272 lines
36 KiB
XML
272 lines
36 KiB
XML
<document id="D3B8357A2012494A2EC672C64A0F354F" ID-CLB-Dataset="2380" ID-DOI="10.5281/zenodo.5739643" ID-GBIF-Dataset="f9439543-76ca-43e9-aca5-0a8d3899a2b1" ID-ISSN="0374-1036" ID-Zenodo-Dep="5739643" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638252251656" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2013" docId="03CEDF09FFFCF349D1A593E9D8A1FC81" docLanguage="en" docName="ActaEntMusNatPra.2013.53.1.177-208.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus sudanensis" docType="treatment" docVersion="7" lastPageNumber="186" masterDocId="FFF7A771FFF4F340D0219333DB72FFF2" masterDocTitle="Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="208" masterPageNumber="177" pageNumber="185" updateTime="1699471184067" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BCAEF504164747A61E5104B5B34DE0D9">Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="48ABD5C7C9734BF5A386122A1224BC9E">Makranczy, György</mods:namePart>
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<mods:affiliation id="5BACAAD6DBCD95A66297F5B8DFC68D0D">Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross utca 13, Hungary</mods:affiliation>
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<mods:nameIdentifier id="2C930300724D7342A0B6239E7EFC6072" type="email">makranczy@zoo.nhmus.hu</mods:nameIdentifier>
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<mods:title id="D4259C4C6D6D4EA09C13C99218EB89EE">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="639AD0B5459972507A805BC865D53126">
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<mods:date id="592968F335459EB1572824E77B7652E7">2013</mods:date>
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<mods:number id="152EFFC371B43FDF78F066226BA1B062">2013-07-15</mods:number>
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<mods:number id="48FEE34936CE1B209CA4AF8DCC0836B2">53</mods:number>
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<mods:identifier id="ACE651C31A78EE54878518CBCD70A9B0" type="DOI">10.5281/zenodo.5739643</mods:identifier>
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<treatment id="03CEDF09FFFCF349D1A593E9D8A1FC81" ID-DOI="http://doi.org/10.5281/zenodo.5739657" ID-GBIF-Taxon="192662471" ID-Zenodo-Dep="5739657" LSID="urn:lsid:plazi:treatment:03CEDF09FFFCF349D1A593E9D8A1FC81" httpUri="http://treatment.plazi.org/id/03CEDF09FFFCF349D1A593E9D8A1FC81" lastPageId="9" lastPageNumber="186" pageId="8" pageNumber="185">
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<subSubSection id="C37D3D94FFFCF348D1A593E9D80BFF01" box="[388,889,218,243]" pageId="8" pageNumber="185" type="nomenclature">
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<paragraph id="8BD86E1FFFFCF348D1A593E9D80BFF01" blockId="8.[388,889,218,243]" box="[388,889,218,243]" pageId="8" pageNumber="185">
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<heading id="D090D973FFFCF348D1A593E9D80BFF01" bold="true" box="[388,889,218,243]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="185" reason="3">
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<taxonomicName id="4C67159CFFFCF348D1A593E9D80BFF01" ID-CoL="56H6D" authority="(Scheerpeltz, 1974)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[388,889,218,243]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">
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<emphasis id="B913B20DFFFCF348D1A593E9D80BFF01" bold="true" box="[388,889,218,243]" pageId="8" pageNumber="185">
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<emphasis id="B913B20DFFFCF348D1A593E9D9E9FF01" bold="true" box="[388,667,218,243]" italics="true" pageId="8" pageNumber="185">Thinodromus sudanensis</emphasis>
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(Scheerpeltz, 1974)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D3D94FFFCF348D23993CAD994FEFE" box="[536,742,249,269]" pageId="8" pageNumber="185" type="description">
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<paragraph id="8BD86E1FFFFCF348D23993CAD994FEFE" blockId="8.[151,1127,249,532]" box="[536,742,249,269]" pageId="8" pageNumber="185">
