
)....
<taxonomicName id="4C624D22E944FFEBF614FA92FA8FFAF2" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1100,1353,1294,1316]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="monachicucullus" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF614FA92FACAFAF5" box="[1100,1292,1294,1315]" italics="true" pageId="26" pageNumber="521">O. monachicucullus</emphasis>
Janák
</taxonomicName>
<taxonomicNameLabel id="A22557C8E944FFEBF716FA92FA5AFAF5" box="[1358,1436,1294,1315]" pageId="26" pageNumber="521" rank="species">
<emphasis id="B916EAB3E944FFEBF716FA92FA50FAF5" bold="true" box="[1358,1430,1294,1315]" pageId="26" pageNumber="521">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFAB7FD60FA96" blockId="26.[151,1437,1000,1521]" box="[151,678,1323,1345]" pageId="26" pageNumber="521">69j (69i) Aedeagus with different internal structures.</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFAD4FA5AFAAD" blockId="26.[151,1437,1000,1521]" pageId="26" pageNumber="521">
69k (69l) Aedeagus short and broad, about 3 times as long as broad, with a pair of elongate internal structures near the apex of median lobe (
<figureCitation id="13592A24E944FFEBF363FAFAFE68FAAD" box="[315,430,1382,1404]" captionStart="FIGURES 8598" captionStartId="18.[151,264,1364,1388]" captionTargetBox="[179,1409,194,1327]" captionTargetId="figure-29@18.[166,1422,181,1340]" captionTargetPageId="18" captionText="FIGURES 8598. 8591: Octavius gxaligengwanus Janák, sp. nov.; 8588, 9091, holotype; 89, paratype. 9298: Octavius monachicullus Janák, sp. nov.; 9295, 9798, holotype; 96, paratype, Injisuthi. 85, 92, habitus; 86, 93, fore parts; 87, 94, fore parts lateral; 8889, 9596, aedeagus ventral; 90, 97, male sternite VIII; 91, 98, male sternite IX. Scale 0.5 mm: 85, 92; scales 0.1 mm: 8687, 9394; 8889, 9596; 9091, 9798; o: ommatidium." figureDoi="http://doi.org/10.5281/zenodo.11064292" httpUri="https://zenodo.org/record/11064292/files/figure.png" pageId="26" pageNumber="521">Figs. 8889</figureCitation>
)......................................................
<taxonomicName id="4C624D22E944FFEBF616FAFAFA80FAAA" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1102,1350,1382,1404]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="gxaligengwanus" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF616FAFAFAC1FAAD" box="[1102,1287,1382,1403]" italics="true" pageId="26" pageNumber="521">O. gxaligengwanus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E944FFEBF708FAFAFA5AFAAD" bold="true" box="[1360,1436,1382,1403]" pageId="26" pageNumber="521">
<taxonomicNameLabel id="A22557C8E944FFEBF708FAFAFA5AFAAD" box="[1360,1436,1382,1403]" pageId="26" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFA1FFA5AFA05" blockId="26.[151,1437,1000,1521]" pageId="26" pageNumber="521">
69l (69k) Aedeagus long and narrow, at least 5 times as long as broad, with a pair of moderately long curved structures in apical two third and with a pair of elongate fields covered by wavy shaped small scales (
<figureCitation id="13592A24E944FFEBF1B5FA3CFBB1FA60" box="[1005,1143,1440,1462]" captionStart="FIGURES 113121" captionStartId="23.[151,264,1358,1382]" captionTargetBox="[212,1375,191,1316]" captionTargetId="figure-15@23.[197,1390,181,1334]" captionTargetPageId="23" captionText="FIGURES 113121. Octavius undulatus Janák, sp. nov.; 113116, 118119, holotype; 117, paratype. 113, habitus; 114, fore parts; 115, fore parts lateral; 116117, aedeagus ventral; 118, male sternite VIII; 119, male sternite IX. Scale 0.5 mm: 113; scales 0.1 mm: 114115; 116117; 118119; o: ommatidium. 120: Distribution of Octavius subgroup A. 121: Distribution of Octavius subgroup B." figureDoi="http://doi.org/10.5281/zenodo.11064300" httpUri="https://zenodo.org/record/11064300/files/figure.png" pageId="26" pageNumber="521">Figs. 116117</figureCitation>
).................................................................................................
<taxonomicName id="4C624D22E944FFEBF6D3FA22FA80FA02" authority="Janak" authorityName="Janak" authorityYear="2024" box="[1163,1350,1470,1492]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="521" phylum="Arthropoda" rank="species" species="undulatus" status="sp. nov.">
<emphasis id="B916EAB3E944FFEBF6D3FA22FAC1FA05" box="[1163,1287,1470,1491]" italics="true" pageId="26" pageNumber="521">O. undulatus</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B916EAB3E944FFEBF708FA22FA5AFA05" bold="true" box="[1360,1436,1470,1491]" pageId="26" pageNumber="521">
<taxonomicNameLabel id="A22557C8E944FFEBF708FA22FA5AFA05" box="[1360,1436,1470,1491]" pageId="26" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDD36A1E944FFEBF2CFFA47FD03FA26" blockId="26.[151,1437,1000,1521]" box="[151,709,1499,1521]" pageId="26" pageNumber="521">70 (69) Pronotum slightly longer than broad (R &lt;1.1).</paragraph>
</subSubSection>
</treatment>
</document>