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<document id="AF5D6BABFA11D2C3B08EACFAA62B01FC" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFFDF30FFF3095A1D1AAF9AB" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes coyotae Bohart 1948" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="15" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="9FF9E97F030BF7E1DD945F91F7EF2F6E">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<mods:namePart id="D1D0AD2663FA00708FE532A64E79B46F">Grandinete, Yuri Campanholo</mods:namePart>
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<mods:namePart id="6601DFE1BD5C2935777D0CB016146069">Noll, Fernando Barbosa</mods:namePart>
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<treatment id="03C5878FFFFDF30FFF3095A1D1AAF9AB" ID-DOI="http://doi.org/10.5281/zenodo.5986372" ID-GBIF-Taxon="148692476" ID-Zenodo-Dep="5986372" LSID="urn:lsid:plazi:treatment:03C5878FFFFDF30FFF3095A1D1AAF9AB" httpUri="http://treatment.plazi.org/id/03C5878FFFFDF30FFF3095A1D1AAF9AB" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C3766512FFFDF30EFF3095A1D1BFFA8F" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BD33699FFFDF30EFF3095A1D12BFAD1" blockId="14.[151,530,1268,1294]" box="[151,530,1268,1294]" pageId="14" pageNumber="15">
<heading id="D09B81F5FFFDF30EFF3095A1D12BFAD1" bold="true" box="[151,530,1268,1294]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C6C4D1AFFFDF30EFF3095A1D12BFAD1" ID-CoL="3CDFN" authority="Bohart, 1948" authorityName="Bohart" authorityYear="1948" box="[151,530,1268,1294]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFDF30EFF3095A1D12BFAD1" bold="true" box="[151,530,1268,1294]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF3095A1D250FAD1" bold="true" box="[151,361,1268,1294]" italics="true" pageId="14" pageNumber="15">Eumenes coyotae</emphasis>
Bohart, 1948
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD33699FFFDF30EFF30946ED1BFFA8F" blockId="14.[151,1146,1338,1514]" box="[151,646,1338,1360]" pageId="14" pageNumber="15">
<taxonomicName id="4C6C4D1AFFFDF30EFF30946ED2E7FA8F" authority="Bohart, 1948" authorityName="Bohart" authorityYear="1948" box="[151,478,1338,1360]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFDF30EFF30946ED2E7FA8F" bold="true" box="[151,478,1338,1360]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF30946ED275FA8F" bold="true" box="[151,332,1339,1360]" italics="true" pageId="14" pageNumber="15">Eumenes coyotae</emphasis>
Bohart, 1948
</emphasis>
</taxonomicName>
, (4) 24 (9): 315.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFDF30EFF60940CD10FFA14" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BD33699FFFDF30EFF60940CD765FAB0" blockId="14.[151,1146,1338,1514]" box="[199,1116,1369,1391]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF60940CD203FAB0" bold="true" box="[199,314,1369,1391]" pageId="14" pageNumber="15">
<typeStatus id="54D7883BFFFDF30EFF60940CD3C5FAB0" box="[199,252,1369,1391]" pageId="14" pageNumber="15">Type</typeStatus>
data:
</emphasis>
<typeStatus id="54D7883BFFFDF30EFEE6940FD24AFAB0" box="[321,371,1370,1391]" pageId="14" pageNumber="15">Type</typeStatus>
male at
<collectionCode id="ED7DAE5CFFFDF30EFE6E940CD2C3FAB0" box="[457,506,1369,1391]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="14" pageNumber="15">CAS</collectionCode>
(n° 5378)
<emphasis id="B918EA8BFFFDF30EFDC2940CD1DDFAB0" bold="true" box="[613,740,1369,1391]" pageId="14" pageNumber="15">(examined).</emphasis>
<typeStatus id="54D7883BFFFDF30EFD4C940FD07AFAB0" box="[747,835,1370,1391]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
male at
<collectionCode id="ED7DAE5CFFFDF30EFC3D940CD0DAFAB0" box="[922,995,1369,1391]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">USNM</collectionCode>
(n° 56539).