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(
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<figureCitation id="135C729AFFFCF348D23F93CAD9F2FEFF" box="[542,640,249,269]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="8" pageNumber="185">Figs 17–18</figureCitation>
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,
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<figureCitation id="135C729AFFFCF348D2A893C9D9D2FEFF" box="[649,672,250,269]" captionStart="Figs 19–27. 19" captionStartId="15.[113,150,1493,1512]" captionTargetBox="[120,1082,219,1480]" captionTargetId="figure-10@15.[119,1082,219,1481]" captionTargetPageId="15" captionText="Figs 19–27. 19 – Thinodromus facilis Gildenkov, 2000, female protarsus; 20 – T. gildenkovi sp. nov., male protarsus; 21 – T. gildenkovi sp. nov., male, fringe of tergite VII. Antennae. 22 – T. sudanensis (Scheerpeltz, 1974), female; 23 – T. meridionalis sp. nov., female; 24–25 – T. gildenkovi sp. nov. (24 – female; 25 – male); 26–27 – T. dasys Gildenkov, 2000 (26 – female; 27 – male) Scale bar: 0.065 mm for 19, 0.08 mm for 20, 0.12 mm for 21, 0.24 mm for 24–25, 0.3 mm for 22 and 26–27, 0.33 mm for 23." figureDoi="http://doi.org/10.5281/zenodo.5739655" httpUri="https://zenodo.org/record/5739655/files/figure.png" pageId="8" pageNumber="185">22</figureCitation>
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,
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<figureCitation id="135C729AFFFCF348D28893CAD9ADFEFE" box="[681,735,249,268]" captionStart="Figs 77–80. 77–78" captionStartId="24.[151,188,1545,1564]" captionTargetBox="[164,1110,235,1477]" captionTargetId="figure-10@24.[162,1114,233,1480]" captionTargetPageId="24" captionText="Figs 77–80. 77–78 – Thinodromus sudanensis (Scheerpeltz, 1974), female (77 – tergite X; 78 – ringstructure and spermatheca); 79 – T. capensis (Bernhauer, 1934), female ringstructure; 80 – T. meridionalis sp. nov., female ringstructure with spermatheca. Scale bar: 0.09 mm for 79–80, 0.11 mm for 78, 0.14 mm for 77." figureDoi="http://doi.org/10.5281/zenodo.5739679" httpUri="https://zenodo.org/record/5739679/files/figure.png" pageId="8" pageNumber="185">77–78</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37D3D94FFFCF348D0B6922ED94AFEBB" pageId="8" pageNumber="185" type="reference_group">
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<paragraph id="8BD86E1FFFFCF348D0B6922ED9DEFEDD" blockId="8.[151,1127,249,532]" box="[151,684,284,303]" pageId="8" pageNumber="185">
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<treatmentCitationGroup id="AB774931FFFCF348D0B6922ED9DEFEDD" box="[151,684,284,303]" pageId="8" pageNumber="185">
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<treatmentCitation id="0AC6480EFFFCF348D0B6922ED9DAFEDD" author="SCHEERPELTZ O." box="[151,680,284,303]" page="5" pageId="8" pageNumber="185" year="1974">
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<taxonomicName id="4C67159CFFFCF348D0B6922ED9DAFEDD" ID-CoL="599H3" authority="Scheerpeltz, 1974 b: 5" authorityName="Scheerpeltz" authorityPageNumber="5" authorityYear="1974" box="[151,680,284,303]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis" subGenus="Carpalimus">
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||
<emphasis id="B913B20DFFFCF348D0B6922EDA7EFEDD" box="[151,268,285,303]" italics="true" pageId="8" pageNumber="185">Trogophloeus</emphasis>
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||
(
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<emphasis id="B913B20DFFFCF348D139922EDA0FFEDD" box="[280,381,285,303]" italics="true" pageId="8" pageNumber="185">Carpalimus</emphasis>
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||
)
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<emphasis id="B913B20DFFFCF348D1A8922EDA95FEDD" box="[393,487,285,303]" italics="true" pageId="8" pageNumber="185">sudanensis</emphasis>
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||
