545 lines
64 KiB
XML
545 lines
64 KiB
XML
<document id="047F455496DE0DC1E3AB8988D6C5CEB7" ID-CLB-Dataset="304560" ID-DOI="10.11646/zootaxa.5519.2.3" ID-GBIF-Dataset="a3266865-054f-4cff-bc0c-faf89ec8074c" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915771" ID-ZooBank="BE0CA3E4-3695-403A-AD98-3C279238518B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1728585742752" checkinUser="plazi" docAuthor="Theischinger, Günther, Kalnins, Martins & Marinov, Milen" docDate="2024" docId="03C587C6CE6D9877FF24829B737EFA78" docLanguage="en" docName="zootaxa.5519.2.3.pdf" docOrigin="Zootaxa 5519 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anax Leach 1815" docType="key" docVersion="3" lastPageNumber="241" masterDocId="FFFCFFBECE74986DFFB3832C7740FF90" masterDocTitle="Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)" masterLastPageNumber="242" masterPageNumber="215" pageNumber="240" updateTime="1729087811061" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DBD062079296F56FF856F2F52E9DE875">
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<mods:title id="5717C66597EA86F10A545EA9AE1C6FF4">Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FFB58437CB06B8E0948B7A13CB1AC26C" type="personal">
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<mods:role id="BC89B31117A7F512811727F6D357DF31">
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<mods:roleTerm id="8C01E1832C9D5EC251E32F317D731CC2">Author</mods:roleTerm>
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<mods:namePart id="45A4D3BD1194A239F635DAB9FC1656F0">Theischinger, Günther</mods:namePart>
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<mods:nameIdentifier id="7067D944BBC480610F5A5263E4D5D44F" type="ORCID">0000-0002-5207-2626</mods:nameIdentifier>
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<mods:affiliation id="65E0D28EC6796FBFCFDB989323111048">Australian Museum, Entomology, 6 College Street, Sydney, NSW 2010</mods:affiliation>
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<mods:nameIdentifier id="34524FBDC385DC8010F7B46CDD772034" type="email">theischingergunther@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="6D3F6A2F1E62D87BB40CF543811E5A8B" type="personal">
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<mods:role id="C484BB54CACCF4009077A9F6F4476A30">
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<mods:roleTerm id="3AA0E2F61802F0659638BD104946322D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1BD1B777C862595D7E95400A1457DF99">Kalnins, Martins</mods:namePart>
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<mods:nameIdentifier id="517C71EDE7BFBB5122E2126C0CE510E1" type="ORCID">0009-0008-8850-4444</mods:nameIdentifier>
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<mods:affiliation id="8EF51F9BA96F9671265C0AA064037EE7">Department of Biosystematics, Institute of Life Sciences and Technology, Daugavpils University, Parādes Street 1 A, Daugavpils, Latvia, LV- 5401</mods:affiliation>
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<mods:nameIdentifier id="51D163ADB68F57B17C9961BA588693D9" type="email">martins.kalnins@biology.lv</mods:nameIdentifier>
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</mods:name>
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<mods:namePart id="168AA99E671E137203921F9B545589E3">Marinov, Milen</mods:namePart>
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<mods:nameIdentifier id="569F63E4AE4C479CA9A4B5145DFFAECD" type="ORCID">0000-0003-3284-2555</mods:nameIdentifier>
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<mods:affiliation id="D35A6285D514EAC283C75882611D2759">Biosecurity Surveillance & Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8544, New Zealand</mods:affiliation>
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<mods:nameIdentifier id="5FD5604910B16273D8595CB7152F3750" type="email">milen.marinov@mpi.govt.nz</mods:nameIdentifier>
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<mods:title id="ADFCE6339EED380A015CE1BACB9A5A2D">Zootaxa</mods:title>
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<mods:part id="26152D9956296CE28E7C5040E9A2B89D">
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<mods:date id="00E8DA322AD6DE62A90565533871E3A0">2024</mods:date>
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<mods:number id="E605BB870EC894E449C21A0FC3FC666A">2024-10-08</mods:number>
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<mods:number id="CEFF59DC831A3028E9693AA89B808E34">5519</mods:number>
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<mods:url id="ED0AD98E295DC4FF88BFFB03BB42F1DC">http://dx.doi.org/10.11646/zootaxa.5519.2.3</mods:url>
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<mods:classification id="31FF6CD55E9F3B339B902582DA48D50D">journal article</mods:classification>
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<mods:identifier id="5E58F9E0C8C44CF3EA2BF65536DD7431" type="DOI">10.11646/zootaxa.5519.2.3</mods:identifier>
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<mods:identifier id="5F238ED01822371B509734CBED5E6F7A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D886D5C3D428D94949709F4055634E9C" type="Zenodo-Dep">13915771</mods:identifier>
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<treatment id="03C587C6CE6D9877FF24829B737EFA78" ID-DOI="http://doi.org/10.5281/zenodo.13940299" ID-GBIF-Taxon="243657024" ID-Zenodo-Dep="13940299" LSID="urn:lsid:plazi:treatment:03C587C6CE6D9877FF24829B737EFA78" httpUri="http://treatment.plazi.org/id/03C587C6CE6D9877FF24829B737EFA78" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240">
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<subSubSection id="C376655BCE6D9874FF24829B75E3FE65" pageId="25" pageNumber="240" type="nomenclature">
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<paragraph id="8BD336D0CE6D9874FF24829B75E3FE65" blockId="25.[151,1436,438,501]" pageId="25" pageNumber="240">
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<heading id="D09B81BCCE6D9874FF24829B75E3FE65" bold="true" centered="true" fontSize="11" level="1" pageId="25" pageNumber="240" reason="1">
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||
<emphasis id="B918EAC2CE6D9874FF24829B75E3FE65" bold="true" pageId="25" pageNumber="240">
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||
Key for species identification (males only) of
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<taxonomicName id="4C6C4D53CE6D9874FD7C829B7450FE41" ID-CoL="VDK" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[719,784,439,465]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
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<emphasis id="B918EAC2CE6D9874FD7C829B7450FE41" bold="true" box="[719,784,439,465]" italics="true" pageId="25" pageNumber="240">Anax</emphasis>
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</taxonomicName>
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from the Australian, the Oceanian and the south of the Indomalayan Biogeographical Realms
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C376655BCE6D9874FF24810F742FFA75" pageId="25" pageNumber="240" type="discussion">
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<paragraph id="8BD336D0CE6D9874FF24810F742FFA75" blockId="25.[151,1437,547,1545]" pageId="25" pageNumber="240">
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As characters such as e. g. ‘presence or absence and shape and definition of the postfrontal T-mark’, ‘colour of occipital triangle and vertex’ and ‘abdominal colour pattern’ have been found by other authors and ourselves as too variable to be used for specific distinction of
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<taxonomicName id="4C6C4D53CE6D9874FD2181477466FD15" box="[658,806,619,645]" pageId="25" pageNumber="240">
