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<document id="379825FE74CFDCE4505F3E0262FB1C0F" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F06DFFB9FF4876E033D12157" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Panghoella VIRAKTAMATH, WEBB &amp; YESHWANTH, 2025, gen. nov." docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="37" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="5C2FB7E16FDE73B88D73169AB0BD3292">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="F2C25B03772533255400704E2036EBC0">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="CF6C0AF25BA0EF131ECC0BE46A90B000">WEBB, M. D.</mods:namePart>
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<subSubSection id="C304BDDEF06DFFB8FF4876E037DA2292" box="[151,407,438,465]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BA1EE55F06DFFB8FF4876E037DA2292" blockId="36.[151,407,438,465]" box="[151,407,438,465]" pageId="36" pageNumber="37">
<heading id="D0E95939F06DFFB8FF4876E037DA2292" bold="true" box="[151,407,438,465]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
<emphasis id="B96A3247F06DFFB8FF4876E037DA2292" bold="true" box="[151,407,438,465]" pageId="36" pageNumber="37">
<taxonomicName id="4C1E95D6F06DFFB8FF4876E0376F2292" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,290,438,465]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96A3247F06DFFB8FF4876E0376F2292" bold="true" box="[151,290,438,465]" italics="true" pageId="36" pageNumber="37">Panghoella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2598F3CF06DFFB8FEF676E137DA2292" box="[297,407,439,465]" pageId="36" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF06DFFB8FF4876AA32CD25D0" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BA1EE55F06DFFB8FF4876AA34942150" blockId="36.[151,729,507,532]" box="[151,729,507,532]" pageId="36" pageNumber="37">
<typeStatus id="54A550F7F06DFFB8FF4876AA36842157" box="[151,201,508,532]" pageId="36" pageNumber="37">Type</typeStatus>
species:
<taxonomicName id="4C1E95D6F06DFFB8FEF976AA34642150" authority="Distant" authorityName="Distant" box="[294,553,507,531]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B96A3247F06DFFB8FEF976AA379A2150" box="[294,471,507,531]" italics="true" pageId="36" pageNumber="37">Banus maculosus</emphasis>
Distant
</taxonomicName>
, here designated.
</paragraph>
<paragraph id="8BA1EE55F06DFFB8FF48751433C92184" blockId="36.[151,1437,577,2008]" pageId="36" pageNumber="37">
<emphasis id="B96A3247F06DFFB8FF48751437442118" box="[151,265,578,603]" italics="true" pageId="36" pageNumber="37">Diagnosis</emphasis>
. Coloration similar to
<taxonomicName id="4C1E95D6F06DFFB8FDC9751434112118" box="[534,604,578,603]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FDC9751434112118" box="[534,604,578,603]" italics="true" pageId="36" pageNumber="37">Banus</emphasis>
</taxonomicName>
. Male pygofer with dorsal process arising from dorsal margin where it articulates with segment IX, directed ventrally (absent in
<taxonomicName id="4C1E95D6F06DFFB8FCC375303527213C" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[796,874,614,639]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nera" status="sp. nov.">
<emphasis id="B96A3247F06DFFB8FCC375303527213C" box="[796,874,614,639]" italics="true" pageId="36" pageNumber="37">P. nera</emphasis>
</taxonomicName>
<emphasis id="B96A3247F06DFFB8FCAD75333581213C" bold="true" box="[882,972,613,639]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A2598F3CF06DFFB8FCAD75333581213C" box="[882,972,613,639]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Subgenital plate with only narrow hair-like setae marginally and short thin setae on ventral surface. Style without subapical lobe. Aedeagus with shaft cylindrical, symmetrical, with one pair of processes arising at base of atrium ventrally and articulated with shaft.
</paragraph>
<paragraph id="8BA1EE55F06DFFB8FF187584330521AF" blockId="36.[151,1437,577,2008]" box="[199,1352,721,748]" pageId="36" pageNumber="37">
<emphasis id="B96A3247F06DFFB8FF187584370621A8" box="[199,331,722,747]" italics="true" pageId="36" pageNumber="37">Description</emphasis>
. Brownish ochraceous with pale and brown markings on head, pronotum, and mesonotum.
</paragraph>
<paragraph id="8BA1EE55F06DFFB8FF1875A3378C27A0" blockId="36.[151,1437,577,2008]" pageId="36" pageNumber="37">
Medium sized leafhoppers measuring
<quantity id="4CE643B0F06DFFB8FDA775A3354C2053" box="[632,769,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.1" metricValueMin="3.5" pageId="36" pageNumber="37" unit="mm" value="4.8" valueMax="6.1" valueMin="3.5">3.56.1 mm</quantity>
long. Head wider than or narrower than pronotum. Crown surface with apical half shagreen, posterior half polished; median sulcus about 0.75 as long as crown; fore margin rounded to face. Eyes large, their outer margin in line with outer margin of crown, obliquely extending over anterolateral angles of pronotum. Ocelli prominent placed a distance equal to own diameter away from eyes. Antennae long, reaching almost half length of body. Face including eyes about as wide as long; frontoclypeus somewhat narrow; clypellus broad apically. Pronotum slightly convex dorsally, about 1.61.9× as long as crown; with lateral margins carinate, surface polished, punctate, posterior margin almost straight. Exposed mesonotum about as long as or distinctly longer than pronotum. Forewing with claval veins separate, inner claval vein connected to claval suture by cross vein, three anteapical and four apical cells, inner anteapical cell open behind, appendix narrow not extending beyond middle of second apical cell. Pro-tibia comparatively long and slender compared to meso-tibia; AV row of setae 1516, conical, short peg-like, in basal 0.66; AM row of setae represented by hair-like AM1, IC row of setae 1415, slender hair-like. Metafemur with 2+2+1 apical macrosetae Meta tibia with PD row macrosetae long, as long or longer than half length of fore tibia. Metabasitarsomere with 3 platellae and one stout tapered seta on either side.
</paragraph>
<paragraph id="8BA1EE55F06DFFB8FF1873BB3454252C" blockId="36.[151,1437,577,2008]" pageId="36" pageNumber="37">
<emphasis id="B96A3247F06DFFB8FF1873BB37222644" box="[199,367,1261,1287]" italics="true" pageId="36" pageNumber="37">Male genitalia</emphasis>
. Pygofer with or without anterior apodemes, tergum short, lobes well produced posteriorly, making tergum deeply U-shaped in dorsal view, with numerous macrosetae, in distal half and row of marginal setae, ventral margin convex, posterodorsal angle subconically rounded dorsal processes (absent in
<taxonomicName id="4C1E95D6F06DFFB8FAC17260333D260C" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[1310,1392,1334,1359]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nera" status="sp. nov.">
<emphasis id="B96A3247F06DFFB8FAC17260333D260C" box="[1310,1392,1334,1359]" italics="true" pageId="36" pageNumber="37">P. nera</emphasis>
</taxonomicName>
<emphasis id="B96A3247F06DFFB8FAA4726336852630" bold="true" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A2598F3CF06DFFB8FAA4726336852630" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) arising from midlength of pygofer dorsal margin and directed ventrally. Valve with posterior margin angularly produced medially. Subgenital plates triangular, elongate, about 2.53.0× as long as basal width, with marginal row of narrow setae in distal 0.66 and narrow short setae on ventral surface or with submarginal row of macrosetae in distal 0.75 region and numerous hair-like setae in distal half. Style without subapical lobe, apophysis pointed and curved near apex, surface transversely corrugated. Connective with stem distinctly shorter than arms. Aedeagus symmetrical, with well-developed dorsal apodeme, shaft slightly curved, apex not produced beyond gonopore; gonopore apical; a pair of articulated basal processes arising from ventral part of atrium. Female valvulae as in the genus
<taxonomicName id="4C1E95D6F06DFFB8FF3F7100376B252C" box="[224,294,1622,1647]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FF3F7100376B252C" box="[224,294,1622,1647]" italics="true" pageId="36" pageNumber="37">Banus</emphasis>
</taxonomicName>
and