<bibRefCitation id="EFF613EEFFFCF348D1CD922FD9DAFEDD" author="SCHEERPELTZ O." box="[492,680,284,303]" pageId="8" pageNumber="185" pagination="3 - 49" refId="ref14513" refString="SCHEERPELTZ O. 1974 b: Coleoptera aus Nordostafrika, Staphylinidae (Zoolocigal Contribution from the Finnish expeditions to the Sudan No. 35). Acta Entomologica Fennica 31: 3 - 49." type="journal article" year="1974">Scheerpeltz, 1974b: 5</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BD86E1FFFFCF348D0B69204D94AFEBB" blockId="8.[151,1127,249,532]" box="[151,568,310,330]" pageId="8" pageNumber="185">
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<treatmentCitationGroup id="AB774931FFFCF348D0B69204D94AFEBB" box="[151,568,310,330]" pageId="8" pageNumber="185">
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<treatmentCitation id="0AC6480EFFFCF348D0B69204D946FEBB" author="GILDENKOV M. & YU" box="[151,564,310,330]" page=")" pageId="8" pageNumber="185" year="2000">
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||
<taxonomicName id="4C67159CFFFCF348D0B69204D946FEBB" ID-CoL="56H6D" authority=": GILDENKOV (2000: 51)" authorityName="GILDENKOV" authorityPageNumber="51" authorityYear="2000" box="[151,564,310,330]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">
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<emphasis id="B913B20DFFFCF348D0B69204DA1EFEBB" box="[151,364,311,329]" italics="true" pageId="8" pageNumber="185">Thinodromus sudanensis</emphasis>
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:
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<bibRefCitation id="EFF613EEFFFCF348D1579205D946FEBB" author="GILDENKOV M. & YU" box="[374,564,310,329]" pageId="8" pageNumber="185" pagination="43 - 56" refId="ref14247" refString="GILDENKOV M. YU. 2000: Novyy podrod i sem' novykh dlya nauki vidov roda Thinodromus iz efiopskoy zoogeograficheskoy oblast'i (Coleoptera, Staphylinidae). [New subgenus and seven species, new to science, of the genus Thinodromus from Ethiopian zoogeographic region (Coleoptera, Staphylinidae)]. Izvestiya Khar ' kovskogo Entomologicheskogo Obshchestva [Kharkov Entomological Society Gazette] 8 (1): 43 - 56 (in Russian)." type="journal article" year="2000">GILDENKOV (2000: 51)</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37D3D94FFFCF348D0B69253D8ACFDE1" pageId="8" pageNumber="185" type="materials_examined">
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<paragraph id="8BD86E1FFFFCF348D0B69253D8ACFDE1" blockId="8.[151,1127,249,532]" pageId="8" pageNumber="185">
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<emphasis id="B913B20DFFFCF348D0B69253DA0CFE81" bold="true" box="[151,382,351,371]" pageId="8" pageNumber="185">Type material examined.</emphasis>
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||
<materialsCitation id="3B0F6442FFFCF348D1A49252D9B8FE31" ID-GBIF-Occurrence="3458217334" collectingDate="1963-02-19" collectionCode="MZHF" collectorName="O. Scheerpeltz" country="Sudan" county="Wau" determinerName="Makranczy" latitude="4.1666665" location="Dahr" longLatPrecision="1307" longitude="10.166667" municipality="Bahr" pageId="8" pageNumber="185" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="54DCD0BDFFFCF348D1A49252DAABFE80" box="[389,473,353,372]" pageId="8" pageNumber="185" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCount id="9D61A596FFFCF348D1C79269DA84FE81" box="[486,502,346,371]" pageId="8" pageNumber="185" type="female">♀</specimenCount>
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, “
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<collectingCountry id="F3702E8FFFFCF348D22D9253D936FE81" box="[524,580,352,371]" name="Sudan" pageId="8" pageNumber="185">Sudan</collectingCountry>
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;
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<location id="8EB838C4FFFCF348D2719253D90FFE81" LSID="urn:lsid:plazi:treatment:03CEDF09FFFCF349D1A593E9D8A1FC81:8EB838C4FFFCF348D2719253D90FFE81" box="[592,637,352,371]" country="Sudan" county="Wau" latitude="4.1666665" longLatPrecision="1307" longitude="10.166667" municipality="Bahr" name="Dahr" pageId="8" pageNumber="185">Dahr</location>
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[