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<emphasis id="B918EAC2CE6D9874FD218147758CFD15" box="[658,716,619,645]" italics="true" pageId="25" pageNumber="240">Anax</emphasis>
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species
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</taxonomicName>
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, the key is primarily based on the shape and proportions of the elements of the male terminalia (epiproct, cerci) that appear in a high degree consistent and thus to be most useful for the identification of taxa of the species group. An illustration of the abdominal patterns (
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<figureCitation id="13572A55CE6D9874FABE819F7221FD5D" box="[1293,1377,690,717]" captionStart="FIGURE 53" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[164,1436,181,1813]" captionTargetId="figure-17@21.[151,1436,181,1833]" captionTargetPageId="21" captionText="FIGURE 53. Anax spp., male terminalia: (a–e, h–p) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, e–j, l–m) modified from Lieftinck (1942); (k) modified from Kennedy (1934)." figureDoi="http://doi.org/10.5281/zenodo.13915810" httpUri="https://zenodo.org/record/13915810/files/figure.png" pageId="25" pageNumber="240">Fig. 53</figureCitation>
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) that may also be useful is presented subsequent to the Key as a further aid to identification. However, it is important to point out inconsistencies and shortcomings in hitherto available keys or similar more or less comprehensive work. These are part of
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<bibRefCitation id="EFFD4B21CE6D9874FED98033754EFCA9" author="Martin, R." box="[362,526,799,825]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
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,
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<bibRefCitation id="EFFD4B21CE6D9874FDAE8033759AFCA9" author="Kennedy, C. H." box="[541,730,799,825]" pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
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||
,
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<bibRefCitation id="EFFD4B21CE6D9874FD5A803374E8FCA9" author="Lieftinck, M. A." box="[745,936,799,825]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
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||
,
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||
<bibRefCitation id="EFFD4B21CE6D9874FC048033733FFCA9" author="Michalski, J." box="[951,1151,799,825]" pageId="25" pageNumber="240" refId="ref12846" refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, & the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp." type="book" year="2012">Michalski (2012)</bibRefCitation>
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,
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<bibRefCitation id="EFFD4B21CE6D9874FB3D803372DCFCA9" author="Orr, A. G. & Kalkman, V. J." box="[1166,1436,798,825]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr & Kalkman (2015)</bibRefCitation>
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||
and
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<bibRefCitation id="EFFD4B21CE6D9874FF7A806F76D7FCCD" author="Seehausen, M." box="[201,407,835,861]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
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.
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<bibRefCitation id="EFFD4B21CE6D9874FE17806F7504FCCD" author="Martin, R." box="[420,580,835,861]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
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presents useful dorsal and lateral view illustrations of the male terminalia of
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<taxonomicName id="4C6C4D53CE6D9874FF0B804B7677FC10" authorityName="Hagen" authorityYear="1867" box="[184,311,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
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<emphasis id="B918EAC2CE6D9874FF0B804B7677FC10" box="[184,311,871,897]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6C4D53CE6D9874FEFB804B768AFC10" authorityName="Selys" authorityYear="1872" box="[328,458,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
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<emphasis id="B918EAC2CE6D9874FEFB804B768AFC10" box="[328,458,871,897]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6C4D53CE6D9874FE68804B7516FC10" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[475,598,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
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<emphasis id="B918EAC2CE6D9874FE68804B7516FC10" box="[475,598,871,897]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6C4D53CE6D9874FDD4804B7441FC11" authorityName="Rambur" authorityYear="1842" box="[615,769,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
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<emphasis id="B918EAC2CE6D9874FDD4804B7441FC11" box="[615,769,871,897]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6C4D53CE6D9874FCA1804B74FEFC11" authorityName="Forster" authorityYear="1898" box="[786,958,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
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<emphasis id="B918EAC2CE6D9874FCA1804B74FEFC11" box="[786,958,871,897]" italics="true" pageId="25" pageNumber="240">A. maclachlani</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6C4D53CE6D9874FC7C804B736EFC11" authorityName="Forster" authorityYear="1900" box="[975,1070,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="selysi">
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<emphasis id="B918EAC2CE6D9874FC7C804B736EFC11" box="[975,1070,871,897]" italics="true" pageId="25" pageNumber="240">A. selysi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6C4D53CE6D9874FBD9804B73BBFC10" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1130,1275,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
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<emphasis id="B918EAC2CE6D9874FBD9804B73BBFC10" box="[1130,1275,871,897]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
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</taxonomicName>
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. He mentions
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<taxonomicName id="4C6C4D53CE6D9874FF2480A77662FC35" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[151,290,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
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<emphasis id="B918EAC2CE6D9874FF2480A77662FC35" box="[151,290,907,933]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
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</taxonomicName>
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only as a race of
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<taxonomicName id="4C6C4D53CE6D9874FE5C80A77529FC34" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[495,617,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
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<emphasis id="B918EAC2CE6D9874FE5C80A77529FC34" box="[495,617,907,933]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
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</taxonomicName>
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and does not refer to
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<taxonomicName id="4C6C4D53CE6D9874FCD680A774A2FC35" authorityName="Hagen" authorityYear="1867" box="[869,994,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
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<emphasis id="B918EAC2CE6D9874FCD680A774A2FC35" box="[869,994,907,933]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
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</taxonomicName>
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at all.
