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<document id="C188AB042DED67CDD84CF3A5A3F88F68" ID-DOI="10.11646/zootaxa.5005.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5141485" ID-ZooBank="86F985AB-BB8D-474A-8A70-19103C316174" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627465624485" checkinUser="plazi" docAuthor="Mawadda, N. V. Ayisha, Kumar, P. Girish &amp; Sureshan, P. M." docDate="2021" docId="03D49323FFF7FF9919C2FE48FC4EFBF9" docLanguage="en" docName="zootaxa.5005.2.6.pdf" docOrigin="Zootaxa 5005 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyroda salai Giner Mari 1945" docType="treatment" docVersion="3" lastPageNumber="208" masterDocId="FFEDEB5BFFF0FF9E1955FF92FFA7FFAD" masterDocTitle="Additions to the knowledge of the genus Lyroda Say (Hymenoptera: Crabronidae) from India with the descriptions of two new species" masterLastPageNumber="217" masterPageNumber="201" pageNumber="208" updateTime="1699025541757" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8E8C9A8BCCB8270BB2EA20A359864070">Additions to the knowledge of the genus Lyroda Say (Hymenoptera: Crabronidae) from India with the descriptions of two new species</mods:title>
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<mods:namePart id="94C6EF44599A57BD0048B5CE3F314039">Mawadda, N. V. Ayisha</mods:namePart>
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<mods:affiliation id="F1C3873A26CE55C15E4607D73EBAD919">ayishakmr @ gmail. com; https: // orcid. org / 0000 - 0003 - 4649 - 9428</mods:affiliation>
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<mods:namePart id="D8E19D53C29C760402F8AB877CC270BE">Kumar, P. Girish</mods:namePart>
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<mods:affiliation id="3601BBCF442BBF025108138EF4924268">kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
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<mods:namePart id="4E1752A2CD89AE39B2F66BA441E39CF1">Sureshan, P. M.</mods:namePart>
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<mods:affiliation id="FB7B195F91EC25D745324BE65C9E1973">pmsuresh 43 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4722 - 6368</mods:affiliation>
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<mods:date id="E3D5AB44351B8C2709BC0F078162D187">2021</mods:date>
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<mods:number id="A2E1027C50A975DFE28C634833F3F21A">5005</mods:number>
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<treatment id="03D49323FFF7FF9919C2FE48FC4EFBF9" ID-DOI="http://doi.org/10.5281/zenodo.5151751" ID-Zenodo-Dep="5151751" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9919C2FE48FC4EFBF9" httpUri="http://treatment.plazi.org/id/03D49323FFF7FF9919C2FE48FC4EFBF9" lastPageNumber="208" pageId="7" pageNumber="208">
<subSubSection id="C36771BEFFF7FF9919C2FE48FDAAFE59" box="[151,525,474,501]" pageId="7" pageNumber="208" type="nomenclature">
<paragraph id="8BC22235FFF7FF9919C2FE48FDAAFE59" blockId="7.[151,525,474,537]" box="[151,525,474,501]" pageId="7" pageNumber="208">
<heading id="D08A9559FFF7FF9919C2FE48FDAAFE59" bold="true" box="[151,525,474,501]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
<taxonomicName id="4C7D59B6FFF7FF9919C2FE48FDAAFE59" ID-CoL="3WTP9" authority="Giner Mari, 1945" authorityName="Giner Mari" authorityYear="1945" box="[151,525,474,501]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="salai">
<emphasis id="B909FE27FFF7FF9919C2FE48FDAAFE59" bold="true" box="[151,525,474,501]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF9919C2FE48FE8AFE58" bold="true" box="[151,301,474,501]" italics="true" pageId="7" pageNumber="208">Lyroda salai</emphasis>
<bibRefCitation id="EFEC5FC4FFF7FF991861FE48FDAAFE59" author="Giner Mari, J." box="[308,525,474,501]" pageId="7" pageNumber="208" pagination="847 - 857" refId="ref11016" refString="Giner Mari, J. (1945) Esfecidos orientales. (Himenopteros de la India inglesa cazados por el P. Ignacio Sala de Castellarnau, S. J. (I). - 2 a serie: Esfecidos). Las Ciencias, 10, 847 - 857." type="journal article" year="1945">Giner Marí, 1945</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FE6DFE80FDB4" box="[151,295,511,537]" pageId="7" pageNumber="208" type="description">