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<collectingMunicipality id="6BBCF465FFFCF348D2A39253D9DFFE81" box="[642,685,352,371]" pageId="8" pageNumber="185">Bahr</collectingMunicipality>
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] el Ghazal.;
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<collectingCounty id="62B91693FFFCF348D3019252D835FE86" box="[800,839,353,372]" pageId="8" pageNumber="185">Wau</collectingCounty>
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[
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<geoCoordinate id="EE5308D8FFFCF348D3759253D8ECFE86" box="[852,926,351,372]" degrees="04" direction="north" minutes="10" orientation="latitude" pageId="8" pageNumber="185" precision="925" value="4.1666665">04°10′N</geoCoordinate>
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,
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<geoCoordinate id="EE5308D8FFFCF348D38B9253D883FE86" box="[938,1009,351,372]" degrees="10" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="185" precision="925" value="10.166667">10°10′E</geoCoordinate>
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] \
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<date id="FFD948DFFFFCF348D42A9253DF11FE81" box="[1035,1123,352,371]" pageId="8" pageNumber="185" value="1963-02-19">
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<collectingDate id="EF9DB137FFFCF348D42A9253DF11FE81" box="[1035,1123,352,371]" pageId="8" pageNumber="185" value="1963-02-19">19.2.1963</collectingDate>
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||
</date>
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||
; [R.] Linnavuori \
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<taxonomicName id="4C67159CFFFCF348D10C9248DAD4FE7C" box="[301,422,379,398]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
; (Carpalimus); sudanensis; nov.spec. \ ex coll.; Scheerpeltz \ Typus;
|
||
<taxonomicName id="4C67159CFFFCF348D3CA9248DF16FE7C" box="[1003,1124,379,398]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
; sudanensis;
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||
<collectorName id="26920BC9FFFCF348D0DE92A5DA0CFE5A" box="[255,382,405,425]" pageId="8" pageNumber="185">O. Scheerpeltz</collectorName>
|
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\
|
||
<typeStatus id="54DCD0BDFFFCF348D1AE92A6DA99FE5A" box="[399,491,405,424]" pageId="8" pageNumber="185" type="holotype">Holotypus</typeStatus>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D1D292A6D91EFE5A" box="[499,620,405,424]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D25492A6D84FFE5A" authority="Scheerpeltz" authorityName="Scheerpeltz" box="[629,829,405,424]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">sudanensis Scheerpeltz</taxonomicName>
|
||
; ver. Makranczy, 2011 \
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||
<taxonomicName id="4C67159CFFFCF348D43192A6DBCFFE36" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D0E89283DAD1FE31" authority="(Scheerpeltz)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[201,419,432,451]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">sudanensis (Scheerpeltz)</taxonomicName>
|
||
; det.
|
||
<determinerName id="7E0482DBFFFCF348D1F79283D945FE31" box="[470,567,432,451]" pageId="8" pageNumber="185">Makranczy</determinerName>
|
||
, 2011” (
|
||
<collectionCode id="ED76F6DAFFFCF348D2A59283D9B6FE31" box="[644,708,432,451]" pageId="8" pageNumber="185">MZHF</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0F6442FFFCF348D2F49282D8A8FDE1" ID-GBIF-Occurrence="3458217341" collectingDate="1963-02-19" collectionCode="NHMW" collectorName="O. Scheerpeltz" country="Sudan" county="Wau" determinerName="Makranczy" location="Dahr" municipality="Bahr" pageId="8" pageNumber="185" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<typeStatus id="54DCD0BDFFFCF348D2F49282D856FE30" box="[725,804,433,452]" pageId="8" pageNumber="185" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
<specimenCount id="9D61A596FFFCF348D3139299D830FE31" box="[818,834,426,451]" pageId="8" pageNumber="185" type="female">♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="F3702E8FFFFCF348D3799283D8E3FE31" box="[856,913,432,451]" name="Sudan" pageId="8" pageNumber="185">Sudan</collectingCountry>