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<bibRefCitation id="EFFD4B21CE6D9874FB8180A773AAFC35" author="Kennedy, C. H." box="[1074,1258,907,933]" pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
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||
includes dorsal and lateral illustrations of the terminalia of
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<taxonomicName id="4C6C4D53CE6D9874FD3E80837452FC58" box="[653,786,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="piraticus">
|
||
<emphasis id="B918EAC2CE6D9874FD3E80837452FC58" box="[653,786,943,969]" italics="true" pageId="25" pageNumber="240">A. piraticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FC91808374EEFC59" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[802,942,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FC91808374EEFC59" box="[802,942,943,969]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FC0D80837318FC59" authorityName="Rambur" authorityYear="1842" box="[958,1112,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
|
||
<emphasis id="B918EAC2CE6D9874FC0D80837318FC59" box="[958,1112,943,969]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FBDA808373A4FC58" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1129,1252,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6D9874FBDA808373A4FC58" box="[1129,1252,943,969]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FB4680837234FC58" authorityName="Hagen" authorityYear="1867" box="[1269,1396,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B918EAC2CE6D9874FB4680837234FC58" box="[1269,1396,943,969]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FA3780837795FC7D" authorityName="Forster" authorityYear="1900" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="selysi">
|
||
<emphasis id="B918EAC2CE6D9874FA3780837795FC7D" italics="true" pageId="25" pageNumber="240">A. selysi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6D9874FEBC80FF76F9FC7D" authorityName="Forster" authorityYear="1898" box="[271,441,979,1005]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
|
||
<emphasis id="B918EAC2CE6D9874FEBC80FF76F9FC7D" box="[271,441,979,1005]" italics="true" pageId="25" pageNumber="240">A. maclachlani</emphasis>
|
||
</taxonomicName>
|
||
. However, what he presents as
|
||
<taxonomicName id="4C6C4D53CE6D9874FC9080FF74FDFC7D" authorityName="Rambur" authorityYear="1842" box="[803,957,979,1005]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
|
||
<emphasis id="B918EAC2CE6D9874FC9080FF74FDFC7D" box="[803,957,979,1005]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
|
||
</taxonomicName>
|
||
is misleading, because it is probably only a different version of what he presents correctly as
|
||
<emphasis id="B918EAC2CE6D9874FD5780DB7426FB80" box="[740,870,1015,1041]" italics="true" pageId="25" pageNumber="240">
|
||
<taxonomicName id="4C6C4D53CE6D9874FD5780DB7422FB80" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[740,866,1015,1041]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">A. guttatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and certainly not
|
||
<taxonomicName id="4C6C4D53CE6D9874FB8B80DB7392FB81" authorityName="Rambur" authorityYear="1842" box="[1080,1234,1015,1041]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
|
||
<emphasis id="B918EAC2CE6D9874FB8B80DB7392FB81" box="[1080,1234,1015,1041]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFFD4B21CE6D9874FB5180DB72DCFB81" author="Lieftinck, M. A." box="[1250,1436,1015,1041]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
|
||
presents excellent drawings of the available species including
|
||
<emphasis id="B918EAC2CE6D9874FCFD87377490FBA5" box="[846,976,1051,1077]" italics="true" pageId="25" pageNumber="240">
|
||
<taxonomicName id="4C6C4D53CE6D9874FCFD8737748CFBA5" authorityName="Hagen" authorityYear="1867" box="[846,972,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">A. fumosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but except
|
||
<taxonomicName id="4C6C4D53CE6D9874FBE287377390FBA4" authorityName="Selys" authorityYear="1872" box="[1105,1232,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
|
||
<emphasis id="B918EAC2CE6D9874FBE287377390FBA4" box="[1105,1232,1051,1077]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B918EAC2CE6D9874FAB5873772DDFBA4" box="[1286,1437,1051,1077]" italics="true" pageId="25" pageNumber="240">
|
||
<taxonomicName id="4C6C4D53CE6D9874FAB5873772D9FBA4" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1286,1433,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">A. papuensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
He adds two taxa of the species group he describes as
|
||
<taxonomicName id="4C6C4D53CE6D9874FD488713742DFBC8" authorityName="Lieftinck" authorityYear="1942" box="[763,877,1087,1113]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="pugnax">
|
||
<emphasis id="B918EAC2CE6D9874FD488713742DFBC8" box="[763,877,1087,1113]" italics="true" pageId="25" pageNumber="240">A. pugnax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6D9874FC17871373D4FBC9" authorityName="Lieftinck" authorityYear="1942" box="[932,1172,1087,1113]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="subSpecies" species="fumosus" subSpecies="celebense">
|
||
<emphasis id="B918EAC2CE6D9874FC17871373D4FBC9" box="[932,1172,1087,1113]" italics="true" pageId="25" pageNumber="240">A. fumosus celebense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFFD4B21CE6D9874FB1187137225FBC9" author="Michalski, J." box="[1186,1381,1087,1113]" pageId="25" pageNumber="240" refId="ref12846" refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, & the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp." type="book" year="2012">Michalski (2012)</bibRefCitation>
|
||
uses the illustrations of
|
||
<bibRefCitation id="EFFD4B21CE6D9874FEDF874F754BFBED" author="Martin, R." box="[364,523,1123,1149]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFD4B21CE6D9874FDF0874F75BCFBED" author="Lieftinck, M. A." box="[579,764,1123,1149]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
|
||
for the presentation of the species of New
|
||
<collectingCountry id="F37B7640CE6D9874FB56874F7279FBED" box="[1253,1337,1123,1149]" name="Guinea-Bissau" pageId="25" pageNumber="240">Guinea</collectingCountry>
|
||
, Maluku and the
|
||
<collectingCountry id="F37B7640CE6D9874FF5D87AB76ECFB31" box="[238,428,1159,1185]" name="Solomon Islands" pageId="25" pageNumber="240">Solomon Islands</collectingCountry>
|
||
.