</paragraph>
<paragraph id="8BD33699FFFDF30EFF60942DD743FA51" blockId="14.[151,1146,1338,1514]" box="[199,1146,1400,1422]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF60942DD263FA51" bold="true" box="[199,346,1400,1422]" pageId="14" pageNumber="15">
<typeStatus id="54D7883BFFFDF30EFF60942DD3C5FA51" box="[199,252,1400,1422]" pageId="14" pageNumber="15">Type</typeStatus>
locality:
</emphasis>
“Coyotae Cove,
<collectingRegion id="49A8F87BFFFDF30EFDAF942DD1B8FA51" box="[520,641,1400,1422]" country="Paraguay" name="Concepcion" pageId="14" pageNumber="15">Concepcion</collectingRegion>
Bay, Lower
<collectingRegion id="49A8F87BFFFDF30EFCA4942DD052FA51" box="[771,875,1400,1422]" country="United States of America" name="California" pageId="14" pageNumber="15">California</collectingRegion>
” (
<typeStatus id="54D7883BFFFDF30EFC25942DD0DBFA51" box="[898,994,1400,1422]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54D7883BFFFDF30EFBB4942CD756FA51" box="[1043,1135,1401,1422]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8BD33699FFFDF30EFF6094C3D2DCFA73" blockId="14.[151,1146,1338,1514]" box="[199,485,1430,1452]" pageId="14" pageNumber="15">
<bibRefCitation id="EFFD4B68FFFDF30EFF6094C3D2BEFA73" box="[199,391,1430,1452]" pageId="14" pageNumber="15" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618.
</paragraph>
<paragraph id="8BD33699FFFDF30EFF6094E0D10FFA14" blockId="14.[151,1146,1338,1514]" box="[199,566,1461,1483]" pageId="14" pageNumber="15">Rodríguez-Palafox, 1996: 479 (list).</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFDF30EFF609481D1B3FA35" box="[199,650,1492,1514]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BD33699FFFDF30EFF609481D1B3FA35" blockId="14.[151,1146,1338,1514]" box="[199,650,1492,1514]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF609481D26AFA35" bold="true" box="[199,339,1492,1514]" pageId="14" pageNumber="15">Distribution:</emphasis>
<collectingRegion id="49A8F87BFFFDF30EFEFE9481D29FFA35" box="[345,422,1492,1514]" country="Mexico" name="Mexico" pageId="14" pageNumber="15">
<collectingCountry id="F37B7609FFFDF30EFEFE9481D29FFA35" box="[345,422,1492,1514]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
</collectingRegion>
(
<collectingRegion id="49A8F87BFFFDF30EFE139481D147FA35" box="[436,638,1492,1514]" country="Mexico" name="Baja California Sur" pageId="14" pageNumber="15">
Baja
<collectingRegion id="49A8F87BFFFDF30EFE4D9481D168FA35" box="[490,593,1492,1514]" country="United States of America" name="California" pageId="14" pageNumber="15">California</collectingRegion>
Sur
</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFDF30EFF309743D041F944" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BD33699FFFDF30EFF309743D041F944" blockId="14.[151,1437,1558,2015]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF309743D2F1F9F0" bold="true" box="[151,456,1558,1583]" pageId="14" pageNumber="15">Comments and diagnosis.</emphasis>
<taxonomicName id="4C6C4D1AFFFDF30EFE689742D007F9F0" authority="Bohart, 1948" authorityName="Bohart" authorityYear="1948" box="[463,830,1558,1583]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFDF30EFE689742D1A1F9F1" box="[463,664,1559,1582]" italics="true" pageId="14" pageNumber="15">Eumenes coyotae</emphasis>
Bohart, 1948
</taxonomicName>
, is identified by the following features: body with predominantly ferrugineous background color; T1 long, slender, and with reduced, almost absent, punctation; and the basal half of T2 also with reduced punctation, while on the apical half the punctation becomes coarse and dense. This species is apparently restricted to
<collectingRegion id="49A8F87BFFFDF30EFD3F97D7D04DF944" box="[664,884,1666,1691]" country="Mexico" name="Baja California Sur" pageId="14" pageNumber="15">
Baja
<collectingRegion id="49A8F87BFFFDF30EFD7597D7D07DF944" box="[722,836,1666,1691]" country="United States of America" name="California" pageId="14" pageNumber="15">California</collectingRegion>
Sur