|
||
;
|
||
<location id="8EB838C4FFFCF348D3BD9283D8BBFE31" LSID="urn:lsid:plazi:treatment:03CEDF09FFFCF349D1A593E9D8A1FC81:8EB838C4FFFCF348D3BD9283D8BBFE31" box="[924,969,432,451]" country="Sudan" county="Wau" municipality="Bahr" name="Dahr" pageId="8" pageNumber="185">Dahr</location>
|
||
[
|
||
<collectingMunicipality id="6BBCF465FFFCF348D3EE9283D888FE31" box="[975,1018,432,451]" pageId="8" pageNumber="185">Bahr</collectingMunicipality>
|
||
] el Ghazal.;
|
||
<collectingCounty id="62B91693FFFCF348D0B692F8DBCCFE2C" box="[151,190,459,478]" pageId="8" pageNumber="185">Wau</collectingCounty>
|
||
\
|
||
<date id="FFD948DFFFFCF348D0F192F8DA5AFE2C" box="[208,296,459,478]" pageId="8" pageNumber="185" value="1963-02-19">
|
||
<collectingDate id="EF9DB137FFFCF348D0F192F8DA5AFE2C" box="[208,296,459,478]" pageId="8" pageNumber="185" value="1963-02-19">19.2.1963</collectingDate>
|
||
</date>
|
||
; [R.] Linnavuori \
|
||
<taxonomicName id="4C67159CFFFCF348D1EB92F8D931FE2C" box="[458,579,459,478]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
; (Carpalimus); sudanensis; n.sp. \ ex coll.; Scheerpeltz \ Typus;
|
||
<taxonomicName id="4C67159CFFFCF348D0B692D6DA62FE0A" box="[151,272,485,504]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
; sudanensis;
|
||
<collectorName id="26920BC9FFFCF348D1A092D5DA8DFE0A" box="[385,511,485,505]" pageId="8" pageNumber="185">O. Scheerpeltz</collectorName>
|
||
\ sudanensis; Schp. \
|
||
<typeStatus id="54DCD0BDFFFCF348D29B92D5D863FE0B" box="[698,785,486,505]" pageId="8" pageNumber="185" type="paratype">Paratypus</typeStatus>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D33892D6D8E0FE0A" box="[793,914,485,504]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D3BA92D6DF11FE0A" authority="Scheerpeltz" authorityName="Scheerpeltz" box="[923,1123,485,504]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">sudanensis Scheerpeltz</taxonomicName>
|
||
; ver. Makranczy, 2011 \
|
||
<taxonomicName id="4C67159CFFFCF348D17E9133DAA7FDE1" authorityName="Kraatz" authorityYear="1857" box="[351,469,512,531]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C67159CFFFCF348D1FC9133D9C6FDE1" authority="(Scheerpeltz)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[477,692,512,531]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="185" phylum="Arthropoda" rank="species" species="sudanensis">sudanensis (Scheerpeltz)</taxonomicName>
|
||
; det.
|
||
<determinerName id="7E0482DBFFFCF348D2C09133D830FDE1" box="[737,834,512,531]" pageId="8" pageNumber="185">Makranczy</determinerName>
|
||
, 2011” (
|
||
<collectionCode id="ED76F6DAFFFCF348D3A89133D8A6FDE1" box="[905,980,512,531]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="185" type="Museum">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D3D94FFFCF348D0B69118DAC3F9D6" pageId="8" pageNumber="185" type="description">
|
||
<paragraph id="8BD86E1FFFFCF348D0B69118DAC3F9D6" blockId="8.[151,1128,555,1572]" pageId="8" pageNumber="185">
|
||
<emphasis id="B913B20DFFFCF348D0B69118DA47FDB1" bold="true" box="[151,309,555,579]" pageId="8" pageNumber="185">Redescription.</emphasis>
|
||
<emphasis id="B913B20DFFFCF348D118911EDABCFDB6" box="[313,462,557,580]" italics="true" pageId="8" pageNumber="185">Measurements</emphasis>
|
||
(in mm, n = 2): HW = 0.63 (0.625–0.63); TW = 0.53 (0.515–0.54); PW = 0.70 (0.68–0.72); SW = 0.81 (0.79–0.83); AW = 0.91 (0.88–0.94); HL = 0.39 (0.37–0.40); EL = 0.25 (0.25–0.255); TL = 0.02 (0.015–0.02); PL = 0.50 (0.48–0.51); SL = 0.79 (0.77–0.80); SC = 0.75 (0.73–0.77); BL = 3.35 (3.19–3.51); FB = 1.74 (1.67–1.80).
|
||
<emphasis id="B913B20DFFFCF348D34B91BEDF53FD56" box="[874,1057,653,676]" italics="true" pageId="8" pageNumber="185">Lustre and colour</emphasis>
|
||
: Body rather shining due to shiny interspaces of not so dense punctation. Forebody and abdomen pitch black with almost absence of brownish or reddish tint. Mouthparts and antennae dark brown, legs reddish medium brown with apices of tibiae and tarsi much lighter, reddish medium brown.