|
||
<bibRefCitation id="EFFD4B21CE6D9874FE0687AB75F5FB31" author="Orr, A. G. & Kalkman, V. J." box="[437,693,1158,1185]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr & Kalkman (2015)</bibRefCitation>
|
||
give excellent colour illustrations including dorsal view terminalia illustrations of New
|
||
<collectingCountry id="F37B7640CE6D9874FE3187877693FB55" box="[386,467,1195,1221]" name="Guinea-Bissau" pageId="25" pageNumber="240">Guinea</collectingCountry>
|
||
species, (but in the detailed line drawings of appendages
|
||
<taxonomicName id="4C6C4D53CE6D9874FBD8878773A7FB55" authorityName="Hagen" authorityYear="1867" box="[1131,1255,1195,1221]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
|
||
<emphasis id="B918EAC2CE6D9874FBD8878773A7FB55" box="[1131,1255,1195,1221]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6D9874FA92878772DCFB54" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1313,1436,1195,1221]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6D9874FA92878772DCFB54" box="[1313,1436,1195,1221]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
are mistakenly interchanged). Finally
|
||
<bibRefCitation id="EFFD4B21CE6D9874FDF387E37448FB79" author="Seehausen, M." box="[576,776,1231,1257]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
|
||
when illustrating
|
||
<taxonomicName id="4C6C4D53CE6D9874FC6187E37321FB78" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[978,1121,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
|
||
<emphasis id="B918EAC2CE6D9874FC6187E37321FB78" box="[978,1121,1231,1257]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FBDC87E373A8FB78" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1135,1256,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6D9874FBDC87E373A8FB78" box="[1135,1256,1231,1257]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FB4587E37236FB78" authorityName="Selys" authorityYear="1872" box="[1270,1398,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
|
||
<emphasis id="B918EAC2CE6D9874FB4587E37236FB78" box="[1270,1398,1231,1257]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C4D53CE6D9874FA3787E37650FA9D" authorityName="Rambur" authorityYear="1842" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
|
||
<emphasis id="B918EAC2CE6D9874FA3787E37650FA9D" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6D9874FEF587DF768EFA9D" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[326,462,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FEF587DF768EFA9D" box="[326,462,1267,1293]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
depicts
|
||
<taxonomicName id="4C6C4D53CE6D9874FD9987DF75E3FA9C" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[554,675,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6D9874FD9987DF75E3FA9C" box="[554,675,1267,1293]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
matching very closely the drawing
|
||
<emphasis id="B918EAC2CE6D9874FB8287DF7381FA9D" box="[1073,1217,1267,1293]" italics="true" pageId="25" pageNumber="240">
|
||
A. f.
|
||
<taxonomicName id="4C6C4D53CE6D9874FBD687DF7381FA9D" authorityName="Hagen" authorityYear="1867" box="[1125,1217,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">fumosus</taxonomicName>
|
||
</emphasis>
|
||
of
|
||
<bibRefCitation id="EFFD4B21CE6D9874FB5687DF72DCFA9D" author="Lieftinck, M. A." box="[1253,1436,1267,1293]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6D9874FF7B863B7603FAA0" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[200,323,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6D9874FF7B863B7603FAA0" box="[200,323,1303,1329]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
(but actually
|
||
<taxonomicName id="4C6C4D53CE6D9874FE50863B7520FAA1" authorityName="Hagen" authorityYear="1867" box="[483,608,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
|
||
<emphasis id="B918EAC2CE6D9874FE50863B7520FAA1" box="[483,608,1303,1329]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
|
||
</taxonomicName>
|
||
) of
|
||
<bibRefCitation id="EFFD4B21CE6D9874FD21863B74DDFAA1" author="Orr, A. G. & Kalkman, V. J." box="[658,925,1302,1329]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr & Kalkman (2015)</bibRefCitation>
|
||
. Seehausen’s drawing of the terminalia of
|
||
<taxonomicName id="4C6C4D53CE6D9874FA37863B7641FAC5" baseAuthorityName="Seehausen" baseAuthorityYear="2017" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FA37863B7641FAC5" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
looks almost like a copy of Lieftinck’s illustration of
|
||
<emphasis id="B918EAC2CE6D9874FCD48617734BFAC5" box="[871,1035,1339,1365]" italics="true" pageId="25" pageNumber="240">