</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFDF30FFF6097F3D3D9FAEF" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="8BD33699FFFDF30EFF6097F3D08DF960" blockId="14.[151,1437,1558,2015]" box="[199,948,1702,1727]" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF6097F3D060F960" bold="true" box="[199,857,1702,1727]" pageId="14" pageNumber="15">
Redescription. Holotype
<taxonomicName id="4C6C4D1AFFFDF30EFE4897F2D060F960" authority="Bohart, 1948" authorityName="Bohart" authorityYear="1948" box="[495,857,1702,1727]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFDF30EFE4897F2D181F961" bold="true" box="[495,696,1703,1726]" italics="true" pageId="14" pageNumber="15">Eumenes coyotae</emphasis>
Bohart, 1948
</taxonomicName>
</emphasis>
(
<figureCitation id="13572A1CFFFDF30EFCCF97F3D095F960" box="[872,940,1702,1727]" captionStart="FIGURES 112" captionStartId="41.[151,264,1386,1408]" captionTargetBox="[151,1436,193,1341]" captionTargetId="figure@41.[151,1436,193,1341]" captionTargetPageId="41" captionText="FIGURES 112. Eumenes species, lateral view. 1) Eumenes americanus de Saussure (female); 2) E. belfragei aureus Isely, 1917 (Holotype, female); 3) E. bollii Cresson, 1872 (Lectotype, female); 4) E. bollii oregonensis Bequaert, 1938 (Holotype, female); 5); E. bequaerti Isely, 1917 (Holotype, male); 6) E. iturbide (= E. consobrinus; Holotype, female); 7) E. coyotae Bohart, 1948 (Holotype, male); 8) E. crucifera Provancher, 1888 (Holotype, female). 9) E. bolliformis Viereck, 1908 (Lectotype, female); 10) E. crucifera var. nearcticus Bequaert, 1944 (female); 11) E. crucifera stricklandi Bequaert, 1944 (Holotype, female); 12) E. cruciferoides Giordani Soika, 1978 (Holotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458304/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="8BD33699FFFDF30FFF60979FD180FF27" blockId="14.[151,1437,1558,2015]" lastBlockId="15.[151,1437,151,1653]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B918EA8BFFFDF30EFF60979FD29CF93C" bold="true" box="[199,421,1738,1763]" pageId="14" pageNumber="15">Male. Color, head:</emphasis>
Yellow marks as follows: mandibles; inter-antennal region; a broad band on inner margin of compound eye that goes from the base of the clypeus to the ocular sinus; a band adjacent to the external margin of the compound eye. Ocelli region with black mark. Frons and vertex (behind the yellow band on gena) ferrugineous. Antenna ferrugineous-yellowish.
<emphasis id="B918EA8BFFFDF30EFD1D9663D078F890" bold="true" box="[698,833,1846,1871]" pageId="14" pageNumber="15">Mesosoma:</emphasis>
predominantly ferrugineous, with yellow marks as follows: broad band along the pronotal carina on dorsal face of pronotum; a mark on upper half of mesepisternum, adjacent to the mesepimeron; parategula; a band on apical third of scutellum; metanotum; marks on femora, almost entirely tibiae and tarsi, although the distal tarsi become brownish darker.
<emphasis id="B918EA8BFFFDF30EFC4F96F7D757F864" bold="true" box="[1000,1134,1954,1979]" pageId="14" pageNumber="15">Metasoma:</emphasis>
ferrugineous, except for a black mark on basal region; with yellow marks as follows: narrow transverse band on apex of T1; broad band on each side of T2 and a band occupying the apical third of it; apical half of S2; T3T7 and S3S7 (with a transverse dark band on apex). Small black mark on the base of T2.
<emphasis id="B918EA8BFFFCF30FFCB991E9D04AFF0A" bold="true" box="[798,883,188,213]" pageId="15" pageNumber="16">Wings:</emphasis>
Hyaline, with costal, medial and submedial cells yellowish. Pterostigma and veins dark brownish.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF609051D78EFE0F" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF609051D299FEC2" bold="true" box="[199,416,260,285]" pageId="15" pageNumber="16">Pubescence, head:</emphasis>
Golden and moderately long pubescence on frons, vertex and occipital region. Clypeus and gena with short and whitish pubescence, better observed in oblique view. Scape with short (as in clypeus) golden pubescence.
<emphasis id="B918EA8BFFFCF30FFE899019D28FFEBA" bold="true" box="[302,438,332,357]" pageId="15" pageNumber="16">Mesosoma:</emphasis>
Pronotum, mesoscutum, scutellum and metanotum with short golden pubescence. Mesepisternum and posterior face of propodeum with whitish pubescence, better observed in oblique view. On posterior face of propodeum the pubescence is longer, as in frons.
<emphasis id="B918EA8BFFFCF30FFC3B90C1D71BFE72" bold="true" box="[924,1058,404,429]" pageId="15" pageNumber="16">Metasoma:</emphasis>
Golden and shortest pubescence along body, except for some scattered bristles on ventral face of T1 and apex of each segment.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF609089D777FD5A" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF609089D112FE2A" bold="true" box="[199,555,476,501]" pageId="15" pageNumber="16">Surface of integument, head:</emphasis>
Clypeus with punctation mostly on basal two-thirds; frons and vertex with coarse and dense punctation, becoming sparser behind the ocelli.
<emphasis id="B918EA8BFFFCF30FFC2590AAD730FDC7" bold="true" box="[898,1033,511,536]" pageId="15" pageNumber="16">Mesosoma:</emphasis>
Coarse and dense punctation as on frons.
<emphasis id="B918EA8BFFFCF30FFF469371D25EFDE2" bold="true" box="[225,359,548,573]" pageId="15" pageNumber="16">Metasoma:</emphasis>
T1 with reduced punctation, at least with some more evident on apex; basal half of T2 with weak punctation, very sparse, while the apical half the punctures becomes coarse and clearly denser. S2 with evident punctation on lateral portions. T3T7 and S3S7 without evident punctation.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF6093DAD7A6FC32" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF6093DAD2B3FD77" bold="true" box="[199,394,655,680]" pageId="15" pageNumber="16">Structure, head:</emphasis>
Clypeus convex, longer than wide. Apex of clypeus concave, forming two projections with carina on apex of each one. Inter-antennal region longitudinally cariniform, narrow than the antennal socket. Lateral ocelli closer to the compound eyes than to each other. Occipital carina well developed, not angled on middle area of gena. F11 short, without longitudinal carina on dorsal surface, with the apex pointed and surpassing the apical edge of F8 (
<figureCitation id="13572A1CFFFCF30FFE06924AD2C1FCE7" box="[417,504,799,824]" captionStart="FIGURES 7792" captionStartId="48.[151,264,1946,1968]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@48.[151,1436,193,1925]" captionTargetPageId="48" captionText="FIGURES 7792. First tergum, dorsal view. 77) Eumenes americanus; 78) E. bequaerti; 79) E. bollii; 80) E. consobrinus; 81) E. coyotae; 82) E. crucifera; 83) E. cubensis; 84) E. flavitinctus; 85) E. fraternus; 86) E. pictus; 87) E. sculleni; 88) E. smithii; 89) E. verticalis; 90) E. filiformis; 91) E. rufomaculactus; 92) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458318/files/figure.png" pageId="15" pageNumber="16">Fig. 81</figureCitation>
). Ventral surface with microscopic bristles on apex.
<emphasis id="B918EA8BFFFCF30FFBFA924AD7DDFCE7" bold="true" box="[1117,1252,799,824]" pageId="15" pageNumber="16">Mesosoma:</emphasis>
Pronotal carina present dorsally, weak developed, while on lateral portion it becomes more developed. Pronotal fovea present. Posterior projection of tegula weak developed, rounded; parategula lamelliform; posterior face of propodeum moderately concave.
<emphasis id="B918EA8BFFFCF30FFE2C92D9D129FC7A" bold="true" box="[395,528,908,933]" pageId="15" pageNumber="16">Metasoma:</emphasis>
T1 more than three times longer than wide, slender, weakly swollen in dorsal view (
<figureCitation id="13572A1CFFFCF30FFF7892FAD20CFC17" box="[223,309,943,968]" captionStart="FIGURES 2742" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 2742. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="15" pageNumber="16">Fig. 31</figureCitation>
). T2 globose, longer than wide (
<figureCitation id="13572A1CFFFCF30FFD0992FAD03DFC17" box="[686,772,943,968]" captionStart="FIGURES 4358" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 4358. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="15" pageNumber="16">Fig. 47</figureCitation>
), with the basal angle (in lateral view) rising up abruptly. Apical lamella present and becoming reduced towards lateral margins. S7 flattened apically.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF6092A2D277FBA3" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF6092A2D245FBCF" bold="true" box="[199,380,1015,1040]" pageId="15" pageNumber="16">Male genitalia:</emphasis>
Aedeagus as on figure 97a, b. Paramere (
<figureCitation id="13572A1CFFFCF30FFCF192A2D08CFBCF" box="[854,949,1015,1040]" captionStart="FIGURES 93101" captionStartId="49.[151,264,1500,1522]" captionTargetBox="[151,1435,193,1478]" captionTargetId="figure@49.[151,1436,193,1478]" captionTargetPageId="49" captionText="FIGURES 93101. Male genitalia: a, b) aedeagus (lateral and ventral view, respectively); c) paramere, internal view. 93) Eumenes americanus; 94) E. bequaerti; 95) E. bollii; 96) E. consobrinus; 97) E. coyotae; 98) E. crucifera; 99) E. cubensis; 100) E. flavitinctus; 101) E. fraternus. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458320/files/figure.png" pageId="15" pageNumber="16">Fig. 97c</figureCitation>
) with long bristles on middle region of the gonostyle; cuspis with thin, long and dense pubescence; digitus slender on apex, with short bristles on ventral surface (
<figureCitation id="13572A1CFFFCF30FFF5F956AD263FB87" box="[248,346,1087,1112]" captionStart="FIGURES 110125" captionStartId="51.[151,264,1899,1921]" captionTargetBox="[163,1425,209,1857]" captionTargetId="figure@51.[150,1437,193,1878]" captionTargetPageId="51" captionText="FIGURES 110125. Male genitalia: digitus. 110) Eumenes americanus; 111) E. bequaerti; 112) E. bollii; 113) E. consobrinus; 114) E. coyotae; 115) E. crucifera; 116) E. cubensis; 117) E. flavitinctus; 118) E. fraternus; 119) E. pictus; 120) E. sculleni; 121) E. smithii; 122) E. verticalis; 123) E. filiformis; 124) E. rufomaculatus; 125) E. versicolor." httpUri="https://zenodo.org/record/1458324/files/figure.png" pageId="15" pageNumber="16">Fig. 114</figureCitation>
); volsella with short (shorter than on cuspis) and denser pubescence; distal lobe with a few erect bristles on apex.
</paragraph>
<paragraph id="8BD33699FFFCF30FFF6095D2D3D9FAEF" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF6095D2D21EFB7F" bold="true" box="[199,295,1159,1184]" pageId="15" pageNumber="16">Female:</emphasis>
Black marks as follows: marks on frons more developed, reaching the vertex; small spot on mesepimeron; narrow, sticklike band longitudinally on mesoscutum and a small spot on the base; broad transverse band between two yellow band on middle area. Pubescence and punctation as on males. Labrum with the apex rounded; clypeus as long as wide; inter-antennal region as long as the antennal socket. All other structure as in males.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFFCF30FFF609469D1AAF9AB" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BD33699FFFCF30FFF609469D1AAF9AB" blockId="15.[151,1437,151,1653]" pageId="15" pageNumber="16">
<emphasis id="B918EA8BFFFCF30FFF609469D24FFA8A" bold="true" box="[199,374,1340,1365]" pageId="15" pageNumber="16">Type material.</emphasis>
The holotype of
<taxonomicName id="4C6C4D1AFFFCF30FFD9C9468D1C6FA8B" authorityName="Bohart" authorityYear="1948" box="[571,767,1341,1364]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFD9C9468D1C6FA8B" box="[571,767,1341,1364]" italics="true" pageId="15" pageNumber="16">Eumenes coyotae</emphasis>
</taxonomicName>
is in good condition and bears the labels: Coyotae Cove \ Concepcion Bay, \ L. Cal.
<date id="FFD21059FFFCF30FFE769435D11FFAA7" box="[465,550,1376,1400]" pageId="15" pageNumber="16" value="1941-10-01">X-1-41</date>
Ross &amp; Bohart \ Collectors [Red Label]
<taxonomicName id="4C6C4D1AFFFCF30FFB8A9434D7ABFAA7" box="[1069,1170,1377,1400]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFFCF30FFB8A9434D7ABFAA7" box="[1069,1170,1377,1400]" italics="true" pageId="15" pageNumber="16">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFFCF30FFB099434D63CFAA7" authorityName="Bohart" authorityYear="1948" box="[1198,1285,1377,1400]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFB099434D63CFAA7" box="[1198,1285,1377,1400]" italics="true" pageId="15" pageNumber="16">coyotae</emphasis>
</taxonomicName>
\ R. Bohart California Academy \ of Sciences \ Type No. 5378. The paratype male (AMNH) is in good condition and bears the labels: Ross &amp; Bohart \ Collectors
<taxonomicName id="4C6C4D1AFFFCF30FFDC394FCD1F0FA1F" box="[612,713,1449,1472]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFFCF30FFDC394FCD1F0FA1F" box="[612,713,1449,1472]" italics="true" pageId="15" pageNumber="16">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFFCF30FFD4794FCD00EFA1F" authorityName="Bohart" authorityYear="1948" box="[736,823,1449,1472]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFD4794FCD00EFA1F" box="[736,823,1449,1472]" italics="true" pageId="15" pageNumber="16">coyotae</emphasis>
</taxonomicName>
\ Bohart \ R. Bohart det.. Two paratype males (UCD) are in good condition and bear the labels: Coyote Cove \ Conception Bay \ L. Cal.
<date id="FFD21059FFFCF30FFBF69499D787FA3B" box="[1105,1214,1484,1508]" pageId="15" pageNumber="16" value="1938-06-29">VI-29-38</date>
Michelbacher &amp; \ Ross Collectors
<taxonomicName id="4C6C4D1AFFFCF30FFEC594A4D2FEF9D7" box="[354,455,1521,1544]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFFCF30FFEC594A4D2FEF9D7" box="[354,455,1521,1544]" italics="true" pageId="15" pageNumber="16">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFFCF30FFE7A94A4D10DF9D7" authorityName="Bohart" authorityYear="1948" box="[477,564,1521,1544]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFE7A94A4D10DF9D7" box="[477,564,1521,1544]" italics="true" pageId="15" pageNumber="16">coyotae</emphasis>
</taxonomicName>
\ R. Bohart; Espiritu Santo \ I. Golf. Calif. \
<date id="FFD21059FFFCF30FFBE794A4D7F4F9D7" box="[1088,1229,1520,1544]" pageId="15" pageNumber="16" value="1921-06-09">June 9 1921</date>
E P Van Duzee \ Collector
<taxonomicName id="4C6C4D1AFFFCF30FFEBD9740D246F9F3" box="[282,383,1557,1580]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFFCF30FFEBD9740D246F9F3" box="[282,383,1557,1580]" italics="true" pageId="15" pageNumber="16">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFFCF30FFE319740D2D4F9F3" authorityName="Bohart" authorityYear="1948" box="[406,493,1557,1580]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFE319740D2D4F9F3" box="[406,493,1557,1580]" italics="true" pageId="15" pageNumber="16">coyotae</emphasis>
</taxonomicName>
\ R. Bohart. The paratype male (USNM) is in good condition and bears the labels: Coyotae Cove, \ Concepcion Bay, \ L. Cal.
<date id="FFD21059FFFCF30FFD2C976DD1D9F98F" box="[651,736,1592,1616]" pageId="15" pageNumber="16" value="1941-10-01">X-1-41</date>
Rose &amp; Bohart \ Collectors
<taxonomicName id="4C6C4D1AFFFCF30FFBE3976CD790F98F" box="[1092,1193,1593,1616]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFFCF30FFBE3976CD790F98F" box="[1092,1193,1593,1616]" italics="true" pageId="15" pageNumber="16">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFFCF30FFB67976CD62EF98F" authorityName="Bohart" authorityYear="1948" box="[1216,1303,1593,1616]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="coyotae">
<emphasis id="B918EA8BFFFCF30FFB67976CD62EF98F" box="[1216,1303,1593,1616]" italics="true" pageId="15" pageNumber="16">coyotae</emphasis>
</taxonomicName>
\ R. Bohart det. [Red Label] Paratype \ 56539 \ USNM.
</paragraph>
</subSubSection>
</treatment>
</document>