<paragraph id="8BC22235FFF7FF9919C2FE6DFE80FDB4" blockId="7.[151,525,474,537]" box="[151,295,511,537]" pageId="7" pageNumber="208">
(
<figureCitation id="13463EB0FFF7FF9919CAFE6DFEB8FDB4" box="[159,287,511,537]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Figs 19–27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FDD1FB78FDF6" box="[151,1247,579,603]" pageId="7" pageNumber="208" type="reference_group">
<paragraph id="8BC22235FFF7FF9919C2FDD1FB78FDF6" blockId="7.[151,1247,579,603]" box="[151,1247,579,603]" pageId="7" pageNumber="208">
<treatmentCitationGroup id="AB6D051BFFF7FF9919C2FDD1FB78FDF6" box="[151,1247,579,603]" pageId="7" pageNumber="208">
<treatmentCitation id="0ADC0424FFF7FF9919C2FDD1FDA4FDF6" author="Giner Mari, J." box="[151,515,579,603]" page="854" pageId="7" pageNumber="208" year="1945">
<taxonomicName id="4C7D59B6FFF7FF9919C2FDD1FDA4FDF6" ID-CoL="3WTP9" authority="Giner Mari, 1945: 854" authorityName="Giner Mari" authorityPageNumber="854" authorityYear="1945" box="[151,515,579,603]" class="Insecta" family="Crabronidae" genus="Lyroda" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="salai">
<emphasis id="B909FE27FFF7FF9919C2FDD1FEB1FDF6" box="[151,278,579,603]" italics="true" pageId="7" pageNumber="208">Lyroda salai</emphasis>
<materialsCitation id="3B152868FFF7FF991848FDD1FDA4FDF6" box="[285,515,579,603]" collectionCode="MNCN" country="India" location="Khandala" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="2" stateProvince="Maharashtra" typeStatus="holotype">
<bibRefCitation id="EFEC5FC4FFF7FF991848FDD1FDA4FDF6" author="Giner Mari, J." box="[285,515,579,603]" pageId="7" pageNumber="208" pagination="847 - 857" refId="ref11016" refString="Giner Mari, J. (1945) Esfecidos orientales. (Himenopteros de la India inglesa cazados por el P. Ignacio Sala de Castellarnau, S. J. (I). - 2 a serie: Esfecidos). Las Ciencias, 10, 847 - 857." type="journal article" year="1945">Giner Marí, 1945: 854</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="9D7BE9BCFFF7FF991B58FDD6FD86FDF6" box="[525,545,580,603]" pageId="7" pageNumber="208" type="female">♀</specimenCount>
.
<typeStatus id="54C69C97FFF7FF991B78FDD1FD28FDF6" box="[557,655,579,603]" pageId="7" pageNumber="208" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D7BE9BCFFF7FF991BCCFDD6FD0BFDF6" box="[665,684,580,603]" pageId="7" pageNumber="208" type="female">♀</specimenCount>
,
<collectingCountry id="F36A62A5FFF7FF991BEDFDD1FD57FDF6" box="[696,752,579,603]" name="India" pageId="7" pageNumber="208">India</collectingCountry>
:
<collectingRegion id="49B9ECD7FFF7FF991BAFFDD1FCDAFDF6" box="[762,893,579,603]" country="India" name="Maharashtra" pageId="7" pageNumber="208">Maharashtra</collectingRegion>
:
<location id="8EA274EEFFF7FF991AD3FDD1FC40FDF6" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9919C2FE48FC4EFBF9:8EA274EEFFF7FF991AD3FDD1FC40FDF6" box="[902,999,579,603]" country="India" name="Khandala" pageId="7" pageNumber="208" stateProvince="Maharashtra">Khandala</location>
near Bombay (MNCN).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF9919C2FD1BFCB6FC32" pageId="7" pageNumber="208" type="diagnosis">
<paragraph id="8BC22235FFF7FF9919C2FD1BFCB6FC32" blockId="7.[151,1437,649,1108]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF9919C2FD1BFEB5FD0E" bold="true" box="[151,274,649,675]" pageId="7" pageNumber="208">Diagnosis.</emphasis>
Anterior margin of clypeus with two teeth medially, two sets of three teeth laterally and two sets of two small intermediate teeth. Median set of clypeal teeth narrow and cuspidate (
<figureCitation id="13463EB0FFF7FF991AA2FD3FFBEAFD6A" box="[1015,1101,685,711]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Figs 21</figureCitation>
&amp;