|
||
<emphasis id="B913B20DFFFCF348D40691DEDA57FCD6" italics="true" pageId="8" pageNumber="185">Shape and sculpture</emphasis>
|
||
: Head (
|
||
<figureCitation id="135C729AFFFCF348D156903FDAB0FCD6" box="[375,450,780,804]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="8" pageNumber="185">Fig. 17</figureCitation>
|
||
) rather transverse, eyes occupy sides, temples barely discernible. Pronotum transverse, sides straight in posterior half, posterior corners rounded, inconspicuous, first half of sides and anterior corners strongly rounded. Sides of a rather deep horseshoe-shaped impression anteriorly with an adjoining shallow depression on lateral parts of disc; center of disc marked by a pair of smaller, similarly shallow impressions. Deflexed margin wider than usual and fully apparent laterally, although more prominent in posterior half. Elytra (
|
||
<figureCitation id="135C729AFFFCF348D433909FDF2DFC36" box="[1042,1119,940,964]" captionStart="Figs 13–18. 13–16" captionStartId="14.[151,187,1532,1551]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@14.[151,1126,219,1497]" captionTargetPageId="14" captionText="Figs 13–18. 13–16 – Thinodromus nigerius Gildenkov, 2000 (13 – head and pronotum;14 – female antenna; 15 – male antenna; 16 – elytron and abdominal base); 17–18 – T. sudanensis (Scheerpeltz, 1974) (17 – head and pronotum; 18 – elytron and abdominal base). Scale bar: 0.5 mm for 13–16, 0.7 mm for 17–18." figureDoi="http://doi.org/10.5281/zenodo.5739653" httpUri="https://zenodo.org/record/5739653/files/figure.png" pageId="8" pageNumber="185">Fig. 18</figureCitation>
|
||
) combined significantly broader than long, gently dilated towards apex; a shallow oval impression posterior of scutellum and a feeble oblique impression across anterior half of disc. Posterior elytral margin with wider membranous lobe not apparent on whole length but conspicuously pulled out near outer corner in a small concavity of posterior margin. Apex of abdominal tergite VII with palisade fringe (widest medially).
|
||
<emphasis id="B913B20DFFFCF348D245977ED8D8FB96" box="[612,938,1101,1124]" italics="true" pageId="8" pageNumber="185">Punctation and microsculpture</emphasis>
|
||
: Head punctation medium large and medium deep, but with rather sparse interspaces roughly equal to puncture diameters. Pronotal punctation with similarly sparse and medium deep, somewhat varying sized punctures, only posterior corners with some discernible scabrous microsculpture. Elytra with very large and deep punctures, more close to each other (interspaces less than 1/3 of puncture diameters), but surface without microsculpture, so shiny. Abdominal tergites with medium deep and medium sparse punctation, interspaces often larger than puncture diameters; traces of microsculpture only in transversal grooves posteriad basal ridges.
|
||
<emphasis id="B913B20DFFFCF348D31E961ED8C8FAB6" box="[831,954,1325,1348]" italics="true" pageId="8" pageNumber="185">Pubescence</emphasis>
|
||
: Forebody, even the eyes, covered by conspicuously long, erect, often strong and dark setae, mostly of uniform size and sparse spacing. Setae on head and posterior part of pronotum appear shorter and more dense. On abdominal terga with setae sparse, long and erect, on posterior edge a row of very long setae.
|
||
<emphasis id="B913B20DFFFCF348D135969ED9C8FA36" box="[276,698,1453,1476]" italics="true" pageId="8" pageNumber="185">Primary and secondary sexual features</emphasis>
|
||
: Female antennae (
|
||
<figureCitation id="135C729AFFFCF348D3B1969FD892FA36" box="[912,992,1452,1476]" captionStart="Figs 19–27. 19" captionStartId="15.[113,150,1493,1512]" captionTargetBox="[120,1082,219,1480]" captionTargetId="figure-10@15.[119,1082,219,1481]" captionTargetPageId="15" captionText="Figs 19–27. 19 – Thinodromus facilis Gildenkov, 2000, female protarsus; 20 – T. gildenkovi sp. nov., male protarsus; 21 – T. gildenkovi sp. nov., male, fringe of tergite VII. Antennae. 22 – T. sudanensis (Scheerpeltz, 1974), female; 23 – T. meridionalis sp. nov., female; 24–25 – T. gildenkovi sp. nov. (24 – female; 25 – male); 26–27 – T. dasys Gildenkov, 2000 (26 – female; 27 – male) Scale bar: 0.065 mm for 19, 0.08 mm for 20, 0.12 mm for 21, 0.24 mm for 24–25, 0.3 mm for 22 and 26–27, 0.33 mm for 23." figureDoi="http://doi.org/10.5281/zenodo.5739655" httpUri="https://zenodo.org/record/5739655/files/figure.png" pageId="8" pageNumber="185">Fig. 22</figureCitation>