|
||
A. f.
|
||
<taxonomicName id="4C6C4D53CE6D9874FC2D8617734BFAC5" box="[926,1035,1339,1365]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="celebense">celebense</taxonomicName>
|
||
</emphasis>
|
||
and not at all similar to Lieftinck’s illustration of
|
||
<taxonomicName id="4C6C4D53CE6D9874FE8A86737684FAE9" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[313,452,1375,1401]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FE8A86737684FAE9" box="[313,452,1375,1401]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
that matches the illustration of the
|
||
<typeStatus id="54D78872CE6D9874FCEF867374CCFAE9" box="[860,908,1375,1401]" pageId="25" pageNumber="240">type</typeStatus>
|
||
of this species by F.M. Carpenter in
|
||
<bibRefCitation id="EFFD4B21CE6D9874FA84867377A4FA0D" author="Kennedy, C. H." pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
|
||
, as well as a photo of this species by Yen Choong Chee and Hagen material of this species in Vienna. This raises doubt as to whether the abdomens figured by
|
||
<bibRefCitation id="EFFD4B21CE6D9874FD6A868B74E0FA51" author="Seehausen, M." box="[729,928,1447,1473]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C6C4D53CE6D9874FC77868B730DFA51" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[964,1101,1447,1473]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FC77868B730DFA51" box="[964,1101,1447,1473]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13572A55CE6D9874FBE8868B7393FA51" box="[1115,1235,1447,1473]" captionStart="FIGURES 18–25" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 18–25. Anax spp., male, dorsal: (18–21) S2–3; (22–25) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." figureDoi="http://doi.org/10.5281/zenodo.13940290" httpUri="https://zenodo.org/record/13940290/files/figure.png" pageId="25" pageNumber="240">Fig. 21 j–l</figureCitation>
|
||
), and in particular if the thick-waisted females (
|
||
<figureCitation id="13572A55CE6D9874FE6E86E7751CFA75" box="[477,604,1483,1509]" captionStart="FIGURES 18–25" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 18–25. Anax spp., male, dorsal: (18–21) S2–3; (22–25) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." figureDoi="http://doi.org/10.5281/zenodo.13940290" httpUri="https://zenodo.org/record/13940290/files/figure.png" pageId="25" pageNumber="240">Fig. 22 j–k</figureCitation>
|
||
), really are
|
||
<taxonomicName id="4C6C4D53CE6D9874FD6C86E77428FA75" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[735,872,1483,1509]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6D9874FD6C86E77428FA75" box="[735,872,1483,1509]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376655BCE6D9874FF7486C376CEF999" box="[199,398,1518,1545]" pageId="25" pageNumber="240" type="multiple">
|
||
<paragraph id="8BD336D0CE6D9874FF7486C376CEF999" blockId="25.[151,1437,547,1545]" box="[199,398,1518,1545]" pageId="25" pageNumber="240">
|
||
<emphasis id="B918EAC2CE6D9874FF7486C377B7F999" bold="true" box="[199,247,1519,1545]" pageId="25" pageNumber="240">Key</emphasis>
|
||
(
|
||
<figureCitation id="13572A55CE6D9874FEB586C3761DF999" box="[262,349,1518,1545]" captionStart="FIGURE 52" captionStartId="19.[151,250,1810,1834]" captionTargetBox="[313,1275,633,1785]" captionTargetId="figure-268@19.[313,1275,633,1786]" captionTargetPageId="19" captionText="FIGURE 52a. Terms used in the descriptions and Key, presented on Anax gibbosulus, male terminalia, dorsal and lateral (modified from Lieftinck 1942): als = apico-lateral spike; bit = basal interior tooth; C = cercus; E = epiproct; S10 = abdominal segment 10; t = tubercles." figureDoi="http://doi.org/10.5281/zenodo.13915806" httpUri="https://zenodo.org/record/13915806/files/figure.png" pageId="25" pageNumber="240">Figs 52</figureCitation>
|
||
,
|
||
<figureCitation id="13572A55CE6D9874FEDA86C276C6F999" box="[361,390,1518,1545]" captionStart="FIGURE 53" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[164,1436,181,1813]" captionTargetId="figure-17@21.[151,1436,181,1833]" captionTargetPageId="21" captionText="FIGURE 53. Anax spp., male terminalia: (a–e, h–p) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, e–j, l–m) modified from Lieftinck (1942); (k) modified from Kennedy (1934)." figureDoi="http://doi.org/10.5281/zenodo.13915810" httpUri="https://zenodo.org/record/13915810/files/figure.png" pageId="25" pageNumber="240">53</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376655BCE6D9877FF10851172D2FB32" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240" type="key">
|
||
<key id="FFFD61E1CE6D9877FF10851172D2FB32" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240">
|
||
<keyStep id="30982E75CE6D9874FF10851172D2F9ED" pageId="25" pageNumber="240">
|
||
<paragraph id="8BD336D0CE6D9874FF10851172D1F9C4" blockId="25.[163,1426,1597,1980]" box="[163,1425,1597,1621]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF10851172D1F9C4" box="[163,1425,1597,1621]" pageId="25" pageNumber="240">
|
||
1 Epiproct dorsally spiny and apically not emarginate (a)
|
||
<taxonomicName id="4C6C4D53CE6D9874FAA3851172D1F9C4" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1296,1425,1597,1620]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
|
||
<emphasis id="B918EAC2CE6D9874FAA3851172D1F9C4" box="[1296,1425,1597,1620]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6D9874FF10854972D2F9ED" blockId="25.[163,1426,1597,1980]" box="[163,1426,1637,1661]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF10854972D2F9ED" box="[163,1426,1637,1661]" pageId="25" pageNumber="240">- Epiproct not spiny and apically slightly to distinctly emarginate (b–p) 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6D9874FF1085A17685F898" pageId="25" pageNumber="240">
|
||