<figureCitation id="13463EB0FFF7FF991D26FD3FFB34FD65" box="[1139,1171,685,712]" captionStart="FIGURES 58–64" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[181,1400,189,1912]" captionTargetId="figure-18@13.[178,1409,181,1918]" captionTargetPageId="13" captionText="FIGURES 58–64. Anterior clypeal margins: 58. Lyroda aurea sp. nov. holotype ♀; 59. Lyroda nuda sp. nov. holotype ♀; 60. Lyroda salai Giner Marí. ♀; 61. Lyroda binghami Tsuneki. ♀; 62. Lyroda binghami Tsuneki. ♂; 63. Lyroda formosa (Smith). ♀; 64. Lyroda formosa (Smith). ♂." figureDoi="http://doi.org/10.5281/zenodo.5141505" httpUri="https://zenodo.org/record/5141505/files/figure.png" pageId="7" pageNumber="208">60</figureCitation>
). Forewing with blackish fascia crossing two submarginal cells and apex of second discoidal cell (
<figureCitation id="13463EB0FFF7FF991AA8FD43FBF4FD41" box="[1021,1107,721,748]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Fig. 25</figureCitation>
). Dense silvery setae present on base of antenna, frons, clypeus, gena, tibiae, femora, and both lateral and sternal areas of thorax (
<figureCitation id="13463EB0FFF7FF991C59FD67FA36FCA2" box="[1292,1425,757,783]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Figs 19–24</figureCitation>
). Propleuron with oblique furrow. Mesopleuron convex. Propodeum with dorsal median furrow which does not extend to declivous area of propodeum (
<figureCitation id="13463EB0FFF7FF991B1CFCAFFD38FCFA" box="[585,671,829,855]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Fig. 23</figureCitation>
). Basal platform of Gt1 approximately triangular. Silvery pile band on Gt1–Gt3, sparsely distributed medio dorsally, appearing somewhat interrupted (
<figureCitation id="13463EB0FFF7FF991D68FCF3FB36FCD1" box="[1085,1169,865,892]" captionStart="FIGURES 19–27" captionStartId="8.[151,264,1944,1968]" captionTargetBox="[185,1398,193,1912]" captionTargetId="figure-17@8.[177,1409,181,1920]" captionTargetPageId="8" captionText="FIGURES 19–27. Lyroda salai Giner Marí. ♀. 19. Habitus, lateral view; 20. Head, frontal view; 21. Lower half of head, fron- tal view; 22. Head &amp; thorax; 23. Propodeum, dorsal view; 24. Thorax, lateral view; 25. Forewing; 26. Gaster, dorsal view; 27. Antennae." figureDoi="http://doi.org/10.5281/zenodo.5141493" httpUri="https://zenodo.org/record/5141493/files/figure.png" pageId="7" pageNumber="208">Fig. 26</figureCitation>
). Hind tibiae with eight spines on posterior external margin, claws without teeth.
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF991992FC3BFE1EFC4A" pageId="7" pageNumber="208" type="discussion">
<paragraph id="8BC22235FFF7FF991992FC3BFE1EFC4A" blockId="7.[151,1437,649,1108]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF991992FC3BFE93FC6E" bold="true" box="[199,308,937,963]" pageId="7" pageNumber="208">Remarks</emphasis>
. According to the original description by
<bibRefCitation id="EFEC5FC4FFF7FF991A5CFC3BFC7CFC69" author="Giner Mari, J." box="[777,987,937,964]" pageId="7" pageNumber="208" pagination="847 - 857" refId="ref11016" refString="Giner Mari, J. (1945) Esfecidos orientales. (Himenopteros de la India inglesa cazados por el P. Ignacio Sala de Castellarnau, S. J. (I). - 2 a serie: Esfecidos). Las Ciencias, 10, 847 - 857." type="journal article" year="1945">Giner Marí (1945)</bibRefCitation>
, six hind tibial spines are present on the posterior external margin.
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF991992FC63FA2DFBA6" box="[199,1418,1009,1036]" pageId="7" pageNumber="208" type="distribution">
<paragraph id="8BC22235FFF7FF991992FC63FA2DFBA6" blockId="7.[151,1437,649,1108]" box="[199,1418,1009,1036]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF991992FC63FEC6FBA6" bold="true" box="[199,353,1009,1035]" pageId="7" pageNumber="208">Distribution.</emphasis>
<collectingCountry id="F36A62A5FFF7FF991832FC63FE02FBA6" box="[359,421,1009,1035]" name="India" pageId="7" pageNumber="208">India</collectingCountry>
:
<collectingRegion id="49B9ECD7FFF7FF9918E5FC63FE5CFBA6" box="[432,507,1009,1035]" country="India" name="Kerala" pageId="7" pageNumber="208">Kerala</collectingRegion>
<emphasis id="B909FE27FFF7FF991B54FC63FD06FBA6" bold="true" box="[513,673,1009,1035]" pageId="7" pageNumber="208">(new record),</emphasis>
<collectingRegion id="49B9ECD7FFF7FF991BFDFC63FC9FFBA6" box="[680,824,1009,1035]" country="India" name="Maharashtra" pageId="7" pageNumber="208">Maharashtra</collectingRegion>
.
<emphasis id="B909FE27FFF7FF991A17FC63FC11FBA6" box="[834,950,1009,1035]" italics="true" pageId="7" pageNumber="208">Elsewhere</emphasis>
:
<collectingCountry id="F36A62A5FFF7FF991A91FC63FBA0FBA6" box="[964,1031,1009,1035]" name="China" pageId="7" pageNumber="208">China</collectingCountry>
(
<bibRefCitation id="EFEC5FC4FFF7FF991D43FC63FB70FBA1" author="Giner Mari, J." box="[1046,1239,1009,1036]" pageId="7" pageNumber="208" pagination="847 - 857" refId="ref11016" refString="Giner Mari, J. (1945) Esfecidos orientales. (Himenopteros de la India inglesa cazados por el P. Ignacio Sala de Castellarnau, S. J. (I). - 2 a serie: Esfecidos). Las Ciencias, 10, 847 - 857." type="journal article" year="1945">Giner Marí 1945</bibRefCitation>
;
<bibRefCitation id="EFEC5FC4FFF7FF991DB7FC63FAD9FBA6" author="Li, T. &amp; Cai, W. &amp; Li, Q." box="[1250,1406,1009,1036]" pageId="7" pageNumber="208" pagination="143 - 147" refId="ref11187" refString="Li, T., Cai, W. &amp; Li, Q. (2009) Key to the Oriental species of Lyroda Say, with description of a new species from China (Hymenoptera: Crabronidae). Entomologica Americana, 115, 143 - 147. https: // doi. org / 10.1664 / 07 - RA- 005.1" type="journal article" year="2009">
Li
<emphasis id="B909FE27FFF7FF991C57FC60FA93FBA6" box="[1282,1332,1009,1035]" italics="true" pageId="7" pageNumber="208">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36771BEFFF7FF991992FB87FC4EFBF9" pageId="7" pageNumber="208" type="materials_examined">
<paragraph id="8BC22235FFF7FF991992FB87FC4EFBF9" blockId="7.[151,1437,649,1108]" pageId="7" pageNumber="208">
<emphasis id="B909FE27FFF7FF991992FB87FE0CFB82" bold="true" box="[199,427,1045,1071]" pageId="7" pageNumber="208">Material examined</emphasis>
.
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<collectingCountry id="F36A62A5FFF7FF9918EEFB84FDA8FB9D" box="[443,527,1046,1072]" name="India" pageId="7" pageNumber="208">INDIA</collectingCountry>
:
<collectingRegion id="49B9ECD7FFF7FF991B4FFB87FDC0FB82" box="[538,615,1045,1071]" country="India" name="Kerala" pageId="7" pageNumber="208">Kerala</collectingRegion>
,
<collectingMunicipality id="6BA6B84FFFF7FF991B21FB87FC8CFB82" box="[628,811,1045,1071]" pageId="7" pageNumber="208">Thrissur district</collectingMunicipality>
,
<location id="8EA274EEFFF7FF991A62FB87FC22FB82" LSID="urn:lsid:plazi:treatment:03D49323FFF7FF9919C2FE48FC4EFBF9:8EA274EEFFF7FF991A62FB87FC22FB82" box="[823,901,1045,1071]" country="India" latitude="10.645833" longLatPrecision="21" longitude="76.37667" municipality="Thrissur district" name="Elanad" pageId="7" pageNumber="208" stateProvince="Kerala">Elanad</location>
(
<specimenCount id="9D7BE9BCFFF7FF991ACDFB87FC67FB9D" box="[920,960,1045,1072]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
) (
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,
<geoCoordinate id="EE4944F2FFF7FF991D2FFB87FAAFFB9D" box="[1146,1288,1045,1072]" degrees="76" direction="east" minutes="22" orientation="longitude" pageId="7" pageNumber="208" precision="15" seconds="36" value="76.37667">76°22’36’’E</geoCoordinate>
),
<date id="FFC304F5FFF7FF991C48FB87FA3FFB82" box="[1309,1432,1045,1071]" pageId="7" pageNumber="208" value="2018-11-19">
<collectingDate id="EF87FD1DFFF7FF991C48FB87FA3FFB82" box="[1309,1432,1045,1071]" pageId="7" pageNumber="208" value="2018-11-19">19.xi.2018</collectingDate>
</date>
,
<collectorName id="268847E3FFF7FF9919C2FBABFE75FBFE" box="[151,466,1081,1108]" pageId="7" pageNumber="208">Coll. N.V. Ayisha Mawadda</collectorName>
, ZSIK
<collectorName id="268847E3FFF7FF991B76FBABFD31FBF9" box="[547,662,1081,1108]" pageId="7" pageNumber="208">Regd. No.</collectorName>
ZSI/ WGRC/IR/INV/115044
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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