|
||
) moderately elongate, middle antennomeres (articles 4–5) about as long as wide, penultimate antennomeres (articles 9–10) slightly transverse. Male: unknown; female: tergite X (
|
||
<figureCitation id="135C729AFFFCF348D3A196DFD8BDF9F6" box="[896,975,1516,1540]" captionStart="Figs 77–80. 77–78" captionStartId="24.[151,188,1545,1564]" captionTargetBox="[164,1110,235,1477]" captionTargetId="figure-10@24.[162,1114,233,1480]" captionTargetPageId="24" captionText="Figs 77–80. 77–78 – Thinodromus sudanensis (Scheerpeltz, 1974), female (77 – tergite X; 78 – ringstructure and spermatheca); 79 – T. capensis (Bernhauer, 1934), female ringstructure; 80 – T. meridionalis sp. nov., female ringstructure with spermatheca. Scale bar: 0.09 mm for 79–80, 0.11 mm for 78, 0.14 mm for 77." figureDoi="http://doi.org/10.5281/zenodo.5739679" httpUri="https://zenodo.org/record/5739679/files/figure.png" pageId="8" pageNumber="185">Fig. 77</figureCitation>
|
||
), spermatheca and ringstructure (
|
||
<figureCitation id="135C729AFFFCF348D177953FDAD7F9D6" box="[342,421,1548,1572]" captionStart="Figs 77–80. 77–78" captionStartId="24.[151,188,1545,1564]" captionTargetBox="[164,1110,235,1477]" captionTargetId="figure-10@24.[162,1114,233,1480]" captionTargetPageId="24" captionText="Figs 77–80. 77–78 – Thinodromus sudanensis (Scheerpeltz, 1974), female (77 – tergite X; 78 – ringstructure and spermatheca); 79 – T. capensis (Bernhauer, 1934), female ringstructure; 80 – T. meridionalis sp. nov., female ringstructure with spermatheca. Scale bar: 0.09 mm for 79–80, 0.11 mm for 78, 0.14 mm for 77." figureDoi="http://doi.org/10.5281/zenodo.5739679" httpUri="https://zenodo.org/record/5739679/files/figure.png" pageId="8" pageNumber="185">Fig. 78</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D3D94FFFDF349D05093E8DA56FE01" pageId="9" pageNumber="186" type="diagnosis">
|
||
<paragraph id="8BD86E1FFFFDF349D05093E8DA56FE01" blockId="9.[113,1089,219,884]" pageId="9" pageNumber="186">
|
||
<emphasis id="B913B20DFFFDF349D05093E8DA18FF01" bold="true" box="[113,362,219,243]" pageId="9" pageNumber="186">Differential diagnosis.</emphasis>
|
||
As the male of
|
||
<taxonomicName id="4C67159CFFFDF349D22D93EFD9E8FF01" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[524,666,220,243]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="sudanensis">
|
||
<emphasis id="B913B20DFFFDF349D22D93EFD9E8FF01" box="[524,666,220,243]" italics="true" pageId="9" pageNumber="186">T. sudanensis</emphasis>
|
||
</taxonomicName>
|
||
is unknown, it is rather difficult to compare it to the other species. Judging by the female genitalia, the most similar species must be
|
||
<taxonomicName id="4C67159CFFFDF349D050922FDB89FEC1" authorityName="Makranczy" authorityYear="2009" box="[113,251,284,307]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="gabonicus">
|
||
<emphasis id="B913B20DFFFDF349D050922FDB89FEC1" box="[113,251,284,307]" italics="true" pageId="9" pageNumber="186">T. gabonicus</emphasis>
|
||
</taxonomicName>
|
||
as both share a more “heart-shaped” ringstructure, with that of
|
||
<taxonomicName id="4C67159CFFFDF349D391922FDF32FEC1" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[944,1088,284,307]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="sudanensis">
|
||
<emphasis id="B913B20DFFFDF349D391922FDF32FEC1" box="[944,1088,284,307]" italics="true" pageId="9" pageNumber="186">T. sudanensis</emphasis>
|
||
</taxonomicName>
|
||
being much wider at the base. The apex of the ringstructure is more square and less broad in
|
||
<taxonomicName id="4C67159CFFFDF349D0AC926FDA64FE81" authorityName="Makranczy" authorityYear="2009" box="[141,278,348,371]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="gabonicus">
|
||
<emphasis id="B913B20DFFFDF349D0AC926FDA64FE81" box="[141,278,348,371]" italics="true" pageId="9" pageNumber="186">T. gabonicus</emphasis>
|
||
</taxonomicName>
|
||
, while rather equally rounded and with the highest density of pores on both sides of the apex in
|
||
<taxonomicName id="4C67159CFFFDF349D163924FDAA2FE61" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1974" box="[322,464,380,403]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="sudanensis">
|
||
<emphasis id="B913B20DFFFDF349D163924FDAA2FE61" box="[322,464,380,403]" italics="true" pageId="9" pageNumber="186">T. sudanensis</emphasis>
|
||
</taxonomicName>
|
||
; the inner arms of the basal loop are also wider in the latter species. These are the only two species known with such heart-shaped ringstructures, with the apex not pointed and not widened, so it is expected that their male genitalia might also bear similarities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D3D94FFFDF349D05092C8D92DFDE1" box="[113,607,507,531]" pageId="9" pageNumber="186" type="distribution">
|
||
<paragraph id="8BD86E1FFFFDF349D05092C8D92DFDE1" blockId="9.[113,1089,219,884]" box="[113,607,507,531]" pageId="9" pageNumber="186">
|
||
<emphasis id="B913B20DFFFDF349D05092C8DA76FDE1" bold="true" box="[113,260,507,531]" pageId="9" pageNumber="186">Distribution.</emphasis>
|
||
Only known from
|
||
<collectingCountry id="F3702E8FFFFDF349D1F192C8D929FDE1" box="[464,603,507,531]" name="South Sudan" pageId="9" pageNumber="186">South Sudan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D3D94FFFDF349D0509128D8A1FC81" pageId="9" pageNumber="186" type="discussion">
|
||
<paragraph id="8BD86E1FFFFDF349D0509128D8A1FC81" blockId="9.[113,1089,219,884]" pageId="9" pageNumber="186">
|
||
<emphasis id="B913B20DFFFDF349D0509128DB92FDC1" bold="true" box="[113,224,539,563]" pageId="9" pageNumber="186">Remarks.</emphasis>
|
||
SCHEERPELTZ (1974) mentioned
|
||
<specimenCount id="9D61A596FFFDF349D212912FD9A3FDC1" box="[563,721,539,563]" pageId="9" pageNumber="186" type="generic">two specimens</specimenCount>
|
||
in the original description, a
|
||
<typeStatus id="54DCD0BDFFFDF349D4299128DBEDFDA6" pageId="9" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
and a
|
||
<typeStatus id="54DCD0BDFFFDF349D0C6910FDA31FDA6" box="[231,323,572,596]" pageId="9" pageNumber="186" type="paratype">paratype</typeStatus>
|
||
with the same collecting data, without noting their sex. Two specimens were found in the Scheerpeltz collection in NHMW, both bearing labels reading “Typus”, and I therefore consider them type specimens. The one intended to be the
|
||
<typeStatus id="54DCD0BDFFFDF349D3B9914FD886FD66" box="[920,1012,636,660]" pageId="9" pageNumber="186" type="paratype">paratype</typeStatus>
|
||
can be recognized by the “bottom label” (i.e. the label which is usually attached to the first specimen of the series retained by Scheerpeltz). The
|
||
<typeStatus id="54DCD0BDFFFDF349D21C9188D9E8FD21" box="[573,666,699,723]" pageId="9" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
does not bear this label, as Scheerpeltz probably intended to return it to MZHF from where the material was on loan. Both type specimens are females and now labelled accordingly. In the catalogue of the MZHF collection by
|
||
<bibRefCitation id="EFF613EEFFFDF349D0509028DA30FCC1" author="SILFVERBERG H." box="[113,322,795,819]" pageId="9" pageNumber="186" pagination="15 - 40" refId="ref14548" refString="SILFVERBERG H. 1988: Lists of the insect types in the Zoological Museum, University of Helsinki. 8. Coleoptera: Staphylinidae. Acta Entomologica Fennica 52: 15 - 40." type="journal article" year="1988">SILFVERBERG (1988)</bibRefCitation>
|
||
, the
|
||
<typeStatus id="54DCD0BDFFFDF349D1509028DABCFCC1" box="[369,462,795,819]" pageId="9" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
is listed as being already present in the MZHF (erroneously listed as “
|
||
<emphasis id="B913B20DFFFDF349D0F9900FDA31FCA1" box="[216,323,828,851]" italics="true" pageId="9" pageNumber="186">sudanicus</emphasis>
|
||
”, the last entry on p. 25.), but this is not true according to the information I got from J. Muona in
|
||
<date id="FFD948DFFFFDF349D1489068D97BFC81" box="[361,521,859,883]" pageId="9" pageNumber="186" value="2013-02">February 2013</date>
|
||
. The
|
||
<typeStatus id="54DCD0BDFFFDF349D2629068D9D2FC81" box="[579,672,859,883]" pageId="9" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
is now forwarded to MZHF.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |