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<document id="88B3725223CDDDA77F9424C80288D9D3" ID-CLB-Dataset="67690" ID-DOI="10.5281/zenodo.6623649" ID-GBIF-Dataset="5be7e52f-9b9c-4c03-99ed-f346bbcde1d0" ID-ISBN="978-84-941892-3-4" ID-Zenodo-Dep="6623649" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654607780702" checkinUser="carolina" docAuthor="Don E. Wilson, Thomas E. Lacher, Jr &amp; Russell A. Mittermeier" docDate="2016" docId="03C5A071FFCBFFFFFADB58F25221F631" docLanguage="en" docName="hbmw_6_Echimyidae_0552.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I, Barcelona: Lynx Edicions" docTitle="Diplomys caniceps" docType="treatment" docVersion="16" lastPageNumber="590" masterDocId="FFFCD809FFEDFFD9FFBC58095A61FF94" masterDocTitle="Echimyidae" masterLastPageNumber="604" masterPageNumber="552" pageNumber="590" updateTime="1699339014676" updateUser="ExternalLinkService">
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<mods:title id="D618ACB887F7A363926B6E8CA5D5D695">Echimyidae</mods:title>
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<mods:namePart id="B4B2D15229F3C0FC666E69185C2DB0A6">Don E. Wilson</mods:namePart>
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<mods:namePart id="202A74FD43F7EE515DB444AD2C275AE6">Thomas E. Lacher, Jr</mods:namePart>
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<mods:namePart id="F4AD3980F3A4C4FE8C92EA8BF88FEB21">Russell A. Mittermeier</mods:namePart>
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<mods:title id="86E1659980D24C3F95A907469992AD89">Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I</mods:title>
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<treatment id="03C5A071FFCBFFFFFADB58F25221F631" ID-DOI="http://doi.org/10.5281/zenodo.6633530" ID-GBIF-Taxon="195810014" ID-Zenodo-Dep="6633530" LSID="urn:lsid:plazi:treatment:03C5A071FFCBFFFFFADB58F25221F631" httpUri="http://treatment.plazi.org/id/03C5A071FFCBFFFFFADB58F25221F631" lastPageNumber="590" pageId="38" pageNumber="590">
<subSubSection id="C37642ECFFCBFFFFFADB58F25FFEFEB9" box="[1383,1439,251,301]" pageId="38" pageNumber="590" type="multiple">
<paragraph id="8BD31167FFCBFFFFFADB58F25FFEFEB9" blockId="38.[1380,2470,251,379]" box="[1383,1439,251,301]" pageId="38" pageNumber="590">
<heading id="D09BA60BFFCBFFFFFADB58F25FFEFEB9" box="[1383,1439,251,301]" pageId="38" pageNumber="590">
<figureCitation id="13570DE2FFCBFFFFFADB58F25FFEFEB9" box="[1383,1439,251,301]" captionStart="Plate 35: Echimyidae" captionStartId="8.[107,137,3323,3348]" captionTargetBox="[14,2758,14,3643]" captionTargetPageId="7" captionText="12. Broad-headed Spiny-rat (Clyomys laticeps), 13. Guiara (Euryzygomatomys spinosus), 14. White-spined Spiny-rat (Irinomys albispinus), 15. Eliass Spiny-rat (Trinomys eliasi), 16. Rigid-spined Atlantic Spiny-rat (Trinomys paratus), 17. Yonenagas Spiny-rat (Trinomys yonenagae), 18. Elegant-spined Atlantic Spiny-rat (Trinomys setosus), 19. Moojens Spiny-rat (Trinomys moojeni), 20. Pau Brasil Spiny-rat (Trinomys mirapitanga), 21. Rio de Janeiro Spiny-rat (Trinomys dimidiatus), 22. Sao Paulo Spiny-rat (Trinomys thering), 23. Gracile Atlantic Spiny-rat (Trinomys gratiosus), 24. Sao Lourenco Punare (Thrichomys laurentius), 25. Jacobina Punare (Thrichomys inermis), 26. Lagoa Santa Punare (Thrichomys apereoides), 27. Pantanal Punare (Thrichomys pachyurus), 28. Fosters Punare (Thrichomys fosteri), 29. Painted Tree-rat (Callistomys pictus), 30. Coypu (Myocastor coypus), 31. Armored Rat (Hoplomys gymnurus)" figureDoi="http://doi.org/10.5281/zenodo.6623784" httpUri="https://zenodo.org/record/6623784/files/figure.png" pageId="38" pageNumber="590">72.</figureCitation>
</heading>
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<subSubSection id="C37642ECFFCBFFFFFA0D58F25D8BFEB9" box="[1457,2026,251,301]" pageId="38" pageNumber="590" type="vernacular_names">
<paragraph id="8BD31167FFCBFFFFFA0D58F25D8BFEB9" blockId="38.[1380,2470,251,379]" box="[1457,2026,251,301]" pageId="38" pageNumber="590">
<heading id="D09BA60BFFCBFFFFFA0D58F25D8BFEB9" box="[1457,2026,251,301]" pageId="38" pageNumber="590">
<vernacularName id="056F6149FFCBFFFFFA0D58F25D8BFEB9" box="[1457,2026,251,301]" pageId="38" pageNumber="590">Colombian Rufous Tree-rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF79258F2530FFEB9" box="[2094,2414,251,301]" pageId="38" pageNumber="590" type="nomenclature">
<paragraph id="8BD31167FFCBFFFFF79258F2530FFEB9" blockId="38.[1380,2470,251,379]" box="[2094,2414,251,301]" pageId="38" pageNumber="590">
<heading id="D09BA60BFFCBFFFFF79258F2530FFEB9" box="[2094,2414,251,301]" pageId="38" pageNumber="590">
<taxonomicName id="4C6C6AE4FFCBFFFFF79258F2530FFEB9" ID-CoL="36M6H" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[2094,2414,251,301]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="species" species="caniceps">
<emphasis id="B918CD75FFCBFFFFF79258F2530FFEB9" box="[2094,2414,251,301]" italics="true" pageId="38" pageNumber="590">Diplomys caniceps</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD9593752B1FEEF" pageId="38" pageNumber="590" type="vernacular_names">
<paragraph id="8BD31167FFCBFFFFFAD9593753C4FEC7" blockId="38.[1380,2470,251,379]" box="[1381,2469,318,339]" pageId="38" pageNumber="590">
<heading id="D09BA60BFFCBFFFFFAD9593753C4FEC7" box="[1381,2469,318,339]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD959375FD0FEC7" bold="true" box="[1381,1457,318,339]" pageId="38" pageNumber="590">French:</emphasis>
<vernacularName id="056F6149FFCBFFFFFA0759375CC9FEC7" box="[1467,1704,318,339]" pageId="38" pageNumber="590">Rat-épineux a téte grise</vernacularName>
/
<emphasis id="B918CD75FFCBFFFFF90159375D79FEC7" bold="true" box="[1725,1816,318,339]" pageId="38" pageNumber="590">German:</emphasis>
<vernacularName id="056F6149FFCBFFFFF89D59375250FEC7" box="[1825,2097,318,339]" pageId="38" pageNumber="590">Graukopf-Baumstachelratte</vernacularName>
/
<emphasis id="B918CD75FFCBFFFFF7FA593752C0FEC7" bold="true" box="[2118,2209,318,339]" pageId="38" pageNumber="590">Spanish:</emphasis>
<vernacularName id="056F6149FFCBFFFFF717593753C4FEC7" box="[2219,2469,318,339]" pageId="38" pageNumber="590">Rata arboricola de Bolivia</vernacularName>
</heading>
</paragraph>
<paragraph id="8BD31167FFCBFFFFFAD9596F52B1FEEF" blockId="38.[1380,2470,251,379]" box="[1381,2256,358,379]" pageId="38" pageNumber="590">
<heading id="D09BA60BFFCBFFFFFAD9596F52B1FEEF" box="[1381,2256,358,379]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD9596F5C3DFEEF" bold="true" box="[1381,1628,358,379]" pageId="38" pageNumber="590">Other common names:</emphasis>
<vernacularName id="056F6149FFCBFFFFF9D8596F5DE8FEEF" box="[1636,1929,358,379]" pageId="38" pageNumber="590">Arboreal Soft-furred Spiny-rat</vernacularName>
,
<vernacularName id="056F6149FFCBFFFFF82A596F52B1FEEF" box="[1942,2256,358,379]" pageId="38" pageNumber="590">Colombian Soft-furred Spiny-rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF87559A65272FE64" pageId="38" pageNumber="590" type="reference_group">
<paragraph id="8BD31167FFCBFFFFF87559A65272FE64" blockId="38.[1993,2587,431,851]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFF87559A65205FE5C" bold="true" box="[1993,2148,431,456]" pageId="38" pageNumber="590">Taxonomy.</emphasis>
<taxonomicName id="4C6C6AE4FFCBFFFFF73F59A6526FFE64" authority="Gunther, 1877" authorityName="Gunther" authorityYear="1877" class="Mammalia" family="Echimyidae" genus="Loncheres" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="species" species="caniceps">Loncheres caniceps Gunther, 1877</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF7A359DA5395FE64" box="[2079,2548,467,496]" pageId="38" pageNumber="590" type="materials_examined">
<paragraph id="8BD31167FFCBFFFFF7A359DA5395FE64" blockId="38.[1993,2587,431,851]" box="[2079,2548,467,496]" pageId="38" pageNumber="590">
<materialsCitation id="3B041B3AFFCBFFFFF7A359DA5391FE64" ID-GBIF-Occurrence="3804303402" box="[2079,2544,467,496]" pageId="38" pageNumber="590">“Medellin,” Antioquia, Colombia</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF87659FF53CEFD1B" pageId="38" pageNumber="590" type="discussion">
<paragraph id="8BD31167FFCBFFFFF87659FF53CEFD1B" blockId="38.[1993,2587,431,851]" pageId="38" pageNumber="590">
<taxonomicName id="4C6C6AE4FFCBFFFFF87659FF52CFFD83" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[1994,2222,502,535]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="species" species="caniceps">Diplomys caniceps</taxonomicName>
has been variously placed in the genera
<taxonomicName id="4C6C6AE4FFCBFFFFF72D5A14536AFDAA" authorityName="Illiger" authorityYear="1811" box="[2193,2315,541,574]" class="Mammalia" family="Echimyidae" genus="Loncheres" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Loncheres</taxonomicName>
or
<taxonomicName id="4C6C6AE4FFCBFFFFF6875A1453D6FDAA" authorityName="Lund" authorityYear="1839" box="[2363,2487,541,574]" class="Mammalia" family="Echimyidae" genus="Phyllomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Phyllomys</taxonomicName>
by 19&quot; century authors; first assigned to
<taxonomicName id="4C6C6AE4FFCBFFFFF61B5A4C507AFDF2" authorityName="Thomas" authorityYear="1916" box="[2471,2587,581,614]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Diplomys</taxonomicName>
by O. Thomas in 1916. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF8765A9A5391FCBF" pageId="38" pageNumber="590" type="distribution">
<caption id="DF1341EFFFCBFFFFF8765A9A5391FCBF" ID-DOI="http://doi.org/10.5281/zenodo.6623832" ID-Zenodo-Dep="6623832" httpUri="https://zenodo.org/record/6623832/files/figure.png" inLine="true" pageId="38" pageNumber="590" startId="38.[1994,2166,659,692]" targetBox="[1379,1972,432,848]" targetPageId="38">
<paragraph id="8BD31167FFCBFFFFF8765A9A5391FCBF" blockId="38.[1993,2587,431,851]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFF8765A9A521BFD20" bold="true" box="[1994,2170,659,692]" pageId="38" pageNumber="590">Distribution.</emphasis>
The Caribe Biogeographical Region of NW Colombia, in the Cauca Valley near Medellin, and possibly the Ser-rania de San Lucas in the Bolivar State.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFF8775B3B523AF85F" pageId="38" pageNumber="590" type="description">
<paragraph id="8BD31167FFCBFFFFF8775B3B523AF85F" blockId="38.[1993,2587,431,851]" lastBlockId="38.[1383,2593,857,2469]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFF8775B3B5284FCC7" bold="true" box="[1995,2277,818,851]" pageId="38" pageNumber="590">Descriptive notes.</emphasis>
Head-body 212-390 mm, tail 178-267 mm. No specific data are available for body weight. The Colombian Rufous Tree-rat is larger than its congener, the Central American Rufous Tree-rat (
<taxonomicName id="4C6C6AE4FFCBFFFFFA4C5BA15C11FC5D" authorityName="Bangs" authorityYear="1901" box="[1520,1648,936,969]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="species" species="labilis">D. labilis</taxonomicName>
). Its fur is stiff but soft, without bristles or spines. Overall upper parts are dull rusty, red-brown slightly streaked with dark brown hairs, but bright red on back one-half of body; forequarters are fulvous in some Colombian Rufous Tree-rats. Nose is grayish brown, and pre-auricular grayish patch extends partially to eyes. Face is marked with small whitish spots over eyes and at bases of vibrissae. A few whitish or yellowish spots are present below ear notches and behind ears. Ears are short, rounded, and divided by small anterior lobe and larger posterior lobe. Vibrissae are long, extending backward to shoulders. Belly and chin are whitish, yellowish, or pale orange; some individuals have rich orange venters. Dark rusty fur often occurs on midline of abdomen, extending posteriorly toward base of tail. Robusttail is 90-105% of head-body length and is covered sparsely by dark brown, bristle-like hairs;tail scales are not well exposed beneath hair cover. Forefeet are whitish-brown or gray-brown. Hindfeet are broad, yellow-brown to brown or gray above. Skulls of both species of
<taxonomicName id="4C6C6AE4FFCBFFFFF9EC5DA05CA7FA5E" authorityName="Thomas" authorityYear="1916" box="[1616,1734,1449,1482]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Diplomys</taxonomicName>
are characteristically convex in outline when viewed laterally. Rostrum is short and narrow; bullae are not inflated but possess well-developed auditory tube thatis oriented anteriorly; and auditory meatus is very close to squamosal bone. Jugal becomes narrow below post-orbital process of zygoma and exhibits an inconspicuous ventral process posterior to dorsal one. Alisphenoid arterial foramen pattern is unique in
<taxonomicName id="4C6C6AE4FFCBFFFFF9605E675D32F91B" authorityName="Thomas" authorityYear="1916" box="[1756,1875,1646,1679]" class="Mammalia" family="Echimyidae" genus="Diplomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Diplomys</taxonomicName>
, with short distance between buccinator foramen and foramen ovale. Hamular process of pterygoid bone has developed anteriorly oriented spur. Mandible has large masseteric ridge and characteristic shallow angle of sigmoid notch between condyloid and angular processes. Mandibular foramenis located anteriorly to condyloid ridge and not in a fossa. Cheekteeth are large and long and are highly convergent in occlusal morphology with those of species of
<taxonomicName id="4C6C6AE4FFCBFFFFF65C5F3A5FD5F8EF" authorityName="Lund" authorityYear="1839" class="Mammalia" family="Echimyidae" genus="Phyllomys" kingdom="Animalia" order="Rodentia" pageId="38" pageNumber="590" phylum="Chordata" rank="genus">Phyllomys</taxonomicName>
. Incisors are strongly curved. Maxillary and mandibular tooth rows are strongly divergent antero-posteriorly, with outward curving of last two molars causing disjunction with straight edge formed by molars M' and M*.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD05FD8533BF78E" pageId="38" pageNumber="590" type="biology_ecology">
<paragraph id="8BD31167FFCBFFFFFAD05FD8533BF78E" blockId="38.[1383,2593,857,2469]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD05FD85FBAF866" bold="true" box="[1388,1499,2001,2034]" pageId="38" pageNumber="590">Habitat.</emphasis>
Caribe Biogeographical Region rainforest of Colombia. Few data on specific habitats of the highly arboreal Colombian Rufous Tree-rat are available.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD0502C5351F7D6" box="[1388,2352,2085,2114]" pageId="38" pageNumber="590" type="food_feeding">
<paragraph id="8BD31167FFCBFFFFFAD0502C5351F7D6" blockId="38.[1383,2593,857,2469]" box="[1388,2352,2085,2114]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD0502C5C10F7D6" bold="true" box="[1388,1649,2085,2114]" pageId="38" pageNumber="590">Food and Feeding.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD0504152D0F7FD" box="[1388,2225,2120,2153]" pageId="38" pageNumber="590" type="breeding">
<paragraph id="8BD31167FFCBFFFFFAD0504152D0F7FD" blockId="38.[1383,2593,857,2469]" box="[1388,2225,2120,2153]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD050415F93F7FD" bold="true" box="[1388,1522,2120,2153]" pageId="38" pageNumber="590">Breeding.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD750665374F704" box="[1387,2325,2159,2192]" pageId="38" pageNumber="590" type="activity">
<paragraph id="8BD31167FFCBFFFFFAD750665374F704" blockId="38.[1383,2593,857,2469]" box="[1387,2325,2159,2192]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD750665C34F704" bold="true" box="[1387,1621,2159,2192]" pageId="38" pageNumber="590">Activity patterns.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD0509F5C20F774" pageId="38" pageNumber="590" type="biology_ecology">
<paragraph id="8BD31167FFCBFFFFFAD0509F5C20F774" blockId="38.[1383,2593,857,2469]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD0509F5255F723" bold="true" box="[1388,2100,2198,2231]" pageId="38" pageNumber="590">Movements, Home range and Social organization.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD150ED52BAF6BA" pageId="38" pageNumber="590" type="conservation">
<paragraph id="8BD31167FFCBFFFFFAD150ED52BAF6BA" blockId="38.[1383,2593,857,2469]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD150ED5CA8F691" bold="true" box="[1389,1737,2276,2309]" pageId="38" pageNumber="590">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. Additional ecologicalstudies of this rare Colombian endemic are needed.
</paragraph>
</subSubSection>
<subSubSection id="C37642ECFFCBFFFFFAD251355221F631" pageId="38" pageNumber="590" type="bibRefCitation_list">
<paragraph id="8BD31167FFCBFFFFFAD251355221F631" blockId="38.[1383,2593,857,2469]" pageId="38" pageNumber="590">
<emphasis id="B918CD75FFCBFFFFFAD251355C67F6C1" bold="true" box="[1390,1542,2364,2389]" pageId="38" pageNumber="590">Bibliography.</emphasis>
Alberico et al. (2000), Allen (1916b), Cuervo et al. (1986), Eisenberg (1989), Emmons (1990, 1997a, 2005), Goldman (1916), Gunther (1877a), Mares (1982), Patton et al. (2015), Thomas (1916a, 1916b), Trouessart (1880, 1904), Woods (1993), Woods &amp; Kilpatrick (2005).
</paragraph>
</subSubSection>
</treatment>
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