<emphasis id="B96A3247F06DFFB8FE8371003454252C" box="[348,537,1622,1647]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="4C1E95D6F06DFFB8FE8371003458252C" authorityName="Dlabola" authorityYear="2025" box="[348,533,1622,1647]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Mavromoustaca</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BA1EE55F06DFFB8FF18712F32CD25D0" blockId="36.[151,1437,577,2008]" box="[199,1152,1657,1684]" pageId="36" pageNumber="37">
<emphasis id="B96A3247F06DFFB8FF18712F370825D0" box="[199,325,1657,1683]" italics="true" pageId="36" pageNumber="37">Etymology.</emphasis>
The generic name is an arbitrary combination of letters, gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF06DFFB9FF1871CB33292396" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BA1EE55F06DFFB8FF1871CB37C624F7" blockId="36.[151,1437,577,2008]" pageId="36" pageNumber="37">
<emphasis id="B96A3247F06DFFB8FF1871CB377925F4" bold="true" box="[199,308,1693,1719]" pageId="36" pageNumber="37">Remarks</emphasis>
. This genus closely externally resembles
<taxonomicName id="4C1E95D6F06DFFB8FCDE71CB35CF25F4" box="[769,898,1693,1719]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FCDE71CB35CF25F4" box="[769,898,1693,1719]" italics="true" pageId="36" pageNumber="37">Monobazus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E95D6F06DFFB8FC6871C8322125F4" authorityName="Dlabola" authorityYear="2025" box="[951,1132,1694,1719]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FC6871C8322125F4" box="[951,1132,1694,1719]" italics="true" pageId="36" pageNumber="37">Mavromoustaca</emphasis>
</taxonomicName>
in coloration and structure, but differs in the male genitalia: pygofer with dorsal process (ventral processes in other mentioned genera), style with subapical lobe absent (prominently developed in other genera), aedeagus with articulated pair of ventral processes arising from base of atrium and articulated with it (lateroventral basal processes in
<taxonomicName id="4C1E95D6F06DFFB8FB38705C33D12460" authorityName="Dlabola" authorityYear="2025" box="[1255,1436,1802,1827]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FB38705C33D12460" box="[1255,1436,1802,1827]" italics="true" pageId="36" pageNumber="37">Mavromoustaca</emphasis>
</taxonomicName>
fused with shaft; ventral processes arising from shaft and fused with it in
<taxonomicName id="4C1E95D6F06DFFB8FC09707B321A2404" box="[982,1111,1837,1863]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FC09707B321A2404" box="[982,1111,1837,1863]" italics="true" pageId="36" pageNumber="37">Monobazus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C1E95D6F06DFFB8FBB0707B32BD2404" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[1135,1264,1837,1863]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FBB0707B32BD2404" box="[1135,1264,1837,1863]" italics="true" pageId="36" pageNumber="37">Panghoella</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="4C1E95D6F06DFFB8FF48700737612428" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,300,1873,1899]" class="Insecta" family="Cicadellidae" genus="Chandibanus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96A3247F06DFFB8FF48700737612428" box="[151,300,1873,1899]" italics="true" pageId="36" pageNumber="37">Chandibanus</emphasis>
</taxonomicName>
<emphasis id="B96A3247F06DFFB8FEE8700737E92428" bold="true" box="[311,420,1873,1899]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A2598F3CF06DFFB8FEE8700737E92428" box="[311,420,1873,1899]" pageId="36" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in having a dorsal marginal pygofer process in the male, but they differ in having the processes directed posteriorly in
<taxonomicName id="4C1E95D6F06DFFB8FDCF702334E824CC" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[528,677,1909,1935]" class="Insecta" family="Cicadellidae" genus="Chandibanus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FDCF702334E824CC" box="[528,677,1909,1935]" italics="true" pageId="36" pageNumber="37">Chandibanus</emphasis>
</taxonomicName>
and ventrally in
<taxonomicName id="4C1E95D6F06DFFB8FCB3702335A024CC" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[876,1005,1909,1935]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FCB3702335A024CC" box="[876,1005,1909,1935]" italics="true" pageId="36" pageNumber="37">Panghoella</emphasis>
</taxonomicName>
and the aedeagus in the former lacks basal atrial processes.
</paragraph>
<paragraph id="8BA1EE55F06DFFB9FF1870EB33292396" blockId="36.[151,1437,577,2008]" lastBlockId="37.[151,1436,150,213]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Based on their male genitalia, two species described in
<taxonomicName id="4C1E95D6F06DFFB8FC8B70EB35902494" authorityName="Ball" authorityYear="1932" box="[852,989,1981,2007]" class="Insecta" family="Cicadellidae" genus="Osbornellus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F06DFFB8FC8B70EB35902494" box="[852,989,1981,2007]" italics="true" pageId="36" pageNumber="37">Osbornellus</emphasis>
</taxonomicName>
from
<collectingCountry id="F309AEC5F06DFFB8FBF770EB32262494" box="[1064,1131,1981,2007]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
belong to this genus, i.e.,
<taxonomicName id="4C1E95D6F06CFFB9FF4877C137CB23F2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[151,390,151,177]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="conoidea">
<emphasis id="B96A3247F06CFFB9FF4877C137CB23F2" box="[151,390,151,177]" italics="true" pageId="37" pageNumber="38">Panghoella conoidea</emphasis>
</taxonomicName>
(Xing &amp; Li in
<bibRefCitation id="EF8F93A4F06CFFB9FDF177C134B323F2" author="Li, Z. Z. &amp; Dai, R. H. &amp; Xing, L. H." box="[558,766,150,177]" pageId="37" pageNumber="38" refId="ref51067" refString="Li, Z. Z., Dai, R. H. &amp; Xing, L. H. (2011) Deltocphalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, China, 336 pp." type="book" year="2011">
Li
<emphasis id="B96A3247F06CFFB9FD9277C134CB23F2" box="[589,646,151,177]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2011: 157
</bibRefCitation>
, figs 5147 17; Dmitriev
<emphasis id="B96A3247F06CFFB9FBF377C1322923F2" box="[1068,1124,151,177]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2024: 276)
<emphasis id="B96A3247F06CFFB9FB3477C1332B23F2" bold="true" box="[1259,1382,151,177]" pageId="37" pageNumber="38">comb. nov</emphasis>
. and
<taxonomicName id="4C1E95D6F06CFFB9FF4877ED370F2397" box="[151,322,187,212]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="suiyongensis">
<emphasis id="B96A3247F06CFFB9FF4877ED370F2397" box="[151,322,187,212]" italics="true" pageId="37" pageNumber="38">P. suiyongensis</emphasis>
</taxonomicName>
(Xing &amp; Li in
<bibRefCitation id="EF8F93A4F06CFFB9FE3477ED34BE2396" author="Li, Z. Z. &amp; Dai, R. H. &amp; Xing, L. H." box="[491,755,186,213]" pageId="37" pageNumber="38" refId="ref51067" refString="Li, Z. Z., Dai, R. H. &amp; Xing, L. H. (2011) Deltocphalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, China, 336 pp." type="book" year="2011">
Li
<emphasis id="B96A3247F06CFFB9FDD577ED34712396" box="[522,572,187,213]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
. 2011: 157158
</bibRefCitation>
, figs 5148 17; Dmitriev
<emphasis id="B96A3247F06CFFB9FBFD77ED32172396" box="[1058,1114,187,213]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2024: 276)
<emphasis id="B96A3247F06CFFB9FB3D77ED33102396" bold="true" box="[1250,1373,187,213]" pageId="37" pageNumber="38">comb. nov</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF06CFFB9FF48767133D12157" pageId="37" pageNumber="38" type="key">
<paragraph id="8BA1EE55F06CFFB9FF487671320A2203" blockId="37.[151,1095,294,321]" box="[151,1095,294,321]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF487671320A2203" bold="true" box="[151,1095,294,321]" pageId="37" pageNumber="38">
Key to species of
<taxonomicName id="4C1E95D6F06CFFB9FEB1767037B42202" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[366,505,294,321]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96A3247F06CFFB9FEB1767037B42202" bold="true" box="[366,505,294,321]" italics="true" pageId="37" pageNumber="38">Panghoella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2598F3CF06CFFB9FDDF767134232202" box="[512,622,295,321]" pageId="37" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
from the Indian subcontinent (males)
</emphasis>
</paragraph>
<key id="FF8FB964F06CFFB9FF48763A33D12157" pageId="37" pageNumber="38">
<keyStep id="30EAF6F0F06CFFB9FF48763A33D1229A" pageId="37" pageNumber="38">
<paragraph id="8BA1EE55F06CFFB9FF48763A33D122DD" blockId="37.[151,1437,363,532]" pageId="37" pageNumber="38">
<keyLead id="30EF4D60F06CFFB9FF48763A33D122DD" pageId="37" pageNumber="38">
1. Pygofer in lateral view less than 1.5× as long as height, dorsal process present or absent, if present more or less straight (
<figureCitation id="1325F2D0F06CFFB9FAAA763A374222DD" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47A</figureCitation>
); aedeagus with basal processes longer than shaft (
<figureCitation id="1325F2D0F06CFFB9FD2976DF350122DD" box="[758,844,393,415]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47G</figureCitation>
) or subequal (
<figureCitation id="1325F2D0F06CFFB9FC0876DF326322DD" box="[983,1070,393,415]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48AI. Panghoella nera sp. nov.: AB, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; GH, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="37" pageNumber="38">Fig. 48H</figureCitation>
)............................ 2
</keyLead>
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF4876F133D1229A" blockId="37.[151,1437,363,532]" pageId="37" pageNumber="38">
<keyLead id="30EF4D60F06CFFB9FF4876F133D1229A" pageId="37" pageNumber="38">
- Pygofer in lateral view more than 1.5× as long as height, dorsal process angularly curved at midlength (
<figureCitation id="1325F2D0F06CFFB9FB0876F0337F22F8" box="[1239,1330,422,444]" captionStart="FIGURES 49" captionStartId="90.[151,264,1860,1884]" captionTargetBox="[156,1430,186,1831]" captionTargetId="figure-26@90.[151,1435,181,1836]" captionTargetPageId="90" captionText="FIGURES 49AF. Panghoella sinuata sp. nov.: A, pygofer, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; D, style, lateral view; EF, aedeagus lateral view at different angles." figureDoi="http://doi.org/10.5281/zenodo.14703877" httpUri="https://zenodo.org/record/14703877/files/figure.png" pageId="37" pageNumber="38">Fig. 49A</figureCitation>
); aedeagus with basal process shorter than shaft (
<figureCitation id="1325F2D0F06CFFB9FD93769234ED229A" box="[588,672,452,474]" captionStart="FIGURES 49" captionStartId="90.[151,264,1860,1884]" captionTargetBox="[156,1430,186,1831]" captionTargetId="figure-26@90.[151,1435,181,1836]" captionTargetPageId="90" captionText="FIGURES 49AF. Panghoella sinuata sp. nov.: A, pygofer, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; D, style, lateral view; EF, aedeagus lateral view at different angles." figureDoi="http://doi.org/10.5281/zenodo.14703877" httpUri="https://zenodo.org/record/14703877/files/figure.png" pageId="37" pageNumber="38">Fig. 49E</figureCitation>
)................................................
<taxonomicName id="4C1E95D6F06CFFB9FB3576923304229A" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[1258,1353,452,473]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sinuata" status="sp. nov.">
<emphasis id="B96A3247F06CFFB9FB3576923304229A" box="[1258,1353,452,473]" italics="true" pageId="37" pageNumber="38">P. sinuata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2598F3CF06CFFB9FA91769233D1229A" box="[1358,1436,452,473]" pageId="37" pageNumber="38" rank="species">
<emphasis id="B96A3247F06CFFB9FA91769233DB229A" bold="true" box="[1358,1430,452,473]" pageId="37" pageNumber="38">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EAF6F0F06CFFB9FF4876B733D12157" pageId="37" pageNumber="38">
<paragraph id="8BA1EE55F06CFFB9FF4876B733D022B4" blockId="37.[151,1437,363,532]" box="[151,1437,481,503]" pageId="37" pageNumber="38">
<keyLead id="30EF4D60F06CFFB9FF4876B733D022B4" box="[151,1437,481,503]" pageId="37" pageNumber="38">
2. Pygofer with dorsal process (
<figureCitation id="1325F2D0F06CFFB9FE2576B7341922B5" box="[506,596,481,503]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47A</figureCitation>
); aedeagal shaft in lateral view curved (
<figureCitation id="1325F2D0F06CFFB9FC0E76B7320422B5" box="[977,1097,481,503]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47G, K</figureCitation>
)..........
<taxonomicName id="4C1E95D6F06CFFB9FB1276B433D022B4" authority="(Distant)" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[1229,1437,481,503]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B96A3247F06CFFB9FB1276B4330C22B5" box="[1229,1345,481,503]" italics="true" pageId="37" pageNumber="38">P. maculosa</emphasis>
(Distant)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF4876A933D12157" blockId="37.[151,1437,363,532]" box="[151,1436,510,532]" pageId="37" pageNumber="38">
<keyLead id="30EF4D60F06CFFB9FF4876A933D12157" box="[151,1436,510,532]" pageId="37" pageNumber="38">
- Pygofer without dorsal process (
<figureCitation id="1325F2D0F06CFFB9FDC676A8343E2150" box="[537,627,510,532]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48AI. Panghoella nera sp. nov.: AB, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; GH, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="37" pageNumber="38">Fig. 48D</figureCitation>
); aedeagal shaft in lateral view more or less straight (
<figureCitation id="1325F2D0F06CFFB9FBAD76A8328D2150" box="[1138,1216,510,532]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48AI. Panghoella nera sp. nov.: AB, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; GH, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="37" pageNumber="38">Fig. 48I</figureCitation>
)....
<taxonomicName id="4C1E95D6F06CFFB9FADA76A933052157" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[1285,1352,511,532]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nera" status="sp. nov.">
<emphasis id="B96A3247F06CFFB9FADA76A933052157" box="[1285,1352,511,532]" italics="true" pageId="37" pageNumber="38">P. nera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2598F3CF06CFFB9FA9176A833D12157" box="[1358,1436,510,532]" pageId="37" pageNumber="38" rank="species">
<emphasis id="B96A3247F06CFFB9FA9176A833DB2150" bold="true" box="[1358,1430,510,531]" pageId="37" pageNumber="38">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>