<paragraph id="8BD336D0CE6D9874FF1085A176EEF952" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF1085A176EEF952" pageId="25" pageNumber="240">
|
||
2 (1) Epiproct much longer than one half of cerci, more than twice as long as wide, apically distinctly
|
||
<taxonomicName id="4C6C4D53CE6D9874FAAD85A272D2F935" authorityName="Selys" authorityYear="1872" box="[1310,1426,1678,1701]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
|
||
<emphasis id="B918EAC2CE6D9874FAAD85A272D2F935" box="[1310,1426,1678,1701]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
|
||
</taxonomicName>
|
||
emarginate (b)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6D9874FF1085FF7685F898" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF1085FF7685F898" pageId="25" pageNumber="240">- Epiproct markedly shorter than one half of cerci, not much longer than wide, apically slightly 3 emarginate (c–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6D9874FF1084347578F803" pageId="25" pageNumber="240">
|
||
<paragraph id="8BD336D0CE6D9874FF10843472D1F8DE" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF10843472D1F8DE" pageId="25" pageNumber="240">
|
||
3 (2) Cerci distinctly increasing in width to markedly less than 1/2 their length; apically simply
|
||
<emphasis id="B918EAC2CE6D9874FACF843572D2F8A0" box="[1404,1426,1817,1840]" italics="true" pageId="25" pageNumber="240">A.</emphasis>
|
||
rounded without apico-lateral spike (c)
|
||
<taxonomicName id="4C6C4D53CE6D9874FAA7841A72D1F8DE" authorityName="Forster" authorityYear="1898" box="[1300,1425,1846,1870]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
|
||
<emphasis id="B918EAC2CE6D9874FAA7841A72D1F8DE" box="[1300,1425,1846,1870]" italics="true" pageId="25" pageNumber="240">maclachlani</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6D9874FF1084727578F803" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF1084727578F803" pageId="25" pageNumber="240">- Cerci distinctly increasing in width to at least 1/2 their length; with apico-lateral spike of 4 variable size and form (d–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6D9877FF108488757EFE90" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240">
|
||
<paragraph id="8BD336D0CE6D9874FF10848872D2F82B" blockId="25.[163,1426,1597,1980]" box="[163,1426,1955,1980]" pageId="25" pageNumber="240">
|
||
<keyLead id="309D95E5CE6D9874FF10848872D2F82B" box="[163,1426,1955,1980]" pageId="25" pageNumber="240">
|
||
4 (3) Cerci widening from base to more than 3/4 length, thence abruptly narrowed (d)
|
||
<taxonomicName id="4C6C4D53CE6D9874FA93848872D2F82B" authorityName="Hagen" authorityYear="1867" box="[1312,1426,1956,1979]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B918EAC2CE6D9874FA93848872D2F82B" box="[1312,1426,1956,1979]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF1083E7757EFE90" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1083E7757EFE90" pageId="26" pageNumber="241">- Cerci widening from base to maximally 2/3 length, thence parallel sided or more or less 5 continuously narrowed (e–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF10823C7557FE1B" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF10823C7697FED6" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10823C7697FED6" pageId="26" pageNumber="241">
|
||
5 (4) Inner edge of cerci straight beyond midlength; ventral edge of cerci without prominent basal
|
||
<taxonomicName id="4C6C4D53CE6E9877FA98823D72D2FEB8" authorityName="Lieftinck" authorityYear="1942" box="[1323,1426,273,296]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="pugnax">
|
||
<emphasis id="B918EAC2CE6E9877FA98823D72D2FEB8" box="[1323,1426,273,296]" italics="true" pageId="26" pageNumber="241">A. pugnax</emphasis>
|
||
</taxonomicName>
|
||
inferior tooth (e–f)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF10827B7557FE1B" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10827B7557FE1B" pageId="26" pageNumber="241">- Inner edge of cerci with some basal convexity or concavity; ventral edge of cerci with prominent 6 basal inferior tooth (g–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF1082B772D2FE4B" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF1082B772D2FE24" blockId="26.[163,1426,203,1186]" box="[163,1426,411,436]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1082B772D2FE24" box="[163,1426,411,436]" pageId="26" pageNumber="241">
|
||
6 (5) Inner edge of cerci widely convex without basal concavity (h)
|
||
<taxonomicName id="4C6C4D53CE6E9877FABB82B072D2FE24" authorityName="Rambur" authorityYear="1842" box="[1288,1426,412,436]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="gibbosulus">
|
||
<emphasis id="B918EAC2CE6E9877FABB82B072D2FE24" box="[1288,1426,412,436]" italics="true" pageId="26" pageNumber="241">A. gibbosulus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF1082E872D2FE4B" blockId="26.[163,1426,203,1186]" box="[163,1426,452,476]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1082E872D2FE4B" box="[163,1426,452,476]" pageId="26" pageNumber="241">- Inner edge of cerci undulate with basal concavity before larger convexity (i–p) 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF1082C1734BFDF7" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF1082C17605FDB1" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1082C17605FDB1" pageId="26" pageNumber="241">7 (6) Cerci approximately three times as long as their greatest width, and apical 1/3 to 1/2 tapered 8 (i–j)</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF10811E734BFDF7" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10811E734BFDF7" pageId="26" pageNumber="241">- Cerci approximately three times as long as their greatest width and apical 1/4 parallel sided (m), 9 or cerci approximately four times as long as their greatest width (k–l, n–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF10815B72D1FD27" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF10815B72D1FD1F" blockId="26.[163,1426,203,1186]" box="[163,1425,631,656]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10815B72D1FD1F" box="[163,1425,631,656]" pageId="26" pageNumber="241">
|
||
8 (7) Cerci greatest width at about 2/3 length and outer margin straight (i)
|
||
<taxonomicName id="4C6C4D53CE6E9877FAA8815472D1FD1F" box="[1307,1425,632,655]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="piraticus">
|
||
<emphasis id="B918EAC2CE6E9877FAA8815472D1FD1F" box="[1307,1425,632,655]" italics="true" pageId="26" pageNumber="241">A. piraticus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF10818C72D1FD27" blockId="26.[163,1426,203,1186]" box="[163,1425,671,696]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10818C72D1FD27" box="[163,1425,671,696]" pageId="26" pageNumber="241">
|
||
- Cerci greatest width at about 1/2 length and outer margin convex (j)
|
||
<taxonomicName id="4C6C4D53CE6E9877FA8E818C72D1FD27" authorityName="Forster" authorityYear="1900" box="[1341,1425,671,695]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="selysi">
|
||
<emphasis id="B918EAC2CE6E9877FA8E818C72D1FD27" box="[1341,1425,671,695]" italics="true" pageId="26" pageNumber="241">A. selysi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF1081E4741AFCD2" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF1081E475F9FD6D" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1081E475F9FD6D" pageId="26" pageNumber="241">9 (7) Cerci with apical 1/4 to 1/3 only slightly tapered and apically distinctly wider than at base, 10 giving them a truncate appearance (k–m)</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF108022741AFCD2" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF108022741AFCD2" pageId="26" pageNumber="241">- Cerci with apical 1/3 more distinctly tapered and apically barely wider than at base, giving them 12 a subtruncate to pointed (n) or rounded (o, p) appearance</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF10807F72D1FC03" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF10807F72D2FCFB" blockId="26.[163,1426,203,1186]" box="[163,1426,851,875]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10807F72D2FCFB" box="[163,1426,851,875]" pageId="26" pageNumber="241">
|
||
10 (9) Cerci inner margin with middle portion not distinctly bulging (k)
|
||
<taxonomicName id="4C6C4D53CE6E9877FAA5807F72D2FCFB" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[1302,1426,851,875]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="panybeus">
|
||
<emphasis id="B918EAC2CE6E9877FAA5807F72D2FCFB" box="[1302,1426,851,875]" italics="true" pageId="26" pageNumber="241">A. panybeus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF10805072D1FC03" blockId="26.[163,1426,203,1186]" box="[163,1425,891,915]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10805072D1FC03" box="[163,1425,891,915]" pageId="26" pageNumber="241">- Cerci inner margin with middle portion distinctly bulging (l–m) 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF10808872D1FC74" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF10808872D1FC2C" blockId="26.[163,1426,203,1186]" box="[163,1425,931,956]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10808872D1FC2C" box="[163,1425,931,956]" pageId="26" pageNumber="241">
|
||
11 (10) Cerci with middle bulge not set off from slightly tapered apical 1/4 (l)
|
||
<taxonomicName id="4C6C4D53CE6E9877FA91808872D1FC2C" authorityName="Hagen" authorityYear="1867" box="[1314,1425,932,956]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="fumosus">
|
||
<emphasis id="B918EAC2CE6E9877FA91808872D1FC2C" box="[1314,1425,932,956]" italics="true" pageId="26" pageNumber="241">A. fumosus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF1080E072D1FC74" blockId="26.[163,1426,203,1186]" box="[163,1425,972,996]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1080E072D1FC74" box="[163,1425,972,996]" pageId="26" pageNumber="241">
|
||
- Cerci with middle bulge set off from parallel sided apical 1/4 (m)
|
||
<taxonomicName id="4C6C4D53CE6E9877FABE80E072D1FC74" authorityName="Lieftinck" authorityYear="1942" box="[1293,1425,972,996]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B918EAC2CE6E9877FABE80E072D1FC74" box="[1293,1425,972,996]" italics="true" pageId="26" pageNumber="241">A. celebensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF1080D876B3FBC2" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF1080D872D2FB9C" blockId="26.[163,1426,203,1186]" box="[163,1426,1012,1036]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1080D872D2FB9C" box="[163,1426,1012,1036]" pageId="26" pageNumber="241">
|
||
12 (9) Cerci with small apico-lateral spike pointing caudo-laterad, making them appear pointed (n)
|
||
<taxonomicName id="4C6C4D53CE6E9877FA9780D872D2FB9C" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1316,1426,1012,1036]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="guttatus">
|
||
<emphasis id="B918EAC2CE6E9877FA9780D872D2FB9C" box="[1316,1426,1012,1036]" italics="true" pageId="26" pageNumber="241">A. guttatus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF10873176B3FBC2" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10873176B3FBC2" pageId="26" pageNumber="241">- Cerci with tiny apico-lateral spike pointing straight backward or slightly incurved, making them 13 appear rounded (o–p)</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30982E75CE6E9877FF10874E72D2FB32" pageId="26" pageNumber="241">
|
||
<paragraph id="8BD336D0CE6E9877FF10874E72D1FBEA" blockId="26.[163,1426,203,1186]" box="[163,1425,1122,1146]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF10874E72D1FBEA" box="[163,1425,1122,1146]" pageId="26" pageNumber="241">
|
||
13 (12) Cerci inner margin with middle portion not distinctly bulging (o)
|
||
<taxonomicName id="4C6C4D53CE6E9877FAA7874E72D1FBEA" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[1300,1425,1122,1146]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
|
||
<emphasis id="B918EAC2CE6E9877FAA7874E72D1FBEA" box="[1300,1425,1122,1146]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF1087A772D2FB32" blockId="26.[163,1426,203,1186]" box="[163,1426,1162,1186]" pageId="26" pageNumber="241">
|
||
<keyLead id="309D95E5CE6E9877FF1087A772D2FB32" box="[163,1426,1162,1186]" pageId="26" pageNumber="241">
|
||
- Cerci inner margin with middle portion distinctly bulging (p)
|
||
<taxonomicName id="4C6C4D53CE6E9877FA8F87A672D2FB32" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[1340,1426,1162,1186]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
|
||
<emphasis id="B918EAC2CE6E9877FA8F87A672D2FB32" box="[1340,1426,1162,1186]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="C376655BCE6E9877FF2487D9737EFA78" pageId="26" pageNumber="241" type="description">
|
||
<paragraph id="8BD336D0CE6E9877FF2487D9748FFA80" blockId="26.[151,975,1269,1296]" box="[151,975,1269,1296]" pageId="26" pageNumber="241">
|
||
<heading id="D09B81BCCE6E9877FF2487D9748FFA80" bold="true" box="[151,975,1269,1296]" fontSize="11" level="1" pageId="26" pageNumber="241" reason="1">
|
||
<emphasis id="B918EAC2CE6E9877FF2487D9748FFA80" bold="true" box="[151,975,1269,1296]" pageId="26" pageNumber="241">
|
||
Abdominal pattern (
|
||
<figureCitation id="13572A55CE6E9877FE2487D976B3FA9F" box="[407,499,1269,1295]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Figs 54</figureCitation>
|
||
,
|
||
<figureCitation id="13572A55CE6E9877FDB387D9755FFA9F" box="[512,543,1269,1295]" captionStart="FIGURE 55" captionStartId="23.[151,250,1850,1874]" captionTargetBox="[161,1422,192,1813]" captionTargetId="figure-17@23.[151,1436,181,1826]" captionTargetPageId="23" captionText="FIGURE 55. Anax spp., males, dorsal, abdominal characters. (a–c) A. insulanus sp. nov.; (d–f) A. tonga sp. nov.: (a, d) abdominal pattern; (b, e) S2–3; (c, f) terminalia; (a, d) illustrated by Albert Orr." figureDoi="http://doi.org/10.5281/zenodo.13915814" httpUri="https://zenodo.org/record/13915814/files/figure.png" pageId="26" pageNumber="241">55</figureCitation>
|
||
) of the
|
||
<taxonomicName id="4C6C4D53CE6E9877FDCE87DA745BFA9F" box="[637,795,1269,1296]" pageId="26" pageNumber="241">
|
||
<emphasis id="B918EAC2CE6E9877FDCE87DA75FEFA80" bold="true" box="[637,702,1270,1296]" italics="true" pageId="26" pageNumber="241">Anax</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(males) keyed
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD336D0CE6E9877FF248612737EFA78" blockId="26.[151,1437,1341,1512]" pageId="26" pageNumber="241">
|
||
Unfortunately it was not possible to present all species in colour, but
|
||
<figureCitation id="13572A55CE6E9877FC0086127361FAC8" box="[947,1057,1341,1368]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Figure 54</figureCitation>
|
||
illustrated the dorsal view of the abdomen of all species, with colours where shown, as known or presumed to occur in life. Because of the small scale and the minute pale pattern elements (barely detectable in preserved specimens) in
|
||
<figureCitation id="13572A55CE6E9877FBD886AA73F9FA30" box="[1131,1209,1413,1440]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Fig. 54</figureCitation>
|
||
the newly described species
|
||
<taxonomicName id="4C6C4D53CE6E9877FF438686763BFA54" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[240,379,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
|
||
<emphasis id="B918EAC2CE6E9877FF438686763BFA54" box="[240,379,1450,1476]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6C4D53CE6E9877FE0686867555FA53" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[437,533,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
|
||
<emphasis id="B918EAC2CE6E9877FE0686867555FA53" box="[437,533,1450,1476]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
|
||
</taxonomicName>
|
||
are also illustrated enlarged as known (
|
||
<taxonomicName id="4C6C4D53CE6E9877FC6986867325FA54" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[986,1125,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
|
||
<emphasis id="B918EAC2CE6E9877FC6986867325FA54" box="[986,1125,1450,1476]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
|
||
</taxonomicName>
|
||
) or presumed (
|
||
<taxonomicName id="4C6C4D53CE6E9877FAA686867235FA53" authorityName="Theischinger & Kalnins & Marinov" authorityYear="2024" box="[1301,1397,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
|
||
<emphasis id="B918EAC2CE6E9877FAA686867235FA53" box="[1301,1397,1450,1476]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
|
||
</taxonomicName>
|
||
) in life, along with enlarged photos of critical characters S2–3 and terminalia (
|
||
<figureCitation id="13572A55CE6E9877FC6E86E27372FA78" box="[989,1074,1485,1512]" captionStart="FIGURE 55" captionStartId="23.[151,250,1850,1874]" captionTargetBox="[161,1422,192,1813]" captionTargetId="figure-17@23.[151,1436,181,1826]" captionTargetPageId="23" captionText="FIGURE 55. Anax spp., males, dorsal, abdominal characters. (a–c) A. insulanus sp. nov.; (d–f) A. tonga sp. nov.: (a, d) abdominal pattern; (b, e) S2–3; (c, f) terminalia; (a, d) illustrated by Albert Orr." figureDoi="http://doi.org/10.5281/zenodo.13915814" httpUri="https://zenodo.org/record/13915814/files/figure.png" pageId="26" pageNumber="241">Fig. 55</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |